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<document id="73D982CA9379B081D4560868AED6B6D6" ID-DOI="10.11646/zootaxa.2279.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5307538" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629653572613" checkinUser="felipe" docAuthor="Brown, John W." docDate="2009" docId="038E7928FFDBFF83FDACE797FC71FC19" docLanguage="en" docName="zt02279p050.pdf" docOrigin="Zootaxa 2279 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2279.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Megalota gutierrezi Brown 2009, new species" docType="treatment" docVersion="5" lastPageNumber="29" masterDocId="FFB70150FFC0FF9FFFBCE420FFFBFFB3" masterDocTitle="The discovery of Megalota in the Neotropics, with a revision of the New World species (Lepidoptera: Tortricidae: Olethreutini)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="28" updateTime="1699089488996" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="66D30C0AB5E451B0EA8014CA92921694">The discovery of Megalota in the Neotropics, with a revision of the New World species (Lepidoptera: Tortricidae: Olethreutini)</mods:title>
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<mods:namePart id="34E1AD92BE5043726DA390CFB3466820">Brown, John W.</mods:namePart>
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<mods:title id="6F0727C6B43335CEB7E8DDA930DA4176">Zootaxa</mods:title>
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<mods:date id="7D27ED0752CFBB0C7D015DB629C9EFC1">2009</mods:date>
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<treatment id="038E7928FFDBFF83FDACE797FC71FC19" ID-DOI="http://doi.org/10.5281/zenodo.6494802" ID-Zenodo-Dep="6494802" LSID="urn:lsid:plazi:treatment:038E7928FFDBFF83FDACE797FC71FC19" httpUri="http://treatment.plazi.org/id/038E7928FFDBFF83FDACE797FC71FC19" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C33D9BB5FFDBFF84FDACE797FC8FFC47" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B98C83EFFDBFF84FDACE797FC8FFC47" blockId="27.[528,1059,951,1012]" pageId="27" pageNumber="28">
<emphasis id="B953142CFFDBFF84FDACE797FBD8FC62" bold="true" box="[528,1059,951,977]" pageId="27" pageNumber="28">
19.
<taxonomicName id="4C27B3BDFFDBFF84FD80E797FC7DFC62" authority="Brown" authorityName="Brown" authorityYear="2009" box="[572,902,951,977]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gutierrezi" status="sp. nov.">
<emphasis id="B953142CFFDBFF84FD80E797FCD2FC62" bold="true" box="[572,809,951,977]" italics="true" pageId="27" pageNumber="28">Megalota gutierrezi</emphasis>
Brown
</taxonomicName>
,
<taxonomicNameLabel id="A260A957FFDBFF84FC2FE797FBD8FC62" box="[915,1059,951,977]" pageId="27" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation id="131CD4BBFFDBFF84FD02E7FAFCE5FC47" box="[702,798,986,1012]" captionStart="FIGURES 1324" captionStartId="37.[151,269,1664,1688]" captionTargetBox="[200,1385,199,1632]" captionTargetId="figure-68@37.[194,1393,194,1640]" captionTargetPageId="37" captionText="FIGURES 1324. Adults of Megalota. 13. M. ricana (holotype), 14. M. ceratovalva (holotype), 15. M. bicolorana (holotype), 16. M. longisetana (holotype), 17. M. deceptana (holotype), 18. M. crassana (holotype), 19. M. gutierrezi (holotype), 20. M. chamelana (holotype), 21. M. beckeri (holotype), 22. M. plenana (holotype), 23. M. flintana (holotype), 24. M. pastranai (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307544" httpUri="https://zenodo.org/record/5307544/files/figure.png" pageId="27" pageNumber="28">Figs. 19</figureCitation>
,
<figureCitation id="131CD4BBFFDBFF84FC97E7FAFCB0FC47" box="[811,843,986,1012]" captionStart="FIGURES 4345" captionStartId="44.[151,269,1979,2003]" captionTargetBox="[190,1397,178,1971]" captionTargetId="figure-56@44.[190,1398,128,1993]" captionTargetPageId="44" captionText="FIGURES 4345. Male genitalia of Megalota; valva spread, phallus removed. 43. M. gutierrezi, image of USNM slide 84,883; 44. M. chamelana, image of USNM slide 84,897; 45. M. beckeri, image of USNM slide 124,145." figureDoi="http://doi.org/10.5281/zenodo.5307560" httpUri="https://zenodo.org/record/5307560/files/figure.png" pageId="27" pageNumber="28">43</figureCitation>
,
<figureCitation id="131CD4BBFFDBFF84FCEBE7FAFC8FFC47" box="[855,884,986,1012]" captionStart="FIGURES 6063" captionStartId="49.[151,269,1907,1931]" captionTargetBox="[218,1369,178,1899]" captionTargetId="figure-57@49.[218,1370,144,1923]" captionTargetPageId="49" captionText="FIGURES 6063. Female genitalia of Megalota. 60. M. longisetana, image of USNM slide 84,894; 61. M. deceptana, image of USNM slide 84,886; 62. M. crassana, image of USNM slide 95,343; 63. M. gutierrezi, image of USNM slide 84,876." figureDoi="http://doi.org/10.5281/zenodo.5339678" httpUri="https://zenodo.org/record/5339678/files/figure.png" pageId="27" pageNumber="28">63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFDBFF84FF2BE00AFEE7FA4F" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8B98C83EFFDBFF84FF2BE00AFEE7FA4F" blockId="27.[151,1436,1066,2012]" pageId="27" pageNumber="28">
<emphasis id="B953142CFFDBFF84FF2BE00AFEF4FBF7" bold="true" box="[151,271,1066,1092]" pageId="27" pageNumber="28">Diagnosis</emphasis>
.
<taxonomicName id="4C27B3BDFFDBFF84FE9FE00AFDEBFBF0" authorityName="Brown" authorityYear="2009" box="[291,528,1066,1091]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gutierrezi">
<emphasis id="B953142CFFDBFF84FE9FE00AFDEBFBF0" box="[291,528,1066,1091]" italics="true" pageId="27" pageNumber="28">Megalota gutierrezi</emphasis>
</taxonomicName>
is a comparatively large species, superficially similar to
<taxonomicName id="4C27B3BDFFDBFF84FB68E00AFA92FBF0" authorityName="Brown" authorityYear="2009" box="[1236,1385,1066,1091]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crassana">
<emphasis id="B953142CFFDBFF84FB68E00AFA92FBF0" box="[1236,1385,1066,1091]" italics="true" pageId="27" pageNumber="28">M. crassana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C27B3BDFFDBFF84FAC0E00AFEECFBDF" authorityName="Brown" authorityYear="2009" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bicolorana">
<emphasis id="B953142CFFDBFF84FAC0E00AFEECFBDF" italics="true" pageId="27" pageNumber="28">M. bicolorana</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C27B3BDFFDBFF84FEE4E073FE0CFBDF" authorityName="Brown" authorityYear="2009" box="[344,503,1107,1132]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deceptana">
<emphasis id="B953142CFFDBFF84FEE4E073FE0CFBDF" box="[344,503,1107,1132]" italics="true" pageId="27" pageNumber="28">M. deceptana</emphasis>
</taxonomicName>
but with the dark mark from mid-termen distinctly triangular. The male genitalia are easily distinguished by the broad basal portion of the valva with a small, angulate, subtriangular process near the venter immediately distad of the basal cavity. Also, the two groups of spiniform setae, usually distinct and separate clusters in congeners, are more or less continuous along the venter of the basal portion of the valva from the subtriangular process to about 0.6 the distance to the apex, where the valva becomes narrower. The phallus of
<taxonomicName id="4C27B3BDFFDBFF84FE20E13BFDCEFA87" authorityName="Brown" authorityYear="2009" box="[412,565,1307,1332]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gutierrezi">
<emphasis id="B953142CFFDBFF84FE20E13BFDCEFA87" box="[412,565,1307,1332]" italics="true" pageId="27" pageNumber="28">M. gutierrezi</emphasis>
</taxonomicName>
is long, slender, and curved throughout, with a single small erect thorn at the tip, whereas the aedeagi of
<taxonomicName id="4C27B3BDFFDBFF84FDBCE163FD5DFAEF" authorityName="Brown" authorityYear="2009" box="[512,678,1347,1372]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bicolorana">
<emphasis id="B953142CFFDBFF84FDBCE163FD5DFAEF" box="[512,678,1347,1372]" italics="true" pageId="27" pageNumber="28">M. bicolorana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C27B3BDFFDBFF84FD63E163FC85FAEF" authorityName="Brown" authorityYear="2009" box="[735,894,1347,1372]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deceptana">
<emphasis id="B953142CFFDBFF84FD63E163FC85FAEF" box="[735,894,1347,1372]" italics="true" pageId="27" pageNumber="28">M. deceptana</emphasis>
</taxonomicName>
are shorter and broader, and have one or more erect thorns from the dorsum before the tip, and
<taxonomicName id="4C27B3BDFFDBFF84FD6AE14BFC93FA37" authorityName="Brown" authorityYear="2009" box="[726,872,1387,1412]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crassana">
<emphasis id="B953142CFFDBFF84FD6AE14BFC93FA37" box="[726,872,1387,1412]" italics="true" pageId="27" pageNumber="28">M. crassana</emphasis>
</taxonomicName>
lacks thorns altogether. The female genitalia of
<taxonomicName id="4C27B3BDFFDBFF84FF2BE1B3FED4FA1F" authorityName="Brown" authorityYear="2009" box="[151,303,1427,1452]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gutierrezi">
<emphasis id="B953142CFFDBFF84FF2BE1B3FED4FA1F" box="[151,303,1427,1452]" italics="true" pageId="27" pageNumber="28">M. gutierrezi</emphasis>
</taxonomicName>
are easily distinguished by the small, deeply U-shaped emargination at the ventro-posterior edge of the ostium and the semi-membranous, bulbous enlargement of the ductus bursae immediately anterad of the colliculum.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFDBFF83FF7AE22AFC9AFE41" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8B98C83EFFDBFF83FF7AE22AFC9AFE41" blockId="27.[151,1436,1066,2012]" lastBlockId="28.[151,1436,152,938]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B953142CFFDBFF84FF7AE22AFEACF997" bold="true" box="[198,343,1546,1572]" pageId="27" pageNumber="28">Description</emphasis>
.
<emphasis id="B953142CFFDBFF84FED5E22BFE53F997" box="[361,424,1547,1572]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
: Vertex reddish brown and creamy white, frons creamy white; labial palpus mixed creamy white and dark brown.
<emphasis id="B953142CFFDBFF84FD9FE213FD83F9FF" box="[547,632,1587,1612]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
: Dorsum pale brown and rust; metascutum with well-developed maroon scale tuft. Hind tibia in male with dense, cream-colored, modified sex scaling and hairpencil. Forewing length
<quantity id="4CDF65DBFFDBFF84FE97E2A2FE43F92F" box="[299,440,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="27" pageNumber="28" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
(mean = 9.1); basal 0.33 somewhat checkered dark brown, pinkish brown, and copper, bordered distally by a pale, ill-defined, oblique fascia; a dark semicircular patch at mid-costa, with a paler, less defined extension to dorsum; small, dark brown, subtriangular patch in subterminal region immediately apicad of distal end of discal cell; tornal region pale brown. Fringe brown. Hindwing brown, anal margin in male with distinct fold bearing long, slender, cream-colored scales.
<emphasis id="B953142CFFDBFF84FC69E303FBB8F88F" box="[981,1091,1827,1852]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
: Brown. Male genitalia (
<figureCitation id="131CD4BBFFDBFF84FAD1E302FF4CF8D7" captionStart="FIGURES 4345" captionStartId="44.[151,269,1979,2003]" captionTargetBox="[190,1397,178,1971]" captionTargetId="figure-56@44.[190,1398,128,1993]" captionTargetPageId="44" captionText="FIGURES 4345. Male genitalia of Megalota; valva spread, phallus removed. 43. M. gutierrezi, image of USNM slide 84,883; 44. M. chamelana, image of USNM slide 84,897; 45. M. beckeri, image of USNM slide 124,145." figureDoi="http://doi.org/10.5281/zenodo.5307560" httpUri="https://zenodo.org/record/5307560/files/figure.png" pageId="27" pageNumber="28">Fig. 43</figureCitation>
;
<figureCitation id="131CD4BBFFDBFF84FF78E36AFF29F8D7" box="[196,210,1866,1892]" captionStart="FIGURES 112" captionStartId="36.[151,269,1661,1685]" captionTargetBox="[199,1384,202,1634]" captionTargetId="figure-73@36.[195,1392,194,1637]" captionTargetPageId="36" captionText="FIGURES 112. Adults of Megalota. 1. M. submicans (holotype), 2. M. synchysis (paratype), 3. M. peruviana (paratype), 4. M. aquilonaris (holotype), 5. M. vulgaris (paratype), 6. M. cacaulana (holotype), 7. M. macrosocia (holotype), 8. M. ochreoapex (holotype), 9. M. spinulosa (holotype), 10. M. simpliciana (paratype), 11. M. delphinosema (Costa Rica), 12. M. jamaicana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307542" httpUri="https://zenodo.org/record/5307542/files/figure.png" pageId="27" pageNumber="28">3</figureCitation>
preparations examined) with tegumen broad, subrectangular, slightly narrowed in dorsal 0.3, with lobelike expansion extremely weak; uncus broad with wide U-shaped notch dorsally, each lobe obovate, densely spined; valva broad in basal 0.6, about twice as broad as distal 0.4; small dense cluster of spiniform setae and accompanying small triangular process near venter at lower-outer edge of basal cavity of valva, with irregularly linear patch of spiniform setae continuous with basal spiniform cluster along venter to abruptly narrowed point of valva; basal process of valva 45 times as long as wide, irregularly expanded beyond middle, lacking distinct elbow, with attenuate tip. Phallus long, ca. 0.65 length of valva, evenly curved throughout, with distal, slightly upturned thorn; vesica with a single slender cornutus. Female genitalia (
<figureCitation id="131CD4BBFFDCFF83FBFDE530FB63FE99" box="[1089,1176,272,298]" captionStart="FIGURES 6063" captionStartId="49.[151,269,1907,1931]" captionTargetBox="[218,1369,178,1899]" captionTargetId="figure-57@49.[218,1370,144,1923]" captionTargetPageId="49" captionText="FIGURES 6063. Female genitalia of Megalota. 60. M. longisetana, image of USNM slide 84,894; 61. M. deceptana, image of USNM slide 84,886; 62. M. crassana, image of USNM slide 95,343; 63. M. gutierrezi, image of USNM slide 84,876." figureDoi="http://doi.org/10.5281/zenodo.5339678" httpUri="https://zenodo.org/record/5339678/files/figure.png" pageId="28" pageNumber="29">Fig. 63</figureCitation>
;
<figureCitation id="131CD4BBFFDCFF83FB19E530FB4FFE99" box="[1189,1204,272,298]" captionStart="FIGURES 112" captionStartId="36.[151,269,1661,1685]" captionTargetBox="[199,1384,202,1634]" captionTargetId="figure-73@36.[195,1392,194,1637]" captionTargetPageId="36" captionText="FIGURES 112. Adults of Megalota. 1. M. submicans (holotype), 2. M. synchysis (paratype), 3. M. peruviana (paratype), 4. M. aquilonaris (holotype), 5. M. vulgaris (paratype), 6. M. cacaulana (holotype), 7. M. macrosocia (holotype), 8. M. ochreoapex (holotype), 9. M. spinulosa (holotype), 10. M. simpliciana (paratype), 11. M. delphinosema (Costa Rica), 12. M. jamaicana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307542" httpUri="https://zenodo.org/record/5307542/files/figure.png" pageId="28" pageNumber="29">2</figureCitation>
preparations examined) with papillae anales simple, unmodified; sterigma in the form of a narrow ill-defined band; ostium with small, deeply U-shaped emargination; colliculum slightly less than 0.5 length of ductus bursae, strongly sclerotized except for narrow longitudinal mesal portion, remainder of ductus bursae membranous, with bulbous enlargement immediately anterad of the colliculum; corpus bursae narrow pear-shaped; signum in the form of a narrow tongue-shaped sclerite from a broad, irregular base.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFDCFF83FF7AE5DFFB8CFD51" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B98C83EFFDCFF83FF7AE5DFFE12FDF1" blockId="28.[151,1436,152,938]" pageId="28" pageNumber="29">
<materialsCitation id="3B4FC263FFDCFF83FF7AE5DFFE12FDF1" collectingDate="1992-03-29" collectingDateMax="1992-03-31" collectingDateMin="1992-03-29" collectionCode="EME" collectorName="S. Meredith &amp; J. Powell" country="Costa Rica" elevation="1400" location="Monteverde" municipality="Male" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="B953142CFFDCFF83FF7AE5DFFEC3FDAA" bold="true" box="[198,312,511,537]" pageId="28" pageNumber="29">
<typeStatus id="549C769CFFDCFF83FF7AE5DFFEC3FDAA" box="[198,312,511,537]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="6BFC5244FFDCFF83FEFAE620FE7EFDA9" box="[326,389,512,538]" pageId="28" pageNumber="29">Male</collectingMunicipality>
,
<collectingCountry id="F33088AEFFDCFF83FE2DE620FDE8FDA9" box="[401,531,512,538]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
,
<collectingRegion id="49E306DCFFDCFF83FDA3E620FD59FDA9" box="[543,674,512,538]" country="Costa Rica" name="Puntarenas" pageId="28" pageNumber="29">Puntarenas</collectingRegion>
,
<location id="8EF89EE5FFDCFF83FD11E620FCC5FDA9" LSID="urn:lsid:plazi:treatment:038E7928FFDBFF83FDACE797FC71FC19:8EF89EE5FFDCFF83FD11E620FCC5FDA9" box="[685,830,512,538]" country="Costa Rica" municipality="Male" name="Monteverde" pageId="28" pageNumber="29" stateProvince="Puntarenas">Monteverde</location>
,
<quantity id="4CDF65DBFFDCFF83FCF5E620FC5FFDA9" box="[841,932,512,538]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">
<elevation id="000A2F0DFFDCFF83FCF5E620FC5FFDA9" box="[841,932,512,538]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FF99EEFEFFDCFF83FC0CE620FB8EFDA9" box="[944,1141,512,538]" pageId="28" pageNumber="29" value="1992-03-29" valueMax="1992-03-31" valueMin="1992-03-29">
<collectingDate id="EFDD1716FFDCFF83FC0CE620FB8EFDA9" box="[944,1141,512,538]" pageId="28" pageNumber="29" value="1992-03-29" valueMax="1992-03-31" valueMin="1992-03-29">2931 Mar 1992</collectingDate>
</date>
,
<collectorName id="26D2ADE8FFDCFF83FB3EE620FAF0FDA9" box="[1154,1291,512,538]" pageId="28" pageNumber="29">S. Meredith</collectorName>
&amp;
<collectorName id="26D2ADE8FFDCFF83FA8CE620FA60FDA9" box="[1328,1435,512,538]" pageId="28" pageNumber="29">J. Powell</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FF23E608FF19FDF1" box="[159,226,552,578]" pageId="28" pageNumber="29">EME</collectionCode>
), USNM slide 84,888.
</materialsCitation>
</paragraph>
<paragraph id="8B98C83EFFDCFF83FF7AE66FFB8CFD51" blockId="28.[151,1436,152,938]" pageId="28" pageNumber="29">
<materialsCitation id="3B4FC263FFDCFF83FF7AE66FFC51FDD9" box="[198,938,591,618]" collectorName="Monteverde" country="Costa Rica" location="Puntarenas" pageId="28" pageNumber="29" specimenCount="12" specimenCount-female="3" specimenCount-male="9" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B953142CFFDCFF83FF7AE66FFEB8FDDA" bold="true" box="[198,323,591,617]" pageId="28" pageNumber="29">
<typeStatus id="549C769CFFDCFF83FF7AE66FFEB8FDDA" box="[198,323,591,617]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D2103B7FFDCFF83FEE8E670FE8CFDDB" box="[340,375,592,618]" pageId="28" pageNumber="29" type="male">9♂</specimenCount>
,
<specimenCount id="9D2103B7FFDCFF83FE39E670FE51FDDA" box="[389,426,591,618]" pageId="28" pageNumber="29" type="female">3♀</specimenCount>
).
<collectingCountry id="F33088AEFFDCFF83FE7EE670FD8AFDD9" box="[450,625,592,618]" name="Costa Rica" pageId="28" pageNumber="29">COSTA RICA</collectingCountry>
:
<emphasis id="B953142CFFDCFF83FDC0E66FFCF0FDDA" bold="true" box="[636,779,591,617]" pageId="28" pageNumber="29">
<collectingRegion id="49E306DCFFDCFF83FDC0E66FFCF0FDDA" box="[636,779,591,617]" country="Costa Rica" name="Puntarenas" pageId="28" pageNumber="29">Puntarenas</collectingRegion>
</emphasis>
:
<collectorName id="26D2ADE8FFDCFF83FCA6E670FC51FDD9" box="[794,938,592,618]" pageId="28" pageNumber="29">Monteverde</collectorName>
</materialsCitation>
,
<materialsCitation id="3B4FC263FFDCFF83FC0AE670FEF9FD21" collectingDate="1987-03-26" collectionCode="USNM" collectorName="Steiner" country="Costa Rica" elevation="1400" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<quantity id="4CDF65DBFFDCFF83FC0AE670FBEAFDD9" box="[950,1041,592,618]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">
<elevation id="000A2F0DFFDCFF83FC0AE670FBEAFDD9" box="[950,1041,592,618]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FF99EEFEFFDCFF83FBA1E670FB4FFDD9" box="[1053,1204,592,618]" pageId="28" pageNumber="29" value="1987-03-26">
<collectingDate id="EFDD1716FFDCFF83FBA1E670FB4FFDD9" box="[1053,1204,592,618]" pageId="28" pageNumber="29" value="1987-03-26">26 Mar 1987</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFDCFF83FB7BE670FB11FDDA" box="[1223,1258,591,618]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
), W. E.
<collectorName id="26D2ADE8FFDCFF83FAF5E670FA67FDD9" box="[1353,1436,592,618]" pageId="28" pageNumber="29">Steiner</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FF23E658FF01FD21" box="[159,250,632,658]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">USNM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFDCFF83FEB3E658FC3AFD21" box="[271,961,631,658]" collectingDate="1990-07-22" collectingDateMax="1990-07-24" collectingDateMin="1990-07-22" collectionCode="EME" collectorName="S. Meredith &amp; Powell" country="Costa Rica" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<date id="FF99EEFEFFDCFF83FEB3E658FE3FFD21" box="[271,452,632,658]" pageId="28" pageNumber="29" value="1990-07-22" valueMax="1990-07-24" valueMin="1990-07-22">
<collectingDate id="EFDD1716FFDCFF83FEB3E658FE3FFD21" box="[271,452,632,658]" pageId="28" pageNumber="29" value="1990-07-22" valueMax="1990-07-24" valueMin="1990-07-22">2224 Jul 1990</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFDCFF83FE64E658FE02FD23" box="[472,505,632,658]" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<specimenCount id="9D2103B7FFDCFF83FDB5E658FDD6FD22" box="[521,557,631,658]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
),
<collectorName id="26D2ADE8FFDCFF83FDFAE658FD2AFD21" box="[582,721,632,658]" pageId="28" pageNumber="29">S. Meredith</collectorName>
&amp; J.
<collectorName id="26D2ADE8FFDCFF83FCAFE658FC9FFD21" box="[787,868,632,658]" pageId="28" pageNumber="29">Powell</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FCC9E658FC43FD21" box="[885,952,632,658]" pageId="28" pageNumber="29">EME</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFDCFF83FC72E658FE81FD09" collectingDate="1992-03-29" collectingDateMax="1992-03-31" collectingDateMin="1992-03-29" collectionCode="EME" collectorName="S. McCarty &amp; Powell" country="Costa Rica" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<date id="FF99EEFEFFDCFF83FC72E658FB69FD21" box="[974,1170,632,658]" pageId="28" pageNumber="29" value="1992-03-29" valueMax="1992-03-31" valueMin="1992-03-29">
<collectingDate id="EFDD1716FFDCFF83FC72E658FB69FD21" box="[974,1170,632,658]" pageId="28" pageNumber="29" value="1992-03-29" valueMax="1992-03-31" valueMin="1992-03-29">2931 Mar 1992</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFDCFF83FB1AE658FB3CFD23" box="[1190,1223,632,658]" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
,
<specimenCount id="9D2103B7FFDCFF83FB6AE658FB00FD22" box="[1238,1275,631,658]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
),
<collectorName id="26D2ADE8FFDCFF83FAA8E658FA67FD21" box="[1300,1436,632,658]" pageId="28" pageNumber="29">S. McCarty</collectorName>
&amp; J.
<collectorName id="26D2ADE8FFDCFF83FF72E680FEE4FD09" box="[206,287,672,698]" pageId="28" pageNumber="29">Powell</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FE92E680FE89FD09" box="[302,370,672,698]" pageId="28" pageNumber="29">EME</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFDCFF83FE3AE680FBFBFD09" box="[390,1024,672,698]" collectingDate="1986-06-08" collectingDateMax="1986-06-10" collectingDateMin="1986-06-08" collectionCode="EME" collectorName="J. Chemsak &amp; Katsura" country="Costa Rica" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<date id="FF99EEFEFFDCFF83FE3AE680FDCBFD09" box="[390,560,672,698]" pageId="28" pageNumber="29" value="1986-06-08" valueMax="1986-06-10" valueMin="1986-06-08">
<collectingDate id="EFDD1716FFDCFF83FE3AE680FDCBFD09" box="[390,560,672,698]" pageId="28" pageNumber="29" value="1986-06-08" valueMax="1986-06-10" valueMin="1986-06-08">810 Jun 1986</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFDCFF83FDFFE680FD9FFD0B" box="[579,612,672,698]" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
),
<collectorName id="26D2ADE8FFDCFF83FDC0E680FCF9FD09" box="[636,770,672,698]" pageId="28" pageNumber="29">J. Chemsak</collectorName>
&amp; H.
<collectorName id="26D2ADE8FFDCFF83FCF6E680FC5EFD09" box="[842,933,672,698]" pageId="28" pageNumber="29">Katsura</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FC08E680FC03FD09" box="[948,1016,672,698]" pageId="28" pageNumber="29">EME</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFDCFF83FBB7E680FECDFD51" country="Costa Rica" location="Monteverde Cloud Forest Reserve" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<location id="8EF89EE5FFDCFF83FBB7E680FA67FD09" LSID="urn:lsid:plazi:treatment:038E7928FFDBFF83FDACE797FC71FC19:8EF89EE5FFDCFF83FBB7E680FA67FD09" box="[1035,1436,672,698]" country="Costa Rica" name="Monteverde Cloud Forest Reserve" pageId="28" pageNumber="29" stateProvince="Puntarenas">Monteverde Cloud Forest Reserve</location>
Headquarters
</materialsCitation>
,
<materialsCitation id="3B4FC263FFDCFF83FEFEE6E8FB89FD51" box="[322,1138,712,738]" collectingDate="1985-05-18" collectionCode="EME" collectorName="J. Powell &amp; P. Opler &amp; Chemsak" country="Costa Rica" elevation="1450" pageId="28" pageNumber="29" specimenCount="6" specimenCount-male="6" stateProvince="Puntarenas" typeStatus="paratype">
<quantity id="4CDF65DBFFDCFF83FEFEE6E8FE67FD51" box="[322,412,712,738]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="28" pageNumber="29" unit="m" value="1450.0">
<elevation id="000A2F0DFFDCFF83FEFEE6E8FE67FD51" box="[322,412,712,738]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="28" pageNumber="29" unit="m" value="1450.0">1450 m</elevation>
</quantity>
,
<date id="FF99EEFEFFDCFF83FE15E6E8FDBEFD51" box="[425,581,712,738]" pageId="28" pageNumber="29" value="1985-05-18">
<collectingDate id="EFDD1716FFDCFF83FE15E6E8FDBEFD51" box="[425,581,712,738]" pageId="28" pageNumber="29" value="1985-05-18">18 May 1985</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFDCFF83FDEAE6E8FD82FD53" box="[598,633,712,738]" pageId="28" pageNumber="29" type="male">6♂</specimenCount>
),
<collectorName id="26D2ADE8FFDCFF83FD2DE6E8FCFBFD51" box="[657,768,712,738]" pageId="28" pageNumber="29">J. Powell</collectorName>
,
<collectorName id="26D2ADE8FFDCFF83FCB0E6E8FC91FD51" box="[780,874,712,738]" pageId="28" pageNumber="29">P. Opler</collectorName>
&amp; J.
<collectorName id="26D2ADE8FFDCFF83FC15E6E8FBEDFD51" box="[937,1046,712,738]" pageId="28" pageNumber="29">Chemsak</collectorName>
(
<collectionCode id="ED3650FBFFDCFF83FB9AE6E8FB91FD51" box="[1062,1130,712,738]" pageId="28" pageNumber="29">EME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFDCFF83FF7AE6CFFDEAFCE9" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8B98C83EFFDCFF83FF7AE6CFFDEAFCE9" blockId="28.[151,1436,152,938]" pageId="28" pageNumber="29">
<emphasis id="B953142CFFDCFF83FF7AE6CFFE01FCBA" bold="true" box="[198,506,751,777]" pageId="28" pageNumber="29">Distribution and Biology</emphasis>
. This species is known only from cloud forest habitat in the vicinity of Monteverde,
<collectingCountry id="F33088AEFFDCFF83FF5BE738FE91FC81" box="[231,362,792,818]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
. Capture records include March (n = 4), May (n = 6), June (n = 1), and July (n = 2). Nothing is known of the early stages.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFDCFF83FF7AE747FC71FC19" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B98C83EFFDCFF83FF7AE747FC71FC19" blockId="28.[151,1436,152,938]" pageId="28" pageNumber="29">
<emphasis id="B953142CFFDCFF83FF7AE747FEB0FC32" bold="true" box="[198,331,871,897]" pageId="28" pageNumber="29">Etymology</emphasis>
. This species is named for Dr. Mario Gutiérrez in recognition of his work in the field of genetics and his generous support of the scientific activities of INBio.
</paragraph>
</subSubSection>
</treatment>
</document>