491 lines
56 KiB
XML
491 lines
56 KiB
XML
<document id="F4134D5A61749F5DB67236496AD0462D" ID-DOI="10.11646/zootaxa.3681.5.4" ID-GBIF-Dataset="36c81ce0-c6ae-498c-b222-022ccfcf5e65" ID-ISSN="1175-5326" ID-Zenodo-Dep="222069" ID-ZooBank="86F99E54-F2F9-4078-8639-8961F38AB75C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460641442902" checkinUser="plazi" docAuthor="Ng, Heok Hee & Kullander, Sven O." docDate="2013" docId="0387B116FFBBFC4CFF641D914F041FD2" docLanguage="en" docName="zt03681p562.pdf" docOrigin="Zootaxa 3681 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyptothorax igniculus Ng & Kullander, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="559" masterDocId="FFBEC96EFFBAFC4BFFF31C054C191E1D" masterDocTitle="Glyptothorax igniculus, a new species of sisorid catfish (Teleostei: Siluriformes) from Myanmar" masterLastPageNumber="562" masterPageNumber="552" pageNumber="553" updateTime="1698335898793" updateUser="plazi">
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<mods:title id="E58A13393CDE45BED35FA7221436B1EF">Glyptothorax igniculus, a new species of sisorid catfish (Teleostei: Siluriformes) from Myanmar</mods:title>
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<mods:namePart id="BB03E5136D70779BA9BC201CAFCA6843">Ng, Heok Hee</mods:namePart>
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<mods:namePart id="8FDDC28D27EC20403C6DA72FBD44C62D">Kullander, Sven O.</mods:namePart>
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<mods:title id="C86FEFF1E8D5BA73E3AD5FB701DC25C4">Zootaxa</mods:title>
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<mods:part id="310EE51D3EAB0B4BF6B6E1F89F0EE6F7">
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<mods:date id="7FCB6C62A0E35B961E2279B4966C21B3">2013</mods:date>
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<mods:number id="6B481543326852692187D63F0C8439BA">3681</mods:number>
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<treatment id="0387B116FFBBFC4CFF641D914F041FD2" ID-DOI="http://doi.org/10.5281/zenodo.5689229" ID-GBIF-Taxon="119571272" ID-Zenodo-Dep="5689229" LSID="urn:lsid:plazi:treatment:0387B116FFBBFC4CFF641D914F041FD2" httpUri="http://treatment.plazi.org/id/0387B116FFBBFC4CFF641D914F041FD2" lastPageId="7" lastPageNumber="559" pageId="1" pageNumber="553">
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<subSubSection id="C334538BFFBBFC4AFF641D914CF21FD3" pageId="1" pageNumber="553" type="nomenclature">
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<paragraph id="8B910000FFBBFC4AFF641D914E551FB0" blockId="1.[151,588,403,462]" box="[151,588,403,430]" pageId="1" pageNumber="553">
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<emphasis id="B95ADC12FFBBFC4AFF641D914E551FB0" bold="true" box="[151,588,403,430]" pageId="1" pageNumber="553">
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<taxonomicName id="4C2E7B83FFBBFC4AFF641D914DB71FB3" ID-CoL="6KM4T" box="[151,430,404,430]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="igniculus" status="sp. nov.">
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<emphasis id="B95ADC12FFBBFC4AFF641D914DB71FB3" bold="true" box="[151,430,404,430]" italics="true" pageId="1" pageNumber="553">Glyptothorax igniculus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2696169FFBBFC4AFE4E1D964E551FB0" box="[445,588,403,429]" pageId="1" pageNumber="553" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B910000FFBBFC4AFF641DB34CF21FD3" blockId="1.[151,588,403,462]" box="[151,235,438,462]" pageId="1" pageNumber="553">
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(
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<figureCitation id="13151C85FFBBFC4AFF6C1DB34CF91FD3" box="[159,224,438,462]" captionStart="FIGURE 1" captionStartId="2.[151,250,1929,1952]" captionTargetBox="[151,1436,375,1905]" captionTargetId="figure@2.[151,1436,375,1905]" captionTargetPageId="2" captionText="FIGURE 1. Glyptothorax igniculus, NRM 64520, holotype, 65.5 mm SL; dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/222070/files/figure.png" pageId="1" pageNumber="553">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C334538BFFBBFC4AFF641DFA4EF31CBB" pageId="1" pageNumber="553" type="materials_examined">
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<paragraph id="8B910000FFBBFC4AFF641DFA4EF31CBB" blockId="1.[151,1436,510,2011]" pageId="1" pageNumber="553">
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<emphasis id="B95ADC12FFBBFC4AFF641DFA4D501C0A" bold="true" box="[151,329,510,535]" pageId="1" pageNumber="553">
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<typeStatus id="5495BEA2FFBBFC4AFF641DFA4CCB1C0A" box="[151,210,511,535]" pageId="1" pageNumber="553">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="5495BEA2FFBBFC4AFEA71DFB4DD91C0B" box="[340,448,510,534]" pageId="1" pageNumber="553" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED3F98C5FFBBFC4AFE231DFA4E091C0B" box="[464,528,511,534]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="553">NRM</collectionCode>
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64520,
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<quantity id="4CD6ADE5FFBBFC4AFD871DFB4EC51C0B" box="[628,732,510,534]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" pageId="1" pageNumber="553" unit="mm" value="65.5">65.5 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFD141DFB4F121C0B" box="[743,779,510,534]" pageId="1" pageNumber="553">SL</collectionCode>
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;
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<collectingCountry id="F3394090FFBBFC4AFCE81DFA4F941C0A" box="[795,909,511,535]" name="Myanmar" pageId="1" pageNumber="553">Myanmar</collectingCountry>
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: Sagaing Region, left bank of Myittha River approximately
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<quantity id="4CD6ADE5FFBBFC4AFECC1E274D611C27" box="[319,376,546,570]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="553" unit="km" value="8.0">8 km</quantity>
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NE by E of Kalaymyo,
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<geoCoordinate id="EE1A66C7FFBBFC4AFCE01E274F911C27" box="[787,904,546,570]" direction="east" orientation="longitude" pageId="1" pageNumber="553" precision="15" value="94.13306">94°7'59"E</geoCoordinate>
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; S.
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<collectingCountry id="F3394090FFBBFC4AFC411E274FD01C27" box="[946,969,546,570]" name="Iceland" pageId="1" pageNumber="553">O</collectingCountry>
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. Kullander & T.-
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<collectingCountry id="F3394090FFBBFC4AFB751E2648851C27" box="[1158,1180,547,570]" name="Italy" pageId="1" pageNumber="553">Y</collectingCountry>
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||
. Liao,
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<date id="FF9026C0FFBBFC4AFB141E27498F1C27" box="[1255,1430,546,570]" pageId="1" pageNumber="553" value="2008-03-30">30 March 2008</date>
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.
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<typeStatus id="5495BEA2FFBBFC4AFF641E424D111C42" box="[151,264,583,607]" pageId="1" pageNumber="553" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED3F98C5FFBBFC4AFEE71E424D4D1C43" box="[276,340,583,606]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="553">NRM</collectionCode>
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64521 (1),
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<quantity id="4CD6ADE5FFBBFC4AFE2E1E434E5D1C43" box="[477,580,582,606]" metricMagnitude="-2" metricUnit="m" metricValue="6.7299999999999995" pageId="1" pageNumber="553" unit="mm" value="67.3">67.3 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFDBF1E434E691C43" box="[588,624,582,606]" pageId="1" pageNumber="553">SL</collectionCode>
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; data as for
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<typeStatus id="5495BEA2FFBBFC4AFCF31E434F7F1C43" box="[768,870,582,606]" pageId="1" pageNumber="553" type="holotype">holotype</typeStatus>
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.
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<collectionCode id="ED3F98C5FFBBFC4AFC801E424FAA1C43" box="[883,947,583,606]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="553">NRM</collectionCode>
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58846 (1),
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<quantity id="4CD6ADE5FFBBFC4AFBCF1E4348BB1C43" box="[1084,1186,582,606]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" pageId="1" pageNumber="553" unit="mm" value="62.5">62.5 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFB581E4348D61C43" box="[1195,1231,582,606]" pageId="1" pageNumber="553">SL</collectionCode>
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;
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<collectionCode id="ED3F98C5FFBBFC4AFB2F1E4249051C43" box="[1244,1308,583,606]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="553">NRM</collectionCode>
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58845 (1),
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<quantity id="4CD6ADE5FFBBFC4AFF641E6F4CE51C9F" box="[151,252,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="7.13" pageId="1" pageNumber="553" unit="mm" value="71.3">71.3 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFEF71E6F4D311C9F" box="[260,296,618,642]" pageId="1" pageNumber="553">SL</collectionCode>
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;
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<collectionCode id="ED3F98C5FFBBFC4AFEC61E6F4D721C9F" box="[309,363,618,642]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="553">ZRC</collectionCode>
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54115 (1),
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<quantity id="4CD6ADE5FFBBFC4AFE031E6F4E4C1C9F" box="[496,597,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" pageId="1" pageNumber="553" unit="mm" value="58.5">58.5 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFDAE1E6F4E981C9F" box="[605,641,618,642]" pageId="1" pageNumber="553">SL</collectionCode>
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;
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||
<collectionCode id="ED3F98C5FFBBFC4AFD7E1E6E4ED41C9F" box="[653,717,619,642]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="553">NRM</collectionCode>
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58928 (1),
|
||
<quantity id="4CD6ADE5FFBBFC4AFCA71E6F4FA01C9F" box="[852,953,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="1" pageNumber="553" unit="mm" value="54.7">54.7 mm</quantity>
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<collectionCode id="ED3F98C5FFBBFC4AFC331E6F4FFD1C9F" box="[960,996,618,642]" pageId="1" pageNumber="553">SL</collectionCode>
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;
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<collectingCountry id="F3394090FFBBFC4AFC021E6E487B1C9E" box="[1009,1122,619,643]" name="Myanmar" pageId="1" pageNumber="553">Myanmar</collectingCountry>
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: Sagaing Region, market in Tamu; S.
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||
<collectingCountry id="F3394090FFBBFC4AFEF11E8B4D001CBB" box="[258,281,654,678]" name="Iceland" pageId="1" pageNumber="553">O</collectingCountry>
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. Kullander & T.-
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||
<collectingCountry id="F3394090FFBBFC4AFE251E8A4DF51CBB" box="[470,492,655,678]" name="Italy" pageId="1" pageNumber="553">Y</collectingCountry>
|
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. Liao,
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||
<date id="FF9026C0FFBBFC4AFDCB1E8B4EFE1CBB" box="[568,743,654,678]" pageId="1" pageNumber="553" value="2008-03-29">29 March 2008</date>
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.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C334538BFFBBFC49FF341EB74D171F22" lastPageId="2" lastPageNumber="554" pageId="1" pageNumber="553" type="diagnosis">
|
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<paragraph id="8B910000FFBBFC4AFF341EB74DE71D63" blockId="1.[151,1436,510,2011]" pageId="1" pageNumber="553">
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<emphasis id="B95ADC12FFBBFC4AFF341EB74D581CD6" bold="true" box="[199,321,690,715]" pageId="1" pageNumber="553">Diagnosis.</emphasis>
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||
<taxonomicName id="4C2E7B83FFBBFC4AFEBA1EB64E541CD6" box="[329,589,691,715]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="igniculus">
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<emphasis id="B95ADC12FFBBFC4AFEBA1EB64E541CD6" box="[329,589,691,715]" italics="true" pageId="1" pageNumber="553">Glyptothorax igniculus</emphasis>
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</taxonomicName>
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is distinguished from all congeners in the Irrawaddy River drainage by its thoracic adhesive apparatus, in which the central depression is almost completely enclosed posteriorly by the skin ridges that make up the apparatus, instead of open caudally or, in
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<taxonomicName id="4C2E7B83FFBBFC4AFC6E1EFE485A1D0E" box="[925,1091,763,787]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="burmanicus">
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||
<emphasis id="B95ADC12FFBBFC4AFC6E1EFE485A1D0E" box="[925,1091,763,787]" italics="true" pageId="1" pageNumber="553">G. burmanicus</emphasis>
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||
</taxonomicName>
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||
, completely closed. It further differs from
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<taxonomicName id="4C2E7B83FFBBFC4AFED01F1A4DDC1D2B" box="[291,453,798,822]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="burmanicus">
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||
<emphasis id="B95ADC12FFBBFC4AFED01F1A4DDC1D2B" box="[291,453,798,822]" italics="true" pageId="1" pageNumber="553">G. burmanicus</emphasis>
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||
</taxonomicName>
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||
in having a larger eye (diameter 10.4–12.7% HL vs. 6.3–10.0), a longer dorsal-fin spine (16.9–21.6% SL vs. 11.9–14.7), and shorter prepectoral (17.1–21.7% SL vs. 22.3–25.4) and prepelvic lengths (47.0–51.3% SL vs. 52.8–59.4).
|
||
</paragraph>
|
||
<paragraph id="8B910000FFBBFC4AFF341F8F4EB5181B" blockId="1.[151,1436,510,2011]" pageId="1" pageNumber="553">
|
||
Besides the morphology of the thoracic adhesive apparatus,
|
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<emphasis id="B95ADC12FFBBFC4AFC951F8E4FFE1DBE" box="[870,999,907,931]" italics="true" pageId="1" pageNumber="553">G. i g n i c u l u s</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFB8E1F8E49031DBE" box="[1149,1306,907,931]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="longicauda">
|
||
<emphasis id="B95ADC12FFBBFC4AFB8E1F8E49031DBE" box="[1149,1306,907,931]" italics="true" pageId="1" pageNumber="553">G. longicauda</emphasis>
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||
</taxonomicName>
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||
in having a longer dorsal-fin spine (16.9–21.6% SL vs. 11.5–14.4), a shorter dorsal-to-adipose distance (19.8–24.6% SL vs. 25.9–26.9), deeper body (depth at anus 15.2–16.4% SL vs. 11.0–12.5) and caudal peduncle (8.2–9.7% SL vs. 5.4– 6.2) and fewer total vertebrae (35–36 vs. 43–44), and from
|
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<taxonomicName id="4C2E7B83FFBBFC4AFCC41FF24FE01A13" box="[823,1017,1014,1038]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="longjiangensis">
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<emphasis id="B95ADC12FFBBFC4AFCC41FF24FE01A13" box="[823,1017,1014,1038]" italics="true" pageId="1" pageNumber="553">G. longjiangensis</emphasis>
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</taxonomicName>
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in having a wider head (20.3–21.8% SL vs. 14.5–19.3), rounded tubercles (vs. large anastomosing plaques bearing parallel unculiferous ridges) on the dorsal surface of the head, deeper body (depth at anus 15.2–16.4% SL vs. 11.7–14.4) and caudal peduncle (8.2– 9.7% SL vs. 4.6–7.6) and fewer total vertebrae (35–36 vs. 38–39).
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFC7F186648881A66" box="[908,1169,1123,1147]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="igniculus">
|
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<emphasis id="B95ADC12FFBBFC4AFC7F186648881A66" box="[908,1169,1123,1147]" italics="true" pageId="1" pageNumber="553">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from
|
||
<emphasis id="B95ADC12FFBBFC4AFF2618824D8F1A82" box="[213,406,1158,1183]" italics="true" pageId="1" pageNumber="553">G. m i n i m a c u l a t u s</emphasis>
|
||
in having fewer total vertebrae (35–36 vs. 40–43), from
|
||
<emphasis id="B95ADC12FFBBFC4AFBD7188248811A82" box="[1060,1176,1159,1183]" italics="true" pageId="1" pageNumber="553">G. m i n u t u s</emphasis>
|
||
in reaching to a much larger size (to at least ca.
|
||
<quantity id="4CD6ADE5FFBBFC4AFE3D18AF4E381ADF" box="[462,545,1194,1218]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="553" unit="mm" value="70.0">70 mm</quantity>
|
||
SL vs. ca.
|
||
<quantity id="4CD6ADE5FFBBFC4AFD5D18AF4F1B1ADF" box="[686,770,1194,1218]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="553" unit="mm" value="27.0">27 mm</quantity>
|
||
SL) and an almost uniform body (vs. with dark saddles overlying a pale body), and from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFDEF18CA4E871AFA" box="[540,670,1231,1255]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBBFC4AFDEF18CA4E871AFA" box="[540,670,1231,1255]" italics="true" pageId="1" pageNumber="553">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
in having a longer nasal barbel (30.1–35.5% HL vs. 15.0–28.7), a wider head (20.3–21.8% SL vs. 13.2–19.7) and a deeper caudal peduncle (8.2–9.7% SL vs. 5.3–6.9).
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFAF518F64CE71B33" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBBFC4AFAF518F64CE71B33" italics="true" pageId="1" pageNumber="553">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFE0219124E411B33" box="[497,600,1302,1326]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="panda">
|
||
<emphasis id="B95ADC12FFBBFC4AFE0219124E411B33" box="[497,600,1302,1326]" italics="true" pageId="1" pageNumber="553">G. panda</emphasis>
|
||
</taxonomicName>
|
||
in having a larger eye (diameter 10.4–12.7% HL vs. 15.7–18.4), longer dorsal-fin spine (16.9–21.6% SL vs. 10.3–15.3), a deeper body (depth at anus 15.2–16.4% SL vs. 10.4–11.8) and a deeper caudal peduncle (8.2–9.7% SL vs. 5.3–6.2), from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFCC0195A4FC51B6A" box="[819,988,1375,1399]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBBFC4AFCC0195A4FC51B6A" box="[819,988,1375,1399]" italics="true" pageId="1" pageNumber="553">G. rugimentum</emphasis>
|
||
</taxonomicName>
|
||
in lacking (vs. with) the ridges of the thoracic adhesive apparatus extending onto the gular region, having a shorter head (25.3–28.2% SL vs. 28.4–31.2) and predorsal distance (35.6–37.4% SL vs. 39.9–42.8), a longer postadipose distance (18.8–20.8% SL vs. 14.4– 17.6) and a deeper caudal peduncle (8.2–9.7% SL vs. 6.1–7.6), and from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFC0A19CE48A21BFE" box="[1017,1211,1483,1507]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="ventrolineatus">
|
||
<emphasis id="B95ADC12FFBBFC4AFC0A19CE48A21BFE" box="[1017,1211,1483,1507]" italics="true" pageId="1" pageNumber="553">G. ventrolineatus</emphasis>
|
||
</taxonomicName>
|
||
in having a longer adipose-fin base (13.9–15.8% SL vs. 9.3–11.1).
|
||
</paragraph>
|
||
<paragraph id="8B910000FFBBFC49FF341A174D171F22" blockId="1.[151,1436,510,2011]" lastBlockId="2.[151,1436,151,319]" lastPageId="2" lastPageNumber="554" pageId="1" pageNumber="553">
|
||
Seven other species of
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFE201A164E711836" box="[467,616,1555,1579]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95ADC12FFBBFC4AFE201A164E711836" box="[467,616,1555,1579]" italics="true" pageId="1" pageNumber="553">Glyptothorax</emphasis>
|
||
</taxonomicName>
|
||
from the Indian subcontinent are known to have the skin ridges of the thoracic adhesive apparatus almost or wholly enclosing the central depression posteriorly, viz.,
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFB221A3249341852" box="[1233,1325,1591,1615]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="cavia">
|
||
<emphasis id="B95ADC12FFBBFC4AFB221A3249341852" box="[1233,1325,1591,1615]" italics="true" pageId="1" pageNumber="553">G. cavia</emphasis>
|
||
</taxonomicName>
|
||
(Ganges- Brahmaputra river system),
|
||
<emphasis id="B95ADC12FFBBFC4AFE221A5E4E49186E" box="[465,592,1627,1651]" italics="true" pageId="1" pageNumber="553">G. g a rh w a l i</emphasis>
|
||
(Ganges River drainage),
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFC8F1A5E481C186E" box="[892,1029,1627,1651]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="jalalensis">
|
||
<emphasis id="B95ADC12FFBBFC4AFC8F1A5E481C186E" box="[892,1029,1627,1651]" italics="true" pageId="1" pageNumber="553">G. jalalensis</emphasis>
|
||
</taxonomicName>
|
||
(Indus River drainage),
|
||
<emphasis id="B95ADC12FFBBFC4AFAEB1A5E4981186E" box="[1304,1432,1627,1651]" italics="true" pageId="1" pageNumber="553">G. j a y a r a m i</emphasis>
|
||
(Kolodyne River drainage),
|
||
<emphasis id="B95ADC12FFBBFC4AFE261A7A4E9B188A" box="[469,642,1662,1687]" italics="true" pageId="1" pageNumber="553">G. k a s h m i re n s i s</emphasis>
|
||
(Indus River drainage),
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFC6E1A7A4820188A" box="[925,1081,1663,1687]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="maceriatus">
|
||
<emphasis id="B95ADC12FFBBFC4AFC6E1A7A4820188A" box="[925,1081,1663,1687]" italics="true" pageId="1" pageNumber="553">G. maceriatus</emphasis>
|
||
</taxonomicName>
|
||
(Surma-Meghna river system), and
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFF341AA64D3118A6" box="[199,296,1699,1723]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="stocki">
|
||
<emphasis id="B95ADC12FFBBFC4AFF341AA64D3118A6" box="[199,296,1699,1723]" italics="true" pageId="1" pageNumber="553">G. stocki</emphasis>
|
||
</taxonomicName>
|
||
(Indus River drainage).
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFDCD1AA64F5A18A6" box="[574,835,1699,1723]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBBFC4AFDCD1AA64F5A18A6" box="[574,835,1699,1723]" italics="true" pageId="1" pageNumber="553">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFC2A1AA6482F18A6" box="[985,1078,1699,1723]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="cavia">
|
||
<emphasis id="B95ADC12FFBBFC4AFC2A1AA6482F18A6" box="[985,1078,1699,1723]" italics="true" pageId="1" pageNumber="553">G. cavia</emphasis>
|
||
</taxonomicName>
|
||
in having the depressed central in the thoracic adhesive apparatus lanceolate (vs. ovoid) and with a single, non-diverging series (vs. double, diverging series) of striae running along its edges, a deeper caudal peduncle (8.2–9.7% SL vs. 4.8–6.4), from
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFA771AEE4CE5193A" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="garhwali">
|
||
<emphasis id="B95ADC12FFBBFC4AFA771AEE4CE5193A" italics="true" pageId="1" pageNumber="553">G. garhwali</emphasis>
|
||
</taxonomicName>
|
||
in having a longer anal-fin base (14.1–17.8% SL vs. 12.7–12.8), and postadipose length (18.8–20.8% SL vs. 16.4–18.0), and from G.
|
||
<taxonomicName id="4C2E7B83FFBBFC4AFE2A1B364E5C1956" box="[473,581,1843,1867]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="jalalensis">
|
||
<emphasis id="B95ADC12FFBBFC4AFE2A1B364E5C1956" box="[473,581,1843,1867]" italics="true" pageId="1" pageNumber="553">jalalensis</emphasis>
|
||
</taxonomicName>
|
||
in having a shorter adipose-fin base (13.9–15.8% SL vs. 19.1) and a more slender caudal peduncle (8.2–9.7% SL vs. 10.9). It is further distinguished from
|
||
<emphasis id="B95ADC12FFBBFC4AFBC61B5248A11972" box="[1077,1208,1879,1903]" italics="true" pageId="1" pageNumber="553">G. j a y a r a m i</emphasis>
|
||
in having a shorter pectoral fin (21.2–26.1% SL vs. 26.3–27.8), lacking (vs. with) plicae on the ventral surfaces of the first pectoral- and pelvic-fin elements, having a deeper body (depth at anus 15.2–16.4% SL vs 13.1–15.0) and caudal peduncle (8.2–9.7% SL vs. 5.5–7.9), from
|
||
<emphasis id="B95ADC12FFBBFC4AFDE51BC64EDC19C6" box="[534,709,1987,2011]" italics="true" pageId="1" pageNumber="553">G. k a s h m i re n s i s</emphasis>
|
||
in having a single, non-diverging series (vs. double, diverging series) of striae running along the edges of the central depression in the thoracic adhesive apparatus, a more slender body (depth at anus 15.2–16.4% SL vs. 17.1–17.7), from
|
||
<taxonomicName id="4C2E7B83FFB8FC49FCC11CB84FD61EC9" box="[818,975,189,212]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="554" phylum="Chordata" rank="species" species="maceriatus">
|
||
<emphasis id="B95ADC12FFB8FC49FCC11CB84FD61EC9" box="[818,975,189,212]" italics="true" pageId="2" pageNumber="554">G. maceriatus</emphasis>
|
||
</taxonomicName>
|
||
in having a deeper body (depth at anus 15.2–16.4% SL vs. 11.3–13.8) and caudal peduncle (8.2–9.7% SL vs. 6.4–7.6), and from
|
||
<emphasis id="B95ADC12FFB8FC49FB621CE548F71EE5" box="[1169,1262,224,248]" italics="true" pageId="2" pageNumber="554">G. s t o c k i</emphasis>
|
||
in lacking (vs. having) both a distinct pale midlateral stripe on the body and distinct dark submarginal stripe along each lobe of the caudal fin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF515088FFB8FC49FF641B8C48CF1982" box="[151,1238,1929,1952]" httpUri="https://zenodo.org/record/222070/files/figure.png" pageId="2" pageNumber="554" targetBox="[151,1436,375,1905]" targetPageId="2">
|
||
<paragraph id="8B910000FFB8FC49FF641B8C48CF1982" blockId="2.[151,1238,1929,1952]" box="[151,1238,1929,1952]" pageId="2" pageNumber="554">
|
||
<emphasis id="B95ADC12FFB8FC49FF641B8C4D0A1982" bold="true" box="[151,275,1929,1952]" pageId="2" pageNumber="554">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C2E7B83FFB8FC49FEEA1B8C4E1A1982" box="[281,515,1929,1951]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="554" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFB8FC49FEEA1B8C4E1A1982" box="[281,515,1929,1951]" italics="true" pageId="2" pageNumber="554">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
, NRM 64520, holotype, 65.5 mm SL; dorsal, lateral and ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C334538BFFB9FC4FFF341C9248041C99" lastPageId="4" lastPageNumber="556" pageId="3" pageNumber="555" type="description">
|
||
<paragraph id="8B910000FFB9FC48FF341C9248391F79" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="555">
|
||
<emphasis id="B95ADC12FFB9FC48FF341C924D401EAD" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="555">Description.</emphasis>
|
||
Morphometric data as in Table 1. Head depressed, body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile flat to anal-fin base, sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Caudal peduncle depth 2.2–2.8 times in its length. Skin almost smooth, with minute, rounded tubercles on sides of body. Lateral line complete, mid-lateral. Vertebrae 18+17=35 (1), 17+19=36 (1) or 18+18=36* (3).
|
||
</paragraph>
|
||
<paragraph id="8B910000FFB9FC48FF341D6A4ECD1FE9" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="555">Head depressed, broad; triangular in lateral view. Snout prominent. Anterior and posterior nares large, separated only by base of nasal barbel. Gill openings broad, extending from immediately ventral to post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, its horizontal axis longest; located entirely in dorsal half of head.</paragraph>
|
||
<paragraph id="8B910000FFB9FC48FF341DFA48701C99" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="555">Barbels in four pairs. Maxillary barbel long slender; extending just beyond base of last pectoral-fin ray. Nasal barbel slender, extending to just beyond anterior orbital margin. Inner mandibular-barbel origin close to midline, extending to two-thirds distance between its base and that of pectoral spine. Outer mandibular barbel originating posterolateral of inner mandibular barbel, extending just beyond base of pectoral spine.</paragraph>
|
||
<paragraph id="8B910000FFB9FC48FF341E8A4EC11CF2" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="555">Mouth inferior, premaxillary tooth band not exposed with mouth closed. Oral teeth small, villiform; in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in a single broad semilunate band. Dentary teeth in narrow crescentic band that does not reach midline.</paragraph>
|
||
<paragraph id="8B910000FFB9FC48FF341EF94D271D62" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="555">
|
||
Thoracic adhesive apparatus present, consisting of ridges of skin (striae) in elliptical field extending from isthmus to level of last pectoral-fin ray base, with lanceolate median depression on posterior half almost enclosed posteriorly by skin ridges (
|
||
<figureCitation id="13151C85FFB9FC48FE1D1F414E271D41" box="[494,574,836,860]" captionStart="FIGURE 2" captionStartId="3.[151,250,1729,1752]" captionTargetBox="[499,1088,949,1708]" captionTargetId="figure@3.[499,1088,949,1708]" captionTargetPageId="3" captionText="FIGURE 2. Thoracic adhesive apparatus of Glyptothorax igniculus, NRM 64520, holotype, 65.5 mm SL." httpUri="https://zenodo.org/record/222071/files/figure.png" pageId="3" pageNumber="555">Fig. 2</figureCitation>
|
||
). Striae uninterrupted; medial striae orientated anteriorly, lateral ones anterolaterally.
|
||
</paragraph>
|
||
<caption id="DF515088FFB9FC48FF641AC448DE18CA" box="[151,1223,1729,1752]" httpUri="https://zenodo.org/record/222071/files/figure.png" pageId="3" pageNumber="555" targetBox="[499,1088,949,1708]" targetPageId="3">
|
||
<paragraph id="8B910000FFB9FC48FF641AC448DE18CA" blockId="3.[151,1223,1729,1752]" box="[151,1223,1729,1752]" pageId="3" pageNumber="555">
|
||
<emphasis id="B95ADC12FFB9FC48FF641AC44D0A18CA" bold="true" box="[151,275,1729,1752]" pageId="3" pageNumber="555">FIGURE 2.</emphasis>
|
||
Thoracic adhesive apparatus of
|
||
<taxonomicName id="4C2E7B83FFB9FC48FDA81AC44F5C18CA" box="[603,837,1729,1751]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFB9FC48FDA81AC44F5C18CA" box="[603,837,1729,1751]" italics="true" pageId="3" pageNumber="555">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
, NRM 64520, holotype, 65.5 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B910000FFB9FC4FFF341AFA4DBE1F00" blockId="3.[151,1436,1791,2032]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="556" pageId="3" pageNumber="555">Pectoral fin with i,8,i (1), i,9* (4) or i,9,i (1) rays; posterior fin margin slightly concave. Pectoral spine very broad and covered with thick skin. Anterior spine margin smooth, its posterior margin with 9–12 serrae. Ventral surface of spine smooth and without plicae. Dorsal fin above anterior third of body, with i,6 (6) rays; fin margin convex; spine broad, straight; with 0–8 serrae on posterior margin. Adipose fin with anterior margin straight or very slightly concave, posterior margin angular. Procurrent rays symmetrical, extending only slightly anterior to fin base. Vertical through anal-fin origin posterior to that through adipose-fin origin. Pelvic fin with slightly convex margin and i,5 (6) rays; tip of adpressed fin not reaching anal-fin origin. Ventral surface of first pelvic-fin ray smooth and without plicae. Vertical through pelvic-fin origin posterior to that through posterior end of dorsal-fin base. Anal fin with straight anterior margin, straight or slightly concave posterior margin; with iii,8,i (2), iv,8 (1), iv,8,i (2) or iv,9* (1) rays. Caudal fin strongly forked, with lower lobe very slightly longer than upper lobe and i,7,8,i (6) principal rays.</paragraph>
|
||
<paragraph id="8B910000FFBEFC4FFF341D22488E1FB1" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="556">
|
||
<emphasis id="B95ADC12FFBEFC4FFF341D224D561F5D" bold="true" box="[199,335,295,320]" pageId="4" pageNumber="556">Coloration.</emphasis>
|
||
In 70% alcohol: dorsal and lateral surfaces of head and body brown fading to cream on ventral surfaces. Faint pale mid-dorsal spot at base of doorsal-fin spine. Dorsal, pectoral, pelvic, anal and caudal fins also brown, particularly on bases and rays. Adipose fin dark brown basally, fading to lighter brown along distal margin. Nasal and maxillary barbels brown dorsally, cream ventrally; all mandibular barbels cream.
|
||
</paragraph>
|
||
<paragraph id="8B910000FFBEFC4FFF341DB248041C99" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="556">
|
||
<emphasis id="B95ADC12FFBEFC4FFF341DB24D331FCD" bold="true" box="[199,298,439,464]" pageId="4" pageNumber="556">Habitat.</emphasis>
|
||
The
|
||
<typeStatus id="5495BEA2FFBEFC4FFE991DBD4D831FCD" box="[362,410,440,464]" pageId="4" pageNumber="556">type</typeStatus>
|
||
locality of
|
||
<emphasis id="B95ADC12FFBEFC4FFDD61DBD4EB31FCD" box="[549,682,440,464]" italics="true" pageId="4" pageNumber="556">G. i g n i c u l u s</emphasis>
|
||
is a wide (ca.
|
||
<quantity id="4CD6ADE5FFBEFC4FFC921DB24FB61FD2" box="[865,943,439,463]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="556" unit="m" value="200.0">200 m</quantity>
|
||
), turbid river with moderate current and a substrate comprised of clay. Associated fishes include
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFD0B1DD948A31FE9" box="[760,1210,476,500]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBEFC4FFD0B1DD948381FE9" box="[760,1057,476,500]" italics="true" pageId="4" pageNumber="556">Crossocheilus burmanicus</emphasis>
|
||
(Cyprinidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFB371DD84D301C0A" class="Actinopterygii" family="Cyprinidae" genus="Danio" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="albolineatus">
|
||
<emphasis id="B95ADC12FFBEFC4FFB371DD849851FE9" box="[1220,1436,476,500]" italics="true" pageId="4" pageNumber="556">
|
||
<collectingCountry id="F3394090FFBEFC4FFB371DD849131FE9" box="[1220,1290,477,500]" name="Denmark" pageId="4" pageNumber="556">Danio</collectingCountry>
|
||
albolineatus
|
||
</emphasis>
|
||
(Cyprinidae)
|
||
</taxonomicName>
|
||
, ‘
|
||
<emphasis id="B95ADC12FFBEFC4FFEB01E054E371C05" box="[323,558,512,536]" italics="true" pageId="4" pageNumber="556">Pethia’ meingangbii</emphasis>
|
||
(
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFDB01DFA4EDF1C0A" box="[579,710,511,535]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFD2C1E0448061C0A" box="[735,1055,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="chola">
|
||
<emphasis id="B95ADC12FFBEFC4FFD2C1E044F661C05" box="[735,895,512,536]" italics="true" pageId="4" pageNumber="556">Puntius chola</emphasis>
|
||
(Cyprinidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFBDC1E0449811C0A" box="[1071,1432,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Raiamas" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="guttatus">
|
||
<emphasis id="B95ADC12FFBEFC4FFBDC1E0448E11C05" box="[1071,1272,513,536]" italics="true" pageId="4" pageNumber="556">Raiamas guttatus</emphasis>
|
||
(Cyprinidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFF641E204DEA1C21" box="[151,499,548,572]" class="Actinopterygii" family="Bagridae" genus="Mystus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="falcarius">
|
||
<emphasis id="B95ADC12FFBEFC4FFF641E204D791C21" box="[151,352,548,572]" italics="true" pageId="4" pageNumber="556">Mystus falcarius</emphasis>
|
||
(Bagridae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFDFD1E204FAD1C21" box="[526,948,548,572]" class="Actinopterygii" family="Heteropneustidae" genus="Heteropneustes" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fossilis">
|
||
<emphasis id="B95ADC12FFBEFC4FFDFD1E204F381C21" box="[526,801,548,572]" italics="true" pageId="4" pageNumber="556">Heteropneustes fossilis</emphasis>
|
||
(Clariidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFC3C1E2149811C21" box="[975,1432,548,572]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBEFC4FFC3C1E21491A1C21" box="[975,1283,548,572]" italics="true" pageId="4" pageNumber="556">Glyptothorax rugimentum</emphasis>
|
||
(Sisoridae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFF641E4D4E1E1C42" box="[151,519,583,608]" class="Actinopterygii" family="Belonidae" genus="Xenentodon" kingdom="Animalia" order="Beloniformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="cancila">
|
||
<emphasis id="B95ADC12FFBEFC4FFF641E4D4D6E1C7D" box="[151,375,584,608]" italics="true" pageId="4" pageNumber="556">Xenentodon cancila</emphasis>
|
||
(Belonidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFDE71E4D4F2A1C42" box="[532,819,583,608]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="ferrarisi">
|
||
<emphasis id="B95ADC12FFBEFC4FFDE71E4D4EA01C7D" box="[532,697,584,608]" italics="true" pageId="4" pageNumber="556">Badis ferrarisi</emphasis>
|
||
(Badidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFCCC1E4D48901C42" box="[831,1161,583,608]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="gachua">
|
||
<emphasis id="B95ADC12FFBEFC4FFCCC1E4D4FEA1C7D" box="[831,1011,584,608]" italics="true" pageId="4" pageNumber="556">Channa gachua</emphasis>
|
||
(Channidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFB661E4D49811C42" box="[1173,1432,583,608]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="panaw">
|
||
<emphasis id="B95ADC12FFBEFC4FFB661E4D491B1C7D" box="[1173,1282,584,608]" italics="true" pageId="4" pageNumber="556">C. panaw</emphasis>
|
||
(Channidae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFF641E694E531C99" box="[151,586,620,644]" class="Actinopterygii" family="Osphronemidae" genus="Trichogaster" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="labiosa">
|
||
<emphasis id="B95ADC12FFBEFC4FFF641E694D991C99" box="[151,384,620,644]" italics="true" pageId="4" pageNumber="556">Trichogaster labiosa</emphasis>
|
||
(Osphronemidae)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFD721E6948011C99" box="[641,1048,620,644]" class="Actinopterygii" family="Tetraodontidae" genus="Tetraodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="cutcutia">
|
||
<emphasis id="B95ADC12FFBEFC4FFD721E694F4D1C99" box="[641,852,620,644]" italics="true" pageId="4" pageNumber="556">Tetraodon cutcutia</emphasis>
|
||
(Tetraodontidae)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334538BFFBEFC4FFF341E954DB91CF2" pageId="4" pageNumber="556" type="etymology">
|
||
<paragraph id="8B910000FFBEFC4FFF341E954DB91CF2" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="556">
|
||
<emphasis id="B95ADC12FFBEFC4FFF341E954D571CB5" bold="true" box="[199,334,656,680]" pageId="4" pageNumber="556">Etymology.</emphasis>
|
||
The specific name comes from the Latin noun
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFC971E8A4FD71CBA" box="[868,974,655,679]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="igniculus">igniculus</taxonomicName>
|
||
, meaning a little flame, in allusion to the lanceolate, flame-shaped central depression in the thoracic adhesive apparatus that is nearly completely enclosed by skin ridges caudally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334538BFFBEFC4CFF341EFE4F041FD2" lastPageId="7" lastPageNumber="559" pageId="4" pageNumber="556" type="distribution">
|
||
<paragraph id="8B910000FFBEFC4FFF341EFE49811D09" blockId="4.[151,1437,151,789]" box="[199,1432,763,789]" pageId="4" pageNumber="556">
|
||
<emphasis id="B95ADC12FFBEFC4FFF341EFE4D791D09" bold="true" box="[199,352,763,788]" pageId="4" pageNumber="556">Distribution.</emphasis>
|
||
Known from the Myittha River, a tributary of the Chindwin River in western
|
||
<collectingCountry id="F3394090FFBEFC4FFB3A1EF8492F1D08" box="[1225,1334,765,789]" name="Myanmar" pageId="4" pageNumber="556">Myanmar</collectingCountry>
|
||
(
|
||
<figureCitation id="13151C85FFBEFC4FFAB51EF949931D09" box="[1350,1418,764,788]" captionStart="FIGURE 3" captionStartId="4.[151,250,1973,1996]" captionTargetBox="[318,1269,845,1952]" captionTargetId="figure@4.[318,1269,841,1952]" captionTargetPageId="4" captionText="FIGURE 3. Map showing collecting localities of Glyptothorax igniculus. Open symbol indicates type locality." httpUri="https://zenodo.org/record/222072/files/figure.png" pageId="4" pageNumber="556">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF515088FFBEFC4FFF641BB048E019D1" box="[151,1273,1973,1996]" httpUri="https://zenodo.org/record/222072/files/figure.png" pageId="4" pageNumber="556" targetBox="[318,1269,845,1952]" targetPageId="4">
|
||
<paragraph id="8B910000FFBEFC4FFF641BB048E019D1" blockId="4.[151,1273,1973,1996]" box="[151,1273,1973,1996]" pageId="4" pageNumber="556">
|
||
<emphasis id="B95ADC12FFBEFC4FFF641BB04D0B19D6" bold="true" box="[151,274,1973,1996]" pageId="4" pageNumber="556">FIGURE 3.</emphasis>
|
||
Map showing collecting localities of
|
||
<taxonomicName id="4C2E7B83FFBEFC4FFD621BB34F6319D1" box="[657,890,1974,1996]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBEFC4FFD621BB34F6319D1" box="[657,890,1974,1996]" italics="true" pageId="4" pageNumber="556">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
. Open symbol indicates type locality.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B910000FFBCFC4DFF341C924F9F1FB1" blockId="6.[151,1436,151,428]" pageId="6" pageNumber="558">
|
||
<emphasis id="B95ADC12FFBCFC4DFF341C924E031EAD" bold="true" box="[199,538,151,176]" pageId="6" pageNumber="558">Comparative morphometry.</emphasis>
|
||
We carried out a morphometric analysis of the four species of
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFAF51C9D49821EAD" box="[1286,1435,152,176]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95ADC12FFBCFC4DFAF51C9D49821EAD" box="[1286,1435,152,176]" italics="true" pageId="6" pageNumber="558">Glyptothorax</emphasis>
|
||
</taxonomicName>
|
||
found sympatrically in the Chindwin (
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFDB31CB84EFB1EC9" box="[576,738,188,212]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBCFC4DFDB31CB84EFB1EC9" box="[576,738,188,212]" italics="true" pageId="6" pageNumber="558">G. burmanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B95ADC12FFBCFC4DFD031CB84F681EC9" box="[752,881,188,212]" italics="true" pageId="6" pageNumber="558">G. i g n i c u l u s</emphasis>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFC711CB8481B1EC9" box="[898,1026,189,212]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBCFC4DFC711CB8481B1EC9" box="[898,1026,189,212]" italics="true" pageId="6" pageNumber="558">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFBCB1CB848C41EC9" box="[1080,1245,189,212]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBCFC4DFBCB1CB848C41EC9" box="[1080,1245,189,212]" italics="true" pageId="6" pageNumber="558">G. rugimentum</emphasis>
|
||
</taxonomicName>
|
||
). In proportional measurements (Table 1),
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFE4B1CE54EA41EE5" box="[440,701,224,248]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBCFC4DFE4B1CE54EA41EE5" box="[440,701,224,248]" italics="true" pageId="6" pageNumber="558">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFCA41CE54FE51EE5" box="[855,1020,224,248]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBCFC4DFCA41CE54FE51EE5" box="[855,1020,224,248]" italics="true" pageId="6" pageNumber="558">G. burmanicus</emphasis>
|
||
</taxonomicName>
|
||
in shorter prepelvic and prepectoral lengths; longer dorsal spine and larger eye; from
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFD231D004F4A1F01" box="[720,851,261,284]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBCFC4DFD231D004F4A1F01" box="[720,851,261,284]" italics="true" pageId="6" pageNumber="558">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
in narrower head and longer nasal barbel; from
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFA771D004D061F5D" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBCFC4DFA771D004D061F5D" italics="true" pageId="6" pageNumber="558">G. rugimentum</emphasis>
|
||
</taxonomicName>
|
||
in shorter predorsal and head lengths, and longer postadipose length; and from
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFB3F1D2D496B1F5D" box="[1228,1394,296,320]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBCFC4DFB3F1D2D496B1F5D" box="[1228,1394,296,320]" italics="true" pageId="6" pageNumber="558">G. burmanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFA771D2D4CE01F79" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBCFC4DFA771D2D4CE01F79" italics="true" pageId="6" pageNumber="558">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFEC41D484DC71F79" box="[311,478,333,356]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBCFC4DFEC41D484DC71F79" box="[311,478,333,356]" italics="true" pageId="6" pageNumber="558">G. rugimentum</emphasis>
|
||
</taxonomicName>
|
||
in deeper caudal peduncle. The SL distribution of the
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFBB71D4848D31F79" box="[1092,1226,332,356]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBCFC4DFBB71D4848D31F79" box="[1092,1226,332,356]" italics="true" pageId="6" pageNumber="558">G. igniculus</emphasis>
|
||
</taxonomicName>
|
||
is too restricted to obtain a strong linear correlation for the caudal peduncle depth, but it is clearly the most important measurement separating from all three species compared with (Table 1, Fig. 4A).
|
||
</paragraph>
|
||
<paragraph id="8B910000FFBCFC4DFF641B6548D419CA" blockId="6.[151,1436,1888,2007]" pageId="6" pageNumber="558">
|
||
<emphasis id="B95ADC12FFBCFC4DFF641B654D0F196B" bold="true" box="[151,278,1888,1911]" pageId="6" pageNumber="558">FIGURE 4.</emphasis>
|
||
Morphometric analysis of
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFDC61B654F3A196B" box="[565,803,1888,1910]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBCFC4DFDC61B654F3A196B" box="[565,803,1888,1910]" italics="true" pageId="6" pageNumber="558">Glyptothorax igniculus</emphasis>
|
||
</taxonomicName>
|
||
and sympatric taxa. A Biplot of caudal peduncle depth (
|
||
<collectingCountry id="F3394090FFBCFC4DFA731B64498D196A" box="[1408,1428,1889,1911]" name="Italy" pageId="6" pageNumber="558">Y</collectingCountry>
|
||
) against Standard length (X) with linear regressions (
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFD571B864F051985" box="[676,796,1922,1944]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="igniculus">
|
||
<emphasis id="B95ADC12FFBCFC4DFD571B864F051985" box="[676,796,1922,1944]" italics="true" pageId="6" pageNumber="558">G. igniculus</emphasis>
|
||
</taxonomicName>
|
||
, 1.14+0.07*SL, R2 = 0.434;
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFBC91B8648D51985" box="[1082,1228,1922,1944]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBCFC4DFBC91B8648D51985" box="[1082,1228,1922,1944]" italics="true" pageId="6" pageNumber="558">G. burmanicus</emphasis>
|
||
</taxonomicName>
|
||
, 1.09+0.09*SL, R2= 0.980;
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFF111BA64D4519A4" box="[226,348,1955,1977]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBCFC4DFF111BA64D4519A4" box="[226,348,1955,1977]" italics="true" pageId="6" pageNumber="558">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
, 0.3+0.07*SL, R2 = 0-985;
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFD7D1BA64F3019A4" box="[654,809,1955,1977]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="rugimentum">
|
||
<emphasis id="B95ADC12FFBCFC4DFD7D1BA64F3019A4" box="[654,809,1955,1977]" italics="true" pageId="6" pageNumber="558">G. rugimentum</emphasis>
|
||
</taxonomicName>
|
||
, 0.41+0.07*SL, R2=0.816); two largest
|
||
<taxonomicName id="4C2E7B83FFBCFC4DFB2B1BA6496919A4" box="[1240,1392,1955,1977]" class="Actinopterygii" family="Cyprinidae" genus="Crossocheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="B95ADC12FFBCFC4DFB2B1BA6496919A4" box="[1240,1392,1955,1977]" italics="true" pageId="6" pageNumber="558">G. burmanicus</emphasis>
|
||
</taxonomicName>
|
||
not included to improve visual clarity in overlapping size range. B–D Plots of scores of principal components.
|
||
</paragraph>
|
||
<paragraph id="8B910000FFBDFC4CFF341C9D4F041FD2" blockId="7.[151,1436,151,463]" pageId="7" pageNumber="559">
|
||
In the PCA (Fig. 4B–D,
|
||
<tableCitation id="C6AC35BBFFBDFC4CFE021C924E521EB2" box="[497,587,151,175]" captionStart="TABLE 2" captionStartId="7.[151,239,511,535]" captionTargetBox="[159,1387,626,1836]" captionTargetPageId="7" captionText="TABLE 2. Variable loadings on principal components I-IV and sheared components II-IV from pooled measurement set of Glyptothorax burmanicus (N = 15), G. igniculus (N = 5), G. ngapang (N = 18), and G. rugimentum (N = 15). Values in boldface highlight the two highest loadings of each component." httpUri="http://table.plazi.org/id/DF515088FFBDFC4CFF641DFA4F241C4A" pageId="7" pageNumber="559" tableUuid="DF515088FFBDFC4CFF641DFA4F241C4A">Table 2</tableCitation>
|
||
), loadings on component I are of roughly the same magnitude, and the variance only 88%, reflecting the similar size of specimens except two large
|
||
<emphasis id="B95ADC12FFBDFC4CFBAF1CB8491A1EC9" box="[1116,1283,188,212]" italics="true" pageId="7" pageNumber="559">G. b u r m a n i c u s</emphasis>
|
||
. Also other components seem to reflect lack of variation, as seen in the only slight accumulation of variance and low eigenvalues. Component II separates
|
||
<taxonomicName id="4C2E7B83FFBDFC4CFDB51D004ED31F01" box="[582,714,261,284]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="559" phylum="Chordata" rank="species" species="ngapang">
|
||
<emphasis id="B95ADC12FFBDFC4CFDB51D004ED31F01" box="[582,714,261,284]" italics="true" pageId="7" pageNumber="559">G. ngapang</emphasis>
|
||
</taxonomicName>
|
||
from the rest (Fig. 4B–C), and is dominated by the nasal and outer mandibular barbels. The third component is dominated by the dorsal spine length, with the nasal barbel and orbital diameter also contributing to the variance. The fourth component is dominated by caudal peduncle depth and orbital diameter. The most distinct clustering is obtained in contrasting Components II and III (Fig. 4C), and the most distinct cluster of
|
||
<emphasis id="B95ADC12FFBDFC4CFE231D904E4A1FB1" box="[464,595,404,428]" italics="true" pageId="7" pageNumber="559">G. i g n i c u l u s</emphasis>
|
||
from the rest is obtained by contrasting Components III and IV (Fig. 5), reflecting the orbital diameter and caudal peduncle depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |