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<document id="FA5BDF051689B2BA86885943BDB924E8" ID-DOI="10.3897/zookeys.1201.120750" ID-ZooBank="23C7E6E7-AE73-4685-AEDA-26DEB0EE0068" ID-publisher-id="120750" URI-arpha="9C1BFC80-2C0B-5936-AEE4-69A1A8247734" URI-zoobank="http://zoobank.org/23C7E6E7-AE73-4685-AEDA-26DEB0EE0068" XM.bibliography_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1715703108403" checkinUser="pensoft" docAuthor="Marques, Mariana P., Parrinha, Diogo, Lopes-Lima, Manuel, Tiutenko, Arthur, Bauer, Aaron M. &amp; Ceríaco, Luis M. P." docDate="2024" docId="FFA1275CDB5553E2BAA430BD690A8FF6" docLanguage="en" docName="ZooKeys 1201: 167-217" docOrigin="ZooKeys 1201" docSource="https://zookeys.pensoft.net/article/120750/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Lycophidion hellmichi Laurent 1964" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="217" masterDocId="9C1BFC802C0B5936AEE469A1A8247734" masterDocTitle="An island in a sea of sand: a first checklist of the herpetofauna of the Serra da Neve inselberg, southwestern Angola" masterLastPageNumber="217" masterPageNumber="167" pageNumber="167" updateTime="1715763917274" updateUser="admin">
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<mods:title id="C487A8A0B1731C0FD214C118CA0A14AC">An island in a sea of sand: a first checklist of the herpetofauna of the Serra da Neve inselberg, southwestern Angola</mods:title>
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<mods:name id="2DB3C4B36370BAF8DC3BCFBDE13915F7" type="personal">
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<mods:namePart id="4E0EF12FA7C664CEE233D7E20ED17299">Marques, Mariana P.</mods:namePart>
<mods:nameIdentifier id="4B9231E60B4AE2125923E193B87E7110" type="ORCID">0000-0002-1712-2632</mods:nameIdentifier>
<mods:affiliation id="575776FEEF9903B53B0FB36280D5B62E">Carnegie Museum of Natural History, 4400 Forbes Avenue, Pittsburgh PA 15213, USA &amp; CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal &amp; Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal</mods:affiliation>
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<mods:namePart id="577FE70E7AA01996FC38C46A50C1395A">Parrinha, Diogo</mods:namePart>
<mods:nameIdentifier id="DEF1133D321C416692FA649660A911EF" type="ORCID">0000-0002-1302-025X</mods:nameIdentifier>
<mods:affiliation id="BC4241C338AC4D9D381CF02AF472A28B">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal &amp; Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal</mods:affiliation>
</mods:name>
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<mods:role id="63760CD93C9FA5A404EA3061FCA84968">
<mods:roleTerm id="DD723FBC5D86B2BA11A905CB6D3F9F02">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DAA4000071372622A489F8279FA4CD87">Lopes-Lima, Manuel</mods:namePart>
<mods:nameIdentifier id="F57A3E23D8B4BB9FE6AE2460BEA3BFA1" type="ORCID">0000-0002-2761-7962</mods:nameIdentifier>
<mods:affiliation id="F6920850AB3FCF0820F7B1642769B5A1">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal</mods:affiliation>
</mods:name>
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<mods:namePart id="5E5E59E5ECB6DBFC4676FA9C199E7867">Tiutenko, Arthur</mods:namePart>
<mods:nameIdentifier id="7A7FD4D90817BE3B3AE5D2354E23482D" type="ORCID">0000-0001-5952-4923</mods:nameIdentifier>
<mods:affiliation id="778B347354C3E3F77CD7274FD9CCD1D7">Friedrich-Alexander-Universität Erlangen-Nürnberg, Schlossplatz 4, 91054 Erlangen, Germany</mods:affiliation>
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<mods:roleTerm id="CCC2AD989C082DA027522F6FE7DA194A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BEB302A698A518ED0ACBB609E94C6F76">Bauer, Aaron M.</mods:namePart>
<mods:nameIdentifier id="539A9C7550757EBF4BEAD23FEF8BC984" type="ORCID">https: // orcid. org / 0000 - 0001 - 6839 - 8025</mods:nameIdentifier>
<mods:affiliation id="D745E99C810CFAA0396E88451CCA7930">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, PA 19085, USA</mods:affiliation>
</mods:name>
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<mods:namePart id="FF1BEF88DCADB45447C998CEE70794C5">Ceríaco, Luis M. P.</mods:namePart>
<mods:nameIdentifier id="41C0EF16B2DD4C64983D6036D1FA859C" type="ORCID">https: // orcid. org / 0000 - 0002 - 0591 - 9978</mods:nameIdentifier>
<mods:affiliation id="9E4028B6EBBEBE33C9DE1B6F914D35DE">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661, Vairão, Portugal &amp; Universidade Federal do Rio de Janeiro, Museu Nacional, Departamento de Vertebrados, Quinta da Boa Vista, São Cristóvão, 20940 - 040 Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:typeOfResource id="2502CF7057DC84F54682065C6B6BCC3F">text</mods:typeOfResource>
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<mods:title id="978BEB19AA327DBCE312F3BDFF4018EA">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="5F85553C0D48D78209A1297530F4C494">
<mods:date id="9E9BDBA2EA7DABDC9CA08158D8B34D54">2024</mods:date>
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<mods:number id="50D0A0D9FC1A57E6F76CA1CA18D1B459">2024-05-14</mods:number>
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<mods:detail id="2CCEA284065C854B3B128A08AF5225DF" type="volume">
<mods:number id="3B2E0CC63CC417B9948A584764EAF96C">1201</mods:number>
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<mods:start id="9C131B5DE6FC1D0E9763BD6FFE9378E7">167</mods:start>
<mods:end id="9EFDE844813DE752CA018B1ED424D1DB">217</mods:end>
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<mods:classification id="E645682428C7B9EC5E73246D46E77723">journal article</mods:classification>
<mods:identifier id="ED054418416E49101A2E280CEB405EC3" type="DOI">10.3897/zookeys.1201.120750</mods:identifier>
<mods:identifier id="57CD8D13A45F3A08392BF98DD70196BA" type="ZooBank">23C7E6E7-AE73-4685-AEDA-26DEB0EE0068</mods:identifier>
</mods:mods>
<treatment id="FFA1275CDB5553E2BAA430BD690A8FF6" ID-arpha="FFA1275C-DB55-53E2-BAA4-30BD690A8FF6" LSID="urn:lsid:plazi:treatment:FFA1275CDB5553E2BAA430BD690A8FF6" httpUri="http://treatment.plazi.org/id/FFA1275CDB5553E2BAA430BD690A8FF6">
<subSubSection id="4EBEDBDB87A09607159F9FE239A50A8E" type="nomenclature">
<paragraph id="9EF69984D3E9451174E1CCC51D1E0FA9">
<taxonomicName id="88CE96746EAF1A2410E95628E19EEF03" ID-CoL="6QPJY" authority="Laurent, 1964" authorityName="Laurent" authorityYear="1964" class="Reptilia" family="Lamprophiidae" genus="Lycophidion" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="hellmichi">
<emphasis id="47EB4B4319BC37B2EE98E4E8A573AF37" italics="true">Lycophidion hellmichi</emphasis>
<bibRefCitation id="58B3B2E3F360C90C8A7DACF4D53FBAA6" author="Laurent RF" journalOrPublisher="Publicações Culturais da Companhia de Diamantes de Angola" pagination="1-165" refId="B81" refString="Laurent RF (1964) Reptiles et amphibiens de l'Angola (Troisieme contribution). Publicacoes Culturais da Companhia de Diamantes de Angola 67: 1 - 165." title="Reptiles et amphibiens de l'Angola (Troisième contribution)." volume="67" year="1964">Laurent, 1964</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="AA2BD3FA900752FB5D52E1000BA177BA" type="description">
<paragraph id="E691AA194F4BAE1CE74655CFF749BDBB">
<figureCitation id="F137BEDD59110E1718CFB4E1E8B5782A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. a Python natalensis (skin) b Bitis (Macrocerastes) gabonica c Causus nasalis d Boaedon variegatus e Hemirhagerrhis viperina f Lycophidion hellmichi g Psammophis subtaeniatus. Photographs by LMPC (a, c, f); AT (d, e, g) and DP (b)." figureDoi="10.3897/zookeys.1201.120750.figure10" httpUri="https://binary.pensoft.net/fig/1048979">Fig. 10 f</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E46DK" type="Records">
<paragraph id="5BDE842E7149F8FAD0921D37927595F5">
<heading id="B1E25BD1CACDFB609D3DBE83358D8B27" reason="title">Records.</heading>
</paragraph>
<paragraph id="263337C626893BF21CFAD59F11D65D4D">
Mauué riparian area [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGAEK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[13.123000,-13.800600]}">
<geoCoordinate id="DE936C2C5CF442210AD27556005FFEEA" degrees="13.8006" direction="south" orientation="latitude" precision="5" value="-13.8006">- 13.8006</geoCoordinate>
,
<geoCoordinate id="6CA642DDE59468B0C752D4BB000A88B7" degrees="13.1230" direction="east" orientation="longitude" precision="5" value="13.123">13.1230</geoCoordinate>
</named-content>
,
<quantity id="773609661B6F2602BB02DA500134AB79" metricMagnitude="2" metricUnit="m" metricValue="7.06" unit="m" value="706.0">706 m</quantity>
] (
<named-content id="9C29BD0BE99AD48B2559AAABB6FF5059" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">
<collectionCode id="0F2430838FEBB2E7B569BE2716A43494" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</named-content>
263033); Catchi surroundings [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESAEK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[13.256900,-13.762000]}">
<geoCoordinate id="98070C633ED8F312B1F400E7E2B33537" degrees="13.7620" direction="south" orientation="latitude" precision="5" value="-13.762">- 13.7620</geoCoordinate>
,
<geoCoordinate id="DCD18CA603F846BA7CCD02E0364A0ED4" degrees="13.2569" direction="east" orientation="longitude" precision="5" value="13.2569">13.2569</geoCoordinate>
</named-content>
,
<quantity id="2290F103704DF1C158EDC225DDEBDF29" metricMagnitude="3" metricUnit="m" metricValue="1.585" unit="m" value="1585.0">1585 m</quantity>
] (MUNHAC /
<collectionCode id="00B310FCED6F373A1B2B59F216A644D4" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB</collectionCode>
03 - 001542, 001543).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXAEK" type="comments">
<paragraph id="E5BD103433684ED849C88C85AA6BFE2F">
<heading id="E4A76F0D43E1F5C256AC6F8A911CFB39" reason="title">Comments.</heading>
</paragraph>
<paragraph id="31FFD4E66AD7FBEB984D3F2734A40EDF">
This recently collected material represents the first Angolan specimens of this species collected since its original description by
<bibRefCitation id="EF7F33B7DA366ABCA4D3061C211CA667" author="Laurent RF" firstAuthor="Laurent" journalOrPublisher="Publicações Culturais da Companhia de Diamantes de Angola" pagination="1-165" refId="B81" refString="Laurent RF (1964) Reptiles et amphibiens de l'Angola (Troisième contribution). Publicações Culturais da Companhia de Diamantes de Angola 67: 1 – 165." title="Reptiles et amphibiens de l'Angola (Troisième contribution)." volume="67" year="1964">Laurent (1964)</bibRefCitation>
. The species appears to be endemic to southwestern
<collectingCountry id="D7CD4BEE94917240EA88686B4E5A4A7E" name="Angola">Angola</collectingCountry>
, from
<collectingRegion id="CAE9499E4C932A00D85DCC22A2FBB214" country="Angola" name="Benguela">Benguela</collectingRegion>
to
<collectingRegion id="230BEF467945887C1D4109B567A1E01B" country="Angola" name="Namibe">Namibe Province</collectingRegion>
and to northwestern
<collectingCountry id="F2B7E28A061FB6A0725B15948A699158" name="Namibia">Namibia</collectingCountry>
. Our specimen from Serra da Neve was collected in the middle of its currently known distribution (
<bibRefCitation id="BCD874CBAE8BBCE5F26A9BC2E32709B6" author="Marques" etAl="et al." firstAuthor="Marques" refId="B86" refString="Marques MP, Ceríaco LMP, Blackburn DC, Bauer AM (2018) Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola – Atlas of Historical and Bibliographic Records (1840 – 2017). Proceedings of the California Academy of Sciences 4 (65, Supplement II): 1 – 501." year="2018">Marques et al. 2018</bibRefCitation>
). A specimen from northwestern
<collectingCountry id="3D1189F4E7B87772FD7D1A87778A33BE" name="Namibia">Namibia</collectingCountry>
was assigned to this species by
<bibRefCitation id="3C864C2911D7F600673B1C8C86139104" author="Broadley" firstAuthor="Broadley" issue="14" journalOrPublisher="Annals of the Transvaal Museum" pagination="209-215" refId="B24" refString="Broadley DG (1991) A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species. Annals of the Transvaal Museum 35 (14): 209 – 215." title="A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species." volume="35" year="1991">Broadley (1991</bibRefCitation>
,
<bibRefCitation id="EE21CBC67BF5DA28A464625A377456B8" author="Broadley" firstAuthor="Broadley" journalOrPublisher="Syntarsus" pagination="1-33" refId="B25" refString="Broadley DG (1996) A revision of the genus Lycophidion Fitzinger (Serpentes: Colubridae) in Africa South of the Equator. Syntarsus 3: 1 – 33." title="A revision of the genus Lycophidion Fitzinger (Serpentes: Colubridae) in Africa South of the Equator." volume="3" year="1996">1996</bibRefCitation>
). A very similar species,
<taxonomicName id="54097B6DAAD57905CB9DECA9BDD02537" authority="Broadley, 1991" authorityName="Broadley" authorityYear="1991" class="Reptilia" family="Lamprophiidae" genus="Lycophidion" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="namibianum">
<emphasis id="C1B111D3ADBB1798CCBDFD7880AD2066" italics="true">L. namibianum</emphasis>
<bibRefCitation id="9650E2228A9C418EE0455870941018F2" author="Broadley DG" issue="14" journalOrPublisher="Annals of the Transvaal Museum" pagination="209-215" refId="B24" refString="Broadley DG (1991) A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species. Annals of the Transvaal Museum 35 (14): 209 - 215." title="A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species." volume="35" year="1991">Broadley, 1991</bibRefCitation>
</taxonomicName>
, which occurs in northwestern
<collectingCountry id="BD1881A1BED860F99B41410A57AD12E5" name="Namibia">Namibia</collectingCountry>
and southern
<collectingRegion id="C8CBEBFDF291E17101002E61158A7E44" inferredBySuffix="true">Namibe Province</collectingRegion>
(
<bibRefCitation id="A2559312296F2EB48BDF0DFECCA867B8" author="Broadley" firstAuthor="Broadley" issue="14" journalOrPublisher="Annals of the Transvaal Museum" pagination="209-215" refId="B24" refString="Broadley DG (1991) A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species. Annals of the Transvaal Museum 35 (14): 209 – 215." title="A review of the Namibian snakes of the genus Lycophidion (Serpentes: Colubridae), with the description of a new endemic species." volume="35" year="1991">Broadley 1991</bibRefCitation>
,
<bibRefCitation id="2365E863D3FE31F4EC99C3746213F58A" author="Broadley" firstAuthor="Broadley" journalOrPublisher="Syntarsus" pagination="1-33" refId="B25" refString="Broadley DG (1996) A revision of the genus Lycophidion Fitzinger (Serpentes: Colubridae) in Africa South of the Equator. Syntarsus 3: 1 – 33." title="A revision of the genus Lycophidion Fitzinger (Serpentes: Colubridae) in Africa South of the Equator." volume="3" year="1996">1996</bibRefCitation>
;
<bibRefCitation id="360DDEACE75382C2B7BDD202E2C487FF" DOI="10.15560/18.2.289" author="Lobon-Rovira" etAl="et al." firstAuthor="Lobon-Rovira" issue="2" journalOrPublisher="Check List" pagination="289-321" refId="B83" refString="Lobón-Rovira J, Pinto PV, Becker FS, Tolley KA, Measey J, Boon B, de Sá S, Conradie W (2022) An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18 (2): 289 – 321. https://doi.org/10.15560/18.2.289" title="An updated herpetofaunal species inventory of Iona National Park in southwestern Angola." volume="18" year="2022">Lobón-Rovira et al. 2022</bibRefCitation>
), is distinguished from
<taxonomicName id="1DEABC153BE5623AC62F6021CE898F99" authorityName="Laurent" authorityYear="1964" class="Reptilia" family="Lamprophiidae" genus="Lycophidion" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="hellmichi">
<emphasis id="25D4BC42F8E81586728D3E81CCE561A2" italics="true">L. hellmichi</emphasis>
</taxonomicName>
by the lack of contact of the post-nasal with the first labial (in contact in
<taxonomicName id="EC0488D827E41A2BE2850F0C49EF338A" authorityName="Laurent" authorityYear="1964" class="Reptilia" family="Lamprophiidae" genus="Lycophidion" kingdom="Animalia" order="Squamata" phylum="Chordata" rank="species" species="hellmichi">
<emphasis id="4575579059A959955CA95B24DFC60F98" italics="true">L. hellmichi</emphasis>
</taxonomicName>
). Our specimens agree with the
<typeStatus id="886945A59794A40132479C36E0E8FA77" type="holotype">holotype</typeStatus>
in both coloration and the contact between post-nasal and first supralabial (
<bibRefCitation id="F685CF70C8CA0B9CD63B79CFBA4E0F6B" author="Laurent" firstAuthor="Laurent" journalOrPublisher="Publicações Culturais da Companhia de Diamantes de Angola" pagination="1-165" refId="B81" refString="Laurent RF (1964) Reptiles et amphibiens de l'Angola (Troisième contribution). Publicações Culturais da Companhia de Diamantes de Angola 67: 1 – 165." title="Reptiles et amphibiens de l'Angola (Troisième contribution)." volume="67" year="1964">Laurent 1964</bibRefCitation>
). The specimens from Catchi are adult male and female collected while copulating.
</paragraph>
</subSubSection>
</treatment>
</document>
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