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<document id="7D7E680E3932872406E49D906A68D0CF" ID-CLB-Dataset="305694" ID-DOI="10.5852/ejt.2024.970.2747" ID-GBIF-Dataset="f18c56fb-614e-4434-b29d-7c5a9533d5d7" ID-ISSN="2118-9773" ID-Zenodo-Dep="14239023" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732791843055" checkinUser="plazi" docAuthor="Bose, Arunima, Niskanen, Tuula, Datta, Sudeshna, Ghosh, Aniket, Vizzini, Alfredo &amp; Das, Kanad" docDate="2024" docId="FE1187D8FF9F6C58FE1061DFFB8EE674" docLanguage="en" docName="EJT.2024.970.38-60.pdf" docOrigin="European Journal of Taxonomy 970" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2747/12589" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cortinarius dhakuricus A. Bose &amp; K. Das 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="53" masterDocId="0228FFA0FF936C57FFE46454FF94E36B" masterDocTitle="Two new species and a new record of Cortinarius subgenus Telamonia (Agaricales) from India" masterLastPageNumber="60" masterPageNumber="38" pageNumber="50" updateTime="1732924269237" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="497CDA8C308EA1A75AC404F183B854D4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3905333C0284F59A66B441E901059BDA">
<mods:title id="811437EB5D54213B06020B3EBDC79044">Two new species and a new record of Cortinarius subgenus Telamonia (Agaricales) from India</mods:title>
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<mods:namePart id="51614B5EBEC218FA697E3DF458A7113F">Bose, Arunima</mods:namePart>
<mods:affiliation id="63BD1A7C3F91B46070A42E4816CE3822">Central National Herbarium, Botanical Survey of India, P. O. - Botanic Garden, Howrah - 711103, India.</mods:affiliation>
<mods:nameIdentifier id="FCB00278DBAAD41C7C9A9B2E1EBCA2A5" type="email">arunimabose96@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FC542EAEAC3C4827F906F24C34ADAC65">Niskanen, Tuula</mods:namePart>
<mods:affiliation id="57B03AE9FE1168C893E54D58C574F10F">Botany and Mycology Unit, Finnish Museum of Natural History, P. O. Box 7, FI- 00014, University of Helsinki, Helsinki, Finland.</mods:affiliation>
<mods:nameIdentifier id="447DF7AE4536F966FDA0D023DE3C52A6" type="email">tuula.niskanen@helsinki.fi</mods:nameIdentifier>
</mods:name>
<mods:name id="A44AFF3934A170EDFD5B902A2FE2FB46" type="personal">
<mods:role id="BCDC8B02B1E02660BC92FA3680F1FC61">
<mods:roleTerm id="1B613035D05399175B908B19DDBC97F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CF9DAB9AABFC9727411DB0BAB74D28C">Datta, Sudeshna</mods:namePart>
<mods:affiliation id="BF24B780A9DDB5930B51B6923C82C878">Central National Herbarium, Botanical Survey of India, P. O. - Botanic Garden, Howrah - 711103, India.</mods:affiliation>
<mods:nameIdentifier id="61E1552A3FBC2DEFAFDCBA085199259F" type="email">datta_su09@yahoo.in</mods:nameIdentifier>
</mods:name>
<mods:name id="D487603581121DCFE2DEA73A283C4AAD" type="personal">
<mods:role id="5D9A063C4ECF140677C876A254A81F52">
<mods:roleTerm id="6D363A18F3975081488BF839AA714D30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50CBC8C531F6DB6B77BDB1F4E432A366">Ghosh, Aniket</mods:namePart>
<mods:affiliation id="8E3B6D70C16633137D5E8C579023AEF5">Department of Biological Sciences, St. Xavier’s college, Maharo, Dumka - 814101, India.</mods:affiliation>
<mods:nameIdentifier id="03A0D9142187EDC7662AC7B6047B2866" type="email">ghosh.aniket87@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8852F07C7C2345550097A710C3E7B36E" type="personal">
<mods:role id="12D8599D07D0E1EA2DEF379EC7809175">
<mods:roleTerm id="56F9CF94FE51627131359E7F9910543F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="660C9AC8C31010C0A0B4601C0585521F">Vizzini, Alfredo</mods:namePart>
<mods:affiliation id="1B31F8EB4749E822926B272F74E9996E">Department of Life Sciences and Systems Biology, University of Torino, 10124 Torino, Italy.</mods:affiliation>
<mods:nameIdentifier id="042A36E712C09511B6BCAB83A23D652D" type="email">alfredo.vizzini@unito.it</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="4FCD978B3A8DC42C33A3075ADECD27C7">Das, Kanad</mods:namePart>
<mods:affiliation id="F332FD0754A267BBFCE43048F37D618D">Central National Herbarium, Botanical Survey of India, P. O. - Botanic Garden, Howrah - 711103, India.</mods:affiliation>
<mods:nameIdentifier id="2AFAABCC31FC76016AA8539C61C01CB5" type="email">daskanadbsi@gmail.com</mods:nameIdentifier>
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<mods:title id="EF045859CF44038ED7FE51715DF5CB25">European Journal of Taxonomy</mods:title>
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<mods:date id="B0112D6A1AFA77043CCF211C42AE65F0">2024</mods:date>
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<mods:number id="FE988AC2575A0B0146A97CD178DC4DC7">2024-11-25</mods:number>
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<mods:number id="37C1A32D9D9BE529DC21B003A0B0F0EC">970</mods:number>
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<mods:identifier id="CAC3E8EAA5C7F4D465D25A8306A51A4F" type="CLB-Dataset">305694</mods:identifier>
<mods:identifier id="C2EE32A63AED9DBA7CEC896735521AA2" type="DOI">10.5852/ejt.2024.970.2747</mods:identifier>
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<mods:identifier id="65E828A98ADF1E2C9F274AC61FD91F5A" type="Zenodo-Dep">14239023</mods:identifier>
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<treatment id="FE1187D8FF9F6C58FE1061DFFB8EE674" ID-DOI="http://doi.org/10.5281/zenodo.14239660" ID-GBIF-Taxon="248070563" ID-Zenodo-Dep="14239660" LSID="urn:lsid:plazi:treatment:FE1187D8FF9F6C58FE1061DFFB8EE674" httpUri="http://treatment.plazi.org/id/FE1187D8FF9F6C58FE1061DFFB8EE674" lastPageId="15" lastPageNumber="53" pageId="12" pageNumber="50" scope_class="Agaricomycetes" scope_order="Agaricales">
<subSubSection id="3EA26545FF9F6C5BFE1061DFFBD4E6CD" box="[500,1088,1419,1446]" pageId="12" pageNumber="50" type="nomenclature">
<paragraph id="760736CEFF9F6C5BFE1061DFFBD4E6CD" blockId="12.[500,1088,1419,1514]" box="[500,1088,1419,1446]" pageId="12" pageNumber="50">
<heading id="2D4F81A2FF9F6C5BFE1061DFFBD4E6CD" box="[500,1088,1419,1446]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="50" reason="2">
<taxonomicName id="B1B84D4DFF9F6C5BFE1061DFFC4AE6CD" authority="A.Bose &amp; K.Das" authorityName="A. Bose &amp; K. Das" authorityYear="2024" box="[500,990,1419,1446]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="12" pageNumber="50" phylum="Basidiomycota" rank="species" species="dhakuricus" status="sp. nov.">
<emphasis id="44CCEADCFF9F6C5BFE1061DFFC84E6CD" bold="true" box="[500,784,1419,1446]" italics="true" pageId="12" pageNumber="50">Cortinarius dhakuricus</emphasis>
A.Bose &amp; K.Das
</taxonomicName>
<taxonomicNameLabel id="5FFF57A7FF9F6C5BFC0261D8FBD4E6CD" box="[998,1088,1420,1446]" pageId="12" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EA26545FF9F6C5BFD6F61F9FCD9E681" pageId="12" pageNumber="50" type="description">
<paragraph id="760736CEFF9F6C5BFD6F61F9FC3CE6A3" blockId="12.[500,1088,1419,1514]" box="[651,936,1453,1480]" pageId="12" pageNumber="50">
<emphasis id="44CCEADCFF9F6C5BFD6F61F9FC80E6AC" bold="true" box="[651,788,1453,1479]" pageId="12" pageNumber="50">MycoBank</emphasis>
MB 851486
</paragraph>
<paragraph id="760736CEFF9F6C5BFD03619BFCD9E681" blockId="12.[500,1088,1419,1514]" box="[743,845,1487,1514]" pageId="12" pageNumber="50">
<figureCitation id="EE832A4BFF9F6C5BFD03619BFCD9E681" box="[743,845,1487,1514]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1869,1895]" captionStartId-1="14.[189,232,1928,1954]" captionTargetBox-0="[189,1398,265,1831]" captionTargetBox-1="[198,1387,482,1888]" captionTargetId-0="figure-21@13.[189,1398,265,1831]" captionTargetId-1="figure-52@14.[189,1398,473,1889]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 4. Cortinarius dhakuricus A.Bose &amp; K.Das sp. nov. (holotype; A. Bose &amp; K. Das, AB23-033, CAL). a–b. Fresh basidiomata. c. Basidia. d–e. Basidiospores (SEM). f. Transverse section through squamules. Scale bars: c–e = 10 µm; f = 20 µm." captionText-1="Fig. 5. Cortinarius dhakuricus A.Bose &amp; K.Das sp. nov. (holotype; A. Bose &amp; K. Das, AB23-033, CAL). a. Basidiospores. b. Basidia. c. Marginal cells. d–e. Transverse section of pileipellis through squamules. Scale bars = 10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.14239031" figureDoi-1="http://doi.org/10.5281/zenodo.14239033" httpUri-0="https://zenodo.org/record/14239031/files/figure.png" httpUri-1="https://zenodo.org/record/14239033/files/figure.png" pageId="12" pageNumber="50">Figs 4–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3EA26545FF9F6C5BFF59625BFCA2E5F0" pageId="12" pageNumber="50" type="diagnosis">
<paragraph id="760736CEFF9F6C5BFF59625BFEA2E542" blockId="12.[189,310,1551,1577]" box="[189,310,1551,1577]" pageId="12" pageNumber="50">
<heading id="2D4F81A2FF9F6C5BFF59625BFEA2E542" bold="true" box="[189,310,1551,1577]" fontSize="11" level="3" pageId="12" pageNumber="50" reason="3">
<emphasis id="44CCEADCFF9F6C5BFF59625BFEA2E542" bold="true" box="[189,310,1551,1577]" pageId="12" pageNumber="50">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9F6C5BFF596269FCA2E5F0" blockId="12.[189,1399,1597,1692]" pageId="12" pageNumber="50">
<taxonomicName id="B1B84D4DFF9F6C5BFF596269FE44E53C" authorityName="Bose &amp; Niskanen &amp; Datta &amp; Ghosh &amp; Vizzini &amp; Das" authorityYear="2024" box="[189,464,1597,1623]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="12" pageNumber="50" phylum="Basidiomycota" rank="species" species="dhakuricus" status="sp. nov.">
<emphasis id="44CCEADCFF9F6C5BFF596269FE44E53C" box="[189,464,1597,1623]" italics="true" pageId="12" pageNumber="50">Cortinarius dhakuricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FFF57A7FF9F6C5BFE3D626AFDA0E533" box="[473,564,1598,1624]" pageId="12" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
is separated from all the known species from
<collectingCountry id="0EAF765EFF9F6C5BFBB06269FB04E53C" box="[1108,1168,1597,1623]" name="India" pageId="12" pageNumber="50">India</collectingCountry>
by possessing very prominent silvery to yellowish white squamules on pileus surface and lamellae that are broadly attached with rare bifurcations and nrITS-based sequence data.
</paragraph>
</subSubSection>
<subSubSection id="3EA26545FF9F6C5BFF596296FBB7E462" box="[189,1059,1730,1802]" pageId="12" pageNumber="50" type="etymology">
<paragraph id="760736CEFF9F6C5BFF596296FED0E5B7" blockId="12.[189,324,1730,1756]" box="[189,324,1730,1756]" pageId="12" pageNumber="50">
<heading id="2D4F81A2FF9F6C5BFF596296FED0E5B7" bold="true" box="[189,324,1730,1756]" fontSize="11" level="3" pageId="12" pageNumber="50" reason="3">
<emphasis id="44CCEADCFF9F6C5BFF596296FED0E5B7" bold="true" box="[189,324,1730,1756]" pageId="12" pageNumber="50">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9F6C5BFF5962BBFBB7E462" blockId="12.[189,1059,1775,1802]" box="[189,1059,1775,1802]" pageId="12" pageNumber="50">
Referring to Dhakuri (of the state of
<collectingRegion id="B47CF82CFF9F6C5BFD8862BBFC94E462" box="[620,768,1775,1801]" country="India" name="Uttarakhand" pageId="12" pageNumber="50">Uttarakhand</collectingRegion>
,
<collectingCountry id="0EAF765EFF9F6C5BFCE862BBFCD9E462" box="[780,845,1775,1801]" name="India" pageId="12" pageNumber="50">India</collectingCountry>
), the
<typeStatus id="A903886CFF9F6C5BFC6F62A4FC2AE461" box="[907,958,1776,1802]" pageId="12" pageNumber="50">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="3EA26545FF9F6C59FF59637BFE07E2FC" lastPageId="14" lastPageNumber="52" pageId="12" pageNumber="50" type="materials_examined">
<paragraph id="760736CEFF9F6C5BFF59637BFEF8E422" blockId="12.[189,364,1839,1865]" box="[189,364,1839,1865]" pageId="12" pageNumber="50">
<heading id="2D4F81A2FF9F6C5BFF59637BFEF8E422" bold="true" box="[189,364,1839,1865]" fontSize="11" level="3" pageId="12" pageNumber="50" reason="3">
<emphasis id="44CCEADCFF9F6C5BFF59637BFEF8E422" bold="true" box="[189,364,1839,1865]" pageId="12" pageNumber="50">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9F6C5BFF596309FE77E483" blockId="12.[188,1399,1885,2024]" pageId="12" pageNumber="50">
<materialsCitation id="C6D03C93FF9F6C5BFF596309FE77E483" ID-GBIF-Occurrence="4986899303" collectedFrom="on the soil under Quercus semecarpifolia" collectingDate="2023-08-13" collectionCode="CAL" collectorName="A. Bose &amp; K. Das" country="India" elevation="2870" latitude="30.0711" location="Dhakuri" longLatPrecision="6" longitude="79.9204" municipality="Bageshwar District" pageId="12" pageNumber="50" specimenCode="CAL 1970" specimenCount="1" stateProvince="Uttarakhand" typeStatus="holotype">
<emphasis id="44CCEADCFF9F6C5BFF596309FEBBE41C" bold="true" box="[189,303,1885,1911]" pageId="12" pageNumber="50">
<typeStatus id="A903886CFF9F6C5BFF596309FEBBE41C" box="[189,303,1885,1911]" pageId="12" pageNumber="50" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="0EAF765EFF9F6C5BFF5963DEFE9BE4CF" box="[189,271,1930,1956]" name="India" pageId="12" pageNumber="50">INDIA</collectingCountry>
•
<collectingRegion id="B47CF82CFF9F6C5BFEC863DEFE56E4CF" box="[300,450,1930,1956]" country="India" name="Uttarakhand" pageId="12" pageNumber="50">Uttarakhand</collectingRegion>
,
<collectingMunicipality id="9663ACB4FF9F6C5BFE2B63DEFD2CE4CF" box="[463,696,1930,1956]" pageId="12" pageNumber="50">Bageshwar District</collectingMunicipality>
,
<location id="73676015FF9F6C5BFD2163DEFCBEE4CF" LSID="urn:lsid:plazi:treatment:FE1187D8FF9F6C58FE1061DFFB8EE674:73676015FF9F6C5BFD2163DEFCBEE4CF" box="[709,810,1930,1956]" country="India" latitude="30.0711" longLatPrecision="6" longitude="79.9204" municipality="Bageshwar District" name="Dhakuri" pageId="12" pageNumber="50" stateProvince="Uttarakhand">Dhakuri</location>
;
<elevation id="FD95D1FDFF9F6C5BFCDC63DEFC41E4CF" box="[824,981,1930,1957]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="12" pageNumber="50" unit="m" value="2870.0">
<quantity id="B1409B2BFF9F6C5BFCDC63DEFC07E4CE" box="[824,915,1930,1957]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="12" pageNumber="50" unit="m" value="2870.0">2870 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="138C5009FF9F6C5BFC0163DEFBE5E4CE" box="[997,1137,1930,1957]" degrees="30.0711" direction="north" orientation="latitude" pageId="12" pageNumber="50" precision="5" value="30.0711">30.0711° N</geoCoordinate>
,
<geoCoordinate id="138C5009FF9F6C5BFB9A63DEFA9DE4CE" box="[1150,1289,1930,1957]" degrees="79.9204" direction="east" orientation="longitude" pageId="12" pageNumber="50" precision="5" value="79.9204">79.9204° E</geoCoordinate>
;
<date id="0206100EFF9F6C5BFAF363DEFF69E4AD" pageId="12" pageNumber="50" value="2023-08-13">
<collectingDate id="1242E9E6FF9F6C5BFAF363DEFF69E4AD" pageId="12" pageNumber="50" value="2023-08-13">13 Aug. 2023</collectingDate>
</date>
;
<emphasis id="44CCEADCFF9F6C5BFEE263F9FDCCE4AD" box="[262,600,1964,1991]" italics="true" pageId="12" pageNumber="50">
<collectorName id="DB4D5318FF9F6C5BFEE263F9FECBE4AD" box="[262,351,1965,1991]" pageId="12" pageNumber="50">A. Bose</collectorName>
&amp;
<collectorName id="DB4D5318FF9F6C5BFE6763F9FE43E4AD" box="[387,471,1965,1990]" pageId="12" pageNumber="50">K. Das</collectorName>
, AB23-033
</emphasis>
;
<collectedFrom id="C5A47533FF9F6C5BFD8363F9FBD1E4AD" box="[615,1093,1964,1991]" pageId="12" pageNumber="50">
on the soil under
<taxonomicName id="B1B84D4DFF9F6C5BFCCA63F9FBD1E4AD" box="[814,1093,1964,1991]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="50" phylum="Tracheophyta" rank="species" species="semecarpifolia">
<emphasis id="44CCEADCFF9F6C5BFCCA63F9FBD1E4AD" box="[814,1093,1964,1991]" italics="true" pageId="12" pageNumber="50">Quercus semecarpifolia</emphasis>
</taxonomicName>
</collectedFrom>
; GenBank no.:
<accessionNumber id="69EBAB2DFF9F6C5BFB1A63F8FAE2E4AD" box="[1278,1398,1964,1990]" pageId="12" pageNumber="50">PP316707</accessionNumber>
(ITS);
<collectionCode id="10A9AE0BFF9F6C5BFEEF639AFED1E483" box="[267,325,1998,2024]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="12" pageNumber="50" type="Herbarium">CAL</collectionCode>
[
<specimenCode id="261E9EB5FF9F6C5BFEB7639AFE4CE483" box="[339,472,1998,2024]" collectionCode="CAL" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="12" pageNumber="50" type="Herbarium">CAL 1970</specimenCode>
].
</materialsCitation>
</paragraph>
<caption id="22C76646FF9E6C5AFF596319FDF1E4C5" ID-DOI="http://doi.org/10.5281/zenodo.14239031" ID-Zenodo-Dep="14239031" httpUri="https://zenodo.org/record/14239031/files/figure.png" pageId="13" pageNumber="51" startId="13.[189,232,1869,1895]" targetBox="[189,1398,265,1831]" targetPageId="13">
<paragraph id="760736CEFF9E6C5AFF596319FDF1E4C5" blockId="13.[188,1399,1869,1966]" pageId="13" pageNumber="51">
<emphasis id="44CCEADCFF9E6C5AFF596319FE9DE40C" bold="true" box="[189,265,1869,1895]" pageId="13" pageNumber="51">Fig. 4.</emphasis>
<taxonomicName id="B1B84D4DFF9E6C5AFEEB631AFD7EE403" authority="A.Bose &amp; K.Das" authorityName="A. Bose &amp; K. Das" authorityYear="2024" box="[271,746,1869,1896]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="13" pageNumber="51" phylum="Basidiomycota" rank="species" species="dhakuricus" status="sp. nov.">
<emphasis id="44CCEADCFF9E6C5AFEEB631AFDB4E40C" box="[271,544,1869,1896]" italics="true" pageId="13" pageNumber="51">Cortinarius dhakuricus</emphasis>
A.Bose &amp; K.Das
</taxonomicName>
<taxonomicNameLabel id="5FFF57A7FF9E6C5AFD14631AFCDDE403" box="[752,841,1870,1896]" pageId="13" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
(holotype;
<emphasis id="44CCEADCFF9E6C5AFC28631AFA88E403" box="[972,1308,1870,1896]" italics="true" pageId="13" pageNumber="51">A. Bose &amp; K. Das, AB23-033</emphasis>
, CAL).
<emphasis id="44CCEADCFF9E6C5AFF586325FF7EE4E0" bold="true" box="[188,234,1905,1931]" pageId="13" pageNumber="51">a–b</emphasis>
. Fresh basidiomata.
<emphasis id="44CCEADCFF9E6C5AFE3D6325FE72E4E0" bold="true" box="[473,486,1905,1931]" pageId="13" pageNumber="51">c</emphasis>
. Basidia.
<emphasis id="44CCEADCFF9E6C5AFDBD6325FD12E4E0" bold="true" box="[601,646,1905,1931]" pageId="13" pageNumber="51">d–e</emphasis>
. Basidiospores (SEM).
<emphasis id="44CCEADCFF9E6C5AFC7F6324FC31E4E1" bold="true" box="[923,933,1904,1930]" pageId="13" pageNumber="51">f</emphasis>
. Transverse section through squamules. Scale bars: c–e = 10 µm; f = 20 µm.
</paragraph>
</caption>
<paragraph id="760736CEFF9D6C59FF59655CFDA0E249" blockId="14.[189,564,264,290]" box="[189,564,264,290]" pageId="14" pageNumber="52">
<heading id="2D4F81A2FF9D6C59FF59655CFDA0E249" bold="true" box="[189,564,264,290]" fontSize="11" level="3" pageId="14" pageNumber="52" reason="3">
<emphasis id="44CCEADCFF9D6C59FF59655CFDA0E249" bold="true" box="[189,564,264,290]" pageId="14" pageNumber="52">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9D6C59FF596563FE07E2FC" blockId="14.[188,1399,311,408]" pageId="14" pageNumber="52">
<materialsCitation id="C6D03C93FF9D6C59FF596563FE1BE2FC" ID-GBIF-Occurrence="4986899307" collectedFrom="on the soil under Q. semecarpifolia" collectingDate="2023-08-14" collectionCode="CAL" collectorName="A. Bose &amp; K. Das" country="India" elevation="2901" latitude="30.0711" location="Dhakuri" longLatPrecision="6" longitude="79.9204" municipality="Bageshwar District" pageId="14" pageNumber="52" specimenCode="CAL 1986" specimenCount="1" stateProvince="Uttarakhand">
<collectingCountry id="0EAF765EFF9D6C59FF596563FE9BE23A" box="[189,271,311,337]" name="India" pageId="14" pageNumber="52">INDIA</collectingCountry>
•
<collectingRegion id="B47CF82CFF9D6C59FEC86563FE56E23A" box="[300,450,311,337]" country="India" name="Uttarakhand" pageId="14" pageNumber="52">Uttarakhand</collectingRegion>
,
<collectingMunicipality id="9663ACB4FF9D6C59FE2B6563FD2CE23A" box="[463,696,311,337]" pageId="14" pageNumber="52">Bageshwar District</collectingMunicipality>
,
<location id="73676015FF9D6C59FD216563FCBEE23A" LSID="urn:lsid:plazi:treatment:FE1187D8FF9F6C58FE1061DFFB8EE674:73676015FF9D6C59FD216563FCBEE23A" box="[709,810,311,337]" country="India" latitude="30.0711" longLatPrecision="6" longitude="79.9204" municipality="Bageshwar District" name="Dhakuri" pageId="14" pageNumber="52" stateProvince="Uttarakhand">Dhakuri</location>
;
<elevation id="FD95D1FDFF9D6C59FCDC6563FC41E23A" box="[824,981,311,338]" metricMagnitude="3" metricUnit="m" metricValue="2.901" pageId="14" pageNumber="52" unit="m" value="2901.0">
<quantity id="B1409B2BFF9D6C59FCDC6563FC07E239" box="[824,915,311,338]" metricMagnitude="3" metricUnit="m" metricValue="2.901" pageId="14" pageNumber="52" unit="m" value="2901.0">2901 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="138C5009FF9D6C59FC016563FBE5E239" box="[997,1137,311,338]" degrees="30.0711" direction="north" orientation="latitude" pageId="14" pageNumber="52" precision="5" value="30.0711">30.0711° N</geoCoordinate>
,
<geoCoordinate id="138C5009FF9D6C59FB9B6563FA9DE239" box="[1151,1289,311,338]" degrees="79.9204" direction="east" orientation="longitude" pageId="14" pageNumber="52" precision="5" value="79.9204">79.9204° E</geoCoordinate>
;
<date id="0206100EFF9D6C59FAF36563FF69E21F" pageId="14" pageNumber="52" value="2023-08-14">
<collectingDate id="1242E9E6FF9D6C59FAF36563FF69E21F" pageId="14" pageNumber="52" value="2023-08-14">14 Aug. 2023</collectingDate>
</date>
;
<emphasis id="44CCEADCFF9D6C59FEE2650FFE45E21F" box="[262,465,347,373]" italics="true" pageId="14" pageNumber="52">
<collectorName id="DB4D5318FF9D6C59FEE2650FFECAE21F" box="[262,350,347,373]" pageId="14" pageNumber="52">A. Bose</collectorName>
&amp;
<collectorName id="DB4D5318FF9D6C59FE65650FFE45E21F" box="[385,465,347,372]" pageId="14" pageNumber="52">K. Das</collectorName>
</emphasis>
,
<emphasis id="44CCEADCFF9D6C59FE3A650EFDC2E21F" box="[478,598,346,372]" italics="true" pageId="14" pageNumber="52">AB23-055</emphasis>
;
<collectedFrom id="C5A47533FF9D6C59FD87650FFC6DE21F" box="[611,1017,346,373]" pageId="14" pageNumber="52">
on the soil under
<taxonomicName id="B1B84D4DFF9D6C59FCCC650FFC6DE21F" box="[808,1017,346,373]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="52" phylum="Tracheophyta" rank="species" species="semecarpifolia">
<emphasis id="44CCEADCFF9D6C59FCCC650FFC6DE21F" box="[808,1017,346,373]" italics="true" pageId="14" pageNumber="52">Q. semecarpifolia</emphasis>
</taxonomicName>
</collectedFrom>
; GenBank no.:
<accessionNumber id="69EBAB2DFF9D6C59FB54650EFABDE21E" box="[1200,1321,346,373]" pageId="14" pageNumber="52">PP345590</accessionNumber>
(ITS);
<collectionCode id="10A9AE0BFF9D6C59FF58652AFF62E2F3" box="[188,246,382,408]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="14" pageNumber="52" type="Herbarium">CAL</collectionCode>
[
<specimenCode id="261E9EB5FF9D6C59FEE7652AFE1CE2FC" box="[259,392,381,408]" collectionCode="CAL" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="14" pageNumber="52" type="Herbarium">CAL 1986</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="22C76646FF9D6C59FF5963DCFE3CE482" ID-DOI="http://doi.org/10.5281/zenodo.14239033" ID-Zenodo-Dep="14239033" httpUri="https://zenodo.org/record/14239033/files/figure.png" pageId="14" pageNumber="52" startId="14.[189,232,1928,1954]" targetBox="[198,1387,482,1888]" targetPageId="14">
<paragraph id="760736CEFF9D6C59FF5963DCFE3CE482" blockId="14.[188,1399,1928,2025]" pageId="14" pageNumber="52">
<emphasis id="44CCEADCFF9D6C59FF5963DCFE9DE4C9" bold="true" box="[189,265,1928,1954]" pageId="14" pageNumber="52">Fig. 5.</emphasis>
<taxonomicName id="B1B84D4DFF9D6C59FEEB63DCFD7DE4C9" authority="A.Bose &amp; K.Das" authorityName="A. Bose &amp; K. Das" authorityYear="2024" box="[271,745,1928,1954]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="14" pageNumber="52" phylum="Basidiomycota" rank="species" species="dhakuricus" status="sp. nov.">
<emphasis id="44CCEADCFF9D6C59FEEB63DCFDB4E4C9" box="[271,544,1928,1954]" italics="true" pageId="14" pageNumber="52">Cortinarius dhakuricus</emphasis>
A.Bose &amp; K.Das
</taxonomicName>
<taxonomicNameLabel id="5FFF57A7FF9D6C59FD0B63DCFCDDE4C9" box="[751,841,1928,1954]" pageId="14" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
(holotype;
<emphasis id="44CCEADCFF9D6C59FC2963DCFA89E4C9" box="[973,1309,1928,1954]" italics="true" pageId="14" pageNumber="52">A. Bose &amp; K. Das, AB23-033</emphasis>
, CAL).
<emphasis id="44CCEADCFF9D6C59FF5863FFFF5FE4AE" bold="true" box="[188,203,1963,1989]" pageId="14" pageNumber="52">a</emphasis>
. Basidiospores.
<emphasis id="44CCEADCFF9D6C59FE6F63FFFE0FE4AE" bold="true" box="[395,411,1963,1989]" pageId="14" pageNumber="52">b</emphasis>
. Basidia.
<emphasis id="44CCEADCFF9D6C59FDEB63FFFD88E4AE" bold="true" box="[527,540,1963,1989]" pageId="14" pageNumber="52">c</emphasis>
. Marginal cells.
<emphasis id="44CCEADCFF9D6C59FD3B63FFFC9FE4AE" bold="true" box="[735,779,1963,1989]" pageId="14" pageNumber="52">d–e</emphasis>
. Transverse section of pileipellis through squamules. Scale bars = 10 µm.
</paragraph>
</caption>
<subSubSection id="3EA26545FF9C6C58FF59655CFEEBE0BD" pageId="15" pageNumber="53" type="description">
<paragraph id="760736CEFF9C6C58FF59655CFEDAE249" blockId="15.[189,334,264,290]" box="[189,334,264,290]" pageId="15" pageNumber="53">
<heading id="2D4F81A2FF9C6C58FF59655CFEDAE249" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="15" pageNumber="53" reason="3">
<emphasis id="44CCEADCFF9C6C58FF59655CFEDAE249" bold="true" box="[189,334,264,290]" pageId="15" pageNumber="53">Description</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9C6C58FF59656CFACCE1FD" blockId="15.[187,1401,312,663]" pageId="15" pageNumber="53">
Pileus
<quantity id="B1409B2BFF9C6C58FEEE656CFE13E239" box="[266,391,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="15" pageNumber="53" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20 mm</quantity>
diam., acutely conical when young, becoming conico-umbonate or obtusely umbonate, with an acute umbo; margin entire; surface grayish brown, covered with very prominent silvery to yellowish white squamules, umbo reddish brown, turning maroon brown with KOH; dry, hygrophanous. Lamellae broadly attached, sinuate, rarely forked; subdistant (8–10/cm at pileus margin, lamellulae in 5 series), rather thick; reddish brown (9D4); edges dentate. Stipe
<quantity id="B1409B2BFF9C6C58FC42659CFBB1E288" box="[934,1061,456,483]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" pageId="15" pageNumber="53" unit="mm" value="44.0" valueMax="50.0" valueMin="38.0">38–50 mm</quantity>
long,
<quantity id="B1409B2BFF9C6C58FB94659CFB45E288" box="[1136,1233,456,483]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="15" pageNumber="53" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
thick at apex, up to
<quantity id="B1409B2BFF9C6C58FEE465B8FEC7E16C" box="[256,339,492,519]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="53" unit="mm" value="10.0">10 mm</quantity>
thick at base, cylindrical to somewhat enlarged below, slightly tapered towards very base, with longitudinal striate; surface yellowish brown, turning light yellow with KOH, dry. Universal veil forming a yellowish white membranous annular zone on upper part of stipe and some incomplete and complete girdles lower down. Basal mycelium yellow-white. Pileus context brown, color unchanging when bruised; stipe context stuffed, consistency brittle. Odor strongly raphanoid. Taste not recorded.
</paragraph>
<paragraph id="760736CEFF9C6C58FF5966EBFEEBE0BD" blockId="15.[189,1400,703,982]" pageId="15" pageNumber="53">
Basidiospores 8.0–8.7–10.0 × 4.1–5.9–6.0 μm, Q = 1.5–1.71–2.12, ellipsoid to obovoid-ellipsoid, (moderately) to strongly verrucose dextrinoid. Basidia 30–37 × 10–20 μm, 4-spored, clavate. Sterile marginal elements 20–30 × 6–10 μm, cylindrical to clavate, colorless and thin-walled, clamped at the base. Pileipellis composed of compactly arranged, cylindrical, repent parallel hyphae giving rise to frequent squamules; each squamule composed of clusters of isodiametric pseudoparenchymatous elements to bulbous (inflated) erect hyphae in clusters that
<taxonomicName id="B1B84D4DFF9C6C58FD146727FCC2E0E6" box="[752,854,883,909]" form="the" pageId="15" pageNumber="53" rank="form">form the</taxonomicName>
squamule tip at regular to irregular intervals; terminal elements of squamules 29–51 × 15–35 μm, subcylindrical to ventricose with rounded or rarely mucronate apex.
</paragraph>
</subSubSection>
<subSubSection id="3EA26545FF9C6C58FF5967AAFB8EE674" pageId="15" pageNumber="53" type="discussion">
<paragraph id="760736CEFF9C6C58FF5967AAFEBBE773" blockId="15.[189,303,1022,1048]" box="[189,303,1022,1048]" pageId="15" pageNumber="53">
<heading id="2D4F81A2FF9C6C58FF5967AAFEBBE773" bold="true" box="[189,303,1022,1048]" fontSize="11" level="3" pageId="15" pageNumber="53" reason="3">
<emphasis id="44CCEADCFF9C6C58FF5967AAFEBBE773" bold="true" box="[189,303,1022,1048]" pageId="15" pageNumber="53">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="760736CEFF9C6C58FF596079FB8EE674" blockId="15.[188,1400,1069,1312]" pageId="15" pageNumber="53">
Based on available sequences in the public database it appears that
<taxonomicName id="B1B84D4DFF9C6C58FC2E6079FBFAE72C" authorityName="Bose &amp; Niskanen &amp; Datta &amp; Ghosh &amp; Vizzini &amp; Das" authorityYear="2024" box="[970,1134,1069,1095]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="53" phylum="Basidiomycota" rank="species" species="dhakuricus" status="sp. nov.">
<emphasis id="44CCEADCFF9C6C58FC2E6079FBFAE72C" box="[970,1134,1069,1095]" italics="true" pageId="15" pageNumber="53">C. dhakuricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FFF57A7FF9C6C58FB916079FB5BE72C" box="[1141,1231,1069,1095]" pageId="15" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
has no known sister taxa in the section. However, because of its conic-umbonate pileus with squamulose surface it partly resembles another Himalayan species
<taxonomicName id="B1B84D4DFF9C6C58FD936021FB61E7E4" authority="K. Das, D. Chakr., A. Ghosh &amp; Vizzini" authorityName="K. Das, D. Chakr., A. Ghosh &amp; Vizzini" box="[631,1269,1141,1168]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="53" phylum="Basidiomycota" rank="species" species="dombangensis">
<emphasis id="44CCEADCFF9C6C58FD936021FCD4E7E4" box="[631,832,1141,1167]" italics="true" pageId="15" pageNumber="53">C. dombangensis</emphasis>
K.Das, D.Chakr., A.Ghosh &amp; Vizzini
</taxonomicName>
(originally reported from
<collectingCountry id="0EAF765EFF9C6C58FE8D60CDFE3EE7D8" box="[361,426,1177,1203]" name="India" pageId="15" pageNumber="53">India</collectingCountry>
), but the latter is segregated from the former by possessing a silky stipe which is encircled with complete to incomplete irregular whitish bands of velar remnants in multiple rows, with a yellowish white membranous annular zone, and occurs under
<taxonomicName id="B1B84D4DFF9C6C58FC4860B5FB11E790" authority="Griff." authorityName="Griff." box="[940,1157,1249,1275]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="53" phylum="Tracheophyta" rank="species" species="densa">
<emphasis id="44CCEADCFF9C6C58FC4860B5FBADE790" box="[940,1081,1249,1275]" italics="true" pageId="15" pageNumber="53">Abies densa</emphasis>
Griff.
</taxonomicName>
;
<taxonomicName id="B1B84D4DFF9C6C58FB7560B6FACFE790" box="[1169,1371,1249,1276]" class="Agaricomycetes" family="Cortinariaceae" genus="Cortinarius" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="53" phylum="Basidiomycota" rank="species" species="dombangensis">
<emphasis id="44CCEADCFF9C6C58FB7560B6FACFE790" box="[1169,1371,1249,1276]" italics="true" pageId="15" pageNumber="53">C. dombangensis</emphasis>
</taxonomicName>
is also segregated based on its nrITS-based sequence data (
<bibRefCitation id="12294B3FFF9C6C58FCB26151FB9AE674" author="Das" box="[854,1038,1285,1311]" etAl="et al." firstAuthor="Das" pageId="15" pageNumber="53" refId="ref9277" refString="Das K., Chakraborty D., Ghosh A. &amp; Vizzini A. 2022. Cortinarius dombangensis sp. nov. and C. longistipitatus (Agaricales, Cortinariaceae) from Indian Himalaya. Nordic Journal of Botany 2022: e 03579. https: // doi. org / 10.1111 / njb. 03579" type="journal volume" year="2022">
Das
<emphasis id="44CCEADCFF9C6C58FC686152FC53E674" box="[908,967,1285,1311]" italics="true" pageId="15" pageNumber="53">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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