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<document id="563A8E961291D18D95FEB227D4870539" ID-DOI="10.1080/00222933.2018.1506521" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747249" ID-ZooBank="1C028676-CCD9-4654-A755-6ACA3BCE3E98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670908261" checkinUser="carolina" docAuthor="Ferretti, Nelson, Cavallo, Patricio, Chaparro, Juan C., Ríos-Tamayo, Duniesky, Seimon, Tracie A. &amp; West, Rick" docDate="2018" docId="FD618785AA59CA1F1BCDCD95FC0164DB" docLanguage="en" docName="JNATHIST.52.29-30.1927-1984.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (29 - 30)" docSource="http://dx.doi.org/10.1080/00222933.2018.1506521" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hapalotremus carabaya Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West 2018, sp. nov." docType="treatment" docUuid="FE87A5DD-3FD1-49D7-A549-B5603FCA3383" docUuidSource="ZooBank" docVersion="4" lastPageNumber="1959" masterDocId="0158FFFDAA44CA3E1A06C90AFFA66264" masterDocTitle="The Neotropical genus Hapalotremus Simon, 1903 (Araneae: Theraphosidae), with the description of seven new species and the highest altitude record for the family" masterLastPageNumber="1984" masterPageNumber="1927" pageNumber="1955" updateTime="1698914576834" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="51F19C899F5B5322A83B4283CA575027">The Neotropical genus Hapalotremus Simon, 1903 (Araneae: Theraphosidae), with the description of seven new species and the highest altitude record for the family</mods:title>
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<mods:namePart id="AB006FAD9EBFBA04C0F5711111B507C4">Ferretti, Nelson</mods:namePart>
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<mods:namePart id="42DD530F8E5FD12CEF5F287483080600">Cavallo, Patricio</mods:namePart>
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<mods:namePart id="0173AE10E65B8F6686E31DDBB4CE6C5D">Chaparro, Juan C.</mods:namePart>
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<mods:namePart id="2CCAA0E2797476E4E697BF1136C7B83B">Ríos-Tamayo, Duniesky</mods:namePart>
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<taxonomicName id="B2C84D10AA59CA231BCDCD95FD5566DC" ID-CoL="3JKLQ" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[459,755,1183,1208]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1955" phylum="Arthropoda" rank="species" species="carabaya" status="sp. nov.">
<emphasis id="47BCEA81AA59CA231BCDCD95FD5566DC" bold="true" box="[459,755,1183,1208]" italics="true" pageId="29" pageNumber="1955">Hapalotremus carabaya</emphasis>
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(
<figureCitation id="EDF32A16AA59CA231BE7CDC8FDDD66BF" box="[481,635,1218,1243]" captionStart="Figure 2" captionStartId="6.[160,225,1289,1312]" captionTargetBox="[163,1152,142,1258]" captionTargetId="figure-139@6.[163,1152,142,1258]" captionTargetPageId="6" captionText="Figure 2. (af) Hapalotremus spp. Habitus. (a, b) Hapalotremus carabaya sp. nov., female holotype. (c, d) Hapalotremus vilcanota sp. nov., female. (e) Hapalotremus vilcanota sp. nov., female and juvenile (white arrow) found under the same stone. (f) Hapalotremus vilcanota sp. nov. burrow. Photos: a, b, df: Tracie Seimon; c: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747253" httpUri="https://zenodo.org/record/4747253/files/figure.png" pageId="29" pageNumber="1955">Figures 2a, b</figureCitation>
,
<figureCitation id="EDF32A16AA59CA23188CCDC8FD1966BF" box="[650,703,1218,1243]" captionStart="Figure 3" captionStartId="7.[160,225,1665,1688]" captionTargetBox="[163,1152,142,1634]" captionTargetId="figure-1@7.[163,1152,142,1634]" captionTargetPageId="7" captionText="Figure 3. (aj) Hapalotremus spp. habitat. (a) Escoipe River, next to route 33, Salta Province, Argentina, type locality of Hapalotremus martinorum. (b, c) Los Sosa River, next to route 307, Tucumán Province, Argentina, habitat of Hapalotremus chasqui sp. nov. (d, f) Cordillera Carabaya, Michi Michani Cunca, type locality of Hapalotremus carabaya sp. nov. (e, gj) Cordillera Vilcanota, Upis, type locality of Hapalotremus vilcanota sp. nov. Photos: a: Patricio Cavallo;b: Gabriel Pompozzi; c: Laura Montes de Oca; di: Tracie Seimon; j: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747255" httpUri="https://zenodo.org/record/4747255/files/figure.png" pageId="29" pageNumber="1955">3d, f</figureCitation>
,
<figureCitation id="EDF32A16AA59CA2318C9CDC8FCB066BF" box="[719,790,1218,1243]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStartId-0="21.[160,225,1303,1326]" captionStartId-1="22.[160,225,1143,1166]" captionTargetBox-0="[168,1151,142,1272]" captionTargetBox-1="[163,1152,142,1112]" captionTargetId-0="figure-66@21.[163,1152,142,1272]" captionTargetId-1="figure-198@22.[163,1152,142,1112]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Figure 17. (ad) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. Scale bar: 5 mm. Photos: Nelson Ferretti." captionText-1="Figure 18. (af) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Eyes, dorsal view. (b) Labium, ventral view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. (f) Spermathecae, ventral view. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi-0="http://doi.org/10.5281/zenodo.4747287" figureDoi-1="http://doi.org/10.5281/zenodo.4747289" httpUri-0="https://zenodo.org/record/4747287/files/figure.png" httpUri-1="https://zenodo.org/record/4747289/files/figure.png" pageId="29" pageNumber="1955">1718</figureCitation>
,
<figureCitation id="EDF32A16AA59CA231923CDC8FCE466BF" box="[805,834,1218,1243]" captionStart="Figure 37" captionStartId="41.[160,225,846,869]" captionTargetBox="[163,1152,142,815]" captionTargetId="figure-275@41.[163,1152,142,815]" captionTargetPageId="41" captionText="Figure 37. Distribution map of Hapalotremus species from Peru." figureDoi="http://doi.org/10.5281/zenodo.4747338" httpUri="https://zenodo.org/record/4747338/files/figure.png" pageId="29" pageNumber="1955">37</figureCitation>
)
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<emphasis id="47BCEA81AA59CA231AA6CC19FEE36748" bold="true" box="[160,325,1299,1324]" italics="true" pageId="29" pageNumber="1955">Type material</emphasis>
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) from
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a
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<materialsCitation id="C5A03CCEAA59CA23195ECC53FCCD6715" box="[856,875,1369,1393]" collectionCode="MUSM" country="Peru" county="Female" elevation="4057" isTailEnumeration="true" latitude="-13.5792" location="Michi Michani Cunca" longLatPrecision="7" longitude="-70.5778" municipality="Quispicanchi" pageId="29" pageNumber="1955" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">.s</materialsCitation>
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<date id="01761053AA59CA231987CC53FBBE6715" box="[897,1048,1369,1393]" pageId="29" pageNumber="1955" value="2006-06-30">30 June 2006</date>
, Severino Crispin Huanca col
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<emphasis id="47BCEA81AA59CA231AA6CCC1FDA36780" bold="true" box="[160,517,1483,1508]" italics="true" pageId="29" pageNumber="1955">Additional material examined</emphasis>
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<materialsCitation id="C5A03CCEAA59CA231AA6CCE5FD5E644D" collectionCode="MUSM" country="Peru" county="Immature" elevation="3947" latitude="-13.5792" location="Michi Michani Cunca" longLatPrecision="7" longitude="-70.5778" municipality="Quispicanchi" pageId="29" pageNumber="1955" specimenCode="ENT 0508678" specimenCount="1" stateProvince="Cusco">
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),
<quantity id="B2309B76AA59CA231888CF1BFD46644D" box="[654,736,1553,1577]" metricMagnitude="3" metricUnit="m" metricValue="3.947" pageId="29" pageNumber="1955" unit="m" value="3947.0">
<elevation id="FEE5D1A0AA59CA231888CF1BFD46644D" box="[654,736,1553,1577]" metricMagnitude="3" metricUnit="m" metricValue="3.947" pageId="29" pageNumber="1955" unit="m" value="3947.0">3947 m</elevation>
</quantity>
a
</materialsCitation>
<materialsCitation id="C5A03CCEAA59CA2318FECF1BFCAD644D" box="[760,779,1553,1577]" collectionCode="MUSM" country="Peru" county="Immature" elevation="3947" isTailEnumeration="true" latitude="-13.5792" location="Michi Michani Cunca" longLatPrecision="7" longitude="-70.5778" municipality="Quispicanchi" pageId="29" pageNumber="1955" specimenCount="1" stateProvince="Cusco">.s</materialsCitation>
.l.,
<date id="01761053AA59CA231919CF1BFC15644D" box="[799,947,1553,1577]" pageId="29" pageNumber="1955" value="2006-06-28">28 June 2006</date>
, Tracie Seimon col
<materialsCitation id="C5A03CCEAA59CA231E86CF1BFB22644D" box="[1152,1156,1553,1577]" collectionCode="MUSM" country="Peru" county="Immature" elevation="3947" isTailEnumeration="true" latitude="-13.5792" location="Michi Michani Cunca" longLatPrecision="7" longitude="-70.5778" municipality="Quispicanchi" pageId="29" pageNumber="1955" specimenCount="1" stateProvince="Cusco">.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3DD26518AA59CA201AA6CF6BFC186476" lastPageId="30" lastPageNumber="1956" pageId="29" pageNumber="1955" type="etymology">
<paragraph id="75773693AA59CA231AA6CF6BFEBB641E" blockId="29.[160,1155,1633,1727]" box="[160,285,1633,1658]" pageId="29" pageNumber="1955">
<emphasis id="47BCEA81AA59CA231AA6CF6BFEBB641E" bold="true" box="[160,285,1633,1658]" italics="true" pageId="29" pageNumber="1955">Etymology</emphasis>
</paragraph>
<paragraph id="75773693AA59CA201AA6CF8FFC186476" blockId="29.[160,1155,1633,1727]" lastBlockId="30.[160,1155,1460,1554]" lastPageId="30" lastPageNumber="1956" pageId="29" pageNumber="1955">
<materialsCitation id="C5A03CCEAA59CA231AA6CF8FFBCE64DB" collectorName="The" country="Peru" location="Cordillera Carabaya" pageId="29" pageNumber="1955" specimenCount="1">
<collectorName id="D83D5345AA59CA231AA6CF8FFF6F64F9" box="[160,201,1669,1693]" pageId="29" pageNumber="1955">The</collectorName>
specific epithet refers to the
<typeStatus id="AA738831AA59CA23181ACF8FFDE864F9" box="[540,590,1669,1693]" pageId="29" pageNumber="1955">type</typeStatus>
locality where the species was found and also to the
<location id="70176048AA59CA231AC9CFADFE0D64DB" LSID="urn:lsid:plazi:treatment:FD618785AA59CA1F1BCDCD95FC0164DB:70176048AA59CA231AC9CFADFE0D64DB" box="[207,427,1703,1727]" country="Peru" name="Cordillera Carabaya" pageId="29" pageNumber="1955">Cordillera Carabaya</location>
which is a mountain chain located in the southeast of
<collectingCountry id="0DDF7603AA59CA231E34CFADFBCE64DB" box="[1074,1128,1703,1727]" name="Peru" pageId="29" pageNumber="1955">Peru</collectingCountry>
</materialsCitation>
.
<materialsCitation id="C5A03CCEAA59CA201E72CFADFCE9678B" collectorName="It &amp; Andes" country="Peru" lastPageId="30" lastPageNumber="1956" location="Eastern Cordillera" pageId="29" pageNumber="1955" specimenCount="1" stateProvince="Puno">
<collectorName id="D83D5345AA59CA231E72CFADFB2564DB" box="[1140,1155,1703,1727]" pageId="29" pageNumber="1955">It</collectorName>
forms part of the
<location id="70176048AA5ACA201B6FCCBFFD9667A9" LSID="urn:lsid:plazi:treatment:FD618785AA59CA1F1BCDCD95FC0164DB:70176048AA5ACA201B6FCCBFFD9667A9" box="[361,560,1461,1485]" country="Peru" name="Eastern Cordillera" pageId="30" pageNumber="1956" stateProvince="Puno">Eastern Cordillera</location>
of the
<collectorName id="D83D5345AA5ACA201885CCBFFD6F67A9" box="[643,713,1461,1485]" pageId="30" pageNumber="1956">Andes</collectorName>
and extends in a northwestsoutheast direction for about
<quantity id="B2309B76AA5ACA201B8DCCDDFE7F678B" box="[395,473,1495,1519]" metricMagnitude="4" metricUnit="m" metricValue="9.5" pageId="30" pageNumber="1956" unit="km" value="95.0">95 km</quantity>
, from the department of
<collectingRegion id="B70CF871AA5ACA201914CCDDFCE9678B" box="[786,847,1495,1519]" country="Peru" name="Puno" pageId="30" pageNumber="1956">Puno</collectingRegion>
</materialsCitation>
. It also presents a glacier surface of
<quantity id="B2309B76AA5ACA201B1ECCF0FECB6476" box="[280,365,1530,1554]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="1956" unit="km" value="100.0">100 km</quantity>
<superScript id="82BD9BDBAA5ACA201B6ACCFEFED06460" attach="left" box="[364,374,1524,1540]" fontSize="7" pageId="30" pageNumber="1956">2</superScript>
with its maximum elevation reaching
<quantity id="B2309B76AA5ACA201936CCF0FC206476" box="[816,902,1530,1554]" metricMagnitude="3" metricUnit="m" metricValue="5.779999999999999" pageId="30" pageNumber="1956" unit="m" value="5780.0">5780 m</quantity>
a
<materialsCitation id="C5A03CCEAA5ACA2019A6CCF0FC156476" box="[928,947,1530,1554]" collectorName="It &amp; Andes" country="Peru" isTailEnumeration="true" location="Eastern Cordillera" pageId="30" pageNumber="1956" specimenCount="1" stateProvince="Puno">.s</materialsCitation>
.l.
</paragraph>
</subSubSection>
<caption id="21B7661BAA5ACA201AA6CC1FFD1D6703" ID-DOI="http://doi.org/10.5281/zenodo.4747313" ID-Zenodo-Dep="4747313" httpUri="https://zenodo.org/record/4747313/files/figure.png" pageId="30" pageNumber="1956" startId="30.[160,225,1301,1324]" targetBox="[163,1151,142,1271]" targetPageId="30">
<paragraph id="75773693AA5ACA201AA6CC1FFD1D6703" blockId="30.[160,1157,1301,1383]" pageId="30" pageNumber="1956">
<emphasis id="47BCEA81AA5ACA201AA6CC1FFEAE6748" bold="true" box="[160,264,1301,1324]" pageId="30" pageNumber="1956">Figure 26.</emphasis>
(ae)
<taxonomicName id="B2C84D10AA5ACA201B4FCC1FFD916748" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[329,567,1301,1324]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="1956" phylum="Arthropoda" rank="species" species="chespiritoi" status="sp. nov.">
<emphasis id="47BCEA81AA5ACA201B4FCC1FFD916748" box="[329,567,1301,1324]" italics="true" pageId="30" pageNumber="1956">Hapalotremus chespiritoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C8F57FAAA5ACA201839CC1CFD296749" box="[575,655,1302,1325]" pageId="30" pageNumber="1956" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype (MUBI). (a) Labium, ventral view. (b) Eyes, dorsal view. (c) Tarsus IV, prolateral view. (d) Tarsus IV, ventral view. (e) Spermathecae, dorsal view. Scale bar: 1 mm. Photos: Nelson Ferretti.
</paragraph>
</caption>
<subSubSection id="3DD26518AA5ACA211AA6CF5EFCE2667E" lastPageId="31" lastPageNumber="1957" pageId="30" pageNumber="1956" type="diagnosis">
<paragraph id="75773693AA5ACA201AA6CF5EFEB26409" blockId="30.[160,1155,1620,1714]" box="[160,276,1620,1645]" pageId="30" pageNumber="1956">
<emphasis id="47BCEA81AA5ACA201AA6CF5EFEB26409" bold="true" box="[160,276,1620,1645]" italics="true" pageId="30" pageNumber="1956">Diagnosis</emphasis>
</paragraph>
<paragraph id="75773693AA5ACA201AA6CF7DFB2564D6" blockId="30.[160,1155,1620,1714]" pageId="30" pageNumber="1956">
Females of
<taxonomicName id="B2C84D10AA5ACA201B2DCF7DFE1564EB" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[299,435,1655,1679]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="1956" phylum="Arthropoda" rank="species" species="carabaya" status="sp. nov.">
<emphasis id="47BCEA81AA5ACA201B2DCF7DFE1564EB" box="[299,435,1655,1679]" italics="true" pageId="30" pageNumber="1956">H. carabaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C8F57FAAA5ACA201BB9CF7DFDBD64EB" box="[447,539,1655,1679]" pageId="30" pageNumber="1956" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from those of all other species of
<taxonomicName id="B2C84D10AA5ACA201B27CF90FE1C64D6" authorityName="Simon" authorityYear="1903" box="[289,442,1690,1714]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="1956" phylum="Arthropoda" rank="genus">
<emphasis id="47BCEA81AA5ACA201B27CF90FE1C64D6" box="[289,442,1690,1714]" italics="true" pageId="30" pageNumber="1956">Hapalotremus</emphasis>
</taxonomicName>
by the spermathecae bearing an additional well-developed
</paragraph>
<caption id="21B7661BAA5BCA211AA6CA09FE9F61F4" ID-DOI="http://doi.org/10.5281/zenodo.4747315" ID-Zenodo-Dep="4747315" httpUri="https://zenodo.org/record/4747315/files/figure.png" pageId="31" pageNumber="1957" startId="31.[160,225,771,794]" targetBox="[163,1151,142,740]" targetPageId="31">
<paragraph id="75773693AA5BCA211AA6CA09FE9F61F4" blockId="31.[160,1157,771,912]" pageId="31" pageNumber="1957">
<emphasis id="47BCEA81AA5BCA211AA6CA09FEA1617E" bold="true" box="[160,263,771,794]" pageId="31" pageNumber="1957">Figure 27.</emphasis>
(ag)
<taxonomicName id="B2C84D10AA5BCA211B4CCA09FDA3617E" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[330,517,771,794]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="1957" phylum="Arthropoda" rank="species" species="kuka" status="sp. nov.">
<emphasis id="47BCEA81AA5BCA211B4CCA09FDA3617E" box="[330,517,771,794]" italics="true" pageId="31" pageNumber="1957">Hapalotremus kuka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C8F57FAAA5BCA21180ACA09FDFD617E" box="[524,603,771,794]" pageId="31" pageNumber="1957" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Eyes, dorsal view. (e) Labium and maxillae, ventral view. (f) Metatarsus and tarsus IV, lateral view. (g) Spermathecae, dorsal view. Arrow indicates the scopulae on ventral metatarsus IV. Scale bars: ac: 5 mm. dg: 1 mm. Photos: Nelson Ferretti.
</paragraph>
</caption>
<paragraph id="75773693AA5BCA211AA6CAEAFCE2667E" blockId="31.[160,1155,992,1050]" pageId="31" pageNumber="1957">
apical median projection with a long thin wavy groove extending downward dorsal median from this projection (
<figureCitation id="EDF32A16AA5BCA211BF6CD08FDD9667E" box="[496,639,1026,1050]" captionStart="Figure 18" captionStartId="22.[160,225,1143,1166]" captionTargetBox="[163,1152,142,1112]" captionTargetId="figure-198@22.[163,1152,142,1112]" captionTargetPageId="22" captionText="Figure 18. (af) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Eyes, dorsal view. (b) Labium, ventral view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. (f) Spermathecae, ventral view. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747289" httpUri="https://zenodo.org/record/4747289/files/figure.png" pageId="31" pageNumber="1957">Figure 18e, f</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="3DD26518AA5BCA1F1AA6CD53FE06678A" lastPageId="33" lastPageNumber="1959" pageId="31" pageNumber="1957" type="description">
<paragraph id="75773693AA5BCA211AA6CD53FE806616" blockId="31.[160,1156,1113,1727]" box="[160,294,1113,1138]" pageId="31" pageNumber="1957">
<emphasis id="47BCEA81AA5BCA211AA6CD53FE806616" bold="true" box="[160,294,1113,1138]" italics="true" pageId="31" pageNumber="1957">Description</emphasis>
</paragraph>
<paragraph id="75773693AA5BCA211AA6CD77FBD966D3" blockId="31.[160,1156,1113,1727]" pageId="31" pageNumber="1957">
<typeStatus id="AA738831AA5BCA211AA6CD77FEA566F1" box="[160,259,1149,1173]" pageId="31" pageNumber="1957" type="holotype">Holotype</typeStatus>
female (MUSMENT 0508676): Total length (not including chelicerae and spinnerets) 36.9. Carapace: length, 15.7; width 13.6. Abdomen: length 20.2. Fovea: procurved, width 2.1.
</paragraph>
<paragraph id="75773693AA5BCA211AC6CDC8FBD96699" blockId="31.[160,1156,1113,1727]" pageId="31" pageNumber="1957">
Colour pattern (live specimen): carapace dark, with pink setae on margins; legs dark with stripes on patellae distinct; abdomen black with red setae on urticating patch (
<figureCitation id="EDF32A16AA5BCA2119F5CDEFFBD56699" box="[1011,1139,1253,1277]" captionStart="Figure 2" captionStartId="6.[160,225,1289,1312]" captionTargetBox="[163,1152,142,1258]" captionTargetId="figure-139@6.[163,1152,142,1258]" captionTargetPageId="6" captionText="Figure 2. (af) Hapalotremus spp. Habitus. (a, b) Hapalotremus carabaya sp. nov., female holotype. (c, d) Hapalotremus vilcanota sp. nov., female. (e) Hapalotremus vilcanota sp. nov., female and juvenile (white arrow) found under the same stone. (f) Hapalotremus vilcanota sp. nov. burrow. Photos: a, b, df: Tracie Seimon; c: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747253" httpUri="https://zenodo.org/record/4747253/files/figure.png" pageId="31" pageNumber="1957">Figure 2a, b</figureCitation>
).
</paragraph>
<paragraph id="75773693AA5BCA211AC6CC0DFB9D6476" blockId="31.[160,1156,1113,1727]" pageId="31" pageNumber="1957">
Eyes: anterior eye row procurved, posterior recurved (
<figureCitation id="EDF32A16AA5BCA211945CC0DFC66677B" box="[835,960,1287,1311]" captionStart="Figure 18" captionStartId="22.[160,225,1143,1166]" captionTargetBox="[163,1152,142,1112]" captionTargetId="figure-198@22.[163,1152,142,1112]" captionTargetPageId="22" captionText="Figure 18. (af) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Eyes, dorsal view. (b) Labium, ventral view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. (f) Spermathecae, ventral view. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747289" httpUri="https://zenodo.org/record/4747289/files/figure.png" pageId="31" pageNumber="1957">Figure 18a</figureCitation>
). Eyes sizes and interdistances: AME 0.19, ALE 0.23, PME 0.28, PLE 0.24, AMEAME 0.30, AMEALE 0.18, PMEPME 0.91, PMEPLE 0.11, ALEPLE 0.24. Ocular tubercle: length, 1.60; width, 1.75. Clypeus: length, 0.25. Labium: length, 2.5; width, 3.0, with 16 cuspules (
<figureCitation id="EDF32A16AA5BCA2119BACC65FB9967E3" box="[956,1087,1391,1415]" captionStart="Figure 17" captionStartId="21.[160,225,1303,1326]" captionTargetBox="[168,1151,142,1272]" captionTargetId="figure-66@21.[163,1152,142,1272]" captionTargetPageId="21" captionText="Figure 17. (ad) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. Scale bar: 5 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747287" httpUri="https://zenodo.org/record/4747287/files/figure.png" pageId="31" pageNumber="1957">Figures 17b</figureCitation>
,
<figureCitation id="EDF32A16AA5BCA211E4ACC65FBD067E3" box="[1100,1142,1391,1415]" captionStart="Figure 18" captionStartId="22.[160,225,1143,1166]" captionTargetBox="[163,1152,142,1112]" captionTargetId="figure-198@22.[163,1152,142,1112]" captionTargetPageId="22" captionText="Figure 18. (af) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Eyes, dorsal view. (b) Labium, ventral view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. (f) Spermathecae, ventral view. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747289" httpUri="https://zenodo.org/record/4747289/files/figure.png" pageId="31" pageNumber="1957">18b</figureCitation>
). Maxillae: right with 112 cuspules, left with 108 cuspules. Sternum: length, 6.4; width, 5.8. Three pairs of sternal circular sigillae, first pair hardly visible, second and third pairs visible, third pair about its length from the margin (
<figureCitation id="EDF32A16AA5BCA2118E2CCDDFCFB678B" box="[740,861,1495,1519]" captionStart="Figure 17" captionStartId="21.[160,225,1303,1326]" captionTargetBox="[168,1151,142,1272]" captionTargetId="figure-66@21.[163,1152,142,1272]" captionTargetPageId="21" captionText="Figure 17. (ad) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. Scale bar: 5 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747287" httpUri="https://zenodo.org/record/4747287/files/figure.png" pageId="31" pageNumber="1957">Figure 17b</figureCitation>
). Chelicerae: with 13 welldeveloped teeth on promargin of furrow and 19 small teeth on the retromargin.
</paragraph>
<paragraph id="75773693AA5BCA211AC6CF17FDEC64DB" blockId="31.[160,1156,1113,1727]" pageId="31" pageNumber="1957">Leg formula: IV&gt; I&gt; II&gt; III. Length of legs and palpal segments (femur, patella, tibia, metatarsus, tarsus, total): I: 11.3, 6.8, 8.0, 6.8, 4.6, 37.5; II: 9.6, 5.8, 6.5, 6.1, 4.5, 32.5; III: 8.3, 5.3, 5.8, 7.8, 5.1, 32.3; IV: 10.7, 6.5, 8.2, 11.0, 5.5, 41.9; palp: 8.0, 5.2, 6.1, , 5.8, 25.1. Field of spiniform setae on basal retrolateral coxal face of palp and legs IIV. Spinnerets: PMS, 1.5 long; PLS, three-segmented, 5.3 long.</paragraph>
<caption id="21B7661BAA64CA1E1AA6CDBCFE75676C" ID-DOI="http://doi.org/10.5281/zenodo.4747317" ID-Zenodo-Dep="4747317" httpUri="https://zenodo.org/record/4747317/files/figure.png" pageId="32" pageNumber="1958" startId="32.[160,225,1206,1229]" targetBox="[163,1152,142,1175]" targetPageId="32">
<paragraph id="75773693AA64CA1E1AA6CDBCFE75676C" blockId="32.[160,1156,1206,1288]" pageId="32" pageNumber="1958">
<emphasis id="47BCEA81AA64CA1E1AA6CDBCFEAD66A9" bold="true" box="[160,267,1206,1229]" pageId="32" pageNumber="1958">Figure 28.</emphasis>
(ad)
<taxonomicName id="B2C84D10AA64CA1E1B54CDBCFDED66A9" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[338,587,1206,1229]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="1958" phylum="Arthropoda" rank="species" species="marcapata" status="sp. nov.">
<emphasis id="47BCEA81AA64CA1E1B54CDBCFDED66A9" box="[338,587,1206,1229]" italics="true" pageId="32" pageNumber="1958">Hapalotremus marcapata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C8F57FAAA64CA1E1850CDBCFD0C66A9" box="[598,682,1206,1229]" pageId="32" pageNumber="1958" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype (MUBI). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. Scale bar: 5 mm. Photos: Nelson Ferretti.
</paragraph>
</caption>
<paragraph id="75773693AA64CA1E1AC6CC47FE436476" blockId="32.[160,1156,1357,1693]" pageId="32" pageNumber="1958">Scopulae: Ventrally, all tarsi densely scopulate and divided by conical long setae increasing in number from tarsus I to IV. Metatarsi I 1/2 scopulate divided by one row of setae, II 1/2 scopulate divided by three rows of setae, III 1/4 scopulate divided by more than three rows of setae, IV with no scopulae. Dorsal face of all tarsi with two parallel rows of non-plumose clavate trichobothria interspersed with filiform, separated by a row of long, thin setae.</paragraph>
<paragraph id="75773693AA64CA1E1AC6CF17FEEA64F9" blockId="32.[160,1156,1357,1693]" pageId="32" pageNumber="1958">Chaetotaxy (left side): Femora: palp 0; I 2d; II 0; III 0; IV 0. Patellae: palp and legs IIV 0. Tibiae: palp 1-2v; I 1v; II 1-2v, 1-1-1p, 1-1r; III 2-1-2v, 1-1-1p, 1-1r; IV 1-2-2v, 1p, 1-1-1r. Metatarsi: I 1-1v; II 1-2v, 1r; III 2d, 1-1-2v, 1-2-2-2-2p, 1-1-1r; IV 2d, 2-1-3v, 1-1-1-2p, 1-1-1- 1r. Tarsi: IIV 0.</paragraph>
<caption id="21B7661BAA65CA1F1AA6CD80FD1D66B8" ID-DOI="http://doi.org/10.5281/zenodo.4747319" ID-Zenodo-Dep="4747319" httpUri="https://zenodo.org/record/4747319/files/figure.png" pageId="33" pageNumber="1959" startId="33.[160,225,1162,1185]" targetBox="[163,1152,142,1131]" targetPageId="33">
<paragraph id="75773693AA65CA1F1AA6CD80FD1D66B8" blockId="33.[160,1157,1162,1244]" pageId="33" pageNumber="1959">
<emphasis id="47BCEA81AA65CA1F1AA6CD80FEA166C5" bold="true" box="[160,263,1162,1185]" pageId="33" pageNumber="1959">Figure 29.</emphasis>
(ae)
<taxonomicName id="B2C84D10AA65CA1F1B4ECD80FD9A66C5" authority="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West, 2018" authorityName="Ferretti &amp; Cavallo &amp; Chaparro &amp; Ríos-Tamayo &amp; Seimon &amp; West" authorityYear="2018" box="[328,572,1162,1185]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="1959" phylum="Arthropoda" rank="species" species="marcapata" status="sp. nov.">
<emphasis id="47BCEA81AA65CA1F1B4ECD80FD9A66C5" box="[328,572,1162,1185]" italics="true" pageId="33" pageNumber="1959">Hapalotremus marcapata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C8F57FAAA65CA1F1845CD80FD3566C5" box="[579,659,1162,1185]" pageId="33" pageNumber="1959" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype (MUBI). (a) Labium, ventral view. (b) Eyes, dorsal view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. Scale bar: 1 mm. Photos: Nelson Ferretti.
</paragraph>
</caption>
<paragraph id="75773693AA65CA1F1AC6CC23FC7267CD" blockId="33.[160,1156,1321,1518]" pageId="33" pageNumber="1959">
Genitalia: Spermathecal receptacle with basal portion much wider than apical; apical median region rounded and apical lateral projections slightly developed; presence of an additional well-developed apical median projection with a long thin wavy groove extending downward dorsal median from this projection (
<figureCitation id="EDF32A16AA65CA1F193FCC9BFC6167CD" box="[825,967,1425,1449]" captionStart="Figure 18" captionStartId="22.[160,225,1143,1166]" captionTargetBox="[163,1152,142,1112]" captionTargetId="figure-198@22.[163,1152,142,1112]" captionTargetPageId="22" captionText="Figure 18. (af) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Eyes, dorsal view. (b) Labium, ventral view. (c) Tarsus IV, ventral view. (d) Tarsus IV, prolateral view. (e) Spermathecae, dorsal view. (f) Spermathecae, ventral view. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747289" httpUri="https://zenodo.org/record/4747289/files/figure.png" pageId="33" pageNumber="1959">Figure 18e, f</figureCitation>
).
</paragraph>
<paragraph id="75773693AA65CA1F1AC6CCBEFE06678A" blockId="33.[160,1156,1321,1518]" pageId="33" pageNumber="1959">
Urticating setae:
<typeStatus id="AA738831AA65CA1F1B78CCBEFE1267A8" box="[382,436,1460,1484]" pageId="33" pageNumber="1959">Type</typeStatus>
III present and arranged in one big dorsal median patch on the abdomen (
<figureCitation id="EDF32A16AA65CA1F1B1BCCDCFE35678A" box="[285,403,1494,1518]" captionStart="Figure 17" captionStartId="21.[160,225,1303,1326]" captionTargetBox="[168,1151,142,1272]" captionTargetId="figure-66@21.[163,1152,142,1272]" captionTargetPageId="21" captionText="Figure 17. (ad) Hapalotremus carabaya sp. nov., female holotype (MUSM-ENT 0508676). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. Scale bar: 5 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747287" httpUri="https://zenodo.org/record/4747287/files/figure.png" pageId="33" pageNumber="1959">Figure 17c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DD26518AA65CA1F1AA6CF16FC0164DB" pageId="33" pageNumber="1959" type="distribution">
<paragraph id="75773693AA65CA1F1AA6CF16FE656451" blockId="33.[160,1155,1564,1727]" box="[160,451,1564,1589]" pageId="33" pageNumber="1959">
<emphasis id="47BCEA81AA65CA1F1AA6CF16FE656451" bold="true" box="[160,451,1564,1589]" italics="true" pageId="33" pageNumber="1959">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="75773693AA65CA1F1AA6CF35FC0164DB" blockId="33.[160,1155,1564,1727]" pageId="33" pageNumber="1959">
Known only from
<typeStatus id="AA738831AA65CA1F1B61CF35FE3E6433" box="[359,408,1599,1623]" pageId="33" pageNumber="1959">type</typeStatus>
locality (
<figureCitation id="EDF32A16AA65CA1F1BFACF35FDC56433" box="[508,611,1599,1623]" captionStart="Figure 37" captionStartId="41.[160,225,846,869]" captionTargetBox="[163,1152,142,815]" captionTargetId="figure-275@41.[163,1152,142,815]" captionTargetPageId="41" captionText="Figure 37. Distribution map of Hapalotremus species from Peru." figureDoi="http://doi.org/10.5281/zenodo.4747338" httpUri="https://zenodo.org/record/4747338/files/figure.png" pageId="33" pageNumber="1959">Figure 37</figureCitation>
). Specimens were found on a hill under different rocks in close proximity to each other at about
<quantity id="B2309B76AA65CA1F18B9CF68FCB2641E" box="[703,788,1634,1658]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="1959" unit="m" value="4000.0">4000 m</quantity>
a.s.l. (
<figureCitation id="EDF32A16AA65CA1F195DCF68FC7B641E" box="[859,989,1634,1658]" captionStart="Figure 3" captionStartId="7.[160,225,1665,1688]" captionTargetBox="[163,1152,142,1634]" captionTargetId="figure-1@7.[163,1152,142,1634]" captionTargetPageId="7" captionText="Figure 3. (aj) Hapalotremus spp. habitat. (a) Escoipe River, next to route 33, Salta Province, Argentina, type locality of Hapalotremus martinorum. (b, c) Los Sosa River, next to route 307, Tucumán Province, Argentina, habitat of Hapalotremus chasqui sp. nov. (d, f) Cordillera Carabaya, Michi Michani Cunca, type locality of Hapalotremus carabaya sp. nov. (e, gj) Cordillera Vilcanota, Upis, type locality of Hapalotremus vilcanota sp. nov. Photos: a: Patricio Cavallo;b: Gabriel Pompozzi; c: Laura Montes de Oca; di: Tracie Seimon; j: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747255" httpUri="https://zenodo.org/record/4747255/files/figure.png" pageId="33" pageNumber="1959">Figure 3d, f</figureCitation>
). Spiders were located at 2:02 pm under misty weather conditions with drizzle. Day temperature on
<date id="01761053AA65CA1F1E6ECF8FFEB464DB" pageId="33" pageNumber="1959" value="2006-06-30">30 June 2006</date>
was 8°C but dropped to an overnight low of about 3.3°C.
</paragraph>
</subSubSection>
</treatment>
</document>