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<document id="29ACCB8A37EE27B85C3F146B48F05561" ID-DOI="10.1080/00222933.2018.1506521" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747249" ID-ZooBank="1C028676-CCD9-4654-A755-6ACA3BCE3E98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670908261" checkinUser="carolina" docAuthor="Ferretti, Nelson, Cavallo, Patricio, Chaparro, Juan C., Ríos-Tamayo, Duniesky, Seimon, Tracie A. &amp; West, Rick" docDate="2018" docId="FD618785AA4ACA2C1BBACC30FC6567BD" docLanguage="en" docName="JNATHIST.52.29-30.1927-1984.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (29 - 30)" docSource="http://dx.doi.org/10.1080/00222933.2018.1506521" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hapalotremus albipes : Simon 1903" docType="treatment" docVersion="3" lastPageNumber="1943" masterDocId="0158FFFDAA44CA3E1A06C90AFFA66264" masterDocTitle="The Neotropical genus Hapalotremus Simon, 1903 (Araneae: Theraphosidae), with the description of seven new species and the highest altitude record for the family" masterLastPageNumber="1984" masterPageNumber="1927" pageNumber="1940" updateTime="1698914576834" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="80901030A7C71C5B5A2FEEF5825BCFDB">The Neotropical genus Hapalotremus Simon, 1903 (Araneae: Theraphosidae), with the description of seven new species and the highest altitude record for the family</mods:title>
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<mods:namePart id="8382116DF9567F4A6AE4F8646E6D6528">Chaparro, Juan C.</mods:namePart>
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<taxonomicName id="B2C84D10AA4ACA301BBACC30FCC06737" ID-CoL="3JKLP" authority="Simon, 1903" authorityName=": Simon" authorityYear="1903" box="[444,870,1338,1363]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1940" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="47BCEA81AA4ACA301BBACC30FD6C6737" bold="true" box="[444,714,1338,1363]" italics="true" pageId="14" pageNumber="1940">Hapalotremus albipes</emphasis>
<bibRefCitation id="11594B62AA4ACA3018D2CC30FCC06737" author="Simon E" box="[724,870,1338,1363]" pageId="14" pageNumber="1940" pagination="669 - 1080" refId="ref24980" refString="Simon E. 1903. Histoire naturelle des araignees. Deuxieme Edition Paris (Roret). 2: 669 - 1080." type="journal article" year="1903">Simon, 1903</bibRefCitation>
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(
<figureCitation id="EDF32A16AA4ACA301827CC57FD3A6712" box="[545,668,1373,1398]" captionStart="Figure 1" captionStartId="4.[160,225,1268,1291]" captionTargetBox="[163,1152,142,1237]" captionTargetId="figure-72@4.[163,1152,142,1237]" captionTargetPageId="4" captionText="Figure 1. (af) Hapalotremus spp. habitus. (a) Hapalotremus albipes, female. (b) Hapalotremus major, female. (c, d) Hapalotremus martinorum. (c) Male. (d) Female. (e) Hapalotremus chasqui sp. nov., juvenile. (f) Hapalotremus marcapata sp. nov., female. Photos: a: Mauricio Pacheco Suarez; b: Juan Chaparro; c, d: Patricio Cavallo; e: Laura Montes de Oca; f: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747251" httpUri="https://zenodo.org/record/4747251/files/figure.png" pageId="14" pageNumber="1940">Figures 1a</figureCitation>
,
<figureCitation id="EDF32A16AA4ACA3018ADCC57FD706712" box="[683,726,1373,1398]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="11.[160,225,1453,1476]" captionStartId-1="12.[160,225,985,1008]" captionStartId-2="13.[160,225,1252,1275]" captionTargetBox-0="[163,1152,142,1423]" captionTargetBox-1="[163,1153,142,952]" captionTargetBox-2="[163,1152,143,1221]" captionTargetId-0="figure-52@11.[163,1152,142,1423]" captionTargetId-1="figure-272@12.[163,1153,142,954]" captionTargetId-2="figure-131@13.[163,1152,142,1221]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." captionText-1="Figure 8. (ad) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). Male left palpal bulb. (a) Prolateral view. (b) Retrolateral view. (c) Dorsal view. (d) Ventral view. A, apical keel; DA, digitiform apophysis; PI, prolateral inferior keel; PS, prolateral superior keel. Scale bar: 1 mm. Photos: Nelson Ferretti." captionText-2="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi-0="http://doi.org/10.5281/zenodo.4747267" figureDoi-1="http://doi.org/10.5281/zenodo.4747269" figureDoi-2="http://doi.org/10.5281/zenodo.4747271" httpUri-0="https://zenodo.org/record/4747267/files/figure.png" httpUri-1="https://zenodo.org/record/4747269/files/figure.png" httpUri-2="https://zenodo.org/record/4747271/files/figure.png" pageId="14" pageNumber="1940">79</figureCitation>
,
<figureCitation id="EDF32A16AA4ACA3018E3CC57FCA46712" box="[741,770,1373,1398]" captionStart="Figure 36" captionStartId="40.[160,225,871,894]" captionTargetBox="[163,1152,143,840]" captionTargetId="figure-287@40.[163,1152,142,841]" captionTargetPageId="40" captionText="Figure 36. Distribution map of Hapalotremus species from Argentina and Bolivia." figureDoi="http://doi.org/10.5281/zenodo.4747336" httpUri="https://zenodo.org/record/4747336/files/figure.png" pageId="14" pageNumber="1940">36</figureCitation>
)
</paragraph>
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<treatmentCitation id="F4691082AA4ACA301AA6CCABFDC367DD" author="Simon E" box="[160,613,1441,1465]" page="930" pageId="14" pageNumber="1940" year="1903">
<taxonomicName id="B2C84D10AA4ACA301AA6CCABFDC367DD" ID-CoL="3JKLP" authority=": Simon, 1903: 930" authorityName=": Simon" authorityPageNumber="930" authorityYear="1903" box="[160,613,1441,1465]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="1940" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="47BCEA81AA4ACA301AA6CCABFE2867DD" box="[160,398,1441,1465]" italics="true" pageId="14" pageNumber="1940">Hapalotremus albipes</emphasis>
:
<bibRefCitation id="11594B62AA4ACA301B9BCCABFDC367DD" author="Simon E" box="[413,613,1441,1465]" pageId="14" pageNumber="1940" pagination="669 - 1080" refId="ref24980" refString="Simon E. 1903. Histoire naturelle des araignees. Deuxieme Edition Paris (Roret). 2: 669 - 1080." type="journal article" year="1903">Simon, 1903: 930</bibRefCitation>
</taxonomicName>
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, fig. 1083.
<treatmentCitation id="F4691082AA4ACA3018E4CCABFEA767BF" author="Gerschman de Pikelin BS &amp; Schiapelli RD" page="72" pageId="14" pageNumber="1940" year="1973">
<bibRefCitation id="11594B62AA4ACA3018E4CCABFEA767BF" author="Gerschman de Pikelin BS &amp; Schiapelli RD" pageId="14" pageNumber="1940" pagination="43 - 77" refId="ref23474" refString="Gerschman de Pikelin BS, Schiapelli RD. 1973. La subfamilia Ischnocolinae (Araneae: Theraphosidae). Rev Mus Argent Cienc Nat. 4: 43 - 77." type="journal article" year="1973">Gerschman De Pikelín and Schiapelli 1973: 72</bibRefCitation>
</treatmentCitation>
, figs 6367.
<treatmentCitation id="F4691082AA4ACA301B8DCCC9FDE967BF" author="Schmidt G" box="[395,591,1475,1499]" page="41" pageId="14" pageNumber="1940" year="1986">
<bibRefCitation id="11594B62AA4ACA301B8DCCC9FDE967BF" author="Schmidt G" box="[395,591,1475,1499]" pageId="14" pageNumber="1940" pagination="1 - 87" refId="ref24633" refString="Schmidt G. 1986. Vogelspinnen: lebensweise, Bestimmungsschlussel, Haltung und Zucht. Minden: Albrecht Philler; p. 1 - 87." type="book chapter" year="1986">Schmidt 1986: 41</bibRefCitation>
</treatmentCitation>
, fig. 12.
<treatmentCitation id="F4691082AA4ACA3018ADCCC9FCD667BF" author="Schmidt G" box="[683,880,1475,1499]" page="10" pageId="14" pageNumber="1940" year="1993">
<bibRefCitation id="11594B62AA4ACA3018ADCCC9FCD667BF" author="Schmidt G" box="[683,880,1475,1499]" pageId="14" pageNumber="1940" pagination="10 - 14" refId="ref24659" refString="Schmidt G. 1993. Das Weibchen von Hapalotremus albipes Simon 1903 (Araneida: theraphosidae). Arachnologischer Anzeiger. 4: 10 - 14." type="journal article" year="1993">Schmidt 1993: 10</bibRefCitation>
</treatmentCitation>
, figs 16.
<treatmentCitation id="F4691082AA4ACA3019D8CCC9FF18679A" author="Schmidt G" page="66" pageId="14" pageNumber="1940" year="1993">
<bibRefCitation id="11594B62AA4ACA3019D8CCC9FF18679A" author="Schmidt G" pageId="14" pageNumber="1940" pagination="10 - 14" refId="ref24659" refString="Schmidt G. 1993. Das Weibchen von Hapalotremus albipes Simon 1903 (Araneida: theraphosidae). Arachnologischer Anzeiger. 4: 10 - 14." type="journal article" year="1993">Schmidt, 1993: 66</bibRefCitation>
</treatmentCitation>
, figs 101103.
<treatmentCitation id="F4691082AA4ACA301B6DCCECFD3D679A" author="Perez-Miles F &amp; Lucas SM &amp; Jpi DS &amp; Bertani R" box="[363,667,1510,1534]" page="52" pageId="14" pageNumber="1940" year="1996">
<bibRefCitation id="11594B62AA4ACA301B6DCCECFD3D679A" author="Perez-Miles F &amp; Lucas SM &amp; Jpi DS &amp; Bertani R" box="[363,667,1510,1534]" pageId="14" pageNumber="1940" pagination="33 - 68" refId="ref24187" refString="Perez-Miles F, Lucas SM, Jpi DS, Bertani R. 1996. Systematic revision and cladistic analysis of Theraphosinae (Araneae: theraphosidae). Mygalomorph. 1: 33 - 68." type="journal article" year="1996">Pérez-Miles et al. 1996: 52</bibRefCitation>
</treatmentCitation>
, fig. 23.
<treatmentCitation id="F4691082AA4ACA3018F8CCECFC68679A" author="Schmidt G" box="[766,974,1510,1534]" page="16" pageId="14" pageNumber="1940" year="1997">
<bibRefCitation id="11594B62AA4ACA3018F8CCECFC68679A" author="Schmidt G" box="[766,974,1510,1534]" pageId="14" pageNumber="1940" pagination="1 - 27" refId="ref24686" refString="Schmidt G. 1997. Bestimmungsschlussel fur die Gattungen der Unterfamilie Theraphosinae (Araneae: Theraphosidae). Arachnologisches Magazin. 3: 1 - 27." type="journal article" year="1997">Schmidt, 1997: 16</bibRefCitation>
</treatmentCitation>
, figs 7679.
<treatmentCitation id="F4691082AA4ACA301E59CCECFEA46444" author="Vol F" page="11" pageId="14" pageNumber="1940" year="1999">
<bibRefCitation id="11594B62AA4ACA301E59CCECFEA46444" author="Vol F" pageId="14" pageNumber="1940" pagination="1 - 13" refId="ref25218" refString="Vol F. 1999. A propos d' une spermatheque inhabituelle. Arachnides. 42: 1 - 13." type="journal article" year="1999">Vol 1999: 11</bibRefCitation>
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, fig. K.
<treatmentCitation id="F4691082AA4ACA301B51CF02FD8A6444" author="Schmidt G" box="[343,556,1544,1568]" page="127" pageId="14" pageNumber="1940" year="2003">
<bibRefCitation id="11594B62AA4ACA301B51CF02FD8A6444" author="Schmidt G" box="[343,556,1544,1568]" pageId="14" pageNumber="1940" pagination="1 - 383" refId="ref24743" refString="Schmidt G. 2003. Die Vogelspinnen: eine weltweite Ubersicht. Hohenwarsleben: Neue Brehm- Bucherei; p. 1 - 383." type="book chapter" year="2003">Schmidt 2003: 127</bibRefCitation>
</treatmentCitation>
, figs 111114.
<treatmentCitation id="F4691082AA4ACA3018D0CF02FB886444" author="Cavallo P &amp; Ferretti N" box="[726,1070,1544,1568]" page="878" pageId="14" pageNumber="1940" year="2015">
<bibRefCitation id="11594B62AA4ACA3018D0CF02FB886444" author="Cavallo P &amp; Ferretti N" box="[726,1070,1544,1568]" pageId="14" pageNumber="1940" pagination="873 - 887" refId="ref23215" refString="Cavallo P, Ferretti N. 2015. The first Hapalotremus Simon, 1903 (Araneae: Theraphosidae) from Argentina: description and natural history of Hapalotremus martinorum sp. nov. J Nat Hist. 49 (15 - 16): 873 - 887." type="journal article" year="2015">Cavallo and Ferretti 2015: 878</bibRefCitation>
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, fig. 3c.
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<emphasis id="47BCEA81AA4ACA301AA6CF6BFEE3641E" bold="true" box="[160,325,1633,1658]" italics="true" pageId="14" pageNumber="1940">Type material</emphasis>
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<paragraph id="75773693AA4ACA301AA6CF8FFEDE64DB" blockId="14.[160,1156,1633,1727]" pageId="14" pageNumber="1940">
<materialsCitation id="C5A03CCEAA4ACA301AA6CF8FFD3764F9" box="[160,657,1669,1693]" collectionCode="MNHN" country="Bolivia" location="Chaco" pageId="14" pageNumber="1940" specimenCount="1" typeStatus="holotype">
Male
<typeStatus id="AA738831AA4ACA301AE6CF8FFEE264F9" box="[224,324,1669,1693]" pageId="14" pageNumber="1940" type="holotype">holotype</typeStatus>
(
<collectionCode id="13D9AE56AA4ACA301B52CF8FFE0764F9" box="[340,417,1669,1693]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="1940">MNHN</collectionCode>
) from
<collectingCountry id="0DDF7603AA4ACA301BF6CF8FFD9A64F9" box="[496,572,1669,1693]" name="Bolivia" pageId="14" pageNumber="1940">Bolivia</collectingCountry>
,
<location id="70176048AA4ACA30184FCF8FFD3764F9" LSID="urn:lsid:plazi:treatment:FD618785AA4ACA2C1BBACC30FC6567BD:70176048AA4ACA30184FCF8FFD3764F9" box="[585,657,1669,1693]" country="Bolivia" name="Chaco" pageId="14" pageNumber="1940">Chaco</location>
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. Male
<typeStatus id="AA738831AA4ACA3018D8CF8FFCE464F9" box="[734,834,1669,1693]" pageId="14" pageNumber="1940" type="holotype">holotype</typeStatus>
not examined, only photos provided by F. Vol.
</paragraph>
<caption id="21B7661BAA4BCA311AA6CC97FBA96468" ID-DOI="http://doi.org/10.5281/zenodo.4747275" ID-Zenodo-Dep="4747275" httpUri="https://zenodo.org/record/4747275/files/figure.png" pageId="15" pageNumber="1941" startId="15.[160,225,1437,1460]" targetBox="[163,1152,142,1406]" targetPageId="15">
<paragraph id="75773693AA4BCA311AA6CC97FBA96468" blockId="15.[160,1156,1437,1548]" pageId="15" pageNumber="1941">
<emphasis id="47BCEA81AA4BCA311AA6CC97FEAC67D0" bold="true" box="[160,266,1437,1460]" pageId="15" pageNumber="1941">Figure 11.</emphasis>
(ad)
<taxonomicName id="B2C84D10AA4BCA311B49CC97FDB167D0" baseAuthorityName="Chamberlin" baseAuthorityYear="1916" box="[335,535,1437,1460]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="1941" phylum="Arthropoda" rank="species" species="major">
<emphasis id="47BCEA81AA4BCA311B49CC97FDB167D0" box="[335,535,1437,1460]" italics="true" pageId="15" pageNumber="1941">Hapalotremus major</emphasis>
</taxonomicName>
, male holotype (MCZ 142). (a, b) Male right palpal bulb. (a) Retrolateral view. (b) Prolateral view. (c. d) Palpal tibia. (c) Palpal tibia, retrolateral view. (d) Palpal tibia, ventral view. Arrows indicate the rounded process. A, apical keel; DA, digitiform apophysis; PI, prolateral inferior keel; PS, prolateral superior keel. Scale bar: 1 mm. Photos: Rick West.
</paragraph>
</caption>
<paragraph id="75773693AA4BCA311AA6CF30FDA36437" blockId="15.[160,1156,1594,1723]" box="[160,517,1594,1619]" pageId="15" pageNumber="1941">
<emphasis id="47BCEA81AA4BCA311AA6CF30FDA36437" bold="true" box="[160,517,1594,1619]" italics="true" pageId="15" pageNumber="1941">Additional material examined</emphasis>
</paragraph>
<paragraph id="75773693AA4BCA311AA6CF57FB2564DF" blockId="15.[160,1156,1594,1723]" pageId="15" pageNumber="1941">
<materialsCitation id="C5A03CCEAA4BCA311AA6CF57FDCE6411" accessionNumber="SMF37093" box="[160,616,1629,1653]" collectionCode="SMF" collectorName="Three" country="Bolivia" location="Bolivia" pageId="15" pageNumber="1941" specimenCount="1">
<collectorName id="D83D5345AA4BCA311AA6CF57FF7B6411" box="[160,221,1629,1653]" pageId="15" pageNumber="1941">Three</collectorName>
females (
<collectionCode id="13D9AE56AA4BCA311B41CF57FEDF6411" box="[327,377,1629,1653]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="15" pageNumber="1941">SMF</collectionCode>
37093) from
<collectingCountry id="0DDF7603AA4BCA311802CF57FDEB6411" box="[516,589,1629,1653]" name="Bolivia" pageId="15" pageNumber="1941">Bolivia</collectingCountry>
, P
</materialsCitation>
. Klass col, only illustrations provided by F.
<materialsCitation id="C5A03CCEAA4BCA311E47CF57FB2664FC" collectionCode="MHNC" collectorName="Miles" country="Bolivia" county="Murillo" elevation="3600" location="Unduavi" municipality="Pongo" pageId="15" pageNumber="1941" specimenCount="1" specimenCount-female="1" stateProvince="La Paz">
Pérez-
<collectorName id="D83D5345AA4BCA311AA6CF8AFF7C64FC" box="[160,218,1664,1688]" pageId="15" pageNumber="1941">Miles</collectorName>
;
<specimenCount id="63CEFD1AAA4BCA311AE7CF8AFEFE64FC" box="[225,344,1664,1688]" pageId="15" pageNumber="1941" type="female">one female</specimenCount>
(
<collectionCode id="13D9AE56AA4BCA311B64CF8AFE0F64FC" box="[354,425,1664,1688]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="15" pageNumber="1941">MHNC</collectionCode>
B) from
<collectingCountry id="0DDF7603AA4BCA31180ECF8AFDF764FC" box="[520,593,1664,1688]" name="Bolivia" pageId="15" pageNumber="1941">Bolivia</collectingCountry>
,
<collectingRegion id="B70CF871AA4BCA31185ECF8AFD3B64FC" box="[600,669,1664,1688]" country="Bolivia" name="La Paz" pageId="15" pageNumber="1941">La Paz</collectingRegion>
,
<collectingCounty id="9C164E1FAA4BCA3118A2CF8AFD5764FC" box="[676,753,1664,1688]" pageId="15" pageNumber="1941">Murillo</collectingCounty>
,
<collectingMunicipality id="9513ACE9AA4BCA3118FECF8AFCE764FC" box="[760,833,1664,1688]" pageId="15" pageNumber="1941">Pongo</collectingMunicipality>
,
<quantity id="B2309B76AA4BCA31194FCF8AFCDB64FC" box="[841,893,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="1941" unit="km" value="3.0">3 km</quantity>
from
<location id="70176048AA4BCA3119BCCF8AFBB164FC" LSID="urn:lsid:plazi:treatment:FD618785AA4ACA2C1BBACC30FC6567BD:70176048AA4BCA3119BCCF8AFBB164FC" box="[954,1047,1664,1688]" country="Bolivia" county="Murillo" municipality="Pongo" name="Unduavi" pageId="15" pageNumber="1941" stateProvince="La Paz">Unduavi</location>
,
<quantity id="B2309B76AA4BCA311E1BCF8AFBCB64FC" box="[1053,1133,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="1941" unit="m" value="3600.0">
<elevation id="FEE5D1A0AA4BCA311E1BCF8AFBCB64FC" box="[1053,1133,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="1941" unit="m" value="3600.0">3600 m</elevation>
</quantity>
a
</materialsCitation>
<materialsCitation id="C5A03CCEAA4BCA311E86CF8AFF1664DF" collectionCode="MHNC" collectorName="Miles" country="Bolivia" county="Murillo" elevation="3600" isTailEnumeration="true" location="Unduavi" municipality="Pongo" pageId="15" pageNumber="1941" specimenCount="1" stateProvince="La Paz">. s</materialsCitation>
.l.,
<date id="01761053AA4BCA311AC1CFA9FEE064DF" box="[199,326,1699,1723]" pageId="15" pageNumber="1941" value="1991-07-07">7 July 1991</date>
, E
<materialsCitation id="C5A03CCEAA4BCA311B67CFA9FDAC64DF" box="[353,522,1699,1723]" collectionCode="MHNC" collectorName="Penaranda" country="Bolivia" county="Murillo" elevation="3600" isTailEnumeration="true" location="Unduavi" municipality="Pongo" pageId="15" pageNumber="1941" specimenCount="1" stateProvince="La Paz">
.
<collectorName id="D83D5345AA4BCA311B6ACFA9FE7964DF" box="[364,479,1699,1723]" pageId="15" pageNumber="1941">Peñaranda</collectorName>
col
</materialsCitation>
<materialsCitation id="C5A03CCEAA4BCA31180DCFA9FB2564DF" box="[523,1155,1699,1723]" collectionCode="MHNC" country="Bolivia" location="Sud Yungas" pageId="15" pageNumber="1941" specimenCount="1" specimenCount-female="1" stateProvince="La Paz">
.;
<specimenCount id="63CEFD1AAA4BCA31181FCFA9FD3964DF" box="[537,671,1699,1723]" pageId="15" pageNumber="1941" type="female">two females</specimenCount>
(
<collectionCode id="13D9AE56AA4BCA3118ABCFA9FD5264DF" box="[685,756,1699,1723]" country="0" name="Museo de Historia Natural de Concepcion (Chile)" pageId="15" pageNumber="1941">MHNC</collectionCode>
B) from
<collectingCountry id="0DDF7603AA4BCA31195CCFA9FC0564DF" box="[858,931,1699,1723]" name="Bolivia" pageId="15" pageNumber="1941">Bolivia</collectingCountry>
,
<collectingRegion id="B70CF871AA4BCA3119A8CFA9FC5164DF" box="[942,1015,1699,1723]" country="Bolivia" name="La Paz" pageId="15" pageNumber="1941">La Paz</collectingRegion>
,
<location id="70176048AA4BCA311E04CFA9FB2564DF" LSID="urn:lsid:plazi:treatment:FD618785AA4ACA2C1BBACC30FC6567BD:70176048AA4BCA311E04CFA9FB2564DF" box="[1026,1155,1699,1723]" country="Bolivia" name="Sud Yungas" pageId="15" pageNumber="1941" stateProvince="La Paz">Sud Yungas</location>
</materialsCitation>
</paragraph>
<caption id="21B7661BAA54CA2E1AA6CA04FEFE6119" ID-DOI="http://doi.org/10.5281/zenodo.4747277" ID-Zenodo-Dep="4747277" httpUri="https://zenodo.org/record/4747277/files/figure.png" pageId="16" pageNumber="1942" startId="16.[160,225,782,805]" targetBox="[163,1152,142,751]" targetPageId="16">
<paragraph id="75773693AA54CA2E1AA6CA04FEFE6119" blockId="16.[160,1156,782,893]" pageId="16" pageNumber="1942">
<emphasis id="47BCEA81AA54CA2E1AA6CA04FEAD6141" bold="true" box="[160,267,782,805]" pageId="16" pageNumber="1942">Figure 12.</emphasis>
(ag)
<taxonomicName id="B2C84D10AA54CA2E1B54CA04FDBD6141" baseAuthorityName="Chamberlin" baseAuthorityYear="1916" box="[338,539,782,805]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1942" phylum="Arthropoda" rank="species" species="major">
<emphasis id="47BCEA81AA54CA2E1B54CA04FDBD6141" box="[338,539,782,805]" italics="true" pageId="16" pageNumber="1942">Hapalotremus major</emphasis>
</taxonomicName>
, female paratype (MCZ 143). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Eyes, dorsal view. (d) Labium and maxillae. (e) Abdomen, dorsal view. (f) Tarsus IV, ventral view. (g) Spermathecae, dorsal view. Scale bars: ab, e: 5 mm; cd, fg: 1 mm. Photos: Rick West.
</paragraph>
</caption>
<paragraph id="75773693AA54CA2E1AA6CAC4FCD8662A" blockId="16.[160,1156,974,1102]" pageId="16" pageNumber="1942">
<materialsCitation id="C5A03CCEAA54CA2E1AA6CAC4FE436182" box="[160,485,974,998]" elevation="3660" location="Unduavi" municipality="Province" pageId="16" pageNumber="1942" specimenCount="1">
<collectingMunicipality id="9513ACE9AA54CA2E1AA6CAC4FF596182" box="[160,255,974,998]" pageId="16" pageNumber="1942">Province</collectingMunicipality>
,
<location id="70176048AA54CA2E1B0ACAC4FECF6182" LSID="urn:lsid:plazi:treatment:FD618785AA4ACA2C1BBACC30FC6567BD:70176048AA54CA2E1B0ACAC4FECF6182" box="[268,361,974,998]" municipality="Province" name="Unduavi" pageId="16" pageNumber="1942">Unduavi</location>
,
<quantity id="B2309B76AA54CA2E1B73CAC4FE6C6182" box="[373,458,974,998]" metricMagnitude="3" metricUnit="m" metricValue="3.66" pageId="16" pageNumber="1942" unit="m" value="3660.0">
<elevation id="FEE5D1A0AA54CA2E1B73CAC4FE6C6182" box="[373,458,974,998]" metricMagnitude="3" metricUnit="m" metricValue="3.66" pageId="16" pageNumber="1942" unit="m" value="3660.0">3660 m</elevation>
</quantity>
a
</materialsCitation>
<materialsCitation id="C5A03CCEAA54CA2E1BE3CAC4FE516182" box="[485,503,974,998]" elevation="3660" isTailEnumeration="true" location="Unduavi" municipality="Province" pageId="16" pageNumber="1942" specimenCount="1">.s</materialsCitation>
.l.,
<date id="01761053AA54CA2E1808CAC4FD7F6182" box="[526,729,974,998]" pageId="16" pageNumber="1942" value="1990-11-01">1 November 1990</date>
, E
<materialsCitation id="C5A03CCEAA54CA2E18F3CAC4FC076182" box="[757,929,974,998]" collectorName="Penaranda" elevation="3660" isTailEnumeration="true" location="Unduavi" municipality="Province" pageId="16" pageNumber="1942" specimenCount="1">
.
<collectorName id="D83D5345AA54CA2E1907CAC4FCD26182" box="[769,884,974,998]" pageId="16" pageNumber="1942">Peñaranda</collectorName>
col
</materialsCitation>
<materialsCitation id="C5A03CCEAA54CA2E19A4CAC4FBC2666D" collectionCode="AMNH" collectorName="Coscapa" country="Bolivia" elevation="3200" location="Oruro" pageId="16" pageNumber="1942" specimenCount="11" specimenCount-female="1" specimenCount-immature="10" stateProvince="Oruro">
.;
<specimenCount id="63CEFD1AAA54CA2E19B4CAC4FB8D6182" box="[946,1067,974,998]" pageId="16" pageNumber="1942" type="female">six females</specimenCount>
and
<specimenCount id="63CEFD1AAA54CA2E1E6ECAC4FEB5666D" pageId="16" pageNumber="1942" type="immature">10 immatures</specimenCount>
(
<collectionCode id="13D9AE56AA54CA2E1B18CAFBFECE666D" box="[286,360,1009,1033]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="1942">AMNH</collectionCode>
) from
<collectingCountry id="0DDF7603AA54CA2E1BAACAFBFE53666D" box="[428,501,1009,1033]" name="Bolivia" pageId="16" pageNumber="1942">Bolivia</collectingCountry>
,
<collectingRegion id="B70CF871AA54CA2E1BFACAFBFD9B666D" box="[508,573,1009,1033]" country="Bolivia" name="Oruro" pageId="16" pageNumber="1942">Oruro</collectingRegion>
, from a point between
<collectorName id="D83D5345AA54CA2E1932CAFBFC28666D" box="[820,910,1009,1033]" pageId="16" pageNumber="1942">Coscapa</collectorName>
to
<collectingRegion id="B70CF871AA54CA2E19A9CAFBFC53666D" box="[943,1013,1009,1033]" country="Bolivia" name="La Paz" pageId="16" pageNumber="1942">La Paz</collectingRegion>
,
<quantity id="B2309B76AA54CA2E19FBCAFBFBEB666D" box="[1021,1101,1009,1033]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="1942" unit="m" value="3200.0">
<elevation id="FEE5D1A0AA54CA2E19FBCAFBFBEB666D" box="[1021,1101,1009,1033]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="1942" unit="m" value="3200.0">3200 m</elevation>
</quantity>
a
</materialsCitation>
<materialsCitation id="C5A03CCEAA54CA2E1E65CAFBFBD3666D" box="[1123,1141,1009,1033]" collectionCode="AMNH" collectorName="Coscapa" country="Bolivia" elevation="3200" isTailEnumeration="true" location="Oruro" pageId="16" pageNumber="1942" specimenCount="1" stateProvince="Oruro">.s</materialsCitation>
.l.,
<date id="01761053AA54CA2E1AA6CD1EFEFB6648" box="[160,349,1044,1068]" pageId="16" pageNumber="1942" value="1984-10-24">24 October 1984</date>
, L
<materialsCitation id="C5A03CCEAA54CA2E1B7ECD1EFE326648" box="[376,404,1044,1068]" collectionCode="AMNH" collectorName="Coscapa" country="Bolivia" elevation="3200" isTailEnumeration="true" location="Oruro" pageId="16" pageNumber="1942" specimenCount="1" stateProvince="Oruro">. E</materialsCitation>
. Peña col
<materialsCitation id="C5A03CCEAA54CA2E1804CD1EFDB3662A" collectionCode="AMNH" country="Bolivia" elevation="3600" latitude="-16.1919" location="Botijlaca" longLatPrecision="7" longitude="-68.1403" municipality="Pacajes" pageId="16" pageNumber="1942" specimenCount="1" specimenCount-male="1" stateProvince="La Paz">
.;
<specimenCount id="63CEFD1AAA54CA2E1814CD1EFDDD6648" box="[530,635,1044,1068]" pageId="16" pageNumber="1942" type="male">one male</specimenCount>
(
<collectionCode id="13D9AE56AA54CA2E188CCD1EFD726648" box="[650,724,1044,1068]" country="USA" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="1942">AMNH</collectionCode>
) from
<collectingCountry id="0DDF7603AA54CA2E1926CD1EFCCF6648" box="[800,873,1044,1068]" name="Bolivia" pageId="16" pageNumber="1942">Bolivia</collectingCountry>
,
<collectingRegion id="B70CF871AA54CA2E1973CD1EFC196648" box="[885,959,1044,1068]" country="Bolivia" name="La Paz" pageId="16" pageNumber="1942">La Paz</collectingRegion>
,
<collectingMunicipality id="9513ACE9AA54CA2E19CDCD1EFBBB6648" box="[971,1053,1044,1068]" pageId="16" pageNumber="1942">Pacajes</collectingMunicipality>
,
<location id="70176048AA54CA2E1E2FCD1EFB256648" LSID="urn:lsid:plazi:treatment:FD618785AA4ACA2C1BBACC30FC6567BD:70176048AA54CA2E1E2FCD1EFB256648" box="[1065,1155,1044,1068]" country="Bolivia" latitude="-16.1919" longLatPrecision="7" longitude="-68.1403" municipality="Pacajes" name="Botijlaca" pageId="16" pageNumber="1942" stateProvince="La Paz">Botijlaca</location>
(-
<geoCoordinate id="10FC5054AA54CA2E1AAACD3CFEBE662A" box="[172,280,1078,1102]" degrees="16.1919" direction="south" orientation="latitude" pageId="16" pageNumber="1942" precision="5" value="-16.1919">16.1919 S</geoCoordinate>
, -
<geoCoordinate id="10FC5054AA54CA2E1B21CD3CFE38662A" box="[295,414,1078,1102]" degrees="68.1403" direction="west" orientation="longitude" pageId="16" pageNumber="1942" precision="5" value="-68.1403">68.1403 W</geoCoordinate>
),
<quantity id="B2309B76AA54CA2E1BAACD3CFE58662A" box="[428,510,1078,1102]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="1942" unit="m" value="3600.0">
<elevation id="FEE5D1A0AA54CA2E1BAACD3CFE58662A" box="[428,510,1078,1102]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="1942" unit="m" value="3600.0">3600 m</elevation>
</quantity>
a
</materialsCitation>
<materialsCitation id="C5A03CCEAA54CA2E1813CD3CFD81662A" box="[533,551,1078,1102]" collectionCode="AMNH" country="Bolivia" elevation="3600" isTailEnumeration="true" latitude="-16.1919" location="Botijlaca" longLatPrecision="7" longitude="-68.1403" municipality="Pacajes" pageId="16" pageNumber="1942" specimenCount="1" stateProvince="La Paz">.s</materialsCitation>
.l.,
<date id="01761053AA54CA2E183DCD3CFD45662A" box="[571,739,1078,1102]" pageId="16" pageNumber="1942" value="1984-10-02">2 October 1984</date>
, L
<materialsCitation id="C5A03CCEAA54CA2E18FDCD3CFCB2662A" box="[763,788,1078,1102]" collectionCode="AMNH" country="Bolivia" elevation="3600" isTailEnumeration="true" latitude="-16.1919" location="Botijlaca" longLatPrecision="7" longitude="-68.1403" municipality="Pacajes" pageId="16" pageNumber="1942" specimenCount="1" stateProvince="La Paz">. E</materialsCitation>
. Peña col.
</paragraph>
</subSubSection>
<subSubSection id="3DD26518AA54CA2F1AA6CD71FD3B6699" lastPageId="17" lastPageNumber="1943" pageId="16" pageNumber="1942" type="etymology">
<paragraph id="75773693AA54CA2E1AA6CD71FEBB66F0" blockId="16.[160,1156,1147,1276]" box="[160,285,1147,1172]" pageId="16" pageNumber="1942">
<emphasis id="47BCEA81AA54CA2E1AA6CD71FEBB66F0" bold="true" box="[160,285,1147,1172]" italics="true" pageId="16" pageNumber="1942">Etymology</emphasis>
</paragraph>
<paragraph id="75773693AA54CA2E1AA6CD95FD696698" blockId="16.[160,1156,1147,1276]" pageId="16" pageNumber="1942">
A compound word whose first element is the Latin adjective
<emphasis id="47BCEA81AA54CA2E195BCD95FC3166D3" box="[861,919,1183,1207]" italics="true" pageId="16" pageNumber="1942">albus</emphasis>
meaning white, and the second element is the genitive Latin noun
<emphasis id="47BCEA81AA54CA2E18D0CDCBFD5C66BD" box="[726,762,1217,1241]" italics="true" pageId="16" pageNumber="1942">pes</emphasis>
meaning foot, that refers to the distinctive white-legged coloration (
<bibRefCitation id="11594B62AA54CA2E183CCDEEFD646698" author="Simon E" box="[570,706,1252,1276]" pageId="16" pageNumber="1942" pagination="669 - 1080" refId="ref24980" refString="Simon E. 1903. Histoire naturelle des araignees. Deuxieme Edition Paris (Roret). 2: 669 - 1080." type="journal article" year="1903">Simon 1903</bibRefCitation>
).
</paragraph>
<paragraph id="75773693AA54CA2E1AA6CC23FE2C6726" blockId="16.[160,1155,1321,1727]" box="[160,394,1321,1346]" pageId="16" pageNumber="1942">
<emphasis id="47BCEA81AA54CA2E1AA6CC23FE2C6726" bold="true" box="[160,394,1321,1346]" italics="true" pageId="16" pageNumber="1942">Amended diagnosis</emphasis>
</paragraph>
<paragraph id="75773693AA54CA2F1AA6CC47FD3B6699" blockId="16.[160,1155,1321,1727]" lastBlockId="17.[160,1156,1079,1277]" lastPageId="17" lastPageNumber="1943" pageId="16" pageNumber="1942">
The following character combination, modified from
<bibRefCitation id="11594B62AA54CA2E190DCC47FC116701" author="Schmidt G" box="[779,951,1357,1381]" pageId="16" pageNumber="1942" pagination="10 - 14" refId="ref24659" refString="Schmidt G. 1993. Das Weibchen von Hapalotremus albipes Simon 1903 (Araneida: theraphosidae). Arachnologischer Anzeiger. 4: 10 - 14." type="journal article" year="1993">Schmidt (1993)</bibRefCitation>
, is diagnostic for
<taxonomicName id="B2C84D10AA54CA2E1AA6CC65FEA167E3" authorityName=": Simon" authorityYear="1903" box="[160,263,1391,1415]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1942" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="47BCEA81AA54CA2E1AA6CC65FEA167E3" box="[160,263,1391,1415]" italics="true" pageId="16" pageNumber="1942">H. albipes</emphasis>
</taxonomicName>
. Carapace and legs entirely covered dorsally with white setae, except femora of legs and palps, which are dark grey (
<figureCitation id="EDF32A16AA54CA2E1836CC98FD3967CE" box="[560,671,1426,1450]" captionStart="Figure 1" captionStartId="4.[160,225,1268,1291]" captionTargetBox="[163,1152,142,1237]" captionTargetId="figure-72@4.[163,1152,142,1237]" captionTargetPageId="4" captionText="Figure 1. (af) Hapalotremus spp. habitus. (a) Hapalotremus albipes, female. (b) Hapalotremus major, female. (c, d) Hapalotremus martinorum. (c) Male. (d) Female. (e) Hapalotremus chasqui sp. nov., juvenile. (f) Hapalotremus marcapata sp. nov., female. Photos: a: Mauricio Pacheco Suarez; b: Juan Chaparro; c, d: Patricio Cavallo; e: Laura Montes de Oca; f: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747251" httpUri="https://zenodo.org/record/4747251/files/figure.png" pageId="16" pageNumber="1942">Figures 1a</figureCitation>
,
<figureCitation id="EDF32A16AA54CA2E18AACC98FD6367CE" box="[684,709,1426,1450]" captionStart="Figure 7" captionStartId="11.[160,225,1453,1476]" captionTargetBox="[163,1152,142,1423]" captionTargetId="figure-52@11.[163,1152,142,1423]" captionTargetPageId="11" captionText="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747267" httpUri="https://zenodo.org/record/4747267/files/figure.png" pageId="16" pageNumber="1942">7a</figureCitation>
,
<figureCitation id="EDF32A16AA54CA2E18D4CC98FD4A67CE" box="[722,748,1426,1450]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="16" pageNumber="1942">9a</figureCitation>
), abdomen dark with rounded dorsal median patch of reddish-orange urticating setae, and two or three white spots dorsally (more noticeably on females), the bigger spot is arrow-shaped, and with three or four white lines posterior laterally in each side of abdomen, reaching the urticating patch (
<figureCitation id="EDF32A16AA54CA2E19C1CCF0FB906476" box="[967,1078,1530,1554]" captionStart="Figure 1" captionStartId="4.[160,225,1268,1291]" captionTargetBox="[163,1152,142,1237]" captionTargetId="figure-72@4.[163,1152,142,1237]" captionTargetPageId="4" captionText="Figure 1. (af) Hapalotremus spp. habitus. (a) Hapalotremus albipes, female. (b) Hapalotremus major, female. (c, d) Hapalotremus martinorum. (c) Male. (d) Female. (e) Hapalotremus chasqui sp. nov., juvenile. (f) Hapalotremus marcapata sp. nov., female. Photos: a: Mauricio Pacheco Suarez; b: Juan Chaparro; c, d: Patricio Cavallo; e: Laura Montes de Oca; f: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747251" httpUri="https://zenodo.org/record/4747251/files/figure.png" pageId="16" pageNumber="1942">Figures 1a</figureCitation>
,
<figureCitation id="EDF32A16AA54CA2E1E45CCF0FBD16476" box="[1091,1143,1530,1554]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="16" pageNumber="1942">9c, d</figureCitation>
). Labium wider than long bearing a reduced number of cuspules (613) (
<figureCitation id="EDF32A16AA54CA2E19C2CF17FB906451" box="[964,1078,1565,1589]" captionStart="Figure 7" captionStartId="11.[160,225,1453,1476]" captionTargetBox="[163,1152,142,1423]" captionTargetId="figure-52@11.[163,1152,142,1423]" captionTargetPageId="11" captionText="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747267" httpUri="https://zenodo.org/record/4747267/files/figure.png" pageId="16" pageNumber="1942">Figures 7e</figureCitation>
,
<figureCitation id="EDF32A16AA54CA2E1E43CF17FBD06451" box="[1093,1142,1565,1589]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="16" pageNumber="1942">9b, f</figureCitation>
). Absence of spiniform setae on pro- and retrolateral coxal faces of legs IIV. Metatarsi IV with no scopulae (
<figureCitation id="EDF32A16AA54CA2E1B15CF68FED8641E" box="[275,382,1634,1658]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="16" pageNumber="1942">Figure 9h</figureCitation>
). Male tibia I very short, more than 20% shorter than metatarsus I (
<figureCitation id="EDF32A16AA54CA2E1AA1CF8FFEB464F9" box="[167,274,1669,1693]" captionStart="Figure 7" captionStartId="11.[160,225,1453,1476]" captionTargetBox="[163,1152,142,1423]" captionTargetId="figure-52@11.[163,1152,142,1423]" captionTargetPageId="11" captionText="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747267" httpUri="https://zenodo.org/record/4747267/files/figure.png" pageId="16" pageNumber="1942">
Figure
<quantity id="B2309B76AA54CA2E1AF0CF8FFEB464F9" box="[246,274,1669,1693]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="16" pageNumber="1942" unit="g" value="7.0">7g</quantity>
</figureCitation>
). Male palpal tibia with retrolateral rounded process developed (
<figureCitation id="EDF32A16AA54CA2E19F3CF8FFBD164F9" box="[1013,1143,1669,1693]" captionStart="Figure 7" captionStartId="11.[160,225,1453,1476]" captionTargetBox="[163,1152,142,1423]" captionTargetId="figure-52@11.[163,1152,142,1423]" captionTargetPageId="11" captionText="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747267" httpUri="https://zenodo.org/record/4747267/files/figure.png" pageId="16" pageNumber="1942">Figure 7h, i</figureCitation>
). Palpal bulb base with a basal digitiform apophysis well-developed (
<figureCitation id="EDF32A16AA54CA2E1988CFADFBB664DB" box="[910,1040,1703,1727]" captionStart="Figure 8" captionStartId="12.[160,225,985,1008]" captionTargetBox="[163,1153,142,952]" captionTargetId="figure-272@12.[163,1153,142,954]" captionTargetPageId="12" captionText="Figure 8. (ad) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). Male left palpal bulb. (a) Prolateral view. (b) Retrolateral view. (c) Dorsal view. (d) Ventral view. A, apical keel; DA, digitiform apophysis; PI, prolateral inferior keel; PS, prolateral superior keel. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747269" httpUri="https://zenodo.org/record/4747269/files/figure.png" pageId="16" pageNumber="1942">Figure 8ad</figureCitation>
); embolus thick and very curved to the retrolateral face and prolateral keels slightly developed; PI bearing a ventral medial crest pointed at distal end and located half way forward of embolus (
<figureCitation id="EDF32A16AA55CA2F1E1ACD50FF0966F1" captionStart="Figure 8" captionStartId="12.[160,225,985,1008]" captionTargetBox="[163,1153,142,952]" captionTargetId="figure-272@12.[163,1153,142,954]" captionTargetPageId="12" captionText="Figure 8. (ad) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). Male left palpal bulb. (a) Prolateral view. (b) Retrolateral view. (c) Dorsal view. (d) Ventral view. A, apical keel; DA, digitiform apophysis; PI, prolateral inferior keel; PS, prolateral superior keel. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747269" httpUri="https://zenodo.org/record/4747269/files/figure.png" pageId="17" pageNumber="1943">Figure 8a, b</figureCitation>
). Female spermathecae consist of one receptacle with basal portion slightly wider than apical, apical lateral projections slightly developed and apical median region depressed (
<figureCitation id="EDF32A16AA55CA2F1AA1CDC8FEAE66BE" box="[167,264,1218,1242]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="17" pageNumber="1943">Figure 9j</figureCitation>
). Urticating setae:
<typeStatus id="AA738831AA55CA2F1BD1CDC8FDA166BE" box="[471,519,1218,1242]" pageId="17" pageNumber="1943">type</typeStatus>
III present on male and female and arranged in a dorsal median patch on the abdomen (
<figureCitation id="EDF32A16AA55CA2F1BF9CDEFFDCA6699" box="[511,620,1253,1277]" captionStart="Figure 7" captionStartId="11.[160,225,1453,1476]" captionTargetBox="[163,1152,142,1423]" captionTargetId="figure-52@11.[163,1152,142,1423]" captionTargetPageId="11" captionText="Figure 7. (ai) Hapalotremus albipes, male from Botijlaca, Bolivia (AMNH). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, ventral view. (i) Palpal tibia, retrolateral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747267" httpUri="https://zenodo.org/record/4747267/files/figure.png" pageId="17" pageNumber="1943">Figures 7c</figureCitation>
,
<figureCitation id="EDF32A16AA55CA2F187ECDEFFD366699" box="[632,656,1253,1277]" captionStart="Figure 9" captionStartId="13.[160,225,1252,1275]" captionTargetBox="[163,1152,143,1221]" captionTargetId="figure-131@13.[163,1152,142,1221]" captionTargetPageId="13" captionText="Figure 9. (aj) Hapalotremus albipes, female from La Paz, Bolivia (MHNCB). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Abdomen, ventral view. (e) Abdomen, lateral view. (f) Labium and maxillae, ventral view. (g) Eyes, dorsal view. (h) Tarsus IV, ventral view. (i) Tarsus IV, prolateral view. (j) Spermathecae, dorsal view. Scale bars: ae: 5 mm; fj: 1 mm. Photos: Nelson Ferretti." figureDoi="http://doi.org/10.5281/zenodo.4747271" httpUri="https://zenodo.org/record/4747271/files/figure.png" pageId="17" pageNumber="1943">9c</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="21B7661BAA55CA2F1AA6CA51FE9F618C" ID-DOI="http://doi.org/10.5281/zenodo.4747279" ID-Zenodo-Dep="4747279" httpUri="https://zenodo.org/record/4747279/files/figure.png" pageId="17" pageNumber="1943" startId="17.[160,225,859,882]" targetBox="[165,1152,142,828]" targetPageId="17">
<paragraph id="75773693AA55CA2F1AA6CA51FE9F618C" blockId="17.[160,1157,859,1000]" pageId="17" pageNumber="1943">
<emphasis id="47BCEA81AA55CA2F1AA6CA51FEA16116" bold="true" box="[160,263,859,882]" pageId="17" pageNumber="1943">Figure 13.</emphasis>
(ai)
<taxonomicName id="B2C84D10AA55CA2F1B47CA51FDE76116" authorityName=": Cavallo and Ferretti" authorityYear="2015" box="[321,577,859,882]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="1943" phylum="Arthropoda" rank="species" species="martinorum">
<emphasis id="47BCEA81AA55CA2F1B47CA51FDE76116" box="[321,577,859,882]" italics="true" pageId="17" pageNumber="1943">Hapalotremus martinorum</emphasis>
</taxonomicName>
, male holotype (MLP 19152). (a) Carapace, dorsal view. (b) Sternum, ventral view. (c) Abdomen, dorsal view. (d) Eyes, dorsal view. (e) Labium and maxillae, ventral view. (f) Tibial apophysis, ventral view. (g) Leg I, lateral view. (h) Palpal tibia, retrolateral view. (i) Palpal tibia, ventral view. Arrows indicate the rounded process. Scale bar: 1 mm. Photos: Nelson Ferretti.
</paragraph>
</caption>
<subSubSection id="3DD26518AA55CA2F1AA6CC23FC4A67A9" pageId="17" pageNumber="1943" type="description">
<paragraph id="75773693AA55CA2F1AA6CC23FD6B6726" blockId="17.[160,1157,1321,1485]" box="[160,717,1321,1346]" pageId="17" pageNumber="1943">
<emphasis id="47BCEA81AA55CA2F1AA6CC23FD6B6726" bold="true" box="[160,717,1321,1346]" italics="true" pageId="17" pageNumber="1943">Variation (from additional material examined)</emphasis>
</paragraph>
<paragraph id="75773693AA55CA2F1AA6CC47FC4A67A9" blockId="17.[160,1157,1321,1485]" pageId="17" pageNumber="1943">
Total length, females (
<emphasis id="47BCEA81AA55CA2F1B9CCC47FE0E6701" box="[410,424,1357,1381]" italics="true" pageId="17" pageNumber="1943">n</emphasis>
= 3): 26.729.8 (mean = 28.3 ± 1.5SD); carapace length, females: 12.413.8 (13.1 ± 0.7); length of palp and legs, females: palp 17.820.4 (18.9 ± 1.3), I 25.4 31.5 (27.9 ± 3.1), II 24.228.3 (26 ± 2.07), III 23.625.5 (24.8 ± 1.04), IV 31.433.4 (32 ± 1.15). Labial cuspules, females 1013, maxillary cuspules, right 91120, left 96118.
</paragraph>
</subSubSection>
<subSubSection id="3DD26518AA55CA2C1AA6CCF3FC6567BD" lastPageId="18" lastPageNumber="1944" pageId="17" pageNumber="1943" type="distribution">
<paragraph id="75773693AA55CA2F1AA6CCF3FE656476" blockId="17.[160,1156,1529,1727]" box="[160,451,1529,1554]" pageId="17" pageNumber="1943">
<emphasis id="47BCEA81AA55CA2F1AA6CCF3FE656476" bold="true" box="[160,451,1529,1554]" italics="true" pageId="17" pageNumber="1943">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="75773693AA55CA2C1AA6CF17FC6567BD" blockId="17.[160,1156,1529,1727]" lastBlockId="18.[160,1155,1265,1497]" lastPageId="18" lastPageNumber="1944" pageId="17" pageNumber="1943">
The
<typeStatus id="AA738831AA55CA2F1AD4CF17FE926451" box="[210,308,1565,1589]" pageId="17" pageNumber="1943" type="holotype">holotype</typeStatus>
male described by
<bibRefCitation id="11594B62AA55CA2F1812CF17FD0E6451" author="Simon E" box="[532,680,1565,1589]" pageId="17" pageNumber="1943" pagination="669 - 1080" refId="ref24980" refString="Simon E. 1903. Histoire naturelle des araignees. Deuxieme Edition Paris (Roret). 2: 669 - 1080." type="journal article" year="1903">Simon (1903)</bibRefCitation>
gives no indication other than Chaco for the type locality and this involves a large region in
<collectingCountry id="0DDF7603AA55CA2F18CACF35FCB06433" box="[716,790,1599,1623]" name="Bolivia" pageId="17" pageNumber="1943">Bolivia</collectingCountry>
. In the same way,
<bibRefCitation id="11594B62AA55CA2F19D8CF35FB226433" author="Schmidt G" box="[990,1156,1599,1623]" pageId="17" pageNumber="1943" pagination="10 - 14" refId="ref24659" refString="Schmidt G. 1993. Das Weibchen von Hapalotremus albipes Simon 1903 (Araneida: theraphosidae). Arachnologischer Anzeiger. 4: 10 - 14." type="journal article" year="1993">Schmidt (1993)</bibRefCitation>
described the female of
<taxonomicName id="B2C84D10AA55CA2F1BABCF68FDB3641E" authorityName=": Simon" authorityYear="1903" box="[429,533,1634,1658]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="1943" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="47BCEA81AA55CA2F1BABCF68FDB3641E" box="[429,533,1634,1658]" italics="true" pageId="17" pageNumber="1943">H. albipes</emphasis>
</taxonomicName>
for the first time but did not provide a locality for those specimens, only Bolivia. A specimen of
<taxonomicName id="B2C84D10AA55CA2F185ECF8FFD1964F9" authorityName=": Simon" authorityYear="1903" box="[600,703,1669,1693]" class="Arachnida" family="Theraphosidae" genus="Hapalotremus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="1943" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="47BCEA81AA55CA2F185ECF8FFD1964F9" box="[600,703,1669,1693]" italics="true" pageId="17" pageNumber="1943">H. albipes</emphasis>
</taxonomicName>
is shown in
<figureCitation id="EDF32A16AA55CA2F194ECF8FFC0964F9" box="[840,943,1669,1693]" captionStart="Figure 1" captionStartId="4.[160,225,1268,1291]" captionTargetBox="[163,1152,142,1237]" captionTargetId="figure-72@4.[163,1152,142,1237]" captionTargetPageId="4" captionText="Figure 1. (af) Hapalotremus spp. habitus. (a) Hapalotremus albipes, female. (b) Hapalotremus major, female. (c, d) Hapalotremus martinorum. (c) Male. (d) Female. (e) Hapalotremus chasqui sp. nov., juvenile. (f) Hapalotremus marcapata sp. nov., female. Photos: a: Mauricio Pacheco Suarez; b: Juan Chaparro; c, d: Patricio Cavallo; e: Laura Montes de Oca; f: Josh Richards." figureDoi="http://doi.org/10.5281/zenodo.4747251" httpUri="https://zenodo.org/record/4747251/files/figure.png" pageId="17" pageNumber="1943">Figure 1a</figureCitation>
and was located in
<collectingRegion id="B70CF871AA55CA2F1AA6CFADFF4364DB" box="[160,229,1703,1727]" country="Bolivia" name="La Paz" pageId="17" pageNumber="1943">La Paz</collectingRegion>
(
<collectingCountry id="0DDF7603AA55CA2F1AF6CFADFE9D64DB" box="[240,315,1703,1727]" name="Bolivia" pageId="17" pageNumber="1943">Bolivia</collectingCountry>
) at about
<quantity id="B2309B76AA55CA2F1BAFCFADFE5C64DB" box="[425,506,1703,1727]" metricMagnitude="3" metricUnit="m" metricValue="4.252" pageId="17" pageNumber="1943" unit="m" value="4252.0">4252 m</quantity>
a.s.l., near the type locality. Here, we provide records of this species for Unduavi, Coscapa and Botijlaca (among 3200 and
<quantity id="B2309B76AA56CA2C198CCDFBFC42676D" box="[906,996,1265,1289]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="1944" unit="m" value="3600.0">3600 m</quantity>
a.s.l.),
<collectingCountry id="0DDF7603AA56CA2C1E3DCDFBFB25676D" box="[1083,1155,1265,1289]" name="Bolivia" pageId="18" pageNumber="1944">Bolivia</collectingCountry>
(
<figureCitation id="EDF32A16AA56CA2C1AA1CC1EFEB56748" box="[167,275,1300,1324]" captionStart="Figure 36" captionStartId="40.[160,225,871,894]" captionTargetBox="[163,1152,143,840]" captionTargetId="figure-287@40.[163,1152,142,841]" captionTargetPageId="40" captionText="Figure 36. Distribution map of Hapalotremus species from Argentina and Bolivia." figureDoi="http://doi.org/10.5281/zenodo.4747336" httpUri="https://zenodo.org/record/4747336/files/figure.png" pageId="18" pageNumber="1944">Figure 36</figureCitation>
). These sites are located east of Cordillera Real on the edge of the Yungas region, which constitutes a transition between the highlands of the Andes and the tropical lowlands of the Amazon rainforests. The annual average temperature of this area is 17.4°C with monthly average values ranging from 15°C in June and 19°C in December. The average annual precipitation is
<quantity id="B2309B76AA56CA2C180FCC94FDCC67D2" box="[521,618,1438,1462]" metricMagnitude="-1" metricUnit="m" metricValue="7.25" pageId="18" pageNumber="1944" unit="mm" value="725.0">725 mm</quantity>
, with a dry season from May to August and a wet season from December to February (www.censosbolivia.ine.gob.bo).
</paragraph>
</subSubSection>
</treatment>
</document>