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<mods:title id="A6055690F68992BD66D5A12C0CD24D2D">A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)</mods:title>
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<mods:namePart id="41D0D474ED75C6E000A8596449D0AE2D">Wood, Hannah M.</mods:namePart>
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<mods:namePart id="31785EE7D49D2DDC18100B9989D346B5">Scharff, Nikolaj</mods:namePart>
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<treatment id="FC10DDF7D7F7D2A95FEE686C466E4E1B" ID-GBIF-Taxon="140245899" LSID="urn:lsid:plazi:treatment:FC10DDF7D7F7D2A95FEE686C466E4E1B" httpUri="http://treatment.plazi.org/id/FC10DDF7D7F7D2A95FEE686C466E4E1B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="A69B593D23789E8C34E0887D53E0CE20" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="6CF0023B9719807409BC75EBDB5828DE" pageId="0" pageNumber="1">
<taxonomicName id="48DDE728091001F6AF8E0D98C7A3D233" ID-CoL="3XDXR" authority="Platnick, 1991" authorityName="Platnick" authorityYear="1991" class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea legendrei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="legendrei">Madagascarchaea legendrei (Platnick, 1991)</taxonomicName>
<taxonomicNameLabel id="02C895E4B977C0DFA51BACC7A7975DF2" pageId="0" pageNumber="1">comb. n.</taxonomicNameLabel>
Figs 24, 33
</paragraph>
</subSubSection>
<subSubSection id="26E7E7E99C0E346A55E46A014F49785C" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="9B9DB7BA25EC9372FE2F46C297CCFC2E" pageId="0" pageNumber="1">
<taxonomicName id="84136BA1E654002E1C8CCC109288269E" class="Arachnida" family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea legendrei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="legendrei">Archaea legendrei</taxonomicName>
Platnick, 1991: 137, figs 7-14.
</paragraph>
<paragraph id="78C75A045B3CD278C956AEE979C50659" pageId="0" pageNumber="1">
<taxonomicName id="2B1C8EA4DD0FC7DBB6194748AF45F797" class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eriauchenius legendrei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="legendrei">Eriauchenius legendrei</taxonomicName>
(Platnick, 1991):
<bibRefCitation id="9334F1FD2ECDD1637F9415CECFB6FB8B" author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
: 286, figs 2E,12B,18A,20
<normalizedToken id="0AC20271DDE466A09C95EBAE90B12349" originalValue="AD">A-D</normalizedToken>
(new combination).
</paragraph>
</subSubSection>
<subSubSection id="BAAF693BE032AC6A0E764C5B1598D687" pageId="0" pageNumber="1" type="type material">
<paragraph id="E81123AE8711F5214B81713DE97C4929" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="8A969A253780A759A1D1A712070B3574" pageId="0" pageNumber="1">
Male holotype: as
<taxonomicName id="24F2C7900894424A543EC14E8FD8992F" class="Arachnida" family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea legendrei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="legendrei">Archaea legendrei</taxonomicName>
Platnick, 1991: Madagascar, Fianarantsoa, 7 km W Ranomafana, 1100 m, 1-7 Nov 1988, montane rainforest, pyrethrin fogging of dead leaves on fallen tree, W.E. Steiner (deposited in USNM, examined; USNMENT00879968)
</paragraph>
</subSubSection>
<subSubSection id="8976B88B2242EF2F499339DCC840A978" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="944BC11AB97A638BB78DFA9C8C82F45C" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="ADFD6B4F1CC6782D7F9576CA8273796A" pageId="0" pageNumber="1">
MADAGASCAR: Female paratype, same data as holotype (USNMENT00879968); Fianarantsoa, Parc National Ranomafana: 2M,8F, 4 eggcases, Vohiparara, Sahamalaotra forest, 41.1 km 54° NE Fianarantsoa,
<geoCoordinate id="155B08F8CB2C0A1BB58E4D491849D79C" direction="south" orientation="latitude" precision="1" value="-21.238861">21°14'19.9&quot;S</geoCoordinate>
,
<geoCoordinate id="C7ECD1F0F2E6E18E9F71B8272734D233" direction="east" orientation="longitude" precision="1" value="47.394222">47°23'39.2&quot;E</geoCoordinate>
, 1200 m, 26 Dec 2005 - 14 Jan 2006, montane rainforest, general collecting day, beating vegetation: in clumps of dead dry foliage, H. Wood, J. Miller, J.J. Rafonomezantsoa, E. Rajeriarison, V. Andriamananony (USNMENT01377217, USNMENT01377218, USNMENT01377219, USNMENT01377210, USNMENT01377211); 2M,10F,16Juvs, 5 eggcases, Talatekely forest, 42.3 km 58° NE Fianarantsoa,
<geoCoordinate id="FEF4E7E9A77B70219BF9CF83051E4DF6" direction="south" orientation="latitude" precision="1" value="-21.257778">21°15'28.0&quot;S</geoCoordinate>
,
<geoCoordinate id="6DE1AA216B1D6BEA4883CC7B3FB6E20A" direction="east" orientation="longitude" precision="1" value="47.42272">47°25'21.8&quot;E</geoCoordinate>
, 1050 m, 24 Dec 2005 - 14 Jan 2006, montane rainforest, general collecting day and night, H. Wood, J. Miller, J.J. Rafonomezantsoa, E. Rajeriarison, V. Andriamananony (USNMENT01377212, USNMENT01377213, USNMENT01377214, USNMENT01377205); 2M,4F,3Juvs,1 eggcase, Vohiparara, 3.6 km W Ranomafana,
<geoCoordinate id="58747FF9DE190710D692E5E94C5A4D0A" direction="south" orientation="latitude" precision="1" value="-21.237383">21°14.243'S</geoCoordinate>
,
<geoCoordinate id="AF94359E6D003A59EBA66998EEE57AB6" direction="east" orientation="longitude" precision="1" value="47.397366">47°23.842'E</geoCoordinate>
, 1150 m, 13-14 Jan 2009, primary montane rainforest, general collecting day and night, C. Griswold, A. Saucedo and H. Wood (USNMENT01377206, USNMENT01377207, USNMENT01377208, USNMENT01377209); 1F, Talatakely,
<geoCoordinate id="596519072950013D5EC0CC4C256999B7" direction="south" orientation="latitude" precision="92" value="-21.248333">21°14.9'S</geoCoordinate>
,
<geoCoordinate id="D984944A5185149BA8A30DA4BDDE6E80" direction="east" orientation="longitude" precision="92" value="47.426666">47°25.6'E</geoCoordinate>
, 5-18 Apr 1998, C. Griswold, D. Kavanaugh, N. Penny, M. Raherilalao, J. Ranorianarisoa, J. Schweikert, D. Ubick (CASENT9012347); 1M, same data as previous except 19-30 Apr 1998 (CASENT9010075); 2M,4F,3Juv, Vatoharanana,
<geoCoordinate id="CE7F42434DACF0FD1BC00CC2FACD5032" direction="south" orientation="latitude" precision="92" value="-21.278334">21°16.7'S</geoCoordinate>
,
<geoCoordinate id="F980BA8C255C356C38DCAAD8EBE67CA7" direction="east" orientation="longitude" precision="92" value="47.435">47°26.1'E</geoCoordinate>
, 1200 m, 15 Apr 1998, primary forest, C. Griswold, D. Kavanaugh, N. Penny, M. Raherilalao, J. Ranorianarisoa, J. Schweikert, D. Ubick (CASENT9012349, CASENT9010076); 1F,1juv, 2.3 km N Vohiparara village,
<geoCoordinate id="C52A91CEB42BA7E1B569D87B5F8BB081" direction="south" orientation="latitude" precision="92" value="-21.213333">21°12.8'S</geoCoordinate>
,
<geoCoordinate id="80AD4B6421966C400893EAD592583059" direction="east" orientation="longitude" precision="92" value="47.383335">47°23.0'E</geoCoordinate>
, 1100 m, 18 Apr 1998, C. Griswold, D. Kavanaugh, N. Penny, M. Raherilalao, E. Rajeriarison, J. Ranorianarisoa, J. Schweikert, D. Ubick (CASENT9012345); 1F, same data as previous except 10-11 Apr 1998 (CASENT9012348); 1M,2F, Vohiparara, Piste Touristique,
<geoCoordinate id="C40B3950615E3AF093FDD5A10191AFF8" direction="south" orientation="latitude" precision="92" value="-21.226667">21°13.6'S</geoCoordinate>
,
<geoCoordinate id="B2938787C154A3FFBD3927010590C4AD" direction="east" orientation="longitude" precision="92" value="47.4">47°24.0'E</geoCoordinate>
, 1000 m, 23 Apr 1998, C. Griswold, D. Kavanaugh, N. Penny, M. Raherilalao, E. Rajeriarison, J. Ranorianarisoa, J. Schweikert, D. Ubick (CASENT9012333); 1M, Talatakely,
<geoCoordinate id="1090EF92C99DABE08CEEE22A1922CFAF" direction="south" orientation="latitude" precision="925" value="-21.25">21°15'S</geoCoordinate>
,
<geoCoordinate id="6127D17CF46C4EBBA8B495DEF2D48D9B" direction="east" orientation="longitude" precision="925" value="47.433334">47°26'E</geoCoordinate>
, 915-1000 m, 30 Oct - 20 Nov 1998, V.F. Lee, K.J. Ribardo (CASENT9010074); 1M, Vatoharanana River, 4.1 km 231° SW Ranomafana,
<geoCoordinate id="8175AF1C0229B750BCA84BBCD53FB7FF" direction="south" orientation="latitude" precision="15" value="-21.289999">21°17'24&quot;S</geoCoordinate>
,
<geoCoordinate id="75D246AFEF16366FA9AB2D67BB3C65FC" direction="east" orientation="longitude" precision="15" value="47.433334">47°26'00&quot;E</geoCoordinate>
, 1100 m, 27-31 Mar 2003, montane rainforest, EB20 yellow pan trap, Griswold, Fisher et al (CASENT9018908); 1M,2F, Talatakely,
<geoCoordinate id="5E54C9C8CF5B48030D5FFC7D451418FB" direction="south" orientation="latitude" precision="925" value="-21.25">21°15'S</geoCoordinate>
,
<geoCoordinate id="C77EED91169532B4F8D0BA668B9FA27B" direction="east" orientation="longitude" precision="925" value="47.416668">47°25'E</geoCoordinate>
, 900 m, 5-7 Dec 1993, N. Scharff, S. Larcher, C. Griswold, and R. Andriamasimanana (CASENT9046586); 1M,1F,2juvs, Trail FF, 14 May 1992, sifting litter 1 hr sample, B. Roth (CASENT9012334); 1F,1juv, 7 km W Ranomafana, 1100 m, 23-28 Feb 1990, montane rainforest, W.E. Steiner (USNMENT00879971); 1F, 7 km W Ranomafana,
<geoCoordinate id="CC29D1640FCC0F18EAD51CF43E5C4FE8" direction="south" orientation="latitude" precision="925" value="-21.2">21°12'S</geoCoordinate>
,
<geoCoordinate id="BD9EF8B18AE3437E6327ACD5B17B0996" direction="east" orientation="longitude" precision="925" value="47.45">47°27'E</geoCoordinate>
, 1000 m, 1-7 Mar 1990, montane rainforest, W.E. Steiner (USNMENT00879987).
</paragraph>
</subSubSection>
<subSubSection id="0C4B41EE5F9A99E8B43B9CDA74C75E50" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="AC4E2F3EF1BD29227A26009483B353FF" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="E82A5BF5C40F99311E306AD36DB8C283" pageId="0" pageNumber="1">
Distinguished from all other archaeids, except
<taxonomicName id="CA63ED99B5B66ECB75728D4848DDC10C" lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
,
<taxonomicName id="CC962D84B2C18F76113F5AF9DE79795F" lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n.,
<taxonomicName id="4B0EF14711A70A704CE866E19A65B794" lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
sp. n., and
<taxonomicName id="EE92CC5CC2AD1AC4E311C9723395B163" lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
(presumably, because the male is unknown) by having a retrolateral apophysis on the distal side of the male pedipalpal femur and patella (see fig. 12B in
<bibRefCitation id="9B2E8F993487F2FA633A8090CFE70B4C" author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
), and by the shape of the FSGP that is highly reduced and lacks
<normalizedToken id="B23A952CD62E58A0A1B590A31BF85C33" originalValue="“wings”">&quot;wings&quot;</normalizedToken>
(Fig. 24B).
<taxonomicName id="DC692061B9B46BDFFAEF2AE491F8908F" lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
is distinguished from
<taxonomicName id="94436370F5A2B5181CD6A627461B8BEB" lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
by having a rounded cephalon with the cephalic spines not on protrusions (Fig. 24A), rather than the larger protrusions seen in
<taxonomicName id="644DF09B9AEF24CB3ECCB8740223F56A" lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
(compare figs 18A and 18C in
<bibRefCitation id="9E159654F581A31AF865AD3067F9D47E" author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
); from
<taxonomicName id="1B2CC9B6F6F393DDA8ECD107096933A9" lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
by having less than 6 teeth on the cheliceral retromargin, by having a rounded bulge that the AME rests upon, and by lacking a bulge on the posterio-basal side of the chelicerae (see fig. 18D in
<bibRefCitation id="499904EAC4B340F98DE18DEEBF37BE7D" author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
); from
<taxonomicName id="E5701DFC6903CB618D414FA3EC5D38D7" lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
sp. n. by lacking a heavily sclerotized rod-shaped sclerite on the palpal bulb (Fig. 23H,J, arrow), and by lacking setae on the retrolateral cymbial protrusion (Figs 23C, 24C, arrow); and from
<taxonomicName id="D032F056DE50B0BF7C5DA43951086A4E" lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n. by having a more rounded cephalon, and by having a comparatively smaller and less sclerotized triangular basal portion of the conductor (compare basal
<normalizedToken id="02BA542DBD577296070EBB6E929EBD4C" originalValue="“c”">&quot;c&quot;</normalizedToken>
in Figs 24G, 25G).
</paragraph>
</subSubSection>
<subSubSection id="18D7F7428779D5555ED15E5EA59CD0A0" pageId="0" pageNumber="1" type="description">
<paragraph id="E0BB93B452186DE243D64A5BB50DD7B7" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="9DAD0BEB547C7AE50C4338B92EB75C91" pageId="0" pageNumber="1">
Male (based on CASENT9012333, from Parc National Ranomafana, Madagascar). Total length 1.67, carapace 0.74 long, 0.58 wide. Abdomen 0.90 long, 1.13 high. Carapace tilt angle 61.9°, tilt height (CtH) 1.42, constriction 0.28, head length 0.58, neck length 0.84. CtH divided by carapace length 1.92. Cephalon with AME on large bulge, and with 6 short post-ocular spines at the crown, not on protrusions, and 1 short spine between the LE and AME (on each side, for a total of 2; see fig. 18A in
<bibRefCitation id="DC4CE05283089697427CE736542266AD" author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
). Chelicerae 1.53 long, and with a small spine 0.34 from base of chelicerae and projecting downward (Fig. 24A). Femur I 2.35 long. Sternum 0.49 long, 0.29 wide. Carapace, chelicerae, sternum and legs reddish brown with white setae. Legs with darker annulations on tibiae and metatarsi. Abdomen dark brown with lighter circular patches throughout, with white setae (Fig. 24A). Pedipalpal bulb of the &quot;
<taxonomicName id="843DB846BB595A5BEB52717056C1A6F0" genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
group&quot; form (Fig. 24
<normalizedToken id="DA083408E6D72B834C85E416C9687C3F" originalValue="DI">D-I</normalizedToken>
): pedipalpal bulbs elongated with conductor swirling around a mostly membraneous embolus; conductor base triangular (Fig. 24D,G), with remainder of conductor elongate and cradling embolus, and with a curved tip; MA translucent and fans out (Fig. 24
<normalizedToken id="C10F1D30525690BA51353FB73426630F" originalValue="EF">E-F</normalizedToken>
,
<normalizedToken id="55E7C2722B48A7BC695A96F1756794F8" originalValue="HI">H-I</normalizedToken>
). Cymbium with small retrolateral protrusion that lacks setae (Fig. 24C, arrow).
</paragraph>
<paragraph id="A71E8D7A58787C80862B5D5065AE82F2" pageId="0" pageNumber="1">
Female (based on CASENT9012333, from Parc National Ranomafana, Madagascar). Total length 1.92, carapace 0.79 long, 0.63 wide. Abdomen 1.13 long, 1.40 high. Carapace tilt angle 54.4°, tilt height (CtH) 1.63, constriction 0.36, head length 0.71, neck length 0.90. CtH divided by carapace length 2.06. Cephalon as in male. Chelicerae 1.61 long, and with short spine 0.34 from base of chelicerae and projecting downward (Fig. 24A). Femur I 2.26 long. Sternum 0.53 long, 0.31 wide. Colors as in male. Genitalia of the &quot;
<taxonomicName id="08C45C050A2AAF7273B592E10FEC585E" genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
-group&quot; form (Fig. 24B): with one group of poreplates on each side of the bursa anterior, and with FSGP highly reduced, lacking
<normalizedToken id="B5EA8611A34F56532478E362C62BF840" originalValue="“wings.”">&quot;wings.&quot;</normalizedToken>
</paragraph>
<caption id="FDE6558E1391141567D642A17267B96D" pageId="0" pageNumber="1">
<paragraph id="FD6660803227483E0B39B2A6A1F5BE69" pageId="0" pageNumber="1">
Figure 24.
<taxonomicName id="A0F4E231CA13229265690A4FBA07DF3D" class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eriauchenius legendrei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="legendrei">Eriauchenius legendrei</taxonomicName>
(Platnick, 1991). A male (USNMENT01377217) habitus, lateral view B female (CASENT9012333) internal genitalia, dorsal view.
<normalizedToken id="06035146DC895546A622F2BAA32A183D" originalValue="CI">C-I</normalizedToken>
male pedipalpal bulbs
<normalizedToken id="15DDF0670A616664D14E733C9AD5117D" originalValue="DF">D-F</normalizedToken>
left bulb (CASENT9012333) C left bulb (CASENT9010074)
<normalizedToken id="D7504E89A50AF3CBB1576FB0503593A5" originalValue="GI">G-I</normalizedToken>
left bulb (USNMENT01377217): C cymbium close-up, dorsal view, arrow showing the small retrolateral cymbium protrusion that lacks setae D, G prolateral view E, H ventral view F, I retrolateral view. Scale bars: 1 mm (A); 0.25 mm (B, D).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4A96F582FE6A0194D004A59E35D0F509" pageId="0" pageNumber="1" type="variation">
<paragraph id="F91B7A46ECE0CCA9C0444649F07DA535" pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph id="7E69D05FC8109603BC88386644C90381" pageId="0" pageNumber="1">Total length 1.46-1.67 (males; n=5), 1.81-2.02 (females; n=5); Carapace length 0.66-0.77 (males; n=5), 0.76-0.82 (females; n=5); Femur I 3.01-3.66 times the length of carapace in males (n=5), 2.71-3.05 in females (n=5); CtH divided by carapace length 1.93-2.24 in males (n=5), 2.01-2.21 in females (n=5); Average femur I length 2.32 in males (n=5), 2.29 in females (n= 5).</paragraph>
</subSubSection>
<subSubSection id="D4E00C32822A0CB403DD1400AE774357" pageId="0" pageNumber="1" type="natural history">
<paragraph id="BFF007E85D075BA60EFA70E9B7906601" pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph id="16FAF1649EE071ED571D1231AEB32F21" pageId="0" pageNumber="1">Specimens were collected in montane rainforest from 915-1200 m in elevation by beating clumps of dead, dry foliage, by sifting litter, in yellow pan traps, and by general collecting day and night.</paragraph>
</subSubSection>
<subSubSection id="CD677A88237A3EA7CC972A7DD45C6B3D" pageId="0" pageNumber="1" type="distribution">
<paragraph id="6CF2E8CECEC2ED7852189627A41F3564" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="F725813B25E67093D1A1283C1ED028E1" pageId="0" pageNumber="1">Known only from Parc National Ranomafana in central eastern Madagascar (Fig. 33).</paragraph>
</subSubSection>
</treatment>
</document>