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<mods:title id="086D6CBAC49FB7E470815CD0A2904439">Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola</mods:title>
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<taxonomicName id="C0FEED589ADE831C743E4F188EE045C4" LSID="http://zoobank.org/681544BA-AA5F-473B-90BD-B2790838BC81" authority="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie, 2021" authorityName="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" rank="species" species="wulfhaackei" status="sp. nov.">Afroedura wulfhaackei</taxonomicName>
<taxonomicNameLabel id="894EA2A414A7B802C69C2DD97227C59B" pageId="0" pageNumber="55">sp. nov.</taxonomicNameLabel>
<figureCitation id="4E01242111D3425D75A7B9A543313AC7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">Angolan Flat Gecko Osga-achatada de Angola Figures 5B</figureCitation>
<figureCitation id="2AC90104EECA4E71278DCA5AF6B19A74" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura wulfhaackei sp. nov. (PEM R 24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55">, 7</figureCitation>
;
<figureCitation id="F2D14F8E7C6731BC164055065F6C29F4" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." pageId="0" pageNumber="55">Tables 3</figureCitation>
,
<figureCitation id="99C2CA9677521FE9B3F114D3587AB57D" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." pageId="0" pageNumber="55">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="405C37C5BEFC5E1B1587DD71D53FB1C7" pageId="0" pageNumber="55" type="notes">
<paragraph id="8BED1554CFF91C9C89C686B14D823FB6" pageId="0" pageNumber="55">Note.</paragraph>
<paragraph id="0A5CE430FF0FDFAF11DBE51258540FC7" pageId="0" pageNumber="55">
The phylogenetic analysis identified four well-defined clades (
<taxonomicName id="81D77896D6281EDBBB3D6656C81FDD8C" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="4AF0DC0AEA60554D13F2165AC6DB8CEE" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 4-7) within the southern inland/highland major clade (see Results). Our data show that this southern major clade clearly represents at least one undescribed species of
<taxonomicName id="4D1EF8AF068DF60EDF125B18F9E3ACDA" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="9A42F9F94F15D2BA29BB3BC7F8213C90" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
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. Despite comparatively-low genetic distances between the four clades recovered (Table
<tableCitation id="0876AC643D3F0A26E3CB06848D6964F5" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of intra- and interclade uncorrected pairwise genetic distances for specimens of Afroedura clades compared to A. loveridgei for 16 S rRNA. Intraclade / intraspecific distances are marked in bold." httpUri="http://table.plazi.org/id/3F87ADCE25A20DA61B3A1568F02AD894" pageId="0" pageNumber="55" tableUuid="3F87ADCE25A20DA61B3A1568F02AD894">2</tableCitation>
), they were all retrieved separately in our phylogenetic analyses and, as our morphological examination failed to separate the four clades, we cannot assign them as separate species. Therefore, we decided to take a conservative approach and to describe only one of the clades as a new species for now and tentatively assign
<taxonomicName id="F298C8C45E9C4521B772C19DAE4AA8A0" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="A28AA2A6DC0AAA7D31DDBBC0140105CA" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
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sp. 5-7 to this species pending further work. Thus, we grouped the material according to these clades and selected the clade called here
<taxonomicName id="43E0DC3706A17CCFB76350B3171B288A" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="E15E63B0EDA1886003D01F52566DE1FA" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 4 (Fig.
<figureCitation id="9D7CB19AB628EC9DE62BD43007E2940B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic tree topology for the 16 S gene (identical for Bayes and ML) using Afroedura loveridgei as outgroup. Support values for posterior probability and bootstraps are indicated by circles (ML:&gt; 70 % / PP:&gt; 0.95)." figureDoi="10.3897/zse.97.57202.figure2" httpUri="https://binary.pensoft.net/fig/496671" pageId="0" pageNumber="55">2</figureCitation>
) as the name-bearing clade for our new species. Material from lower-altitude localities near Bocoio in Benguela Province groups morphologically with our new species, but needs to be tested in a phylogenetic framework.
</paragraph>
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<paragraph id="DFB19BA4BDD3A60FBFBC9F1033790ADD" pageId="0" pageNumber="55">Synonym.</paragraph>
<paragraph id="2BA3492354B85CC93E3CE3FBC399B037" pageId="0" pageNumber="55">
<taxonomicName id="60BBD3B3BF8F7D2DF8E55D8E4A0625C2" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="E9E9BF59941F14A5A887264A26F72C91" italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
</taxonomicName>
-
<bibRefCitation id="6A8FD8BEC9E8C49408F86D3946D5963D" DOI="https://doi.org/10.3897/zse.93.10915" author="Branch, WR" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="55" pagination="157 - 166" refId="B13" refString="Branch, WR, Haacke, W, Vaz Pinto, P, Conradie, W, Baptista, N, Verburgt, L, Verissimo, L, 2017b. Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution 93 (1): 157 - 166, DOI: https://doi.org/10.3897/zse.93.10915" title="Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status." url="https://doi.org/10.3897/zse.93.10915" volume="93" year="2017 b">Branch et al. 2017b</bibRefCitation>
:157 (part);
<bibRefCitation id="70343F3B4F90C813FFE0B78244D3676E" author="Marques, MP" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" refId="B49" refString="Marques, MP, Ceriaco, LMP, Blackburn, DC, Bauer, AM, 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840-2017). Proceedings of the California Academy of Science, Series 4 65(Supplement II): 1-501." title="Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola. Atlas of Historical and Bibliographic Records (1840 - 2017). Proceedings of the California Academy of Science, Series 4 65 (Supplement II): 1 - 501." year="2018">Marques et al. 2018</bibRefCitation>
: 177 (part);
<bibRefCitation id="AF7147293CD094E8DF0FC6D0A7DF2391" DOI="https://doi.org/10.1007/978-3-030-03083-4_13" author="Branch, WR" editor="Huntley, BJ" journalOrPublisher="Science and Conservation: A Modern Synthesis. Springer Open, Cham" pageId="0" pageNumber="55" pagination="283 - 326" refId="B9" refString="Branch, WR, Baptista, N, Vaz Pinto, P, Conradie, W, 2019a. The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research. In: Huntley, BJ, Ferrand, N, Eds., Biodiversity of Angola. Science and Conservation: A Modern Synthesis. Springer Open, Cham: 283 - 326, DOI: https://doi.org/10.1007/978-3-030-03083-4_13" title="The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research." url="https://doi.org/10.1007/978-3-030-03083-4_13" volumeTitle="Biodiversity of Angola." year="2019 a">Branch et al. 2019a</bibRefCitation>
: 287 (part);
<taxonomicName id="4F761690D8A5E8DD5E60D4034794027D" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura bogerti" order="Squamata" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="5E15D431992760CFCD4BD5A97B5FF63F" italics="true" pageId="0" pageNumber="55">Afroedura bogerti</emphasis>
</taxonomicName>
(clade 4) -
<bibRefCitation id="A6126423D2C47FB19BD570D88C002D9E" DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
:147.
</paragraph>
</subSubSection>
<subSubSection id="BCE8A94D8E73EC1500A044A4DF96FE4C" pageId="0" pageNumber="55" type="holotype">
<materialsCitation id="7E0471446301AA577B88BA70D4071DAA" collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch &amp; Pedro Vaz Pinto" country="Angola" elevation="0" latitude="-12.17056" location="Maka-Mombolo" longLatPrecision="1" longitude="14.88167" specimenCode="PEM R24234" specimenCount="1" stateProvince="Benguela Province" typeStatus="holotype">
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<typeStatus id="9585EB8ABF484525F4C3D6B5DB9C5D01">Holotype</typeStatus>
.
</paragraph>
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<specimenCode id="723C42A08B700F8E9AB76449D95CA42E">PEM R24234</specimenCode>
,
<specimenCount id="68681704925A0E3987164727EFABBCBA" type="adult">adult male</specimenCount>
, collected
<locationDeviation id="7DABE78E75A9647CB4B54124A9F224E0">3 km north</locationDeviation>
of
<location id="A49F307E44A4A9D6E452283B24042C3F" LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:A49F307E44A4A9D6E452283B24042C3F" country="Angola" latitude="-12.17056" longLatPrecision="1" longitude="14.88167" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
(
<geoCoordinate id="744BE481A632010B5146359C8BA5D5CB" degrees="12.17056" direction="south" orientation="latitude" precision="1" value="-12.17056">-12.17056</geoCoordinate>
,
<geoCoordinate id="C6AAFFACA49D99C4712345BEA5B994CF" degrees="14.88167" direction="east" orientation="longitude" precision="1" value="14.88167">14.88167</geoCoordinate>
,
<elevation id="C1F371F5D253224C2F9202C800B8DD3E">1756 m</elevation>
a.s.l.),
<collectingRegion id="4D8856BB4276559FAFACA7433F94B4C4">Benguela Province</collectingRegion>
,
<collectingCountry id="9A00A77DD7374698197CE4437778031C">Angola</collectingCountry>
, by
<collectorName id="CDA351F5A3C1B8E9F4E144CDFA919219">William R. Branch</collectorName>
and
<collectorName id="5FA78E986A128955219E3C40EE16D5E2">Pedro Vaz Pinto</collectorName>
on
<collectingDate id="7D2E1AC294B9191BB036E46A6E9B7EF0">15 November 2016</collectingDate>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="0BFE4FF0E30B008DC7BE48EE67C282E1" pageId="0" pageNumber="55" type="paratypes">
<materialsCitation id="11859081EF5132E2B1707F846FE4BD20" collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch &amp; Pedro Vaz Pinto" country="Angola" elevation="0" latitude="-12.17056" location="Maka-Mombolo" longLatPrecision="1" longitude="14.88167" specimenCode="PEM R24232-3" specimenCount="1" stateProvince="Benguela Province" typeStatus="paratype">
<paragraph id="ADFDFA9A4227D07327CA2E72DE06EE3C" pageId="0" pageNumber="55">
<typeStatus id="4D87206B7AE99C9A836D9F2D2593E4F9">Paratypes</typeStatus>
.
</paragraph>
<paragraph id="E12FBAEDCBD2772F3C04F5A3804C8BA5" pageId="0" pageNumber="55">
<specimenCode id="7222552E7DB3BF164F9828A768257171">PEM R24232-3</specimenCode>
,
<specimenCount id="525913B8B7A1694CE0C66DDAB8078DD8" type="adult">adult females</specimenCount>
, collected
<locationDeviation id="795DBE3377CBBA7AB54F39FC36B66CBD">3 km north</locationDeviation>
of
<location id="217E3B2B28CBABB33505FFC92C4BB1E6" LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:217E3B2B28CBABB33505FFC92C4BB1E6" country="Angola" latitude="-12.17056" longLatPrecision="1" longitude="14.88167" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
(
<geoCoordinate id="77062C77B968A348C7354796AD4A909C" degrees="12.17056" direction="south" orientation="latitude" precision="1" value="-12.17056">-12.17056</geoCoordinate>
,
<geoCoordinate id="9AFEF80C7506B45D04A452925D4E2429" degrees="14.88167" direction="east" orientation="longitude" precision="1" value="14.88167">14.88167</geoCoordinate>
,
<elevation id="3115A59BCC6586803F424EB375CD1EBF">1756 m</elevation>
a.s.l),
<collectingRegion id="42B395BAC924D397D9775380873281C8">Benguela Province</collectingRegion>
,
<collectingCountry id="561E45BB5ED1F1E1A2FB702881EDA2EF">Angola</collectingCountry>
, by
<collectorName id="280950EF10DA9347D6FDE5FE3F6D87E2">William R. Branch</collectorName>
and
<collectorName id="56716EE95817D20211D4246A14F1F4B7">Pedro Vaz Pinto</collectorName>
on
<collectingDate id="681B578CF78C6E6AC31F8DBF848C7E90">15 November 2016</collectingDate>
;
<materialsCitation id="85A6F03C62040A7432636AF70544DEFA" collectingDate="1944-01-01" collectingDateMax="1944-12-31" collectingDateMin="1944-01-01" collectionCode="PEM" collectorName="William R. Branch &amp; Pedro Vaz Pinto" elevation="0" latitude="-12.19833" location="Maka-Mombolo" longLatPrecision="1" longitude="14.86833" specimenCode="PEM R24236" specimenCount="1" stateProvince="Benguela Province" typeStatus="paratype">
<specimenCode id="330CFD91B8B0E56A401D56698AA47841">PEM R24236</specimenCode>
,
<specimenCount id="A0B1D82F0D25AF8699739CA87A3323F8" type="adult">adult male</specimenCount>
, collected above
<location id="3B3E0EC6BB5C8A70E906F23B4B9A765F" LSID="urn:lsid:plazi:treatment:FB80B6B01ADB502A8B2FB84ECB88CDE0:3B3E0EC6BB5C8A70E906F23B4B9A765F" latitude="-12.19833" longLatPrecision="1" longitude="14.86833" name="Maka-Mombolo" stateProvince="Benguela Province">Maka-Mombolo</location>
,
<locationDeviation id="F9E57240E29E30E7AB7C63D526729A7E">north-east of Balombo</locationDeviation>
(
<geoCoordinate id="63C3F44D3DD86AFAB3C695EA8D93106B" degrees="12.19833" direction="south" orientation="latitude" precision="1" value="-12.19833">-12.19833</geoCoordinate>
,
<geoCoordinate id="CDCA43E8A252C8959A6DB692A201A36B" degrees="14.86833" direction="east" orientation="longitude" precision="1" value="14.86833">14.86833</geoCoordinate>
,
<elevation id="8ADEC4AF3E718CF0B2D596296F10CC08">1857 m</elevation>
a.s.l.),
<collectingRegion id="E6332681E2F0B9660FA041A959B53D6C">Benguela Province</collectingRegion>
, Angola, by
<collectorName id="FF2268FAAA3088CF0B9255A749F9E9A2">William R. Branch</collectorName>
and
<collectorName id="44B7A01D729430BB1D2F55B1FA746A94">Pedro Vaz Pinto</collectorName>
on
<collectingDate id="625CA95CA82CA61E4071533226CD6E40">15 November 2016</collectingDate>
</materialsCitation>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="4B4B9EA708ABE1D7276B575DC33CB855" pageId="0" pageNumber="55" type="materials_examined">
<paragraph id="68AB3B2CA4683652DF690F706D8E1E58" pageId="0" pageNumber="55">Additional referred material</paragraph>
<paragraph id="D603998F2FD149DD5EF11489FF3BE463" pageId="0" pageNumber="55">
<emphasis id="19B0A4C816089D812B7AC73BA530D391" bold="true" pageId="0" pageNumber="55">(not examined).</emphasis>
NB 817-9, collected at Morro do Moco, camp near Canjonde (
<geoCoordinate id="77F20C0ED8624156A142FCABAE90F64A" degrees="12.42611" direction="south" orientation="latitude" precision="1" value="-12.42611">-12.42611</geoCoordinate>
,
<geoCoordinate id="7A17C4156A45E5584EC14AD5F7ACC867" degrees="15.14778" direction="east" orientation="longitude" precision="1" value="15.14778">15.14778</geoCoordinate>
, 1931 m a.s.l), Huambo Province, Angola, by Pedro Vaz Pinto on 13 November 2017 (genetic samples included in this study).
</paragraph>
</subSubSection>
<subSubSection id="8EDF4630FC3BC5B645A1389B8DDFC8FA" pageId="0" pageNumber="55" type="materials_examined">
<paragraph id="605F2A8E015B2A7BBA96B83F1F9C73A3" pageId="0" pageNumber="55">Tentative referred additional material examined.</paragraph>
<paragraph id="8D5744311B3DB559876BBA026CE5D38C" pageId="0" pageNumber="55">
<taxonomicName id="6C9E81CD0802C480965D2147B074A87F" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="50AA09C3CAE2F1249DAB159B96D002F1" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 5 (
<emphasis id="A8CA06699910674C103422BEE5212C6F" italics="true" pageId="0" pageNumber="55">Males</emphasis>
): PEM R24192-4, collected at William
<normalizedToken id="1D40CB7AC473DF21E53BFABFCB5F2D63" originalValue="Chapmans">Chapman's</normalizedToken>
Farm Victoria-Verdun (Sandula), (
<geoCoordinate id="4A5F61863A121B3B4FD90405F3FB5222" degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
,
<geoCoordinate id="9FCBED5216AC3B9A1B4636C3D9BF85E3" degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
<taxonomicName id="86AE2C6CBCF34ED14D1DD036B82BBD7C" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="8AD6901E7AA57AFCFC91FAC62FE92D9A" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 5 (
<emphasis id="97F10FD27ACF08D8FEFF52221EC4E5DE" italics="true" pageId="0" pageNumber="55">Females</emphasis>
): PEM R24190-1, PEM R24195-6, PEM R24199, collected at William
<normalizedToken id="ED62857B13A3F55AF5CA61CEA78B65E3" originalValue="Chapmans">Chapman's</normalizedToken>
Farm Victoria-Verdun (Sandula), (
<geoCoordinate id="C7D9253C15D5FD3D781A9EEB0F539A55" degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
,
<geoCoordinate id="143E6AB3C89F0A17931A515A8B79651C" degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
<taxonomicName id="E0D7E65F1B9D0D88D18ECD2C8BAF5DA8" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="571660E087BD7B1FE8DDEE6EDE8A213A" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 5 (
<emphasis id="7A27751DAC8A99777A8742BC7C587440" italics="true" pageId="0" pageNumber="55">Juveniles</emphasis>
): PEM R24197-8, collected at William
<normalizedToken id="23FA7AFAE58E8798F97F45E2D7E4D90E" originalValue="Chapmans">Chapman's</normalizedToken>
Farm Victoria-Verdun (Sandula), (
<geoCoordinate id="DD026696287B456259DE2F284F760157" degrees="12.17194" direction="south" orientation="latitude" precision="1" value="-12.17194">-12.17194</geoCoordinate>
,
<geoCoordinate id="128D77B9CC8E436205D6904CAA3379C5" degrees="15.02667" direction="east" orientation="longitude" precision="1" value="15.02667">15.02667</geoCoordinate>
, 1834 m a.s.l.), Cuanza-Sul Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 6 November 2016.
<taxonomicName id="D48FF1AB10C0A6E9F024A1FDD9360A95" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="D474657FE1F76BC26F44F7A3FFB06593" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 6 (
<emphasis id="3329474C209C04586E7BA815FCB70CB1" italics="true" pageId="0" pageNumber="55">
<normalizedToken id="C22A928720F040889311F888396CA106" originalValue="Мales">Males</normalizedToken>
</emphasis>
): PEM R24201, collected near Rio Chicanda, 5 km southwest of Lepi (
<geoCoordinate id="82623817901A33B51CA8EFD2E0DFB257" degrees="12.90861" direction="south" orientation="latitude" precision="1" value="-12.90861">-12.90861</geoCoordinate>
,
<geoCoordinate id="C0A223025427072CAE88C6062EC9998D" degrees="15.36472" direction="east" orientation="longitude" precision="1" value="15.36472">15.36472</geoCoordinate>
, 1534 m a.s.l.), Huambo Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 8 November 2016.
<taxonomicName id="3CC27624D57630789ACCCE47AF0371C1" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="01B6597A13157466595C34996EF6D1BD" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 7 (
<emphasis id="F6E86073FD0B040908E11A6A34838A71" italics="true" pageId="0" pageNumber="55">
<normalizedToken id="7042946F06CECC29618C38EFA8B53E2B" originalValue="Мales">Males</normalizedToken>
</emphasis>
): TM 45382, TM 45387-8, TM 45392, TM 45396, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
<geoCoordinate id="1F9BACCBEFB4EA8F46792728CB70BDE3" degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
,
<geoCoordinate id="3DE5EAA7B31047D7D39F2FA1CB06C029" degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
<taxonomicName id="F875ED8F679EF36FB540212E24C89B34" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="A18F38DD650F81857D9C97F01277E9E5" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 7 (
<emphasis id="DBBA06B440FC9325071A5EB3DF754EC2" italics="true" pageId="0" pageNumber="55">Females</emphasis>
): PEM R22490-1, collected 1 km west of Candumbo on road to Boas
<normalizedToken id="4E3CF07D5C90CD83BF6D0B1D6C73C0CE" originalValue="Águas">Aguas</normalizedToken>
(
<geoCoordinate id="7AE33FD31C3F5FADCE13C0CA3AD1BF96" degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
,
<geoCoordinate id="10AE63397B72354BFCD093ABD038C8E1" degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
, 1760 m a.s.l), Huambo Province, Angola, by Luke Verburgt on 11 March 2016; TM 45383, TM 45390, TM 45393-4, TM 45397, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
<geoCoordinate id="A9AA77154FFCF7048DBC315BDE6585C4" degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
,
<geoCoordinate id="0765DA621759460FA3E6F2DA0BC5E6A1" degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
<taxonomicName id="F40A870A42189928B85973909CD7E44C" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="56DFB6528228452F271781030F2C3CEB" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 7 (
<emphasis id="FA91F0B083E976B1FC24A3E5684E0B6D" italics="true" pageId="0" pageNumber="55">Juveniles</emphasis>
): PEM R24200, collected at Candumbo Rocks Memorial, 20 km east Humana to Cuito (
<geoCoordinate id="B76FD20D211FC05E1B67C4896DD04433" degrees="12.73722" direction="south" orientation="latitude" precision="1" value="-12.73722">-12.73722</geoCoordinate>
,
<geoCoordinate id="E252262DBBEEF7A205900BA846E28390" degrees="15.97333" direction="east" orientation="longitude" precision="1" value="15.97333">15.97333</geoCoordinate>
, 1750 m a.s.l), Huambo Province, Angola, by William R. Branch, Ninda L. Baptista and Pedro Vaz Pinto on 7 November 2016; TM 45386, TM 45389, TM 45391, TM 45395, collected at Candumbo Rocks, 16 km west of Vila Nova = Huambo (
<geoCoordinate id="F7D455CAB8CF90B74B57B6749F151ACF" degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
,
<geoCoordinate id="5B2B11EED3799F20F71C31CB56597128" degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971.
<emphasis id="897C58612464B30396C524C2DF5F2CBE" italics="true" pageId="0" pageNumber="55">Unassigned clades&gt;</emphasis>
(
<emphasis id="5C5D2E1534D0F0A9A78500F6E7B0A3CE" italics="true" pageId="0" pageNumber="55">Females</emphasis>
): TM 45374, collected 1 km south of Luimbale (
<geoCoordinate id="38DB4706077A705EE23DB97A1B46F963" degrees="12.25367" direction="south" orientation="latitude" precision="1" value="-12.25367">-12.25367</geoCoordinate>
,
<geoCoordinate id="5F4B19D23B42E325E6283A92E75F91D6" degrees="15.31694" direction="east" orientation="longitude" precision="1" value="15.31694">15.31694</geoCoordinate>
, 1591 m a.s.l.), Huambo Province, Angola, collected by Wulf Haacke on 10 May 1971; TM 45367-8, collected 10 km west of Soque (
<geoCoordinate id="4A61E368F37B16DF3BBCC96A3EC42961" degrees="12.34590" direction="south" orientation="latitude" precision="1" value="-12.3459">-12.34590</geoCoordinate>
,
<geoCoordinate id="91A7660FF241B7F267487F2D76856519" degrees="15.01180" direction="east" orientation="longitude" precision="1" value="15.0118">15.01180</geoCoordinate>
, 1974 m a.s.l), Benguela Province, Angola, collected by Wulf Haacke on 10 May 1971; PEM R24743, collected at Morro do Pundo (
<geoCoordinate id="8FC82AE9BF146DACEA6C6D19B21C069B" degrees="12.44389" direction="south" orientation="latitude" precision="1" value="-12.44389">-12.44389</geoCoordinate>
,
<geoCoordinate id="696F36BD15A2F6E1F8A135CF1DBA1F28" degrees="13.92250" direction="east" orientation="longitude" precision="1" value="13.9225">13.92250</geoCoordinate>
, 939 m a.s.l.), Benguela Province, Angola, by Pedro Vaz Pinto on 6 June 2018; TM 46587-8, TM 465890, collected 3 km west of Bocoio (
<geoCoordinate id="90D10A5822BE528D964BC758286FB012" degrees="12.46605" direction="south" orientation="latitude" precision="1" value="-12.46605">-12.46605</geoCoordinate>
,
<geoCoordinate id="8268314C4820EA0EDD74A2BD265E351C" degrees="14.10694" direction="east" orientation="longitude" precision="1" value="14.10694">14.10694</geoCoordinate>
, 926 m a.s.l.), Benguela Province, Angola by Wulf Haacke on 25 May 1971.
<emphasis id="87698B17247792376359B22A672B0A08" italics="true" pageId="0" pageNumber="55">Unassigned clades</emphasis>
(
<emphasis id="F80EF38C1C73D61D1A49165113F49F2B" italics="true" pageId="0" pageNumber="55">Males</emphasis>
): TM 45366 collected 10 km west of Soque (
<geoCoordinate id="BEB2780BD8CB12CD9EA91857D074B9C1" degrees="12.34590" direction="south" orientation="latitude" precision="1" value="-12.3459">-12.34590</geoCoordinate>
,
<geoCoordinate id="F3F740A7E12D6551379EE23B8C6E869F" degrees="15.01180" direction="east" orientation="longitude" precision="1" value="15.0118">15.01180</geoCoordinate>
, 1974 m a.s.l), Benguela Province, Angola, by Wulf Haacke on 10 May 1971, TM 46589, adult male, collected 3 km west of Bocoio (
<geoCoordinate id="E25D53AD975C5837447E905F450B2869" degrees="12.46605" direction="south" orientation="latitude" precision="1" value="-12.46605">-12.46605</geoCoordinate>
,
<geoCoordinate id="C3E3F8F180FA4574C4134C5FDFC28E1B" degrees="14.10694" direction="east" orientation="longitude" precision="1" value="14.10694">14.10694</geoCoordinate>
, 926 m a.s.l.), Benguela Province, Angola, by Wulf Haacke on 25 May 1971.
</paragraph>
</subSubSection>
<subSubSection id="002BADAC1D6AD202400421C221DFE0E4" pageId="0" pageNumber="55" type="materials_examined">
<paragraph id="EDF32001479B33CE03A0DCFA8365B774" pageId="0" pageNumber="55">Tentative referred additional material not examined.</paragraph>
<paragraph id="CBEFC3F9C83314500D97A509FE3508E6" pageId="0" pageNumber="55">
TM 45381, 45384-5, TM 45398, collected at Candumbo Rocks, about 25 km east of the city of Huambo (
<geoCoordinate id="E8F71A31B5B080D1623A94FD0A634065" degrees="12.73614" direction="south" orientation="latitude" precision="1" value="-12.73614">-12.73614</geoCoordinate>
,
<geoCoordinate id="C9C7AFEEE6456EAEDF3FB3CA305F4072" degrees="15.97442" direction="east" orientation="longitude" precision="1" value="15.97442">15.97442</geoCoordinate>
, 1760 m a.s.l), Huambo Province, Angola, by Wulf Haacke on 11 May 1971 (placed in
<taxonomicName id="3149051532BA75510FE2A04E73E9E3B2" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="84CF54016843C76C4148003DDEEDAF98" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 7, based on same geographical area as genetically-assigned material); FKH 0239, collected at Monte Verde-Sandula (
<geoCoordinate id="0388BEEAF9BD9A52F60ED988E4BAF087" degrees="12.17924" direction="south" orientation="latitude" precision="1" value="-12.17924">-12.17924</geoCoordinate>
,
<geoCoordinate id="983EBB0F2306A368B66C003FF2D98534" degrees="15.03086" direction="east" orientation="longitude" precision="1" value="15.03086">15.03086</geoCoordinate>
, 2055 m a.s.l.), Cuanza-Sul Province, Angola, by Pedro Vaz Pinto on 29 May 2019 (genetic sample included in this study placed it in
<taxonomicName id="87113D38828460665656BA0800BC655D" authorityName="Loveridge" authorityYear="1944" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura" order="Squamata" pageId="0" pageNumber="55" rank="genus">
<emphasis id="B7823E3F54DC594846F17CB85DC2A2C7" italics="true" pageId="0" pageNumber="55">Afroedura</emphasis>
</taxonomicName>
sp. 5).
</paragraph>
</subSubSection>
<subSubSection id="6CF8FD9C31F3F8AE32AE3CC4862EF1B1" pageId="0" pageNumber="55" type="etymology">
<paragraph id="27F417AB267B9D65C0E9690A13626AD2" pageId="0" pageNumber="55">Etymology.</paragraph>
<paragraph id="AD7474984109B3598A334E3F0D098781" pageId="0" pageNumber="55">The new species is named in honour of Wulf Haacke, retired curator of the herpetology collection at the former Transvaal Museum (now Ditsong National Museum of Natural History). His herpetological expeditions to Angola in the early 1970s paved the way for this study and much of the material used in this study resulted from his expeditions. The name is constructed in the masculine singular genitive.</paragraph>
</subSubSection>
<subSubSection id="9FC42F9389AF89DC7D6F6F9E464A4CB4" pageId="0" pageNumber="55" type="diagnosis">
<paragraph id="84E76B85D8489A012C84BBDC2CD26D5A" pageId="0" pageNumber="55">Diagnosis.</paragraph>
<paragraph id="0AE0986FBF6D4520155F75A3B5A79F90" pageId="0" pageNumber="55">
A member of the greater '
<taxonomicName id="6907D5EB9510B925581FBC7EB661CF30" baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" rank="species" species="transvaalica">
<emphasis id="BA934118A28DD1CF3C0461853C6506B1" italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
</taxonomicName>
' group in possessing two pairs of enlarged scansors per digit and a strongly verticillate and flattened tail (
<bibRefCitation id="A84E235B3F62429C181CA80BB04859FB" DOI="https://doi.org/10.11646/zootaxa.3846.4.1" author="Jacobsen, NHG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="451 - 501" refId="B41" refString="Jacobsen, NHG, Kuhn, AL, Jackman, TR, Bauer, AM, 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451 - 501, DOI: https://doi.org/10.11646/zootaxa.3846.4.1" title="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa." url="https://doi.org/10.11646/zootaxa.3846.4.1" volume="3846" year="2014">Jacobsen et al. 2014</bibRefCitation>
). It is part of the
<taxonomicName id="ED9FCA56D972F98F0FF1F1202D308AE0" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="934A5B7C9471405916339CBB228D32A1" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
-group which differs from other members of the '
<taxonomicName id="5AAEB8C039B59F62649CED78E276F46B" baseAuthorityName="Hewitt" baseAuthorityYear="1925" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura transvaalica" order="Squamata" pageId="0" pageNumber="55" rank="species" species="transvaalica">
<emphasis id="C72FD265BB400D486DD2CA5975A27B36" italics="true" pageId="0" pageNumber="55">Afroedura transvaalica</emphasis>
</taxonomicName>
' group by having less than 88 mid-body scale rows (vs. 97-102 in
<taxonomicName id="3A58038D78BC7B9827D01C078D4589FD" lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
<emphasis id="13239EAE89EB3F30355C0C728DF8495F" italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
</taxonomicName>
, 113-120 in
<taxonomicName id="0123C3D02CD35B671E3A30C42703B12D" lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
<emphasis id="CD6EB087E7FB9FD01C1593939F1A9200" italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
</taxonomicName>
, 102-119 in
<taxonomicName id="913FA545248727C244052CBFE980DAAC" lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
<emphasis id="4339079053310D75FFCC2F21F88A57A4" italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
</taxonomicName>
); by the rostral bordering the nostril (nostril excluded from rostral in
<taxonomicName id="EDD4B8CDD40450AF2AC996C3F7EE10B9" lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
<emphasis id="B499FF17A39856135F8ADBB9B174E4AD" italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
</taxonomicName>
); by the anterior nasals being mostly in contact ~ 68% (separated by 1-3 granules in
<taxonomicName id="5B4AAFF3776E4993946FACF30CDCA68F" lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
<emphasis id="1D98E359E4AA03516B372A95B9690687" italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
</taxonomicName>
; always in broad contact in
<taxonomicName id="6C93016E6A399688CB1E0F73AD3188DA" lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
<emphasis id="4C414FC8C35E97A0304901EACE1EC9D9" italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
</taxonomicName>
; usually in broad contact in
<taxonomicName id="7D7576A13386EAAF362664046023AF29" lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
<emphasis id="C3AAD64B114DBC476368364FD61F37B8" italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
</taxonomicName>
~ 3-18%); and in having 11-16 scales between the anterior borders of the eyes (19-22 in
<taxonomicName id="19D7A21FC77AD780ABE872AC6C352E60" lsidName="A. gorongosa" pageId="0" pageNumber="55" rank="species" species="gorongosa">
<emphasis id="EBFA6F7EDCC4E07C471D7D19208F93D9" italics="true" pageId="0" pageNumber="55">A. gorongosa</emphasis>
</taxonomicName>
; 15-19 in
<taxonomicName id="D205BF2E7D9B8D1E00C03B52992D6C26" lsidName="A. loveridgei" pageId="0" pageNumber="55" rank="species" species="loveridgei">
<emphasis id="0F7833F89DE3C9702F7A37C209DC9BC4" italics="true" pageId="0" pageNumber="55">A. loveridgei</emphasis>
</taxonomicName>
; 15-20 in
<taxonomicName id="FC007A453C292BA652F184637710F35D" lsidName="A. transvaalica" pageId="0" pageNumber="55" rank="species" species="transvaalica">
<emphasis id="A493DE89F8B5936622C49848857F4702" italics="true" pageId="0" pageNumber="55">A. transvaalica</emphasis>
</taxonomicName>
) (comparative data
<emphasis id="64EC97809447A3C1171C9E920CB48854" italics="true" pageId="0" pageNumber="55">fide</emphasis>
<bibRefCitation id="E1C4229258762585938D6273AA1D6991" DOI="https://doi.org/10.11646/zootaxa.4324.1.8" author="Branch, WR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="55" pagination="142 - 160" refId="B12" refString="Branch, WR, Guyton, JA, Schmitz, A, Barej, MF, Naskrecki, P, Farooq, H, Verburgt, L, Roedel, M-O, 2017a. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142 - 160, DOI: https://doi.org/10.11646/zootaxa.4324.1.8" title="Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique." url="https://doi.org/10.11646/zootaxa.4324.1.8" volume="4324" year="2017 a">Branch et al. 2017a</bibRefCitation>
).
</paragraph>
<paragraph id="BE6B93B747DE6D024897C1044572A1E7" pageId="0" pageNumber="55">
<taxonomicName id="B15E88E986CA0C164D2D6ECF72308FE3" authorityName="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
<emphasis id="FE72501614680A9009C271C6011185B9" italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
</taxonomicName>
sp. nov. differs from other members of the
<taxonomicName id="C30CBBD2D6A4DE4347D180D81AE10E6F" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="96313808F840EF627618011A336FD1DE" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
-group by a combination of the following characters (see Tables
<tableCitation id="A8F588D51A28F3CF16402B3DB3428B87" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Summary of morphological data for Afroedura bogerti, A. wulfhaackei sp. nov. (including the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades), A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov. Values are given as a range with mean values in parenthesis. M = male, F = female, n = sample size." httpUri="http://table.plazi.org/id/742172291B68E9AFACEE2244550D3FEF" pageId="0" pageNumber="55" tableUuid="742172291B68E9AFACEE2244550D3FEF">3</tableCitation>
,
<tableCitation id="2969BE8B9090F17AE0BE28951BA5F987" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Tukey multiple comparisons of means of the four species combinations of Angolan Afroedura (values for A. wulfhaackei sp. nov. include the members of the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Bold values indicate significant pairwise comparisons. SVL - snout-vent length; HL - head length; HW - head width; ED - eye-diameter; EE - ear-eye length; IN - internostril distance; MSR - mid-body scale rows." httpUri="http://table.plazi.org/id/C30887A74A39988C01C45C883AEB5E57" pageId="0" pageNumber="55" tableUuid="C30887A74A39988C01C45C883AEB5E57">4</tableCitation>
): 76-88 (mean 79.3) mid-body scale rows (69-77 [mean 73.5] in
<taxonomicName id="4FFA8012EB204B3D59CA70AFE6B8F4C6" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="E182D6C9EA996EB397F53913A3812BE2" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
, 64-78 [mean 72.8] in
<taxonomicName id="23A0AC80E97763759F941A96DE8686FF" lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
<emphasis id="D621BEA3C4F3E577852A5734A540B160" italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
</taxonomicName>
sp. nov., 73-86 [mean 80.3] in
<taxonomicName id="53A047D456682E834AB62C47F13E57AF" lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
<emphasis id="666022FD541DDD78D64EA6B621E38625" italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
</taxonomicName>
sp. nov., 73-78 [mean 74.8] in
<taxonomicName id="3CC36100190CD8ADC150BDC92ED05FA7" lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
<emphasis id="38392AED87417FB10082A9668A7E4A13" italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
</taxonomicName>
sp. nov.); by the anterior nasals being mostly (~ 68% of the time) in contact (~ 33% of the time in contact in
<taxonomicName id="2A2FF7C1FE24C8BDAC3ABE4CB6C2D3C4" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="15A876D0AB14D3A6B62020A7043609D7" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
; always in contact in
<taxonomicName id="AAF0646531B6C29CDB2FD607F1669EA2" lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
<emphasis id="82F699C5F22CA9BE579F5D12DC373C56" italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="7485C8888DA2BADB85F3E11BEE1B9B4F" lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
<emphasis id="42007C35325F168D2180071B80B16122" italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="526708D63D3EDB4D6A858377668110F9" lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
<emphasis id="428727515718FF289739E0B37ED0BFEB" italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
</taxonomicName>
sp. nov.); each verticil comprising 4-5 (mean 4.0) ventral and 5-6 (mean 5.1) dorsal rows of scales (4 and 5 in
<taxonomicName id="A292CFD335F13895484155B6E6B6EFA5" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="7F73799508B7367BAB1CFFE405B8C8F2" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
and
<taxonomicName id="DF15199504CBB8B6380D2B039A904DDF" lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
<emphasis id="4922444C08E4AB952344027E6E7D967A" italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
</taxonomicName>
sp. nov.; 5-6 [mean 5.5] and 6-7 [mean 6.6] in
<taxonomicName id="B494C88D7121AAAB6186EFFD3FD6A76A" lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
<emphasis id="7CB2520FC3F02900BB7CBAAC491E0F39" italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
</taxonomicName>
sp. nov.; 5-6 [mean 5.0] and 6-7 [mean 6.1]
<taxonomicName id="25D7C34474268EF38D24A296BD7A5CF4" lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
<emphasis id="2846D6499728250E842ED4A26E7F29CC" italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
</taxonomicName>
sp. nov.); ventral surfaces greyish with scattered small black spots (similar to
<taxonomicName id="3C561B1F058491271369E2F6DFD3F4CA" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="1C57C5B84C35232EFE46F5389763D12A" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
and
<taxonomicName id="C5BBBC213CFA3289A90A9D5EB9D89C86" lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
<emphasis id="E9F897EF4973263C7422D18BF864EA13" italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
</taxonomicName>
sp. nov., immaculate in
<taxonomicName id="9E7525AF48EEA59988154D082B586BC2" lsidName="A. donveae" pageId="0" pageNumber="55" rank="species" species="donveae">
<emphasis id="CE23F1DBF74CB3D50F79BFA562F2F981" italics="true" pageId="0" pageNumber="55">A. donveae</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="093CAC4CBA115AA47621580B0C39E7CC" lsidName="A. vazpintorum" pageId="0" pageNumber="55" rank="species" species="vazpintorum">
<emphasis id="0A03B93FDA90E8A62C46401B6B48EC21" italics="true" pageId="0" pageNumber="55">A. vazpintorum</emphasis>
</taxonomicName>
sp. nov.).
<taxonomicName id="939DC462A3F23D5A2776043B7CA94857" authorityName="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
<emphasis id="283E113AE3D31DC468B000AF13667EBB" italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
</taxonomicName>
sp. nov. differs more specifically from its sister highland species
<taxonomicName id="0784D600129B235B7F32850EDA61FD2C" lsidName="A. bogerti" pageId="0" pageNumber="55" rank="species" species="bogerti">
<emphasis id="AF3CBFCB73C39EF6F1F9D7F27B11C7AE" italics="true" pageId="0" pageNumber="55">A. bogerti</emphasis>
</taxonomicName>
in having a higher number of mid-body scale counts (76-88 [mean 79.3] versus 69-77 [mean 73.5]) and differs from
<taxonomicName id="E9C8199D5527EBCC2C7CA27BDADCF4DD" lsidName="A. praedicta" pageId="0" pageNumber="55" rank="species" species="praedicta">
<emphasis id="C0767C4E9484F20E8E6239E814C6615A" italics="true" pageId="0" pageNumber="55">A. praedicta</emphasis>
</taxonomicName>
sp. nov. in that the nasals are separated by smaller granules (versus always in contact).
</paragraph>
</subSubSection>
<subSubSection id="24BEA700E8082F35E419A5A8DFB7B597" pageId="0" pageNumber="55" type="description">
<paragraph id="05AFFDD89D63CFEA24A578DD4A8C2477" pageId="0" pageNumber="55">Holotype description.</paragraph>
<paragraph id="ADF6AE55083FB58217213368F155C34C" pageId="0" pageNumber="55">
Adult male; SVL 51.4 mm; tail 47.1 mm (full original tail), with a small mid-ventral incision for the removal of liver sample. Measurements and meristic characters of holotype presented in Table
<tableCitation id="DBE03115B8F24650AA6411C6A6DE87B3" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" tableUuid="8580963F746364997A5BD1D5F11DD033">5</tableCitation>
. Head and body dorsoventrally compressed; HL 11.9 mm, HW 9.0 mm, broadest at posterior level of eye and 1.32 times longer than wide. Eye large (2.3 mm wide), pupil vertical with indented margins; circumorbital scales small and smooth, elongate at upper anterior margin, upper three posterior scales with small upward pointing spines. Snout rounded, 4.9 mm long, slightly longer than distance between eye and ear openings (4.0 mm). Scales on top of snout smooth, rounded, scales to the edge larger than central ones, with no intervening minute granules. Scales on snout slightly subequal in size to those on the back of head or the nape. Scales on eyelids larger than those on the crown, five scales deep from circumorbital scale to crown. Circumorbital scales are separated by a row of smaller scales from the larger scales on eyelids. Nostril pierced between rostral and three nasal scales; 1st supralabial narrowly excluded from nostril; the supranasal being much larger than the subequal postnasals and separated from each other by two smaller scales. Nostrils slightly elevated. Rostral roughly rectangular, but with its upper edges elongated due to extensions into the nostril. Eight supralabials on each side, the labial margin flexing upwards at the rictus (approx. mid-orbital position), with 3-4 minute scales proximal to the flexure. Ten infralabials on either side, with a small scale proximal to the flexure. At the lip, mental slightly narrower than adjacent infralabial, only three quarters the width of rostral and in contact with two distinctly elongate postmental scales. Scales on throat much smaller than those on belly; scales touching infralabials larger. Fifteen scales across the crown at level of front of eyes; 16 scales from ear to eye; 83 scales around mid-body. Ear opening deep, oblique and roughly oval, only half as high as wide (0.3
<normalizedToken id="D58D13117C06F51EB1B3C950988D1971" originalValue="×">x</normalizedToken>
0.5 mm). Scales on dorsum smooth, non-overlapping, largest at mid-body, smaller on nape and tail base. Scales on ventrum flattened, not overlapping, more-or-less ovate at mid-ventrum, twice the size of lateral granules and 1.5 times those along the backbone. Original tail slightly dorsoventrally flattened and distinctly verticillate (11 verticils in total), with obvious lateral constrictions that are not that distinct to tip of tail; each verticil comprising 5 imbricate rows of scales dorsally and 4 imbricate scale rows ventrally and with ventral scales approximately twice the size of those on the dorsal surface. Limbs well developed, hindlimbs slightly longer than forelimbs, mite pockets (dermal crevices inhabited by small ectoparasitic mites) only present at anterior margin of hindlimbs. All digits with a large pair of distal scansors, separated by a large, curved claw and followed by a large gap (twice the length of terminal scansor) by a smaller pair of scansors; infero-median row of digital scales enlarged transversely, particularly towards the scansors, where the terminal scale adjoining the first pair of scansors may be medially constricted, swollen and scansor-like; enlarged subdigital lamellae on 4th toe 9. Precloacal pores 9.
</paragraph>
<caption id="8580963F746364997A5BD1D5F11DD033" ID-Table-UUID="8580963F746364997A5BD1D5F11DD033" httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" start="Table 5" startId="T5">
<paragraph id="4584E984CB6DEA437D218BFE26C6EA25" pageId="0" pageNumber="55">
<emphasis id="F8987267F913A8F525BA9797C881AF09" bold="true" pageId="0" pageNumber="55">Table 5.</emphasis>
Measurements (in mm) and scale counts for the type series of
<taxonomicName id="932A405F2F542D0C083C2F0F447DEE1D" authorityName="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
<emphasis id="DCF7E9D118AEF29FEB23F3A12D230814" italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph id="34A61433AA304637B87E3ADC8F98A1DD" pageId="0" pageNumber="55">
<table id="D844ED80A53B02B556D8FAF9EC082865" pageId="0" pageNumber="55">
<tr id="EE9D688E4971F80350122E6E01FEBBA1" pageId="0" pageNumber="55">
<th id="E25452B78BB286918D23A6352A3AFAD2" colspan="1" pageId="0" pageNumber="55" rowspan="1">Catalogue number</th>
<th id="F9EAD3BF21D8140497F5D77577314723" colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24234</th>
<th id="034217B3A692F2D3F20CFA3977991CB0" colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24232</th>
<th id="2580CF72163D7E813DCDA13372A102E3" colspan="1" pageId="0" pageNumber="55" rowspan="1">PEM R24233</th>
</tr>
<tr id="48ADEF4C9031A42C268FE5A62E678E34" pageId="0" pageNumber="55">
<td id="A4FCBB5FF471D4B55011F27BB95AB02B" colspan="1" pageId="0" pageNumber="55" rowspan="1">Type Status</td>
<td id="580C433A6E1032D486416639B269BB0B" colspan="1" pageId="0" pageNumber="55" rowspan="1">Holotype</td>
<td id="9DA15C7FEBEA07A8D97C95BD5158FAE5" colspan="1" pageId="0" pageNumber="55" rowspan="1">Paratype</td>
<td id="B0A2A293628588A3EEB053CA9D9889F0" colspan="1" pageId="0" pageNumber="55" rowspan="1">Paratype</td>
</tr>
<tr id="686FFF8F733AC1C88E1183E1D7731273" pageId="0" pageNumber="55">
<td id="5DCAC903BDFA07B1664368FA147A849E" colspan="1" pageId="0" pageNumber="55" rowspan="1">Sex</td>
<td id="C1EA893CB453ACCC0D205300756A6309" colspan="1" pageId="0" pageNumber="55" rowspan="1">Male</td>
<td id="197B24263D80D5B98295083C29162CD3" colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
<td id="BC9724F0DCAB30D6DD2E0A01A4F99888" colspan="1" pageId="0" pageNumber="55" rowspan="1">Female</td>
</tr>
<tr id="5400A80C53B6053E53409E4A1977C749" pageId="0" pageNumber="55">
<td id="8B2B8343EE2585DF86E25A212D0853E1" colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout vent length</td>
<td id="D5DE03D9B9039573D618C2F876BF25BD" colspan="1" pageId="0" pageNumber="55" rowspan="1">51.4</td>
<td id="7770D912EDDD306273DA7F5605ED3BFB" colspan="1" pageId="0" pageNumber="55" rowspan="1">49.1</td>
<td id="F0F6878EC02881F139E5796E81D838F7" colspan="1" pageId="0" pageNumber="55" rowspan="1">55.7</td>
</tr>
<tr id="9ADF103C8DB94249767B489B11C2531B" pageId="0" pageNumber="55">
<td id="75D74BF72C1C5B22748430C44AB373D2" colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail length</td>
<td id="2DEADAFBADE0F92A1FAF0922D5CEB9D5" colspan="1" pageId="0" pageNumber="55" rowspan="1">47.1</td>
<td id="E2416DBB844B207CF18666015F1A484F" colspan="1" pageId="0" pageNumber="55" rowspan="1">35.4</td>
<td id="82FB9AE852F092B5479D7360A2054878" colspan="1" pageId="0" pageNumber="55" rowspan="1">-57.5</td>
</tr>
<tr id="7DE56C8D241B557A5E400888161847A9" pageId="0" pageNumber="55">
<td id="022E4CA3EE1B707B6E549F0ED396A80B" colspan="1" pageId="0" pageNumber="55" rowspan="1">Tail condition</td>
<td id="5A4DFF8682CE8A5AD84B01D6C4F8AE68" colspan="1" pageId="0" pageNumber="55" rowspan="1">Original</td>
<td id="BAB3E91B140AE3E6D962BAFA1F94D3BD" colspan="1" pageId="0" pageNumber="55" rowspan="1">Regenerated</td>
<td id="6B03EFA370B2D1A1037E81B86AD06583" colspan="1" pageId="0" pageNumber="55" rowspan="1">Regenerated</td>
</tr>
<tr id="5EEEED7DFF9F3B83936C2BCE12B02BED" pageId="0" pageNumber="55">
<td id="2AE6C96046ABDB62CC58EFD99FA9C17A" colspan="1" pageId="0" pageNumber="55" rowspan="1">Head length</td>
<td id="C047FAC47D82EA87B2B018AD330383A2" colspan="1" pageId="0" pageNumber="55" rowspan="1">11.9</td>
<td id="83B37683450BAF45DBDEA3B4D433887C" colspan="1" pageId="0" pageNumber="55" rowspan="1">10.9</td>
<td id="1FADBDF31D88E39F8B7DA6C070F4D199" colspan="1" pageId="0" pageNumber="55" rowspan="1">12.3</td>
</tr>
<tr id="ECC410FC9EC33E5A01F4C3F22F14088E" pageId="0" pageNumber="55">
<td id="E7CE91EAC6E97FFEB3A6649BD42D6DC8" colspan="1" pageId="0" pageNumber="55" rowspan="1">Head width</td>
<td id="F6057FF9EDE345A91B1DEE2694C846B8" colspan="1" pageId="0" pageNumber="55" rowspan="1">9.0</td>
<td id="A6078BEA0C518BB3FBDCA69091A2046F" colspan="1" pageId="0" pageNumber="55" rowspan="1">7.5</td>
<td id="B414B0110E763A8A36BB87EBA7109033" colspan="1" pageId="0" pageNumber="55" rowspan="1">8.5</td>
</tr>
<tr id="56FEE3B2DCBD5040C407D4E3F7B3A7A9" pageId="0" pageNumber="55">
<td id="C2A377999CA539BC02BCDA6AC901F69C" colspan="1" pageId="0" pageNumber="55" rowspan="1">Snout length</td>
<td id="5BFCE8476E32B6784F92ECDCDD5AA880" colspan="1" pageId="0" pageNumber="55" rowspan="1">4.9</td>
<td id="D201781D933E3249C1C05948749A802E" colspan="1" pageId="0" pageNumber="55" rowspan="1">4.5</td>
<td id="ADF4F25F7F45D89FFEA2B5C681732089" colspan="1" pageId="0" pageNumber="55" rowspan="1">5.1</td>
</tr>
<tr id="F4DD8DFBC781C52650ACB5EC6EF614FB" pageId="0" pageNumber="55">
<td id="233D056C4C789DCF914E6ACF07165501" colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye distance</td>
<td id="233F9F253F4A929700BBA034A811722E" colspan="1" pageId="0" pageNumber="55" rowspan="1">2.3</td>
<td id="DD041054D66A379FD9BDDAC8FC035C6B" colspan="1" pageId="0" pageNumber="55" rowspan="1">2.2</td>
<td id="A7B0DA019E8AEE5F4AF7558BD31CAC4C" colspan="1" pageId="0" pageNumber="55" rowspan="1">2.6</td>
</tr>
<tr id="3170FEBE55DF9B1282483ECD9627AFBD" pageId="0" pageNumber="55">
<td id="6B97B576291733A8A2325F5D046E097C" colspan="1" pageId="0" pageNumber="55" rowspan="1">Eye-Ear distance</td>
<td id="A77C0C16FA944DC6CC7E48D0952C9001" colspan="1" pageId="0" pageNumber="55" rowspan="1">4.0</td>
<td id="57D0431615CD31BCE86A21FBE2003CD6" colspan="1" pageId="0" pageNumber="55" rowspan="1">3.5</td>
<td id="D980092451EE20696E2E2B0AF52A6989" colspan="1" pageId="0" pageNumber="55" rowspan="1">4.3</td>
</tr>
<tr id="3E6DA46D30096A774CA51CD633814BAE" pageId="0" pageNumber="55">
<td id="53758550F27EAF70F156994DDF609D9E" colspan="1" pageId="0" pageNumber="55" rowspan="1">Precloacal pores (males)</td>
<td id="02B3B475046D1B85963FFD28DF01A5EA" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
<td id="C1E50C8B494FC3183C0F053786085BB3" colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
<td id="F44B363216074B5D22B3E3FA9F293A58" colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
</tr>
<tr id="0697EAD3A616E7A0256B9655D83A7326" pageId="0" pageNumber="55">
<td id="929D1967D8BC53061AC98096000C900D" colspan="1" pageId="0" pageNumber="55" rowspan="1">Dorsal rows per tail verticil</td>
<td id="07EA2667E329D6D43C439F1C3AE94676" colspan="1" pageId="0" pageNumber="55" rowspan="1">5</td>
<td id="2D6FA67C6E149D9CA449E0516F8D2DC0" colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
<td id="4FE5A19746548BD125018FCCB3F24907" colspan="1" pageId="0" pageNumber="55" rowspan="1">5</td>
</tr>
<tr id="17A6A2836153969E151814C56E54D830" pageId="0" pageNumber="55">
<td id="CFFE9537777B1A153355A83B3437B06A" colspan="1" pageId="0" pageNumber="55" rowspan="1">Ventral rows per tail verticil</td>
<td id="614FBC58DFE9FFF86CCB116B2E5A708D" colspan="1" pageId="0" pageNumber="55" rowspan="1">4</td>
<td id="5E8BD4BFAA14A0CB24E6E97EEDDC4222" colspan="1" pageId="0" pageNumber="55" rowspan="1">-</td>
<td id="4DC7D3F72B3228D899AD61922C0A7BE0" colspan="1" pageId="0" pageNumber="55" rowspan="1">4</td>
</tr>
<tr id="C2F88D035675F36421EF0877E38D66F7" pageId="0" pageNumber="55">
<td id="FB28ECE3408FBE434E3C97C98EF69C7D" colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales below 4th toe</td>
<td id="65A06E9B1FF3D07068D962491DDFECC3" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
<td id="EFE1DCC3F4D22E746C19979A5F1C8677" colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
<td id="F86574E2C4D5BFB549C233B133BC1FCA" colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
</tr>
<tr id="83850711FFF9D7F3496A1ABBDBC33814" pageId="0" pageNumber="55">
<td id="ACF7BAA83522F09456F8943C12EF2492" colspan="1" pageId="0" pageNumber="55" rowspan="1">Mid-body scale rows</td>
<td id="57CBB6FFB6457D7F55B0129880EC7F4E" colspan="1" pageId="0" pageNumber="55" rowspan="1">83</td>
<td id="CDD10D7A2D3BB1AEEF70F66B35BCBB42" colspan="1" pageId="0" pageNumber="55" rowspan="1">82</td>
<td id="56CAE86F6B9EB7B1E315C7448852154D" colspan="1" pageId="0" pageNumber="55" rowspan="1">76</td>
</tr>
<tr id="CEE73097F14A5ACA88252DB5B28AD583" pageId="0" pageNumber="55">
<td id="EB5B911CBE4F5D66192CD4CEF308D8F0" colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales between eyes</td>
<td id="2B0E00830689C20459773B6CD2ACA7E5" colspan="1" pageId="0" pageNumber="55" rowspan="1">15</td>
<td id="68BEBAE246141BEE561F67FB689FF150" colspan="1" pageId="0" pageNumber="55" rowspan="1">13</td>
<td id="A312E846075B953A5575638B1D2E1FCA" colspan="1" pageId="0" pageNumber="55" rowspan="1">14</td>
</tr>
<tr id="D66B0027647EB801835C5B6178E9E14E" pageId="0" pageNumber="55">
<td id="B3825946EF3A30C58C1D4A6102C8E6D1" colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: nostril to eye</td>
<td id="95163B6D33A8B07222AE15EC0D4CD521" colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
<td id="563336CC7B71CC6F62961C2897D55CCB" colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
<td id="91EA7B07D0443BAC18CD4BBE16AAB2EC" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
</tr>
<tr id="647066257DBB17AF4255FF7C0968312D" pageId="0" pageNumber="55">
<td id="FCFB89EF7BF9D8F17A5F027CC4DBB752" colspan="1" pageId="0" pageNumber="55" rowspan="1">Scales: ear to eye</td>
<td id="2EE7F4733DA59E2BA53B53B2BA70850E" colspan="1" pageId="0" pageNumber="55" rowspan="1">16</td>
<td id="B7E39EC35515EE58E07179FAD4C0A717" colspan="1" pageId="0" pageNumber="55" rowspan="1">14</td>
<td id="2041831745CA8D7EA3AB7C1B35F3BEA0" colspan="1" pageId="0" pageNumber="55" rowspan="1">17</td>
</tr>
<tr id="F19B01E7A0932C9B0966DA4BC1E6DEB8" pageId="0" pageNumber="55">
<td id="7F75EBC139EBB53C69883926635E7732" colspan="1" pageId="0" pageNumber="55" rowspan="1">Anterior nasals in contact</td>
<td id="A486BBC5E37AB199CB4EC836E8BAC916" colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
<td id="A6CD4D2C92CADA423733D067A9E02CBB" colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
<td id="EBAC9C38488722AE9CC14113E65B8618" colspan="1" pageId="0" pageNumber="55" rowspan="1">No</td>
</tr>
<tr id="449BF1735CECB647D3768481EFE89FDB" pageId="0" pageNumber="55">
<td id="64322C180F3337BC39225F01C2F4CA5A" colspan="1" pageId="0" pageNumber="55" rowspan="1">Supralabials</td>
<td id="7AA8BF1157203CAE93691DF7A3AB0772" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
<td id="0FFB59E701087009255DE68EB3B540DD" colspan="1" pageId="0" pageNumber="55" rowspan="1">7</td>
<td id="F15525830DE2FDA05D4485DD39BE82A8" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
</tr>
<tr id="6E83F2AB2A4BE9D61A12BEB78156F7B5" pageId="0" pageNumber="55">
<td id="336CB68FD08DC06A4B26B371CED0728A" colspan="1" pageId="0" pageNumber="55" rowspan="1">Infralabials</td>
<td id="C0F037E0530095BE9840B455E036A0C9" colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
<td id="E7AADD2C9D6AAE1BD2DB6D94D96AA420" colspan="1" pageId="0" pageNumber="55" rowspan="1">9</td>
<td id="AE552396F5353ED92D5329E6470BAF86" colspan="1" pageId="0" pageNumber="55" rowspan="1">8</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="BCFD63116187A5699782A12C18C9740C" pageId="0" pageNumber="55" type="paratype variation">
<paragraph id="BD222BB4ADAF20A22BB794C411249CA6" pageId="0" pageNumber="55">Paratype variation</paragraph>
<paragraph id="1E72B5E7296A90DB4A3C4D652DD054AB" pageId="0" pageNumber="55">
<emphasis id="56D5955963E403A3DCD1E8B1F93FD8EF" bold="true" pageId="0" pageNumber="55">
(see Table
<tableCitation id="03DB1D1F1F94D8343EC046D58B035FB9" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) and scale counts for the type series of Afroedura wulfhaackei sp. nov." httpUri="http://table.plazi.org/id/8580963F746364997A5BD1D5F11DD033" pageId="0" pageNumber="55" tableUuid="8580963F746364997A5BD1D5F11DD033">5</tableCitation>
for more measurements and scale counts of type series).
</emphasis>
SVL varied from 49.1-55.7 mm; head length 1.34-1.45 times the head width; snout 1.99 times the diameter of eye. Supranasals always separated by smaller granules, usually with a single large granule in contact with the rostral between the supranasals, followed by 1-2 smaller granules in lateral contact; the first upper labial and rostral always enters the nostril and the width of the rostral at the lip margin is always wider than that of the mental; 2-3 postmental scales; supralabials 7-9, infralabials 8-9; scales between anterior edges of eyes 13-14; scales between nostril and anterior edge of orbit 7-9; scales between anterior edge of ear and rear margin of orbit 14-17; scales around mid-body 76-82; subdigital lamellae on 4th toe 7-8; dorsal scales per tail verticil 5; ventral scales per tail verticil 4. Precloacal pores 9.
</paragraph>
</subSubSection>
<subSubSection id="4AD6A963A64419E2717A52DA39FC37FB" pageId="0" pageNumber="55" type="materials_examined">
<paragraph id="7425516F75DF462798E95290A8A4DC7F" pageId="0" pageNumber="55">Additional material variation.</paragraph>
<paragraph id="05875B1EF7E6141435DDE52EE0D4DC05" pageId="0" pageNumber="55">SVL 36.4-59.6 mm; original tail length 36.4-57.8 mm, 0.94 times SVL; head length 1.19-1.75 times head width; snout 1.91 times diameter of eye. The supranasals in contact in 22 specimens and separated by granules in eight specimens, usually with a single large granule in contact with the rostral between the supranasals, followed by 1-2 smaller granules in lateral contact; the first upper labial and rostral always enters the nostril and the width of the rostral at the lip margin is always wider than that of the mental; 2-4 postmental scales; supralabials 8-9; infralabials 8-9; scales between anterior edge of eye 12-16; scales between nostril and anterior edge of orbit 7-11; scales between anterior edge of ear and rear margin of orbit 15-18; scales around mid-body 75-88; subdigital lamellae on 4th toe 7-9; dorsal scales per tail verticil 5 (TM 45366, TM 46588 and PEM R24743 with 6); ventral scales per tail verticil 4 (TM 46588 with 5). Precloacal pores 9-12.</paragraph>
</subSubSection>
<subSubSection id="8E6826E996A4FB45383A5A0DA9D8E37A" pageId="0" pageNumber="55" type="colouration">
<paragraph id="F08F32140C752894CDDC131E20B4EBB9" pageId="0" pageNumber="55">Colouration.</paragraph>
<paragraph id="2EA23AC53AC1473BC64FB8FFDFE88892" pageId="0" pageNumber="55">
<emphasis id="B15187FE2F80051C7ED520C9B11F9B04" italics="true" pageId="0" pageNumber="55">In life</emphasis>
(paratype PEM R24232, Fig.
<figureCitation id="8331D5C1416A814E8D4AE60CD88E34CF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live photos of Angolan flat geckos: A. Afroedura bogerti (PEM R 24187); B. Afroedura wulfhaackei sp. nov. (paratype PEM R 24232); C. Afroedura praedicta sp. nov. (holotype NB 854); D. Afroedura donveae sp. nov. (holotype PEM R 17937); E. Afroedura vazpintorum sp. nov. (holotype PEM R 24118); F. Afroedura vazpintorum sp. nov. (NB 0745); Photos: A, B, D, E. William R. Branch; C, F. Pedro Vaz Pinto." figureDoi="10.3897/zse.97.57202.figure5" httpUri="https://binary.pensoft.net/fig/496674" pageId="0" pageNumber="55">5B</figureCitation>
): Greyish above with five irregularly-spaced darker crossbars from the occiput to the sacrum, each crossbar consisting posteriorly out of three to four black scales wide forming a W-shape; anterior to W-shape are 8-10 scales deep with a mix of dark grey and mustard colours; each dark crossbar separated by light grey to beige blotches; head with irregular dark grey-mustard blotches on the crown with intervening light grey colouration; dark mustard to dark grey bar from nostril to the anterior margins of the ear opening; a vague, thin pale grey canthal stripe, extends on both sides from the nasal region to anterior margins of eye; upper and lower labials grey with diffuse mustard edges; lateral sides of the body with a mix of dark grey and yellow-mustard colouration; limbs greyish above with scattered darker grey markings with intervening yellow-mustard colouration; tail (regenerated) with irregular grey-mustard mottling; iris golden with a black narrow elliptic pupil with crenulated edge and black reticulation with light grey intervening blotches; ventrum uniform greyish with scattered black specks; ventral limbs with scattered black specks, more prominent than on the ventrum.
<emphasis id="3A996BDF5419E446CBF0E57E86EF4FF0" italics="true" pageId="0" pageNumber="55">In preservative</emphasis>
(holotype PEM R24234, Fig.
<figureCitation id="27F8742F66D5391AFB2FF763E4621890" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Afroedura wulfhaackei sp. nov. (PEM R 24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie." figureDoi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55">7</figureCitation>
): Dorsum with five irregularly-spaced dark grey W-shaped crossbars from the occiput to the sacrum with beige intervening blotches; ventrum is beige with numerous small scattered black specks on each scale, more prominent posteriorly.
<emphasis id="E684BB3E89C53FCDE9CD82AC73A1EC7D" italics="true" pageId="0" pageNumber="55">Variation</emphasis>
: Greyish to brownish above with five to six irregularly-spaced darker grey-brown W-shaped crossbars from the occiput to the sacrum, limbs and tail with grey blotches; ventrum uniform greyish with scattered black specks. Juveniles have sharper patterns and colours.
</paragraph>
<caption id="0D90428AD3C5D894CEB6F032783B8683" doi="10.3897/zse.97.57202.figure7" httpUri="https://binary.pensoft.net/fig/496676" pageId="0" pageNumber="55" start="Figure 7" startId="F7">
<paragraph id="9C3CA4A236B65897F733C444424CDF49" pageId="0" pageNumber="55">
<emphasis id="28332E3ED78D10606E07A4595D190DC7" bold="true" pageId="0" pageNumber="55">Figure 7.</emphasis>
Holotype of
<taxonomicName id="874CAFB08BD0A745BA0C0E05C817B3F4" authorityName="Branch &amp; Schmitz &amp; Lobón-Rovira &amp; Baptista &amp; António &amp; Conradie" authorityYear="2021" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" lsidName="Afroedura wulfhaackei" order="Squamata" pageId="0" pageNumber="55" rank="species" species="wulfhaackei">
<emphasis id="CFE9F5979FB00D71E43B7F16D8C2168F" italics="true" pageId="0" pageNumber="55">Afroedura wulfhaackei</emphasis>
</taxonomicName>
sp. nov. (PEM R24234) from Maka-Mombolo, Benguela Province, Angola. Photos: Werner Conradie.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8CC73773D240149883A117C86D8ACB12" pageId="0" pageNumber="55" type="natural history">
<paragraph id="333218BAEBD318B53CA9071973C959CD" pageId="0" pageNumber="55">Natural history and habitat</paragraph>
<paragraph id="6480D5988DCC4A2157F4E73474246A81" pageId="0" pageNumber="55">
<emphasis id="592A36F8281F5D7BA1D9B1CA3E74F607" bold="true" pageId="0" pageNumber="55">
(Fig.
<figureCitation id="53F068300D92C72D8F34B560365A24CC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Typical habitat of Angolan Afroedura. A. Namba area (A. bogerti); B. Mt Sandula (Afroedura sp. 5); C. Serra da Neve (A. praedicta sp. nov.); D. Omauha Lodge (A. donveae sp. nov.); E. Farm Mucungo (A. vazpintorum sp. nov.); F. Bimbe (A. vazpintorum sp. nov.). Photos: A. Javier Lobon-Rovira; B, D, E. William R. Branch; C. Pedro Vaz Pinto; F. Ninda L. Baptista." figureDoi="10.3897/zse.97.57202.figure4" httpUri="https://binary.pensoft.net/fig/496673" pageId="0" pageNumber="55">4B</figureCitation>
).
</emphasis>
An exclusively rupicolous species living in crevices between rocks or under flakes of exfoliating rocks in boulders at elevations of 920-2,055 m a.s.l. Specimens were collected in cracks of relatively small- to medium-sized boulders of carbonatitic origin surrounded by montane grassland. Some individuals were also found under flakes in large granite boulders and often in steep vertical rock faces and overhangs.
</paragraph>
</subSubSection>
<subSubSection id="8A10D4BEE4043BC7DD87016CF00EFF34" pageId="0" pageNumber="55" type="distribution">
<paragraph id="08206ACABC6670F92C846F57CD745B05" pageId="0" pageNumber="55">Distribution and conservation.</paragraph>
<paragraph id="5E007B3D48DAFAE63A450B44A1AEC86C" pageId="0" pageNumber="55">
This species is currently known from southern Cuanza-Sul, central Huambo and northern Benguela Provinces, Angola (Fig.
<figureCitation id="E3C9066620F84185A68C59BD2EAA25A2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Digital elevation map showing the predicted distribution of the five newly-recognised species of Afroedura occurring in Angola (data for A. wulfhaackei sp. nov. also contain the morphologically-indistinguishable Afroedura sp. 5 - 7 clades). Confirmed locality records are indicated by solid circles, with type localities indicated by stars. Coloured polygons indicate potential distribution areas with more than 90 % of climatic habitat suitability." figureDoi="10.3897/zse.97.57202.figure1" httpUri="https://binary.pensoft.net/fig/496670" pageId="0" pageNumber="55">1</figureCitation>
). It appears to have a relatively large, but patchy, distribution on the Angolan highlands and may extend its range into neighbouring provinces. Although sometimes locally common, it appears to be absent from vast areas in-between, where the species would be expected to occur. Populations in isolated granite outcrops may be threatened by removal of rock flakes for construction of homes and other buildings.
</paragraph>
</subSubSection>
</treatment>
</document>