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<document id="19CD795687F748B340D7F373674BC5BE" ID-DOI="10.1111/j.1096-3642.2008.00406.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630460862998" checkinUser="felipe" docAuthor="Davis, Steve R., Gentili-Poole, Patricia &amp; Mitter, Charles" docDate="2008" docId="FB73C54D5509FFF08D9BF41CFC509C73" docLanguage="en" docName="j.1096-3642.2008.00406.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00406.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Spinulata oblongata Davis &amp; Gentili-Poole &amp; Mitter 2008, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="255" masterDocId="074ABD355528FFD18D65F619FFEE9A39" masterDocTitle="A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae)" masterLastPageNumber="277" masterPageNumber="222" pageNumber="255" updateTime="1699279176942" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="AB82DF4282103EAD6D40BD0B92B47B0A">A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae)</mods:title>
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<mods:namePart id="8C123F230C3387DE1C44B8055EB78DDE">Davis, Steve R.</mods:namePart>
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<mods:namePart id="579CED7DB768A45E97EA17FF3B00A4FC">Gentili-Poole, Patricia</mods:namePart>
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<mods:namePart id="D75165FF37FA7849CCC43F035B4422A5">Mitter, Charles</mods:namePart>
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<mods:title id="2F9748696570DD8502C4BAA9CE4855C5">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="564221E38E8144B18CDEA1A5E30E4244">2008</mods:date>
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<treatment id="FB73C54D5509FFF08D9BF41CFC509C73" ID-DOI="http://doi.org/10.5281/zenodo.5492584" ID-Zenodo-Dep="5492584" LSID="urn:lsid:plazi:treatment:FB73C54D5509FFF08D9BF41CFC509C73" httpUri="http://treatment.plazi.org/id/FB73C54D5509FFF08D9BF41CFC509C73" lastPageNumber="255" pageId="33" pageNumber="255">
<subSubSection id="3BC027D05509FFF08D9BF41CFD669822" box="[254,648,517,539]" pageId="33" pageNumber="255" type="nomenclature">
<paragraph id="7365745B5509FFF08D9BF41CFD669822" blockId="33.[254,648,517,539]" box="[254,648,517,539]" pageId="33" pageNumber="255">
<heading id="282DC3375509FFF08D9BF41CFD669822" box="[254,648,517,539]" centered="true" fontSize="9" level="2" pageId="33" pageNumber="255" reason="2">
<taxonomicName id="B4DA0FD85509FFF08D9BF41CFDF79823" authority="Davis &amp; Gentili-Poole &amp; Mitter, 2008" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" box="[254,537,517,538]" class="Insecta" family="Cossidae" genus="Spinulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="oblongata" status="sp. nov.">SPINULATA OBLONGATA</taxonomicName>
<emphasis id="41AEA8495509FFF08F46F410FD669822" bold="true" box="[547,648,518,539]" pageId="33" pageNumber="255">
<taxonomicNameLabel id="5A9D15325509FFF08F46F410FD669822" box="[547,648,518,539]" pageId="33" pageNumber="255" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3BC027D05509FFF08C48F435FC509C73" pageId="33" pageNumber="255" type="description">
<paragraph id="7365745B5509FFF08C48F435FDB7987B" blockId="33.[301,601,556,578]" box="[301,601,556,578]" pageId="33" pageNumber="255">
(
<figureCitation id="EBE168DE5509FFF08C50F435FE7F9878" box="[309,401,556,577]" captionStart="Figures 2031" captionStartId="16.[164,254,1377,1396]" captionTargetBox="[323,1257,198,1342]" captionTargetId="figure-167@16.[323,1283,196,1347]" captionTargetPageId="16" captionText="Figures 2031. Adults: 20, male, Biocellata alfarae; 21, female, Biocellata alfarae; 22, male, Biocellata bifida; 23, male, Biocellata davisorum; 24, male, Spinulata acutipennis; 25, female, Spinulata acutipennis; 26, male, Spinulata manes; 27, female, Spinulata manes; 28, male, Spinulata oblongata; 29, male, Spinulata quasivinnea; 30, female, Spinulata quasivinnea; 31, male, Simplicivalva ampliophilobia. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5446389" httpUri="https://zenodo.org/record/5446389/files/figure.png" pageId="33" pageNumber="255">FIGS 28</figureCitation>
,
<figureCitation id="EBE168DE5509FFF08CFAF435FE05987B" box="[415,491,556,578]" captionStart="Figures 5364" captionStartId="19.[144,234,1385,1404]" captionTargetBox="[303,1261,198,1349]" captionTargetId="figure-232@19.[303,1263,196,1355]" captionTargetPageId="19" captionText="Figures 5364. Male genitalia. Biocellata alfarae: 53, genital capsule; 54, aedoeagus. Biocellata bifida: 55, genital capsule; 56, aedoeagus. Biocellata davisorum: 57, genital capsule; 58, aedoeagus. Spinulata acutipennis: 59, genital capsule; 60, aedoeagus. Spinulata manes: 61, genital capsule; 62, aedoeagus. Spinulata oblongata: 63, genital capsule; 64, aedoeagus." figureDoi="http://doi.org/10.5281/zenodo.5446395" httpUri="https://zenodo.org/record/5446395/files/figure.png" pageId="33" pageNumber="255">63, 64</figureCitation>
,
<figureCitation id="EBE168DE5509FFF08C9CF435FDF69878" box="[505,536,556,577]" captionStart="Figures 8994" captionStartId="22.[164,254,1364,1383]" captionTargetBox="[389,1219,196,1330]" captionTargetId="figure-201@22.[387,1221,194,1333]" captionTargetPageId="22" captionText="Figures 8994. Male genitalia, lateral view of genital capsule: 89, Biocellata alfarae; 90, Biocellata bifida; 91, Biocellata davisorum; 92, Spinulata acutipennis; 93, Spinulata manes; 94, Spinulata oblongata. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5446403" httpUri="https://zenodo.org/record/5446403/files/figure.png" pageId="33" pageNumber="255">94</figureCitation>
,
<figureCitation id="EBE168DE5509FFF08F43F435FDBE987B" box="[550,592,556,578]" captionStart="Figures 135140" captionStartId="30.[164,254,1467,1486]" captionText="Figures 135140. Species distribution maps in Costa Rica." pageId="33" pageNumber="255">138</figureCitation>
)
</paragraph>
<paragraph id="7365745B5509FFF08DF5F44AFD8B9851" blockId="33.[144,613,595,617]" box="[144,613,595,617]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F44AFF269851" box="[144,200,595,616]" italics="true" pageId="33" pageNumber="255">Male</emphasis>
(
<figureCitation id="EBE168DE5509FFF08DBFF44AFEC79850" box="[218,297,595,617]" captionStart="Figures 2031" captionStartId="16.[164,254,1377,1396]" captionTargetBox="[323,1257,198,1342]" captionTargetId="figure-167@16.[323,1283,196,1347]" captionTargetPageId="16" captionText="Figures 2031. Adults: 20, male, Biocellata alfarae; 21, female, Biocellata alfarae; 22, male, Biocellata bifida; 23, male, Biocellata davisorum; 24, male, Spinulata acutipennis; 25, female, Spinulata acutipennis; 26, male, Spinulata manes; 27, female, Spinulata manes; 28, male, Spinulata oblongata; 29, male, Spinulata quasivinnea; 30, female, Spinulata quasivinnea; 31, male, Simplicivalva ampliophilobia. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5446389" httpUri="https://zenodo.org/record/5446389/files/figure.png" pageId="33" pageNumber="255">Fig. 28</figureCitation>
): Forewing length:
<quantity id="B422D9BE5509FFF08F75F44AFD8F9851" box="[528,609,595,616]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="33" pageNumber="255" unit="mm" value="17.0">17 mm</quantity>
.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F489FD8C98DA" blockId="33.[144,759,656,739]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F489FF3D989C" box="[144,211,656,677]" italics="true" pageId="33" pageNumber="255">Head:</emphasis>
Antenna pale brown; labial palpi cream ventrally, brown dorsally; frons cream; vertex with scales cream basally and rusty-brown apically.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F513FE079F10" blockId="33.[144,759,778,1321]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F513FF069926" box="[144,232,778,799]" italics="true" pageId="33" pageNumber="255">Thorax:</emphasis>
Patagia same colour as vertex. Dorsum and tegula same as patagia except scales brown to rustybrown apically; metathorax brown; pleura creamy white; venter creamy white, lightly irrorated with brown, with a small brown patch directly behind head. Legs with femur and tibia cream mesally and cream irrorated with brown laterally; tarsi brown, irrorated with cream. Forewing greyish brown dorsally; one large, dark rusty-brown spot immediately before middle, and several smaller spots of same colour dispersed around larger spot; several striae traversing distal half of wing, alternating between a darker and lighter brown; basal two-thirds of costal margin with short fuscous striae; fringe brown to cream; forewing brown ventrally with faint brown, transverse striae; cream to pale brown apically. Hindwing brown dorsally and ventrally; fringe mostly cream with some pale brown.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F349FE229FBD" blockId="33.[144,759,1360,1413]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F349FEEF9F5C" box="[144,257,1360,1381]" italics="true" pageId="33" pageNumber="255">Abdomen:</emphasis>
Rusty-brown dorsally; pale brown and cream ventrally; A8 cream.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F3B5FD799C8E" blockId="33.[144,759,1452,1719]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F3B5FED29FF8" box="[144,316,1452,1473]" italics="true" pageId="33" pageNumber="255">Male genitalia</emphasis>
(
<figureCitation id="EBE168DE5509FFF08C37F3B5FE379FFB" box="[338,473,1452,1474]" captionStart="Figures 5364" captionStartId="19.[144,234,1385,1404]" captionTargetBox="[303,1261,198,1349]" captionTargetId="figure-232@19.[303,1263,196,1355]" captionTargetPageId="19" captionText="Figures 5364. Male genitalia. Biocellata alfarae: 53, genital capsule; 54, aedoeagus. Biocellata bifida: 55, genital capsule; 56, aedoeagus. Biocellata davisorum: 57, genital capsule; 58, aedoeagus. Spinulata acutipennis: 59, genital capsule; 60, aedoeagus. Spinulata manes: 61, genital capsule; 62, aedoeagus. Spinulata oblongata: 63, genital capsule; 64, aedoeagus." figureDoi="http://doi.org/10.5281/zenodo.5446395" httpUri="https://zenodo.org/record/5446395/files/figure.png" pageId="33" pageNumber="255">Figs 63, 64</figureCitation>
,
<figureCitation id="EBE168DE5509FFF08C8EF3B5FDE49FFB" box="[491,522,1452,1474]" captionStart="Figures 8994" captionStartId="22.[164,254,1364,1383]" captionTargetBox="[389,1219,196,1330]" captionTargetId="figure-201@22.[387,1221,194,1333]" captionTargetPageId="22" captionText="Figures 8994. Male genitalia, lateral view of genital capsule: 89, Biocellata alfarae; 90, Biocellata bifida; 91, Biocellata davisorum; 92, Spinulata acutipennis; 93, Spinulata manes; 94, Spinulata oblongata. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5446403" httpUri="https://zenodo.org/record/5446403/files/figure.png" pageId="33" pageNumber="255">94</figureCitation>
): Valva rounded at apex; length of valva approximately 0.7¥ length of genital capsule; gnathosuncus expanse approximately 0.5¥ length of genital capsule. Saccular process with an oblong patch of spines. Juxta process elongate. Gnathos broadly curved. Uncus elongate and digitiform, with acute apex. Aedoeagus slightly curved, tapering to an acute rostellum as viewed laterally; apex subacute as viewed ventrally.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F0C7FE849CCD" blockId="33.[144,362,1758,1780]" box="[144,362,1758,1780]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F0C7FF079CCA" box="[144,233,1758,1779]" italics="true" pageId="33" pageNumber="255">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="7365745B5509FFF08DF5F105FF1A9D56" blockId="33.[144,759,1820,1903]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08DF5F105FECF9D08" box="[144,289,1820,1841]" italics="true" pageId="33" pageNumber="255">Distribution:</emphasis>
Known only from the provinces of
<collectingRegion id="B11EBAB95509FFF08FA5F105FF099D69" country="Costa Rica" name="Puntarenas" pageId="33" pageNumber="255">Puntarenas</collectingRegion>
and
<collectingRegion id="B11EBAB95509FFF08C40F123FE409D76" box="[293,430,1850,1871]" country="Costa Rica" name="Guanacaste" pageId="33" pageNumber="255">Guanacaste</collectingRegion>
,
<collectingCountry id="0BCD34CB5509FFF08CD9F123FDD29D69" box="[444,572,1850,1872]" name="Costa Rica" pageId="33" pageNumber="255">Costa Rica</collectingCountry>
, and
<collectingRegion id="B11EBAB95509FFF08F1AF123FD1C9D69" box="[639,754,1850,1872]" country="Ecuador" name="Pichincha" pageId="33" pageNumber="255">Pichincha</collectingRegion>
,
<collectingCountry id="0BCD34CB5509FFF08DF5F140FF1F9D56" box="[144,241,1881,1903]" name="Ecuador" pageId="33" pageNumber="255">Ecuador</collectingCountry>
.
</paragraph>
<paragraph id="7365745B5509FFF08E43F6DFFC6C9B01" blockId="33.[806,1421,197,312]" pageId="33" pageNumber="255">
<materialsCitation id="C3B27E065509FFF08E43F6DFFC6C9B01" collectingDate="1995-07-23" collectingDateMax="1995-08-06" collectingDateMin="1995-07-23" collectionCode="INBio" collectorName="Volcan Rincon de la Vieja &amp; K. Taylor" country="Costa Rica" elevation="800" location="Volcan Rincon de la Vieja" municipality="Sector Las Pailas" pageId="33" pageNumber="255" specimenCode="USNM 85637" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="41AEA8495509FFF08E43F6DFFC7E9AE2" box="[806,912,198,219]" italics="true" pageId="33" pageNumber="255">
<typeStatus id="AC61CAF95509FFF08E43F6DFFC649AE2" box="[806,906,198,219]" pageId="33" pageNumber="255" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="65DCBFD25509FFF08EFCF6DCFC5C9AE2" box="[921,946,197,219]" pageId="33" pageNumber="255" type="male"></specimenCount>
;
<collectingCountry id="0BCD34CB5509FFF08EA7F6DFFB8C9AE2" box="[962,1122,198,219]" name="Costa Rica" pageId="33" pageNumber="255">COSTA RICA</collectingCountry>
:
<collectingRegion id="B11EBAB95509FFF0890BF6DFFAED9AE2" box="[1134,1283,198,219]" country="Costa Rica" name="Guanacaste" pageId="33" pageNumber="255">GUANACASTE</collectingRegion>
:
<collectingMunicipality id="9301EE215509FFF08876F6DFFC819AC3" pageId="33" pageNumber="255">Sector Las Pailas</collectingMunicipality>
, 4.5 km SW del
<collectorName id="DE2F118D5509FFF0894BF6FDFAB69AC3" box="[1070,1368,228,250]" pageId="33" pageNumber="255">
<location id="760522805509FFF0894BF6FDFAB69AC3" LSID="urn:lsid:plazi:treatment:FB73C54D5509FFF08D9BF41CFC509C73:760522805509FFF0894BF6FDFAB69AC3" box="[1070,1368,228,250]" country="Costa Rica" municipality="Sector Las Pailas" name="Volcan Rincon de la Vieja" pageId="33" pageNumber="255" stateProvince="Guanacaste">Volcán Rincón de la Vieja</location>
</collectorName>
,
<quantity id="B422D9BE5509FFF08801F6FCFCD09B20" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="255" unit="m" value="800.0">
<elevation id="F8F793685509FFF08801F6FCFCD09B20" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="255" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="0764529B5509FFF08E2AF71AFBF29B20" box="[847,1052,259,281]" pageId="33" pageNumber="255" value="1995-07-23" valueMax="1995-08-06" valueMin="1995-07-23">
<collectingDate id="1720AB735509FFF08E2AF71AFBF29B20" box="[847,1052,259,281]" pageId="33" pageNumber="255" value="1995-07-23" valueMax="1995-08-06" valueMin="1995-07-23">23.vii.6.viii.1995</collectingDate>
</date>
,
<collectorName id="DE2F118D5509FFF08949F71DFB729B20" box="[1068,1180,259,281]" pageId="33" pageNumber="255">K. Taylor</collectorName>
, slide
<specimenCode id="237CDC205509FFF0898BF71AFA639B20" box="[1262,1421,259,281]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="255" type="Museum">USNM 85637</specimenCode>
(
<collectionCode id="15CBEC9E5509FFF08E4BF73BFC989B01" box="[814,886,290,312]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="33" pageNumber="255">INBio</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="7365745B5509FFF08E43F771FC63982E" blockId="33.[806,1422,360,535]" pageId="33" pageNumber="255">
<materialsCitation id="C3B27E065509FFF08E43F771FA649B82" collectingDate="1997-05-07" collectingDateMax="1997-05-09" collectingDateMin="1997-05-07" collectionCode="USNM" collectorName="Sendero El Ripario &amp; A. P. Luz" country="Costa Rica" elevation="1300" location="Progreso" municipality="Sendero El Ripario" pageId="33" pageNumber="255" specimenCode="USNM 85784" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="41AEA8495509FFF08E43F771FC709B44" box="[806,926,360,381]" italics="true" pageId="33" pageNumber="255">
<typeStatus id="AC61CAF95509FFF08E43F771FC799B44" box="[806,919,360,381]" pageId="33" pageNumber="255" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="0BCD34CB5509FFF08ED0F771FB8D9B44" box="[949,1123,360,381]" name="Costa Rica" pageId="33" pageNumber="255">COSTA RICA</collectingCountry>
:
<collectingRegion id="B11EBAB95509FFF0891BF771FAFC9B45" box="[1150,1298,360,381]" country="Costa Rica" name="Puntarenas" pageId="33" pageNumber="255">PUNTARENAS</collectingRegion>
:
<collectorName id="DE2F118D5509FFF08854F771FC4A9BA5" pageId="33" pageNumber="255">
<collectingMunicipality id="9301EE215509FFF08854F771FC4A9BA5" pageId="33" pageNumber="255">Sendero El Ripario</collectingMunicipality>
</collectorName>
<quantity id="B422D9BE5509FFF08ED3F79FFC009BA5" box="[950,1006,390,412]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="255" unit="km" value="3.0">3 km</quantity>
NE del
<location id="760522805509FFF0890CF79EFB3E9BA5" LSID="urn:lsid:plazi:treatment:FB73C54D5509FFF08D9BF41CFC509C73:760522805509FFF0890CF79EFB3E9BA5" box="[1129,1232,391,412]" country="Costa Rica" municipality="Sendero El Ripario" name="Progreso" pageId="33" pageNumber="255" stateProvince="Puntarenas">Progreso</location>
,
<quantity id="B422D9BE5509FFF08981F79FFAD59BA5" box="[1252,1339,390,412]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="255" unit="m" value="1300.0">
<elevation id="F8F793685509FFF08981F79FFAD59BA5" box="[1252,1339,390,412]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="255" unit="m" value="1300.0">1300 m</elevation>
</quantity>
:
<specimenCount id="65DCBFD25509FFF0882AF79FFA699BA5" box="[1359,1415,390,412]" pageId="33" pageNumber="255" type="male">1 ♂</specimenCount>
,
<date id="0764529B5509FFF08E43F7BCFC4F9B83" box="[806,929,421,442]" pageId="33" pageNumber="255" value="1997-05-07" valueMax="1997-05-09" valueMin="1997-05-07">
<collectingDate id="1720AB735509FFF08E43F7BCFC4F9B83" box="[806,929,421,442]" pageId="33" pageNumber="255" value="1997-05-07" valueMax="1997-05-09" valueMin="1997-05-07">79.v.1997</collectingDate>
</date>
,
<collectorName id="DE2F118D5509FFF08ED5F7BCFBCD9B83" box="[944,1059,421,442]" pageId="33" pageNumber="255">A. P. Luz</collectorName>
, slide
<specimenCode id="237CDC205509FFF08910F7BCFAF99B82" box="[1141,1303,421,443]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="255" type="Museum">USNM 85784</specimenCode>
, (
<collectionCode id="15CBEC9E5509FFF0884BF7BCFA6C9B82" box="[1326,1410,421,443]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="255" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C3B27E065509FFF08E43F7DDFC029BC1" collectingDate="1985-05-19" collectionCode="V" country="Ecuador" elevation="700" location="Tinalandia" pageId="33" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
<collectingCountry id="0BCD34CB5509FFF08E43F7DDFC459BE0" box="[806,939,452,473]" name="Ecuador" pageId="33" pageNumber="255">ECUADOR</collectingCountry>
:
<collectingRegion id="B11EBAB95509FFF08EDAF7DDFBD39BE1" box="[959,1085,452,473]" country="Ecuador" name="Pichincha" pageId="33" pageNumber="255">PICHINCHA</collectingRegion>
:
<location id="760522805509FFF08931F7DDFB3D9BE3" LSID="urn:lsid:plazi:treatment:FB73C54D5509FFF08D9BF41CFC509C73:760522805509FFF08931F7DDFB3D9BE3" box="[1108,1235,452,474]" country="Ecuador" name="Tinalandia" pageId="33" pageNumber="255" stateProvince="Pichincha">Tinalandia</location>
,
<quantity id="B422D9BE5509FFF0898DF7DDFAD49BE0" box="[1256,1338,452,473]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="255" unit="m" value="700.0">
<elevation id="F8F793685509FFF0898DF7DDFAD49BE0" box="[1256,1338,452,473]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="255" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<specimenCount id="65DCBFD25509FFF0882AF7DDFA699BE0" box="[1359,1415,451,473]" pageId="33" pageNumber="255" type="male">1 ♂</specimenCount>
,
<date id="0764529B5509FFF08E43F7FBFC7D9BC1" box="[806,915,482,504]" pageId="33" pageNumber="255" value="1985-05-19">
<collectingDate id="1720AB735509FFF08E43F7FBFC7D9BC1" box="[806,915,482,504]" pageId="33" pageNumber="255" value="1985-05-19">19.v.1985</collectingDate>
</date>
, C.
<collectionCode id="15CBEC9E5509FFF08EBFF7FAFC029BC1" box="[986,1004,483,504]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="33" pageNumber="255" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3B27E065509FFF08960F7FBFC67982E" collectionCode="USNM" collectorName="Covell Jr." country="Ecuador" pageId="33" pageNumber="255" specimenCode="USNM 85726" specimenCount="1" stateProvince="Pichincha" typeStatus="paratype">
<collectorName id="DE2F118D5509FFF08960F7FBFB6E9BC1" box="[1029,1152,482,504]" pageId="33" pageNumber="255">Covell Jr.</collectorName>
, slide
<specimenCode id="237CDC205509FFF08980F7FBFA639BC1" box="[1253,1421,482,504]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="255" type="Museum">USNM 85726</specimenCode>
(
<collectionCode id="15CBEC9E5509FFF08E4BF418FC6C982E" box="[814,898,513,535]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="255" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7365745B5509FFF08E43F451FC0C9864" blockId="33.[806,994,583,605]" box="[806,994,583,605]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08E43F451FC8F9864" box="[806,865,584,605]" italics="true" pageId="33" pageNumber="255">Host:</emphasis>
Unknown.
</paragraph>
<paragraph id="7365745B5509FFF08E43F494FB79989B" blockId="33.[806,1175,653,675]" box="[806,1175,653,675]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08E43F494FC2A989B" box="[806,964,653,674]" italics="true" pageId="33" pageNumber="255">Flight period:</emphasis>
May and August.
</paragraph>
<paragraph id="7365745B5509FFF08E43F4CAFB25997C" blockId="33.[806,1421,723,837]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08E43F4CAFC4A98D1" box="[806,932,723,744]" italics="true" pageId="33" pageNumber="255">Etymology:</emphasis>
Derived from the Latin oblongus meaning longer than broad.
<taxonomicName id="B4DA0FD85509FFF089FEF4EBFA63993E" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" box="[1179,1421,754,775]" class="Insecta" family="Cossidae" genus="Spinulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="oblongata">
<emphasis id="41AEA8495509FFF089FEF4EBFA63993E" box="[1179,1421,754,775]" italics="true" pageId="33" pageNumber="255">Spinulata oblongata</emphasis>
</taxonomicName>
refers to the elongate, oblong shape of the spine patch on the valvae in the male genitalia.
</paragraph>
<paragraph id="7365745B5509FFF08E43F56FFA639EC3" blockId="33.[806,1421,885,1611]" pageId="33" pageNumber="255">
<emphasis id="41AEA8495509FFF08E43F56FFC4999B2" box="[806,935,886,907]" italics="true" pageId="33" pageNumber="255">Discussion:</emphasis>
This species can be differentiated from other members of the
<taxonomicName id="B4DA0FD85509FFF0894DF58DFB6F9990" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" baseAuthorityName="Schaus" baseAuthorityYear="1901" box="[1064,1153,916,937]" class="Insecta" family="Cossidae" genus="Costria" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="maruga">
<emphasis id="41AEA8495509FFF0894DF58DFB6F9990" box="[1064,1153,916,937]" italics="true" pageId="33" pageNumber="255">maruga</emphasis>
</taxonomicName>
species group by characters of the male genitalia and wing pattern. The hindwing of this species is entirely medium to dark brown in colour, in contrast to other members in this complex, which have cream or light brown hindwings, or in a few specimens, shaded partially with cream and partially with light brown. This species also posesses a greater number of dark brown spots on the forewing than any other species in the complex. The arrangement of spots in
<taxonomicName id="B4DA0FD85509FFF08904F2BEFB3F9E85" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" box="[1121,1233,1191,1212]" class="Insecta" family="Cossidae" genus="Spinulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="oblongata">
<emphasis id="41AEA8495509FFF08904F2BEFB3F9E85" box="[1121,1233,1191,1212]" italics="true" pageId="33" pageNumber="255">oblongata</emphasis>
</taxonomicName>
also differs in possessing a greater number of spots in the longitudinal row nearest the inner margin than in
<taxonomicName id="B4DA0FD85509FFF08848F2FCFA689EC3" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" baseAuthorityName="Schaus" baseAuthorityYear="1901" box="[1325,1414,1253,1274]" class="Insecta" family="Cossidae" genus="Costria" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="maruga">
<emphasis id="41AEA8495509FFF08848F2FCFA689EC3" box="[1325,1414,1253,1274]" italics="true" pageId="33" pageNumber="255">maruga</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7365745B5509FFF08E5AF31AFC509C73" blockId="33.[806,1421,885,1611]" pageId="33" pageNumber="255">
The male genitalia exhibit several characters that distinguish
<taxonomicName id="B4DA0FD85509FFF08ED2F338FBC99F0F" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" box="[951,1063,1313,1334]" class="Insecta" family="Cossidae" genus="Spinulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="oblongata">
<emphasis id="41AEA8495509FFF08ED2F338FBC99F0F" box="[951,1063,1313,1334]" italics="true" pageId="33" pageNumber="255">oblongata</emphasis>
</taxonomicName>
from other members of the
<taxonomicName id="B4DA0FD85509FFF08E43F358FC919F6F" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" baseAuthorityName="Schaus" baseAuthorityYear="1901" box="[806,895,1345,1366]" class="Insecta" family="Cossidae" genus="Costria" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="maruga">
<emphasis id="41AEA8495509FFF08E43F358FC919F6F" box="[806,895,1345,1366]" italics="true" pageId="33" pageNumber="255">maruga</emphasis>
</taxonomicName>
species group. Most notable is the shape of the spine patch protruding from the sacculus on the male valvae (
<figureCitation id="EBE168DE5509FFF08EDAF364FBFE9FAA" box="[959,1040,1405,1427]" captionStart="Figures 5364" captionStartId="19.[144,234,1385,1404]" captionTargetBox="[303,1261,198,1349]" captionTargetId="figure-232@19.[303,1263,196,1355]" captionTargetPageId="19" captionText="Figures 5364. Male genitalia. Biocellata alfarae: 53, genital capsule; 54, aedoeagus. Biocellata bifida: 55, genital capsule; 56, aedoeagus. Biocellata davisorum: 57, genital capsule; 58, aedoeagus. Spinulata acutipennis: 59, genital capsule; 60, aedoeagus. Spinulata manes: 61, genital capsule; 62, aedoeagus. Spinulata oblongata: 63, genital capsule; 64, aedoeagus." figureDoi="http://doi.org/10.5281/zenodo.5446395" httpUri="https://zenodo.org/record/5446395/files/figure.png" pageId="33" pageNumber="255">Fig. 63</figureCitation>
). This patch is oblong and nearly a figure of eight in shape, with the middle of the patch slightly constricted. The saccular processes of the
<taxonomicName id="B4DA0FD85509FFF08E43F3C3FC919FD6" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" baseAuthorityName="Schaus" baseAuthorityYear="1901" box="[806,895,1498,1519]" class="Insecta" family="Cossidae" genus="Costria" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="maruga">
<emphasis id="41AEA8495509FFF08E43F3C3FC919FD6" box="[806,895,1498,1519]" italics="true" pageId="33" pageNumber="255">maruga</emphasis>
</taxonomicName>
group differ in being larger, usually with many acute angles that are often elongate. The lateral margins of the spine patch are more angular in
<emphasis id="41AEA8495509FFF08E22F02CFC509C73" box="[839,958,1589,1610]" italics="true" pageId="33" pageNumber="255">
<taxonomicName id="B4DA0FD85509FFF08E22F02CFC549C73" authorityName="Davis &amp; Gentili-Poole &amp; Mitter" authorityYear="2008" box="[839,954,1589,1610]" class="Insecta" family="Cossidae" genus="Spinulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="255" phylum="Arthropoda" rank="species" species="oblongata">oblongata</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>