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<document id="F93DD404C3023D75CCE6FC209C53C29C" ID-DOI="10.11646/zootaxa.3664.4.8" ID-GBIF-Dataset="b1ee41b8-57cb-476c-9416-4b9ca47fbe56" ID-ISSN="1175-5326" ID-Zenodo-Dep="215621" ID-ZooBank="17E68A54-CA49-4F05-BB31-B0E0DBDE3FA4" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460620788622" checkinUser="plazi" docAuthor="Klotz, Werner &amp; Rintelen, Kristina Von" docDate="2013" docId="FB3BD455FFD0B567FF52FE59FD97322C" docLanguage="en" docName="zt03664p570.pdf" docOrigin="Zootaxa 3664 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caridina boehmei Klotz &amp; Rintelen, 2013, n. sp." docType="treatment" docVersion="10" lastPageNumber="564" masterDocId="0702AC2DFFD1B56DFFC5FFAEFFA2303B" masterDocTitle="Three new species of Caridina (Decapoda: Atyidae) from Central Sulawesi and Buton Island, Indonesia, and a checklist of the islands’ endemic species" masterLastPageNumber="570" masterPageNumber="554" pageNumber="555" updateTime="1698333876466" updateUser="plazi">
<mods:mods id="DC4C8D779BAD995E38D47DB49AC55BEF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A27D139AF99D91D2C9D5DE94EBFC51D8">
<mods:title id="98EA32433AF4179D532623E4CC9CF703">Three new species of Caridina (Decapoda: Atyidae) from Central Sulawesi and Buton Island, Indonesia, and a checklist of the islands’ endemic species</mods:title>
</mods:titleInfo>
<mods:name id="2DB8C3D62631B135890F802A7187B212" type="personal">
<mods:role id="28A3E65E11DCE407F652BE5DB4813225">
<mods:roleTerm id="EB6E1FC0EB19BE7F580558B0964A6C28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA80735AC40CAA69CBC5D08EA3B420A4">Klotz, Werner</mods:namePart>
</mods:name>
<mods:name id="FF58A3A30631419501805970B2832019" type="personal">
<mods:role id="4FAFDA9D002178D4CF4FAC8A7A13821B">
<mods:roleTerm id="C555BC27495D17BB1B19E77D432989C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC3E758462121DC60AFDAC7A495A001A">Rintelen, Kristina Von</mods:namePart>
</mods:name>
<mods:typeOfResource id="9D35B5B0080667C8D2B956936FEC63B5">text</mods:typeOfResource>
<mods:relatedItem id="55339A43F506A2B74188F91F9B266A74" type="host">
<mods:titleInfo id="7A5785071E905B4D69350D72566CF93D">
<mods:title id="A8EE7E8C199FC40B545A264D6EA2E331">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D4174240F8BA9D92C6FFBCC0CFA57883">
<mods:date id="DEE650FAD5BC876572262E7042EB2B24">2013</mods:date>
<mods:detail id="D4176EE63BF9E181D7E3516DA48955CF" type="volume">
<mods:number id="DBB7C241D2A951635B139615C32D8026">3664</mods:number>
</mods:detail>
<mods:detail id="DF53606BFAC6EF6D9ADE8BA266D950FB" type="issue">
<mods:number id="8688771E8EC81BB8477CC073E968BDF2">4</mods:number>
</mods:detail>
<mods:extent id="D71367E195E1FBC6130963266323EE03" unit="page">
<mods:start id="4CBA1C0E87AEACAF3E45BDFFACF5B2AA">554</mods:start>
<mods:end id="AEA58FEDB08147A98698A257610471B3">570</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="40FE13CA61AEF110E7C01DF891BCD7FA">journal article</mods:classification>
<mods:identifier id="32FB0A07DC844C56B8E9609C2693B387" type="DOI">10.11646/zootaxa.3664.4.8</mods:identifier>
<mods:identifier id="52A9F98651E0D9692A2B8C06AD16E1E5" type="GBIF-Dataset">b1ee41b8-57cb-476c-9416-4b9ca47fbe56</mods:identifier>
<mods:identifier id="210AA3F9E385F28E0CA94560F400559F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5A6E6FEF4F975BB91CC142D3F56FD431" type="Zenodo-Dep">215621</mods:identifier>
<mods:identifier id="E80FDEA5D1F496D1BB15BC8E6C7233E7" type="ZooBank">17E68A54-CA49-4F05-BB31-B0E0DBDE3FA4</mods:identifier>
</mods:mods>
<treatment id="FB3BD455FFD0B567FF52FE59FD97322C" ID-DOI="http://doi.org/10.5281/zenodo.5657524" ID-GBIF-Taxon="119569042" ID-Zenodo-Dep="5657524" LSID="urn:lsid:plazi:treatment:FB3BD455FFD0B567FF52FE59FD97322C" httpUri="http://treatment.plazi.org/id/FB3BD455FFD0B567FF52FE59FD97322C" lastPageId="10" lastPageNumber="564" pageId="1" pageNumber="555">
<subSubSection id="3B8836C8FFD0B56CFF52FE59FD8F3209" pageId="1" pageNumber="555" type="nomenclature">
<paragraph id="732D6543FFD0B56CFF52FE59FE18322A" blockId="1.[151,557,503,562]" box="[151,442,503,529]" pageId="1" pageNumber="555">
<heading id="2865D22FFFD0B56CFF52FE59FE18322A" bold="true" box="[151,442,503,529]" fontSize="11" level="1" pageId="1" pageNumber="555" reason="1">
<emphasis id="41E6B951FFD0B56CFF52FE59FE18322A" bold="true" box="[151,442,503,529]" pageId="1" pageNumber="555">
<taxonomicName id="B4921EC0FFD0B56CFF52FE59FED2322A" ID-CoL="8VK4R" box="[151,368,503,529]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="555" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD0B56CFF52FE59FED2322A" bold="true" box="[151,368,503,529]" italics="true" pageId="1" pageNumber="555">Caridina boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD0B56CFEB2FE59FE18322A" box="[375,442,503,529]" pageId="1" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="732D6543FFD0B56CFF52FDB4FD8F3209" blockId="1.[151,557,503,562]" box="[151,557,538,562]" pageId="1" pageNumber="555">
(
<figureCitation id="EBA979C6FFD0B56CFF5AFDB4FF453209" box="[159,231,538,562]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Figs. 1</figureCitation>
A–Q, 2A–D, color plate 1A)
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD0B56CFF52FDF5FC9F357F" pageId="1" pageNumber="555" type="materials_examined">
<paragraph id="732D6543FFD0B56CFF52FDF5FCB53383" blockId="1.[151,1437,603,2032]" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF52FDF5FE22324F" bold="true" box="[151,384,603,628]" pageId="1" pageNumber="555">Material examined.</emphasis>
<typeStatus id="AC29DBE1FFD0B56CFE42FDF2FE4D324F" box="[391,495,604,628]" pageId="1" pageNumber="555" type="holotype">Holotype</typeStatus>
male, cl
<quantity id="B46AC8A6FFD0B56CFD90FDF2FD1B324F" box="[597,697,604,628]" metricMagnitude="-3" metricUnit="m" metricValue="3.27" pageId="1" pageNumber="555" unit="mm" value="3.27">3.27 mm</quantity>
(MZB Cru 3179),
<collectingCountry id="0B8525D3FFD0B56CFC48FDF2FC5C324F" box="[909,1022,604,628]" name="Indonesia" pageId="1" pageNumber="555">Indonesia</collectingCountry>
: Central
<taxonomicName id="B4921EC0FFD0B56CFBA4FDF2FB6B324F" box="[1121,1225,604,628]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Tomori area, road Beteleme-Tomata,
<geoCoordinate id="16A60384FFD0B56CFEA8FDD1FE4E32AC" box="[365,492,639,663]" direction="south" orientation="latitude" pageId="1" pageNumber="555" precision="1" value="-2.0995333">02°5.972´S</geoCoordinate>
<geoCoordinate id="16A60384FFD0B56CFE30FDD1FD2132AC" box="[501,643,639,663]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="1" value="121.1612">121°9.672´E</geoCoordinate>
(Loc. 25-05, on mixed substrate), leg. T. von Rintelen,
<date id="072C4383FFD0B56CFAC6FDD1FA3B32AC" box="[1283,1433,639,663]" pageId="1" pageNumber="555" value="2005-01-12">12 Jan. 2005</date>
.
<typeStatus id="AC29DBE1FFD0B56CFF52FD0BFEAA3286" box="[151,264,677,701]" pageId="1" pageNumber="555" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female, cl
<quantity id="B46AC8A6FFD0B56CFDD8FD0AFDD53287" box="[541,631,676,700]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="555" unit="mm" value="3.5">3.5 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFD40FD0AFD543287" box="[645,758,676,700]" pageId="1" pageNumber="555" type="female">4 females</specimenCount>
, cl 3.0–
<quantity id="B46AC8A6FFD0B56CFC94FD0AFC0E3287" box="[849,940,676,700]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="555" unit="mm" value="4.4">4.4 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFC7CFD0AFBB73287" box="[953,1045,676,700]" pageId="1" pageNumber="555" type="male">3 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFBFAFD0AFB6B3287" box="[1087,1225,676,700]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="1" pageNumber="555" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7–3.2 mm</quantity>
(MZB Cru 3180), same data as
<typeStatus id="AC29DBE1FFD0B56CFEF7FD69FE3532E4" box="[306,407,711,735]" pageId="1" pageNumber="555" type="holotype">holotype</typeStatus>
; 1 ovigerous female, cl
<quantity id="B46AC8A6FFD0B56CFD6BFD69FCAA32DB" box="[686,776,711,736]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="555" unit="mm" value="3.8">3.8 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFCD3FD69FC2732E4" box="[790,901,711,735]" pageId="1" pageNumber="555" type="female">4 females</specimenCount>
cl 3.2–4.0 mm,
<specimenCount id="6594AECAFFD0B56CFB80FD69FB3F32E4" box="[1093,1181,711,735]" pageId="1" pageNumber="555" type="male">4 males</specimenCount>
cl
<quantity id="B46AC8A6FFD0B56CFB01FD69FAEF32DB" box="[1220,1357,711,736]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="1" pageNumber="555" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4 mm</quantity>
(ZMB 29308), same data as
<typeStatus id="AC29DBE1FFD0B56CFE4FFD42FE4D333F" box="[394,495,748,772]" pageId="1" pageNumber="555" type="holotype">holotype</typeStatus>
. Others: 5 ovigerous females, cl
<quantity id="B46AC8A6FFD0B56CFCA4FD42FC48333F" box="[865,1002,748,772]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="1" pageNumber="555" unit="mm" value="4.0" valueMax="4.1" valueMin="3.9">3.9–4.1 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFC33FD42FBC4333F" box="[1014,1126,748,772]" pageId="1" pageNumber="555" type="female">4 females</specimenCount>
, cl 3.0–4.0 mm,
<specimenCount id="6594AECAFFD0B56CFAE6FD42FADE333F" box="[1315,1404,748,772]" pageId="1" pageNumber="555" type="male">3 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFF52FCA1FEBD3313" box="[151,287,783,808]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="1" pageNumber="555" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.1–3.3 mm</quantity>
(MZB Cru 3181); 8 ovigerous females, cl
<quantity id="B46AC8A6FFD0B56CFCC6FCA1FC2F3313" box="[771,909,783,808]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.6" metricValueMin="3.7" pageId="1" pageNumber="555" unit="mm" value="4.15" valueMax="4.6" valueMin="3.7">3.7–4.6 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFC5CFCA1FBAB331C" box="[921,1033,783,807]" pageId="1" pageNumber="555" type="female">8 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFBF4FCA1FB183313" box="[1073,1210,783,808]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.4" metricValueMin="3.3" pageId="1" pageNumber="555" unit="mm" value="3.85" valueMax="4.4" valueMin="3.3">3.3–4.4 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFB02FCBFFA82331C" box="[1223,1312,783,808]" pageId="1" pageNumber="555" type="male">7 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFA8DFCA1FF673377" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.7" metricValueMin="2.9" pageId="1" pageNumber="555" unit="mm" value="3.3" valueMax="3.7" valueMin="2.9">2.9–3.7 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFF15FC9AFEE03377" box="[208,322,820,844]" pageId="1" pageNumber="555" type="juvenile">1 juvenile</specimenCount>
, cl 2.0 mm (ZMB 29449),
<collectingCountry id="0B8525D3FFD0B56CFDA8FC9AFD7C3377" box="[621,734,820,844]" name="Indonesia" pageId="1" pageNumber="555">Indonesia</collectingCountry>
: Central
<taxonomicName id="B4921EC0FFD0B56CFC84FC9AFC0B3377" box="[833,937,820,844]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Tomori area, Puawu River, cascades at road Beteleme-Tomata, west of Wawopada,
<geoCoordinate id="16A60384FFD0B56CFD94FCF9FD723354" box="[593,720,855,879]" direction="south" orientation="latitude" pageId="1" pageNumber="555" precision="1" value="-2.0219">02°1.314´S</geoCoordinate>
<geoCoordinate id="16A60384FFD0B56CFD1DFCF9FCD63354" box="[728,884,855,879]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="1" value="121.2007">121°12.042´E</geoCoordinate>
(Loc. 221-05, on roots), leg. T. von Rintelen,
<date id="072C4383FFD0B56CFA45FCF9FEB833AF" pageId="1" pageNumber="555" value="2005-10-21">21 Oct. 2005</date>
; several specimen from commercial aquarium import sold as “
<taxonomicName id="B4921EC0FFD0B56CFBBDFCD2FB7D33AF" box="[1144,1247,892,916]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
Bee Shrimp” (OUMNH.ZC.2012-05-006, OUMNH.ZC.2012-05-007).
</paragraph>
<paragraph id="732D6543FFD0B56CFF0CFC6DFEB434AB" blockId="1.[151,1437,603,2032]" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF0CFC6DFDF633E7" bold="true" box="[201,596,963,988]" pageId="1" pageNumber="555">Comparative material examined.</emphasis>
<taxonomicName id="B4921EC0FFD0B56CFD99FC6AFCFA33E7" authority="Cai" authorityName="Cai" box="[604,856,964,988]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="555" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD0B56CFD99FC6AFC8B33E7" box="[604,809,964,988]" italics="true" pageId="1" pageNumber="555">Caridina sulawesi</emphasis>
Cai
</taxonomicName>
&amp; Ng, 2009.
<typeStatus id="AC29DBE1FFD0B56CFC31FC6BFBC733E6" box="[1012,1125,965,989]" pageId="1" pageNumber="555" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="6594AECAFFD0B56CFBB5FC6AFB6833E7" box="[1136,1226,964,988]" pageId="1" pageNumber="555" type="male">2 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFB37FC6AFA3533E7" box="[1266,1431,964,988]" metricMagnitude="-3" metricUnit="m" metricValue="3.525" metricValueMax="3.75" metricValueMin="3.3" pageId="1" pageNumber="555" unit="mm" value="3.525" valueMax="3.75" valueMin="3.3">3.30–3.75 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFF52FC46FEA4343B" box="[151,262,1000,1024]" pageId="1" pageNumber="555" type="female">2 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFEEEFC46FE6C343B" box="[299,462,1000,1024]" metricMagnitude="-3" metricUnit="m" metricValue="3.79" metricValueMax="3.98" metricValueMin="3.6" pageId="1" pageNumber="555" unit="mm" value="3.79" valueMax="3.98" valueMin="3.6">3.60–3.98 mm</quantity>
(ZRC), Sungei Beru, Kappang, Maros,
<taxonomicName id="B4921EC0FFD0B56CFC4EFC46FC52343B" box="[907,1008,1000,1024]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
Selatan, leg. P. Leclerc,
<date id="072C4383FFD0B56CFAC7FC46FA3A343B" box="[1282,1432,1000,1024]" pageId="1" pageNumber="555" value="1990-08-12">12 Aug.1990</date>
;
<specimenCount id="6594AECAFFD0B56CFF52FBA2FF4D341F" box="[151,239,1036,1060]" pageId="1" pageNumber="555" type="male">2 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFED3FBA2FE18341F" box="[278,442,1036,1060]" metricMagnitude="-3" metricUnit="m" metricValue="3.065" metricValueMax="3.18" metricValueMin="2.95" pageId="1" pageNumber="555" unit="mm" value="3.065" valueMax="3.18" valueMin="2.95">2.95–3.18 mm</quantity>
,
<collectingCountry id="0B8525D3FFD0B56CFE03FBA2FD95341F" box="[454,567,1036,1060]" name="Indonesia" pageId="1" pageNumber="555">Indonesia</collectingCountry>
: SW-Sulawesi, Bantaeng, near Loka village,
<geoCoordinate id="16A60384FFD0B56CFBEAFBA2FB1A341F" box="[1071,1208,1036,1060]" direction="south" orientation="latitude" pageId="1" pageNumber="555" precision="1" value="-5.425667">05°25.540'S</geoCoordinate>
<geoCoordinate id="16A60384FFD0B56CFB7AFBA2FAFB341F" box="[1215,1369,1036,1060]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="1" value="119.87095">119°52.257'E</geoCoordinate>
,
<quantity id="B46AC8A6FFD0B56CFAA1FBA2FF633473" metricMagnitude="2" metricUnit="m" metricValue="8.223504" pageId="1" pageNumber="555" unit="ft" value="2698.0">2698 feet</quantity>
asl, leg. H. G. Evers,
<date id="072C4383FFD0B56CFE7FFB9EFDFB3473" box="[442,601,1072,1096]" pageId="1" pageNumber="555" value="2010-09-21">21 Sept. 2010</date>
;
<specimenCount id="6594AECAFFD0B56CFDA3FB9EFD163473" box="[614,692,1072,1096]" pageId="1" pageNumber="555" type="male">1 male</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFD18FB9EFC9F3473" box="[733,829,1072,1096]" metricMagnitude="-3" metricUnit="m" metricValue="4.08" pageId="1" pageNumber="555" unit="mm" value="4.08">4.08mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFC8DFB9EFC0F3473" box="[840,941,1072,1096]" pageId="1" pageNumber="555" type="female">1 female</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFC13FB9EFB9F3473" box="[982,1085,1072,1096]" metricMagnitude="-3" metricUnit="m" metricValue="5.12" pageId="1" pageNumber="555" unit="mm" value="5.12">5.12 mm</quantity>
, personal collection of the first author (59-10),
<collectingCountry id="0B8525D3FFD0B56CFE8CFBFAFE183457" box="[329,442,1108,1132]" name="Indonesia" pageId="1" pageNumber="555">Indonesia</collectingCountry>
: South
<taxonomicName id="B4921EC0FFD0B56CFDCAFBFAFDD53457" box="[527,631,1108,1132]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Tilanga,
<geoCoordinate id="16A60384FFD0B56CFD23FBFAFCCD3457" box="[742,879,1108,1132]" direction="south" orientation="latitude" pageId="1" pageNumber="555" precision="1" value="-3.0354333">03°02.126'S</geoCoordinate>
<geoCoordinate id="16A60384FFD0B56CFCB2FBFAFBB33457" box="[887,1041,1108,1132]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="1" value="119.8872">119°53.232'E</geoCoordinate>
,
<quantity id="B46AC8A6FFD0B56CFBD8FBFAFB253457" box="[1053,1159,1108,1132]" metricMagnitude="2" metricUnit="m" metricValue="8.586216" pageId="1" pageNumber="555" unit="ft" value="2817.0">2817 feet</quantity>
asl, leg. H. G. Evers,
<date id="072C4383FFD0B56CFA45FBFAFEB034AB" pageId="1" pageNumber="555" value="2010-09-24">24 Sept. 2010</date>
.
</paragraph>
<paragraph id="732D6543FFD0B56CFF0CFB32FC9F357F" blockId="1.[151,1437,603,2032]" pageId="1" pageNumber="555">
<taxonomicName id="B4921EC0FFD0B56CFF0CFB32FDCC348F" authority="J. Roux, 1904" authorityName="J. Roux" authorityYear="1904" box="[201,622,1180,1204]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="555" phylum="Arthropoda" rank="species" species="parvidentata">
<emphasis id="41E6B951FFD0B56CFF0CFB32FE67348F" box="[201,453,1180,1204]" italics="true" pageId="1" pageNumber="555">Caridina parvidentata</emphasis>
J. Roux, 1904
</taxonomicName>
.
<typeStatus id="AC29DBE1FFD0B56CFDBFFB33FD4C348E" box="[634,750,1181,1205]" pageId="1" pageNumber="555" type="lectotype">Lectotype</typeStatus>
: ovigerous female, cl 4.0 mm (BNHM 8Ia), Malawa Quelle, Celebes (=
<taxonomicName id="B4921EC0FFD0B56CFED5FB6EFED934E3" box="[272,379,1216,1240]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
), leg. G. D. Sarasin.
<typeStatus id="AC29DBE1FFD0B56CFDAEFB6EFCB134E3" box="[619,787,1216,1240]" pageId="1" pageNumber="555" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="6594AECAFFD0B56CFCDAFB6EFCD834E3" box="[799,890,1216,1240]" pageId="1" pageNumber="555" type="male">3 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFC60FB6EFBEE34E3" box="[933,1100,1216,1240]" metricMagnitude="-3" metricUnit="m" metricValue="2.765" metricValueMax="2.93" metricValueMin="2.6" pageId="1" pageNumber="555" unit="mm" value="2.765" valueMax="2.93" valueMin="2.6">2.60–2.93 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFB9FFB6EFB6934E3" box="[1114,1227,1216,1240]" pageId="1" pageNumber="555" type="female">2 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFB30FB6EFA2234E3" box="[1269,1408,1216,1240]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" pageId="1" pageNumber="555" unit="mm" value="3.2" valueMax="3.8" valueMin="2.6">2.6–3.8 mm</quantity>
, 1 ovigerous female, cl 4.0 mm (BNHM 8Ia), Malawa Quelle, Celebes (5
<taxonomicName id="B4921EC0FFD0B56CFC10FB4AFB9C34C7" box="[981,1086,1252,1276]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
), leg. G. D. Sarasin. Others:
<specimenCount id="6594AECAFFD0B56CFA4BFB4AFF79351B" pageId="1" pageNumber="555" type="male">4 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFECBFAA6FE1B351B" box="[270,441,1288,1312]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" metricValueMax="3.05" metricValueMin="2.87" pageId="1" pageNumber="555" unit="mm" value="2.96" valueMax="3.05" valueMin="2.87">2.87–3.05 mm</quantity>
,
<specimenCount id="6594AECAFFD0B56CFE0EFAA6FDE2351B" box="[459,576,1288,1312]" pageId="1" pageNumber="555" type="female">3 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD0B56CFDB6FAA6FCBE351B" box="[627,796,1288,1312]" metricMagnitude="-3" metricUnit="m" metricValue="3.47" metricValueMax="4.03" metricValueMin="2.91" pageId="1" pageNumber="555" unit="mm" value="3.47" valueMax="4.03" valueMin="2.91">2.91–4.03 mm</quantity>
(OUMNH.ZC.2012-05-008),
<collectingCountry id="0B8525D3FFD0B56CFBB9FAA6FB4F351B" box="[1148,1261,1288,1312]" name="Indonesia" pageId="1" pageNumber="555">Indonesia</collectingCountry>
: SW-Sulawesi, Rantepao, tributary of Jalan Pongtiku, leg. R. Masche, 2005.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD0B56FFF02FAE1FD0635E1" lastPageId="2" lastPageNumber="556" pageId="1" pageNumber="555" type="description">
<paragraph id="732D6543FFD0B56CFF02FAE1FEFB3553" blockId="1.[151,1437,603,2032]" box="[199,345,1359,1384]" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF02FAE1FEFB3553" bold="true" box="[199,345,1359,1384]" pageId="1" pageNumber="555">Description.</emphasis>
</paragraph>
<paragraph id="732D6543FFD0B56CFF0CFADDFE2C3697" blockId="1.[151,1437,603,2032]" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF0CFADDFD1735B7" bold="true" box="[201,693,1395,1420]" pageId="1" pageNumber="555">Cephalothorax and cephalic appendages.</emphasis>
Carapace length 2.0–
<quantity id="B46AC8A6FFD0B56CFC6FFADAFBA235B7" box="[938,1024,1396,1420]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="555" unit="mm" value="4.6">4.6 mm</quantity>
(median
<quantity id="B46AC8A6FFD0B56CFBAFFADAFB6735B7" box="[1130,1221,1396,1420]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="555" unit="mm" value="3.5">3.5 mm</quantity>
). Rostrum straight, short and pointed (
<figureCitation id="EBA979C6FFD0B56CFEB6FA36FE14358B" box="[371,438,1432,1456]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
A), not reaching to end of basal segment of antennular peduncle mostly not reaching beyond length of eyes, 0.23–0.27 (median 0.25) times as long as carapace, unarmed or armed with 0–2 teeth on dorsal and ventral margin. Rostrum formula 0 + 0–2(0–1) / 0–2(0–1). Inferior orbital angle with a small antennal spine. Pterygostomian angle broadly rounded. Eyes well developed with globular cornea. Antennular peduncle 0.59–0.77 (median 0.64) times as long as carapace, first segment 1.97–2.2 (median 2.12) times as long as second segment, second segment 1.68–2.0 (median 1.95) times longer than third segment. Stylocerite reaching to 0.87– 0.96 (median 0.91) times of basal segment of antennular peduncle. Scaphocerite (
<figureCitation id="EBA979C6FFD0B56CFBE6F9DEFBC336B3" box="[1059,1121,1648,1672]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
D) 2.88–3.40 (median 3.10) times as long as wide.
</paragraph>
<paragraph id="732D6543FFD0B56CFF02F919FE0C3793" blockId="1.[151,1437,603,2032]" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF02F919FD0D36EB" bold="true" box="[199,687,1719,1744]" pageId="1" pageNumber="555">Abdominal somites, telson and uropods.</emphasis>
Sixth abdominal somite 0.49–0.54 (median 0.53) times carapace length, 1.47–1.79 (median 1.63) times as long as fifth somite, 0.87–0.94 (median 0.90) times as long as telson. Telson length (
<figureCitation id="EBA979C6FFD0B56CFEFAF8AEFEDD3723" box="[319,383,1792,1816]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
B, C) 2.90–3.02 (median 2.96) times as long as proximal wide, distal margin convex without median projection, with 3–4 pairs of dorsal spinules and one pair of dorsolateral spinules; distal end with 8–10 spines, sublateral pair distinctly shorter than lateral and inner spines. Preanal carina (
<figureCitation id="EBA979C6FFD0B56CFB8CF8E6FB25375B" box="[1097,1159,1864,1888]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
F) low, rounded, lacking a tooth or spine with few setae. Uropodal diaeresis (
<figureCitation id="EBA979C6FFD0B56CFD18F8C2FCB937BF" box="[733,795,1900,1924]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
E) with 15–16 movable spinules, outermost ones slightly longer than lateral angle.
</paragraph>
<paragraph id="732D6543FFD0B56FFF02F81AFEE831BD" blockId="1.[151,1437,603,2032]" lastBlockId="2.[151,1437,151,2034]" lastPageId="2" lastPageNumber="556" pageId="1" pageNumber="555">
<emphasis id="41E6B951FFD0B56CFF02F81AFDB737F7" bold="true" box="[199,533,1971,1996]" pageId="1" pageNumber="555">Mouthparts and branchiae.</emphasis>
Incisor process of mandible ending in irregular teeth, molar process truncated (
<figureCitation id="EBA979C6FFD0B56CFF5AF876FF4037CB" box="[159,226,2008,2032]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="1" pageNumber="555">Fig. 1</figureCitation>
G). Lower lacinia of maxillula broadly rounded, upper lacinia elongate, with numerous distinct teeth on inner margin, palp slender with few simple setae at tip (
<figureCitation id="EBA979C6FFD3B56FFCC5FF39FC9C3094" box="[768,830,151,175]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
H). Upper endites of maxilla subdivided, palp slender, scaphognathite tapering posteriorly, fringed with long, curved setae at posterior margin (
<figureCitation id="EBA979C6FFD3B56FFB61FF15FB4A30E8" box="[1188,1256,187,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">
Fig.
<date id="072C4383FFD3B56FFB19FF15FB4A30E8" box="[1244,1256,187,211]" pageId="2" pageNumber="556">1</date>
</figureCitation>
I). Palp of first maxilliped ending in a fingerlike extension (
<figureCitation id="EBA979C6FFD3B56FFD46FF71FD6030CC" box="[643,706,223,247]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
J). Podobranch on second maxilliped reduce to a lamina with few finger-like projections (
<figureCitation id="EBA979C6FFD3B56FFE6FFEADFE4D3120" box="[426,495,259,283]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
K). Third maxilliped with 2 arthrobranches, ultimate segment about as long as penultimate segment (
<figureCitation id="EBA979C6FFD3B56FFE53FE89FE783104" box="[406,474,295,319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
L). First pereiopod with an arthrobranch. Pleurobranchs present on all pereiopods. Well-developed (with hooks on distal end) epipods present on third maxilliped and first 2 pereiopods, lacking on pereiopods 3–5.
</paragraph>
<paragraph id="732D6543FFD3B56FFF0CFE3FFD62351C" blockId="2.[151,1437,151,2034]" pageId="2" pageNumber="556">
<emphasis id="41E6B951FFD3B56FFF0CFE3FFEF63191" bold="true" box="[201,340,401,426]" pageId="2" pageNumber="556">Pereiopods.</emphasis>
Chela and carpus of first pereiopod stouter and broader than chela and carpus of second pereiopod (
<figureCitation id="EBA979C6FFD3B56FFF5AFE18FF4231F5" box="[159,224,438,462]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
M, N); chela of first pereiopod 2.10–2.32 (median 2.11) times as long as wide, 1.12–1.14 (median 1.13) times length of carpus; tips of fingers rounded, without hook; dactylus 1.20–1.38 (median 1.36) times as long as palm; carpus just slightly excavated distally 2.35–2.57 (median 2.40) times as long as wide, 1.27–1.34 (median 1.32) times length of merus. Merus 2.42–2.52 (median 2.48) times as long as wide, shorter than ischium. Chela of second pereiopod 2.78–3.06 (median 2.89) times as long as wide, 0.73–0.85 (median 0.76) times length of carpus; tips of fingers rounded, without hook, dactylus 1.15–1.32 (median 1.29) times as long as palm; carpus 4.81–5.91 (median 5.22) times as long as wide, 1.26–1.46 (median 1.39) times as long as merus; merus 4.22–4.65 (median 4.40) times as long as wide, shorter than ischium. Third pereiopod slender (
<figureCitation id="EBA979C6FFD3B56FFC3EFD01FB9932FC" box="[1019,1083,687,711]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
O–Q), not sexually dimorphic, dactylus 3.46–3.62 (median 3.50) times as long as wide (terminal claw and spines on flexor margin included), terminating in one large claw with five accessory spines on flexor margin; propodus 10.07–11.00 (median 10.67) times as long as wide, 3.40–3.60 (median 3.44) times as long as dactylus; carpus 5.00–5.47 (median 5.06) times as long as wide, 0.59–0.67 (median 0.60) times as long as propodus, 0.51–0.59 (median 0.55) times as long as merus; merus 6.73–7.71 (median 7.17) times as long as wide, 1.68–1.95 (median 1.81) times as long as carpus, bearing 3 small movable spinules on posterior margin of outer surface. Ischium with a small spinule. Fifth pereiopod slender (
<figureCitation id="EBA979C6FFD3B56FFF5AFC04FF4133F9" box="[159,227,938,962]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
A, B), dactylus 4.00–4.57 (median 4.12) times as long as wide (terminal claw and spines on flexor margin included), terminating in one large claw with 50–55 (
<quantity id="B46AC8A6FFD3B56FFCC4FC60FC9C33DD" box="[769,830,974,998]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="2" pageNumber="556" unit="in" value="28.0">28 in</quantity>
one male specimen) spinules on flexor margin, first spinule distinctly stronger than posterior ones; propodus 9.71–11.73 (median 10.42) times as long as wide, 2.47– 2.89 (median 2.61) times length of dactylus, carpus 4.29–5.11 (median 4.93) times as long as wide, 0.51–0.57 (median 0.54) times as long as propodus, 0.66–0.71 (median 0.68) times as long as merus; merus 6.48–7.00 (median 6.83) times as long as wide, 1.41–1.51 (median 1.48) times length of carpus, bearing one or two very small movable spinules on posterior margin of outer surface.
<emphasis id="41E6B951FFD3B56FFCF1FB2FFC0434A2" bold="true" box="[820,934,1153,1177]" pageId="2" pageNumber="556">Pleopods.</emphasis>
Endopod of male first pleopod (
<figureCitation id="EBA979C6FFD3B56FFAF2FB2EFA2234A3" box="[1335,1408,1152,1176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
C) triangularly elongated, 2.28–2.44 times as long as proximal width (n=2), 0.38–0.39 times as long as exopod, without appendix interna. Appendix masculina on male second pleopod (
<figureCitation id="EBA979C6FFD3B56FFC0FFB66FBAE34DB" box="[970,1036,1224,1248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
D) slender (6.91–8.11 times as long as distal wide), saccate, with some long spines on inner and numerous long spines distal margin, appendix interna reaching to about 0.8–0.9 of appendix masculina.
</paragraph>
<paragraph id="732D6543FFD3B56FFF02FA9CFF683554" blockId="2.[151,1437,151,2034]" pageId="2" pageNumber="556">
<emphasis id="41E6B951FFD3B56FFF02FA9CFE6F3570" bold="true" box="[199,461,1330,1355]" pageId="2" pageNumber="556">Reproductive biology.</emphasis>
Ovigerous females with few eggs; size of eggs with developed 0.96–1.03 x
<quantity id="B46AC8A6FFD3B56FFAE9FA9DFF643554" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.4" metricValueMin="6.8" pageId="2" pageNumber="556" unit="mm" value="0.71" valueMax="0.74" valueMin="0.68">0.68–0.74 mm</quantity>
.
</paragraph>
<paragraph id="732D6543FFD3B56FFF02FAD4FD0635E1" blockId="2.[151,1437,151,2034]" pageId="2" pageNumber="556">
<emphasis id="41E6B951FFD3B56FFF02FAD4FEFD35A8" bold="true" box="[199,351,1402,1427]" pageId="2" pageNumber="556">Colouration.</emphasis>
Body semi-transparent. Three broad transverse blotches present on median part of the carapace, and on the third and the sixth abdominal segments. Remaining segments of abdomen and anterior part of the carapace with scattered white chromatophores.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD3B56FFF02FA48FD14361A" pageId="2" pageNumber="556" type="etymology">
<paragraph id="732D6543FFD3B56FFF02FA48FD14361A" blockId="2.[151,1437,151,2034]" pageId="2" pageNumber="556">
<emphasis id="41E6B951FFD3B56FFF02FA48FEEC35C5" bold="true" box="[199,334,1510,1534]" pageId="2" pageNumber="556">Etymology.</emphasis>
<emphasis id="41E6B951FFD3B56FFE9FFA48FD9335C5" box="[346,561,1510,1534]" italics="true" pageId="2" pageNumber="556">
<taxonomicName id="B4921EC0FFD3B56FFE9FFA48FD8F35C5" box="[346,557,1510,1534]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">Caridina boehmei</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="5AD5042AFFD3B56FFDFBFA49FD6F35C5" box="[574,717,1510,1534]" pageId="2" pageNumber="556" rank="species">new species</taxonomicNameLabel>
, is named after Joachim Böhme for his kind support of our research on
<taxonomicName id="B4921EC0FFD3B56FFEDBF9A7FE25361A" box="[286,391,1545,1569]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
´s freshwater shrimp fauna.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD3B56BFF02F980FCEA322C" lastPageId="6" lastPageNumber="560" pageId="2" pageNumber="556" type="discussion">
<paragraph id="732D6543FFD3B56FFF02F980FED537C9" blockId="2.[151,1437,151,2034]" pageId="2" pageNumber="556">
<emphasis id="41E6B951FFD3B56FFF02F980FE99367D" bold="true" box="[199,315,1582,1606]" pageId="2" pageNumber="556">Remarks.</emphasis>
<emphasis id="41E6B951FFD3B56FFE84F983FDB0367E" box="[321,530,1581,1605]" italics="true" pageId="2" pageNumber="556">
<taxonomicName id="B4921EC0FFD3B56FFE84F983FDAD367E" box="[321,527,1581,1605]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">Caridina boehmei</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="5AD5042AFFD3B56FFDDCF980FDF4367E" box="[537,598,1582,1605]" pageId="2" pageNumber="556" rank="species">n. sp.</taxonomicNameLabel>
, is similar to
<taxonomicName id="B4921EC0FFD3B56FFD2EF980FC38367E" authority="Cai" authorityName="Cai" box="[747,922,1581,1605]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFD2EF980FCC9367E" box="[747,875,1581,1605]" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
Cai
</taxonomicName>
&amp; Ng, 2009 showing a short and less strongly dentate rostrum and large-sized eggs. The new species can be distinguished from
<taxonomicName id="B4921EC0FFD3B56FFB8FF9FCFB6F3652" box="[1098,1229,1617,1641]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFB8FF9FCFB6F3652" box="[1098,1229,1617,1641]" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
by the shape and spinulation of the appendix masculina of the male second pleopod. This appendage appears saccate, i.e. 6.91–8.11 times as long as distal wide with numerous long spines on inner and distal margins, respectively in
<taxonomicName id="B4921EC0FFD3B56FFB3BF937FADD368B" box="[1278,1407,1688,1712]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD3B56FFB3BF937FADD368B" box="[1278,1407,1688,1712]" italics="true" pageId="2" pageNumber="556">C. boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD3B56FFA42F937FF1536EF" pageId="2" pageNumber="556" rank="species">n. sp.</taxonomicNameLabel>
vs. very slender (10.73–12.38 times as long as distal wide), stick-like and bearing few strong, short spinules arragned rougly in two rows along inner margin in
<taxonomicName id="B4921EC0FFD3B56FFD11F94FFCF636C3" box="[724,852,1760,1784]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFD11F94FFCF636C3" box="[724,852,1760,1784]" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
(
<figureCitation id="EBA979C6FFD3B56FFCA1F94EFC0A36C3" box="[868,936,1760,1784]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">
Fig.
<date id="072C4383FFD3B56FFC5DF94EFC0A36C3" box="[920,936,1760,1784]" pageId="2" pageNumber="556">2</date>
</figureCitation>
I). The endopod of male first pleopod is subtriangularly elongated, 2.28–2.44 times as long as wide and 0.38–0.39 times as long as endopod vs. sub-rectangular with distal margin angular, 2.87–3.27 times as long as wide and 0.47–0.65 times as long as exopod in
<taxonomicName id="B4921EC0FFD3B56FFADFF886FA3E3704" box="[1306,1436,1831,1855]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFADFF886FA3E3704" box="[1306,1436,1831,1855]" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
(
<figureCitation id="EBA979C6FFD3B56FFF5AF8E5FF443758" box="[159,230,1867,1891]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
H). Further, the two species can be distinguished by the shape of the preanal carina (low vs. high in
<taxonomicName id="B4921EC0FFD3B56FFA47F8E2FF5537BC" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFA47F8E2FF5537BC" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
), more slender carpus of first cheliped (2.35–2.57 times as long as wide vs. 1.51–2.06 times as long as wide in
<taxonomicName id="B4921EC0FFD3B56FFF36F83DFED63790" box="[243,372,1939,1963]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFF36F83DFED63790" box="[243,372,1939,1963]" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
) and by lacking sexual dimorphism of propodus of third pereiopod (with few spines arranged in 2 rows in males and females (
<figureCitation id="EBA979C6FFD3B56FFDD5F818FDF237F5" box="[528,592,1974,1998]" captionStart="FIGURE 1" captionStartId="3.[151,250,1916,1939]" captionTargetBox="[191,1376,193,1867]" captionTargetId="figure@3.[191,1378,193,1867]" captionTargetPageId="3" captionText="FIGURE 1. Caridina boehmei, n. sp., paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod." httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
Q) vs. propodus of third pereiopods with numerous spines in males of
<taxonomicName id="B4921EC0FFD3B56FFA47F819FF5537C9" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="556" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD3B56FFA47F819FF5537C9" italics="true" pageId="2" pageNumber="556">C. sulawesi</emphasis>
</taxonomicName>
(
<figureCitation id="EBA979C6FFD3B56FFEC3F874FEE837C9" box="[262,330,2010,2034]" captionStart="FIGURE 2" captionStartId="4.[151,250,1906,1929]" captionTargetBox="[208,1376,193,1889]" captionTargetId="figure@4.[208,1376,193,1889]" captionTargetPageId="4" captionText="FIGURE 2. Caridina boehmei, n. sp., paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod; Caridina sulawesi, male, cl 4.08 mm, South Sulawesi, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
G)).
</paragraph>
<caption id="27ED35CBFFD2B56EFF52F8D2FB9F37D5" httpUri="https://zenodo.org/record/215622/files/figure.png" pageId="3" pageNumber="557" targetBox="[191,1376,193,1867]" targetPageId="3">
<paragraph id="732D6543FFD2B56EFF52F8D2FB9F37D5" blockId="3.[151,1436,1916,2031]" pageId="3" pageNumber="557">
<emphasis id="41E6B951FFD2B56EFF52F8D2FEB037AA" bold="true" box="[151,274,1916,1939]" pageId="3" pageNumber="557">FIGURE 1.</emphasis>
<taxonomicName id="B4921EC0FFD2B56EFEDDF8D2FE6D37A9" box="[280,463,1916,1938]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="557" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD2B56EFEDDF8D2FE6D37A9" box="[280,463,1916,1938]" italics="true" pageId="3" pageNumber="557">Caridina boehmei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AD5042AFFD2B56EFE1EF8D3FDB037A9" box="[475,530,1917,1938]" pageId="3" pageNumber="557" rank="species">n. sp.</taxonomicNameLabel>
, paratype male, cl 3.1 mm, ZMB 29308; (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped; (M) first pereiopod; (N) second pereiopod; (O) third pereiopod; (P) dactylus of third pereiopod; (Q) inner margin of propodus of third pereiopod.
</paragraph>
</caption>
<caption id="27ED35CBFFD5B569FF52F8DCFAB637DF" httpUri="https://zenodo.org/record/215623/files/figure.png" pageId="4" pageNumber="558" targetBox="[208,1376,193,1889]" targetPageId="4">
<paragraph id="732D6543FFD5B569FF52F8DCFAB637DF" blockId="4.[151,1436,1906,2021]" pageId="4" pageNumber="558">
<emphasis id="41E6B951FFD5B569FF52F8DCFEB437B3" bold="true" box="[151,278,1906,1929]" pageId="4" pageNumber="558">FIGURE 2.</emphasis>
<taxonomicName id="B4921EC0FFD5B569FEE5F8DDFE7937B2" box="[288,475,1907,1929]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="558" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD5B569FEE5F8DDFE7937B2" box="[288,475,1907,1929]" italics="true" pageId="4" pageNumber="558">Caridina boehmei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AD5042AFFD5B569FE2EF8DAFD8537B2" box="[491,551,1908,1929]" pageId="4" pageNumber="558" rank="species">n. sp.</taxonomicNameLabel>
, paratype male, cl 3.1 mm (ZMB 29308); (A) fifth pereiopod; (B) dactylus of fifth pereiopod; (C) endopod of male first pleopod; (D) appendix masculina on male second pleopod;
<taxonomicName id="B4921EC0FFD5B569FB47F83FFA9E379C" box="[1154,1340,1937,1959]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="558" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD5B569FB47F83FFA9E379C" box="[1154,1340,1937,1959]" italics="true" pageId="4" pageNumber="558">Caridina sulawesi</emphasis>
</taxonomicName>
, male, cl 4.08 mm, South
<taxonomicName id="B4921EC0FFD5B569FE89F81EFE0837FD" box="[332,426,1968,1990]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Tilanga; (E) Cephalothorax and cephalic appendages; (F) first pereiopod; (G) inner margin of propodus of third pereiopod; (H) endopod of male first pleopod; (I) appendix masculina on male second pleopod.
</paragraph>
</caption>
<paragraph id="732D6543FFD4B568FF52F947FE753725" blockId="5.[151,1436,1769,1822]" pageId="5" pageNumber="559">
<emphasis id="41E6B951FFD4B568FF52F947FEDF36C4" bold="true" box="[151,381,1769,1792]" pageId="5" pageNumber="559">COLOUR PLATE 1.</emphasis>
Life coloration of (A)
<taxonomicName id="B4921EC0FFD4B568FDB5F944FC88373B" box="[624,810,1770,1792]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD4B568FDB5F944FC88373B" box="[624,810,1770,1792]" italics="true" pageId="5" pageNumber="559">Caridina boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD4B568FCF7F945FCC9373B" box="[818,875,1771,1792]" pageId="5" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
and (B)
<taxonomicName id="B4921EC0FFD4B568FC0FF944FB38373B" box="[970,1178,1770,1792]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="butonensis" status="sp. nov.">
<emphasis id="41E6B951FFD4B568FC0FF944FB38373B" box="[970,1178,1770,1792]" italics="true" pageId="5" pageNumber="559">Caridina butonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD4B568FB66F945FB7E373B" box="[1187,1244,1771,1792]" pageId="5" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
(latter photograph courtesy of Hans Georg Evers).
</paragraph>
<paragraph id="732D6543FFD4B568FF0CF8E8FAB937D5" blockId="5.[151,1436,1862,2030]" pageId="5" pageNumber="559">
The proposed new species can be differentiated from
<taxonomicName id="B4921EC0FFD4B568FC89F8E8FB5D3765" authority="J. Roux, 1904" authorityName="J. Roux" authorityYear="1904" box="[844,1279,1862,1886]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="parvidentata">
<emphasis id="41E6B951FFD4B568FC89F8E8FBEE3765" box="[844,1100,1862,1886]" italics="true" pageId="5" pageNumber="559">Caridina parvidentata</emphasis>
J. Roux, 1904
</taxonomicName>
by the more reduced epipodial apparatus (well developed epipods on first and second pereiopods vs. on first 3 pereiopods in
<taxonomicName id="B4921EC0FFD4B568FA47F8C4FE84379D" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="parvidentata">
<emphasis id="41E6B951FFD4B568FA47F8C4FE84379D" italics="true" pageId="5" pageNumber="559">C. parvidentata</emphasis>
</taxonomicName>
), the stouter scaphocerite (2.88–3.40 times as long as wide vs. 3.8 times in
<taxonomicName id="B4921EC0FFD4B568FB41F821FA97379D" box="[1156,1333,1934,1958]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="parvidentata">
<emphasis id="41E6B951FFD4B568FB41F821FA97379D" box="[1156,1333,1934,1958]" italics="true" pageId="5" pageNumber="559">C. parvidentata</emphasis>
</taxonomicName>
), a more slender carpus on first cheliped (2.35–2.57 times as long as wide vs. 1.8 times as long as wide in
<taxonomicName id="B4921EC0FFD4B568FB27F81CFA3037F1" box="[1250,1426,1970,1994]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="parvidentata">
<emphasis id="41E6B951FFD4B568FB27F81CFA3037F1" box="[1250,1426,1970,1994]" italics="true" pageId="5" pageNumber="559">C. parvidentata</emphasis>
</taxonomicName>
) and by the shape and spinulation of the appendix masculina (same differences as found in
<taxonomicName id="B4921EC0FFD4B568FB4FF879FAA937D5" box="[1162,1291,2006,2030]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD4B568FB4FF879FAA937D5" box="[1162,1291,2006,2030]" italics="true" pageId="5" pageNumber="559">C. sulawesi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="732D6543FFD7B56BFF0CFF39FCEA322C" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="560">
The specimens collected from Puawu River show some differences compared to the
<typeStatus id="AC29DBE1FFD7B56BFBBFFF36FB08308B" box="[1146,1194,152,176]" pageId="6" pageNumber="560">type</typeStatus>
material: the rostrum is longer (reaching to the end of the first segment of the antennular peduncle, 0.29–0.40 times as long as carapace vs. not reaching to the end of the basal segment of the antennular peduncle, mostly not reaching beyond the eyes, 0.23–0.27 times as long as carapace), armed with more teeth (rostral formula 0+2–8(4–6)/0–5(1–3) vs. 0+0–2(0–1) /0–2(0–1)), the podobranch of the second maxilliped reduce to a simple lamina vs. reduce to a lamina with few finger-like projections, the movable spinules on the posterior margin of the merus of the third and the fourth pereiopods are well developed (vs. small) and the first cheliped with a somewhat shorter carpus (0.76–0.87 times as long as chela vs. 0.87–0.90 times as long in the
<typeStatus id="AC29DBE1FFD7B56BFD6EFE3BFD793196" box="[683,731,405,429]" pageId="6" pageNumber="560">type</typeStatus>
specimen). Although there are some morphological variations, however, the general morphological characters of these Puawu River specimens are agree well with those of the
<typeStatus id="AC29DBE1FFD7B56BFF52FE73FF6531CE" box="[151,199,477,501]" pageId="6" pageNumber="560">type</typeStatus>
specimens along with low genetic differences in a mDNA analysis (von Rintelen et al. 2010). Therefore, we tentatively assign these specimens to
<taxonomicName id="B4921EC0FFD7B56BFDFFFDAEFCA73223" box="[570,773,512,536]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD7B56BFDFFFDAEFCA73223" box="[570,773,512,536]" italics="true" pageId="6" pageNumber="560">Caridina boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD7B56BFCC9FDAEFCEA322C" box="[780,840,512,535]" pageId="6" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD7B56BFF02FD8DFED93329" pageId="6" pageNumber="560" type="distribution">
<paragraph id="732D6543FFD7B56BFF02FD8DFBF132BF" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="560">
<emphasis id="41E6B951FFD7B56BFF02FD8DFE513207" bold="true" box="[199,499,547,572]" pageId="6" pageNumber="560">Distribution and habitat.</emphasis>
<taxonomicName id="B4921EC0FFD7B56BFE3FFD8AFD643207" box="[506,710,548,572]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="41E6B951FFD7B56BFE3FFD8AFD643207" box="[506,710,548,572]" italics="true" pageId="6" pageNumber="560">Caridina boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD7B56BFD0BFD8BFCA93207" box="[718,779,549,572]" pageId="6" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
is only known from two small rivers in Central
<taxonomicName id="B4921EC0FFD7B56BFAF5FD8AFA3B3207" box="[1328,1433,548,572]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
. At the
<typeStatus id="AC29DBE1FFD7B56BFF22FDE6FEB5325B" box="[231,279,584,608]" pageId="6" pageNumber="560">type</typeStatus>
locality the shrimp was observed on rocks, which is a rather unusual substrate for riverine
<taxonomicName id="B4921EC0FFD7B56BFAF5FDE6FA37325B" box="[1328,1429,584,608]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="41E6B951FFD7B56BFAF5FDE6FA37325B" box="[1328,1429,584,608]" italics="true" pageId="6" pageNumber="560">Caridina</emphasis>
</taxonomicName>
. The additional material was collected among roots near cascades of the Puawu River.
</paragraph>
<paragraph id="732D6543FFD7B56BFF52FD79FED93329" blockId="6.[151,428,727,786]" pageId="6" pageNumber="560">
<taxonomicName id="B4921EC0FFD7B56BFF52FD79FEC532CA" box="[151,359,727,753]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD7B56BFF52FD79FEC532CA" bold="true" box="[151,359,727,753]" italics="true" pageId="6" pageNumber="560">Caridina laroeha</emphasis>
</taxonomicName>
<emphasis id="41E6B951FFD7B56BFEABFD77FE0E32CA" bold="true" box="[366,428,729,753]" pageId="6" pageNumber="560">
<taxonomicNameLabel id="5AD5042AFFD7B56BFEABFD77FE0E32CA" box="[366,428,729,753]" pageId="6" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="EBA979C6FFD7B56BFF5AFD54FF483329" box="[159,234,762,786]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Figs. 3</figureCitation>
A–L, 4A–H)
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD7B56BFF0CFCECFC363585" pageId="6" pageNumber="560" type="materials_examined">
<paragraph id="732D6543FFD7B56BFF0CFCECFEBD33D1" blockId="6.[151,1437,834,2010]" pageId="6" pageNumber="560">
<emphasis id="41E6B951FFD7B56BFF0CFCECFE103360" bold="true" box="[201,434,834,859]" pageId="6" pageNumber="560">Material examined.</emphasis>
<typeStatus id="AC29DBE1FFD7B56BFE7CFCECFD833361" box="[441,545,834,858]" pageId="6" pageNumber="560" type="holotype">Holotype</typeStatus>
male, cl 4.0 mm (MZB Cru 3770),
<collectingCountry id="0B8525D3FFD7B56BFC75FCECFB833361" box="[944,1057,834,858]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
:
<taxonomicName id="B4921EC0FFD7B56BFBEEFCECFB313361" box="[1067,1171,834,858]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
,
<taxonomicName id="B4921EC0FFD7B56BFB5BFCECFB5F3361" box="[1182,1277,834,858]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="568" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">Laroeha</taxonomicName>
,
<geoCoordinate id="16A60384FFD7B56BFACDFCECFA353361" box="[1288,1431,834,858]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="1" value="-2.4704332">02°28.226’S</geoCoordinate>
,
<geoCoordinate id="16A60384FFD7B56BFF52FCC8FE943345" box="[151,310,870,894]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="1" value="121.06875">121°04.125’E</geoCoordinate>
, leg. F. Herder, 0
<date id="072C4383FFD7B56BFE30FCC8FDDD3345" box="[501,639,870,895]" pageId="6" pageNumber="560" value="2004-05-05">5 May 2004</date>
.
<typeStatus id="AC29DBE1FFD7B56BFD4CFCC9FD583344" box="[649,762,871,895]" pageId="6" pageNumber="560" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female, cl
<quantity id="B46AC8A6FFD7B56BFBC7FCC8FBF83345" box="[1026,1114,870,894]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="560" unit="mm" value="4.7">4.7 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFBA0FCC8FB763345" box="[1125,1236,870,894]" pageId="6" pageNumber="560" type="female">2 females</specimenCount>
, cl 4.0–
<quantity id="B46AC8A6FFD7B56BFAE9FCC8FA213345" box="[1324,1411,870,894]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="560" unit="mm" value="4.6">4.6 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFA4BFCC8FF7A3399" pageId="6" pageNumber="560" type="male">5 males</specimenCount>
cl
<quantity id="B46AC8A6FFD7B56BFF3CFC24FE223399" box="[249,384,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.5" metricValueMin="3.7" pageId="6" pageNumber="560" unit="mm" value="4.1" valueMax="4.5" valueMin="3.7">3.7–4.5 mm</quantity>
(MZB Cru 3771), same data as
<typeStatus id="AC29DBE1FFD7B56BFD21FC24FCE83399" box="[740,842,906,930]" pageId="6" pageNumber="560" type="holotype">holotype</typeStatus>
; 1 ovigerous female, cl
<quantity id="B46AC8A6FFD7B56BFB94FC24FB0B3399" box="[1105,1193,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="560" unit="mm" value="4.2">4.2 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFB71FC24FABD3399" box="[1204,1311,906,930]" pageId="6" pageNumber="560" type="female">1 females</specimenCount>
cl
<quantity id="B46AC8A6FFD7B56BFA85FC24FA353399" box="[1344,1431,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="560" unit="mm" value="4.6">4.6 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFF52FC00FF4D33FD" box="[151,239,942,966]" pageId="6" pageNumber="560" type="male">6 males</specimenCount>
, cl
<quantity id="B46AC8A6FFD7B56BFED3FC00FE3F33FD" box="[278,413,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.2" metricValueMin="3.3" pageId="6" pageNumber="560" unit="mm" value="3.75" valueMax="4.2" valueMin="3.3">3.3–4.2 mm</quantity>
(ZMB 28093) same data as
<typeStatus id="AC29DBE1FFD7B56BFD1FFC00FCE233FD" box="[730,832,942,966]" pageId="6" pageNumber="560" type="holotype">holotype</typeStatus>
. Others: numerous specimen (ZMB 28093) same data as
<typeStatus id="AC29DBE1FFD7B56BFF70FC7CFEB933D1" box="[181,283,978,1002]" pageId="6" pageNumber="560" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="732D6543FFD7B56BFF0CFC58FC363585" blockId="6.[151,1437,834,2010]" pageId="6" pageNumber="560">
<emphasis id="41E6B951FFD7B56BFF0CFC58FDF83434" bold="true" box="[201,602,1014,1039]" pageId="6" pageNumber="560">Comparative material examined.</emphasis>
<taxonomicName id="B4921EC0FFD7B56BFDA1FC58FC473435" authority="J. Roux, 1904" authorityName="J. Roux" authorityYear="1904" box="[612,997,1014,1038]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD7B56BFDA1FC58FC943435" box="[612,822,1014,1038]" italics="true" pageId="6" pageNumber="560">Caridina opaensis</emphasis>
J. Roux, 1904
</taxonomicName>
.
<specimenCount id="6594AECAFFD7B56BFC36FC58FBC43435" box="[1011,1126,1014,1038]" pageId="6" pageNumber="560" type="female">2 females</specimenCount>
, cl 3.4 and
<quantity id="B46AC8A6FFD7B56BFB36FC58FAEE3435" box="[1267,1356,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="560" unit="mm" value="3.7">3.7 mm</quantity>
(ZMB 29338),
<collectingCountry id="0B8525D3FFD7B56BFF33FBB4FEC53409" box="[246,359,1050,1074]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: Southeast
<taxonomicName id="B4921EC0FFD7B56BFE2CFBB4FDF33409" box="[489,593,1050,1074]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, stream in Pruiala (Mulyocinta), N of Aopa area,
<geoCoordinate id="16A60384FFD7B56BFB49FBB4FB5C3409" box="[1164,1278,1050,1074]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="9" value="-4.061167">04°3.67’S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFAC3FBB4FA3A3409" box="[1286,1432,1050,1074]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="1" value="122.11408">122°6.845’E</geoCoordinate>
, leg. M. Glaubrecht &amp; K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFD5EFB90FC9A346D" box="[667,824,1086,1111]" pageId="6" pageNumber="560" value="2005-05-31">31 May 2005</date>
;
<specimenCount id="6594AECAFFD7B56BFC80FB90FC34346D" box="[837,918,1086,1110]" pageId="6" pageNumber="560" type="male">1 male</specimenCount>
, cl
<quantity id="B46AC8A6FFD7B56BFC04FB90FBB9346D" box="[961,1051,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="560" unit="mm" value="3.1">3.1 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFBEDFB90FB38346D" box="[1064,1178,1086,1110]" pageId="6" pageNumber="560" type="female">3 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD7B56BFB01FB90FAEF346D" box="[1220,1357,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="6" pageNumber="560" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
(ZMB 29008),
<collectingCountry id="0B8525D3FFD7B56BFF30FBCCFEC43441" box="[245,358,1122,1146]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: Southeast
<taxonomicName id="B4921EC0FFD7B56BFE23FBCCFDEC3441" box="[486,590,1122,1146]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Benua River, northern arm,
<geoCoordinate id="16A60384FFD7B56BFC53FBCCFB803441" box="[918,1058,1122,1146]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="1" value="-4.223133">04°13.388’S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFBEFFBCCFB1E3441" box="[1066,1212,1122,1146]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="1" value="122.10661">122°6.397’E</geoCoordinate>
, leg. M. Glaubrecht &amp; K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFE77FB28FDEE34A5" box="[434,588,1158,1183]" pageId="6" pageNumber="560" value="2005-05-31">31 May 2005</date>
;
<specimenCount id="6594AECAFFD7B56BFD9DFB28FD1034A5" box="[600,690,1158,1182]" pageId="6" pageNumber="560" type="male">3 males</specimenCount>
, cl 3.8–4.0 mm,
<specimenCount id="6594AECAFFD7B56BFCB4FB28FC4234A5" box="[881,992,1158,1182]" pageId="6" pageNumber="560" type="female">3 females</specimenCount>
,
<quantity id="B46AC8A6FFD7B56BFC29FB28FBD634A5" box="[1004,1140,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="6" pageNumber="560" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5 mm</quantity>
(ZMB 29349),
<collectingCountry id="0B8525D3FFD7B56BFAEDFB28FA3B34A5" box="[1320,1433,1158,1182]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: Southeast
<taxonomicName id="B4921EC0FFD7B56BFED6FB04FED934F9" box="[275,379,1194,1218]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Poniponiki, NW of Raterate, road Kendari-Kolaka,
<geoCoordinate id="16A60384FFD7B56BFBC6FB04FB2334F9" box="[1027,1153,1194,1218]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="1" value="-4.03525">04°2.115’S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFB54FB04FA9234F9" box="[1169,1328,1194,1218]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="1" value="121.880264">121°52.816’E</geoCoordinate>
, leg. M. Glaubrecht &amp; K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFDF4FB60FD6834DD" box="[561,714,1230,1255]" pageId="6" pageNumber="560" value="2005-05-30">30 May 2005</date>
.
<taxonomicName id="B4921EC0FFD7B56BFD11FB60FB4334DD" authority="Cai, Wowor &amp; Choy 2009" authorityName="Cai, Wowor &amp; Choy" authorityYear="2009" box="[724,1249,1230,1254]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="560" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD7B56BFD11FB60FC1234DD" box="[724,944,1230,1254]" italics="true" pageId="6" pageNumber="560">Caridina mahalona</emphasis>
Cai, Wowor &amp; Choy 2009
</taxonomicName>
.
<specimenCount id="6594AECAFFD7B56BFB2EFB60FAE634DD" box="[1259,1348,1230,1254]" pageId="6" pageNumber="560" type="male">3 males</specimenCount>
, cl 3.8– 4.0 mm (ZMB 29085),
<collectingCountry id="0B8525D3FFD7B56BFE61FB5CFDB73531" box="[420,533,1266,1290]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: South
<taxonomicName id="B4921EC0FFD7B56BFDAEFB5CFD713531" box="[619,723,1266,1290]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Larona catchment, Patingko River, road to Tabarano, off road Malili-Soroako N of Balambano,
<geoCoordinate id="16A60384FFD7B56BFDCBFAB8FD393515" box="[526,667,1302,1326]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="1" value="-2.6370335">02°38.222’S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFD67FAB8FC933515" box="[674,817,1302,1326]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="9" value="121.24267">121°14.56’E</geoCoordinate>
(Loc. 14-03), leg. K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFB3FFAB8FA353515" box="[1274,1431,1302,1326]" pageId="6" pageNumber="560" value="2003-09-15">15 Sept. 2003</date>
;
<specimenCount id="6594AECAFFD7B56BFF52FA94FF473569" box="[151,229,1338,1362]" pageId="6" pageNumber="560" type="male">1 male</specimenCount>
, cl
<quantity id="B46AC8A6FFD7B56BFEC9FA94FED33569" box="[268,369,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="4.11" pageId="6" pageNumber="560" unit="mm" value="4.11">4.11 mm</quantity>
,
<specimenCount id="6594AECAFFD7B56BFEB9FA94FE4E3569" box="[380,492,1338,1362]" pageId="6" pageNumber="560" type="female">3 females</specimenCount>
, cl 4.0–
<quantity id="B46AC8A6FFD7B56BFD81FA94FD393569" box="[580,667,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="560" unit="mm" value="4.5">4.5 mm</quantity>
, 2 ovig. Females, cl
<quantity id="B46AC8A6FFD7B56BFC46FA94FBAE3569" box="[899,1036,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.9" metricValueMin="4.2" pageId="6" pageNumber="560" unit="mm" value="4.55" valueMax="4.9" valueMin="4.2">4.2–4.9 mm</quantity>
,
<collectingCountry id="0B8525D3FFD7B56BFBD2FA94FB2A3569" box="[1047,1160,1338,1362]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: South
<taxonomicName id="B4921EC0FFD7B56BFB1EFA94FAE13569" box="[1243,1347,1338,1362]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, at road Malili-Soroako, in
<collectingCountry id="0B8525D3FFD7B56BFEA8FAF1FE04354C" box="[365,422,1375,1399]" name="Togo" pageId="6" pageNumber="560">Togo</collectingCountry>
village,
<geoCoordinate id="16A60384FFD7B56BFDCDFAF0FD25354D" box="[520,647,1374,1398]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="9" value="-2.6088333">02°36.53´S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFD4AFAF0FCBC354D" box="[655,798,1374,1398]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="9" value="121.25933">121°15.56´E</geoCoordinate>
(loc. 34-03), leg. K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFB21FAF0FA20354D" box="[1252,1410,1374,1398]" pageId="6" pageNumber="560" value="2003-09-15">15 Sept. 2003</date>
;
<specimenCount id="6594AECAFFD7B56BFA4BFAF0FF5335A1" pageId="6" pageNumber="560" type="female">3 females</specimenCount>
, cl
<quantity id="B46AC8A6FFD7B56BFEDBFA2CFE0A35A1" box="[286,424,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.3" metricValueMin="4.2" pageId="6" pageNumber="560" unit="mm" value="4.75" valueMax="5.3" valueMin="4.2">4.2–5.3 mm</quantity>
(ZMB 29210),
<collectingCountry id="0B8525D3FFD7B56BFDA1FA2CFD7735A1" box="[612,725,1410,1434]" name="Indonesia" pageId="6" pageNumber="560">Indonesia</collectingCountry>
: South
<taxonomicName id="B4921EC0FFD7B56BFCEBFA2CFC3435A1" box="[814,918,1410,1434]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
, Towuti catchment, Bombongan,
<geoCoordinate id="16A60384FFD7B56BFAD8FA2CFA3935A1" box="[1309,1435,1410,1434]" direction="south" orientation="latitude" pageId="6" pageNumber="560" precision="9" value="-2.8730001">02°52.38’S</geoCoordinate>
<geoCoordinate id="16A60384FFD7B56BFF52FA08FE843585" box="[151,294,1446,1470]" direction="east" orientation="longitude" pageId="6" pageNumber="560" precision="9" value="121.37566">121°22.54’E</geoCoordinate>
(Loc. 49-03), leg. K. &amp; T. von Rintelen,
<date id="072C4383FFD7B56BFD37FA08FC333585" box="[754,913,1446,1470]" pageId="6" pageNumber="560" value="2003-09-20">20 Sept. 2003</date>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD7B564FF02FA64FE7C34D3" lastPageId="9" lastPageNumber="563" pageId="6" pageNumber="560" type="description">
<paragraph id="732D6543FFD7B56BFF02FA64FB023739" blockId="6.[151,1437,834,2010]" pageId="6" pageNumber="560">
<emphasis id="41E6B951FFD7B56BFF02FA64FCED35D8" bold="true" box="[199,847,1482,1507]" pageId="6" pageNumber="560">Description. Cephalothorax and cephalic appendages.</emphasis>
Carapace length
<quantity id="B46AC8A6FFD7B56BFBD3FA64FB1835D9" box="[1046,1210,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.56" metricValueMin="3.3" pageId="6" pageNumber="560" unit="mm" value="3.9299999999999997" valueMax="4.56" valueMin="3.3">3.30–4.56 mm</quantity>
(median
<quantity id="B46AC8A6FFD7B56BFAE2FA64FA3235D9" box="[1319,1424,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.82" pageId="6" pageNumber="560" unit="mm" value="3.82">3.82 mm</quantity>
). Rostrum (
<figureCitation id="EBA979C6FFD7B56BFECAFA40FEFA363D" box="[271,344,1518,1542]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Fig. 3</figureCitation>
A) long and slender, slightly sigmoid and upturned in distal part, reaching to or slightly overreaching distal end of scaphocerite, 0.8–1.2 (median 1.1) times as long as carapace, 0.4 distal dorsal margin unarmed, distal teeth of ventral margin more widely spaced. Rostrum formula 2–3 + 8–15 / 7–13. Antennal spine situated below inferior orbital angle. Pterygostomian angle rounded. Eyes well developed with globular cornea. Antennular peduncle 0.84–1.05 times as long as carapace in males, 0.70–0.83 times as long as carapace in females, first segment 1.45–1.68 (median 1.63) times as long as second segment, second segment 1.91–2.40 (median 2.18) times as long as third segment. Stylocerite reaching to, and 0.83–0.88 (median 0.88) times basal segment of antennular peduncle. Scaphocerite (
<figureCitation id="EBA979C6FFD7B56BFDE2F944FDCA3739" box="[551,616,1770,1794]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Fig. 3</figureCitation>
D) 3.42–4.08 (median 3.69) times as long as wide.
</paragraph>
<paragraph id="732D6543FFD7B56BFF0CF8A0FBAD37E1" blockId="6.[151,1437,834,2010]" pageId="6" pageNumber="560">
<emphasis id="41E6B951FFD7B56BFF0CF8A0FD12371C" bold="true" box="[201,688,1806,1831]" pageId="6" pageNumber="560">Abdominal somites, telson and uropods.</emphasis>
Sixth abdominal somite 0.68–0.78 (median 0.75) times carapace length, 1.69–1.96 (median 1.74) times as long as fifth somite, 0.94–1.12 (median 0.98) times as long as telson. Telson length (
<figureCitation id="EBA979C6FFD7B56BFEFAF8F8FE203755" box="[319,386,1878,1902]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Fig. 3</figureCitation>
B, C) 3.07–3.25 (median 3.23) times as long as proximal wide, distal margin convex without median projection, with 4–6 pairs of dorsal spinules and one pair of dorsolateral spinules; distal end with 7–9 rather short spines, lateral pair longer than inner spines. Preanal carina (
<figureCitation id="EBA979C6FFD7B56BFCBDF830FC18378D" box="[888,954,1950,1974]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Fig. 3</figureCitation>
F) with a distinct hook. Uropodal diaeresis (
<figureCitation id="EBA979C6FFD7B56BFF5AF86CFF4337E1" box="[159,225,1986,2010]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="6" pageNumber="560">Fig. 3</figureCitation>
E) with 9–12 movable spinules, outermost one shorter than lateral angle.
</paragraph>
<caption id="27ED35CBFFD6B56AFF52F8C4FBA53785" httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="7" pageNumber="561" targetBox="[151,1435,193,1876]" targetPageId="7">
<paragraph id="732D6543FFD6B56AFF52F8C4FBA53785" blockId="7.[151,1436,1898,1983]" pageId="7" pageNumber="561">
<emphasis id="41E6B951FFD6B56AFF52F8C4FEB137BB" bold="true" box="[151,275,1898,1921]" pageId="7" pageNumber="561">FIGURE 3.</emphasis>
<taxonomicName id="B4921EC0FFD6B56AFEDCF8C5FE6B37BA" box="[281,457,1899,1921]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="561" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD6B56AFEDCF8C5FE6B37BA" box="[281,457,1899,1921]" italics="true" pageId="7" pageNumber="561">Caridina laroeha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AD5042AFFD6B56AFE13F8C2FDAC37BA" box="[470,526,1900,1921]" pageId="7" pageNumber="561" rank="species">n. sp.</taxonomicNameLabel>
, paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped.
</paragraph>
</caption>
<caption id="27ED35CBFFD9B565FF52F904FC6936C6" httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="8" pageNumber="562" targetBox="[188,1398,193,1684]" targetPageId="8">
<paragraph id="732D6543FFD9B565FF52F904FC6936C6" blockId="8.[151,1436,1706,1790]" pageId="8" pageNumber="562">
<emphasis id="41E6B951FFD9B565FF52F904FEB136FB" bold="true" box="[151,275,1706,1729]" pageId="8" pageNumber="562">FIGURE 4.</emphasis>
<taxonomicName id="B4921EC0FFD9B565FEDFF904FE6936FB" box="[282,459,1706,1728]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="562" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD9B565FEDFF904FE6936FB" box="[282,459,1706,1728]" italics="true" pageId="8" pageNumber="562">Caridina laroeha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AD5042AFFD9B565FE1DF905FDB336FB" box="[472,529,1707,1728]" pageId="8" pageNumber="562" rank="species">n. sp.</taxonomicNameLabel>
, paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod.
</paragraph>
</caption>
<paragraph id="732D6543FFD9B565FF0CF88BFA3E37CA" blockId="8.[151,1436,1829,2034]" pageId="8" pageNumber="562">
<emphasis id="41E6B951FFD9B565FF0CF88BFDB43705" bold="true" box="[201,534,1829,1854]" pageId="8" pageNumber="562">Mouthparts and branchiae.</emphasis>
Incisor process of mandible ending in irregular teeth, molar process truncated (
<figureCitation id="EBA979C6FFD9B565FF5AF8E7FF47375A" box="[159,229,1865,1889]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">Fig. 3</figureCitation>
G). Lower lacinia of maxillula broadly rounded, upper lacinia elongate, with numerous distinct teeth on inner margin, palp slender with few simple setae (
<figureCitation id="EBA979C6FFD9B565FD21F8C3FC8837BE" box="[740,810,1901,1925]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">Fig. 3</figureCitation>
H). Upper endites of maxilla subdivided palp slender, scaphognathite tapering posteriorly, fringed with long, curved setae at posterior margin (
<figureCitation id="EBA979C6FFD9B565FB61F83FFB493792" box="[1188,1259,1937,1961]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">
Fig.
<date id="072C4383FFD9B565FB19F83FFB493792" box="[1244,1259,1937,1961]" pageId="8" pageNumber="562">3</date>
</figureCitation>
I). Palp of first maxilliped ending in a broad triangular extension (
<figureCitation id="EBA979C6FFD9B565FD1CF81BFCBE37F6" box="[729,796,1973,1997]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">Fig. 3</figureCitation>
J). Podobranch on second maxilliped reduce to a lamina (
<figureCitation id="EBA979C6FFD9B565FF5AF877FF4637CA" box="[159,228,2009,2033]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">Fig. 3</figureCitation>
K). Third maxilliped with 2 arthrobranches, ultimate segment about as long as penultimate segment (
<figureCitation id="EBA979C6FFD9B565FAAAF877FA3E37CA" box="[1391,1436,2009,2033]" captionStart="FIGURE 3" captionStartId="7.[151,250,1898,1921]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 3. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) Cephalothorax and cephalic appendages; (B) telson; (C) distal end of telson; (D) scaphocerite; (E) uropodal diaeresis; (F) preanal carina; (G) mandible; (H) maxillula; (I) maxilla; (J) first maxilliped; (K) second maxilliped; (L) third maxilliped." httpUri="https://zenodo.org/record/215624/files/figure.png" pageId="8" pageNumber="562">Fig.</figureCitation>
</paragraph>
<paragraph id="732D6543FFD8B564FF52FF39FC3130CC" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="563">3L). First pereiopod with an arthrobranch, pleurobranchs present on all pereiopods. Well developed (with hooks on distal end) epipods present on third maxilliped and first 2 pereiopods, reduced epipod (short, without distal hook) present on third pereiopod and epipod absent on pereiopods 4 and 5.</paragraph>
<paragraph id="732D6543FFD8B564FF0CFEADFB563447" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="563">
<emphasis id="41E6B951FFD8B564FF0CFEADFEF63127" bold="true" box="[201,340,259,284]" pageId="9" pageNumber="563">Pereiopods.</emphasis>
Chela and carpus of first pereiopod stouter and broader than chela and carpus of second pereiopod (
<figureCitation id="EBA979C6FFD8B564FF5AFE89FF443104" box="[159,230,295,319]" captionStart="FIGURE 4" captionStartId="8.[151,250,1706,1729]" captionTargetBox="[188,1398,193,1684]" captionTargetId="figure@8.[188,1398,193,1685]" captionTargetPageId="8" captionText="FIGURE 4. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="9" pageNumber="563">Fig. 4</figureCitation>
A, B); chela of first pereiopod 2.19–2.32 (median 2.23) times as long as wide, 1.20–1.34 (median 1.30) times length of carpus; tips of fingers rounded, without hook; dactylus 0.97–1.24 (median1.20) times as long as palm; carpus scarcely excavated distally, 2.00–2.66 (median 2.26) times as long as wide, 1.09–1.20 (median 1.14) times length of merus. Merus 2.42–2.84 (median 2.68) times as long as wide, as long as ischium. Chela of second pereiopod 2.83–3.09 (median 3.04) times as long as wide, 0.78–0.85 (median 0.79) times length of carpus; tips of fingers rounded, without hook, dactylus 1.18–1.57 (median 1.39) times as long as palm; carpus 4.83–5.73 (median 5.24) times as long as wide, 1.30–1.45 (median 1.40) times as long as merus; merus 4.42–4.76 (median 4.48) times as long as wide, as long as ischium. Third pereiopod slender (
<figureCitation id="EBA979C6FFD8B564FC96FD8AFC3A3207" box="[851,920,548,572]" captionStart="FIGURE 4" captionStartId="8.[151,250,1706,1729]" captionTargetBox="[188,1398,193,1684]" captionTargetId="figure@8.[188,1398,193,1685]" captionTargetPageId="8" captionText="FIGURE 4. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="9" pageNumber="563">Fig. 4</figureCitation>
C, D), not sexually dimorphic, dactylus 3.94– 4.29 (median 4.16) times as long as wide (terminal claw and spines on flexor margin included), terminating in one large claw with 6–9 accessory spines on flexor margin; propodus 11.61–14.00 (median 12.41) times as long as wide, 3.41–3.92 (median 3.57) times as long as dactylus; carpus 5.44–5.62 (median 5.53) times as long as wide, 0.56–0.63 (median 0.60) times as long as propodus, 0.46–0.55 (median 0.51) times as long as merus; merus 8.77– 9.17 (median 8.88) times as long as wide, 1.82–2.18 (median 1.99) times as long as carpus, bearing 2 or 3 movable spinules on posterior margin of outer surface. Ischium with a small spinule. Fifth pereiopod slender (
<figureCitation id="EBA979C6FFD8B564FAD2FD52FAFE332F" box="[1303,1372,764,788]" captionStart="FIGURE 4" captionStartId="8.[151,250,1706,1729]" captionTargetBox="[188,1398,193,1684]" captionTargetId="figure@8.[188,1398,193,1685]" captionTargetPageId="8" captionText="FIGURE 4. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="9" pageNumber="563">Fig. 4</figureCitation>
E, F), dactylus 3.75–5.20 (median 4.84) times as long as wide (terminal claw and spines on flexor margin included), terminating in one large claw with 33–60 spinules on flexor margin, propodus 11.45–13.00 (median 12.52) times as long as wide, 2.80–3.65 (median 2.97) times length of dactylus, carpus 4.60–6.28 (median 4.88) times as long as wide, 0.48–0.54 (median 0.50) times as long as propodus, 0.57–0.61 (median 0.61) times as long as merus; merus 7.04–9.35 (median 7.76) times as long as wide, 1.63–1.77 (median 1.65) times length of carpus, bearing 3 or 4 movable spinules on posterior margin of outer surface.
<emphasis id="41E6B951FFD8B564FCF6FC7AFC0733D7" bold="true" box="[819,933,980,1004]" pageId="9" pageNumber="563">Pleopods.</emphasis>
Endopod of male first pleopod (
<figureCitation id="EBA979C6FFD8B564FAF0FC7AFADC33D7" box="[1333,1406,980,1004]" captionStart="FIGURE 4" captionStartId="8.[151,250,1706,1729]" captionTargetBox="[188,1398,193,1684]" captionTargetId="figure@8.[188,1398,193,1685]" captionTargetPageId="8" captionText="FIGURE 4. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="9" pageNumber="563">Fig. 4</figureCitation>
G) subtriangular, long setae fringe along outer and distal margins, few medium-length setae fringe along inner margin, 2.07–2.18 times as long as proximal width (n=2), 0.26–0.27 times as long as exopod, without appendix interna. Appendix masculina on male second pleopod (
<figureCitation id="EBA979C6FFD8B564FD61FB91FD45346C" box="[676,743,1087,1111]" captionStart="FIGURE 4" captionStartId="8.[151,250,1706,1729]" captionTargetBox="[188,1398,193,1684]" captionTargetId="figure@8.[188,1398,193,1685]" captionTargetPageId="8" captionText="FIGURE 4. Caridina laroeha, n. sp., paratype male, cl 3.6 mm (ZMB 28093, Präp. Nr. 5028) (A) first pereiopod; (B) second pereiopod; (C) third pereiopod; (D) dactylus of third pereiopod; (E) fifth pereiopod; (F) dactylus of fifth pereiopod; (G) endopod of male first pleopod; (H) appendix masculina on male second pleopod." httpUri="https://zenodo.org/record/215625/files/figure.png" pageId="9" pageNumber="563">Fig. 4</figureCitation>
H) slender, 7.8 times as long as distal width, rod-shaped, with some long spines on inner and distal margins, appendix interna 0.85 as long as appendix masculina.
</paragraph>
<paragraph id="732D6543FFD8B564FF0CFB29FDB334FF" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="563">
<emphasis id="41E6B951FFD8B564FF0CFB29FE72349B" bold="true" box="[201,464,1159,1184]" pageId="9" pageNumber="563">Reproductive biology.</emphasis>
Ovigerous females with few eggs; size of eggs (from one female with undeveloped eggs) 0.94–0.99 x
<quantity id="B46AC8A6FFD8B564FEA3FB02FDAE34FF" box="[358,524,1196,1220]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.0" metricValueMin="5.699999999999999" pageId="9" pageNumber="563" unit="mm" value="0.585" valueMax="0.6" valueMin="0.57">0.57–0.60 mm</quantity>
.
</paragraph>
<paragraph id="732D6543FFD8B564FF0CFB61FE7C34D3" blockId="9.[151,1437,151,2013]" box="[201,478,1231,1256]" pageId="9" pageNumber="563">
<emphasis id="41E6B951FFD8B564FF0CFB61FEC334D3" bold="true" box="[201,353,1231,1256]" pageId="9" pageNumber="563">Colouration.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD8B564FF0CFB5AFAE13537" box="[201,1347,1268,1293]" pageId="9" pageNumber="563" type="etymology">
<paragraph id="732D6543FFD8B564FF0CFB5AFAE13537" blockId="9.[151,1437,151,2013]" box="[201,1347,1268,1293]" pageId="9" pageNumber="563">
<emphasis id="41E6B951FFD8B564FF0CFB5AFEF23537" bold="true" box="[201,336,1268,1292]" pageId="9" pageNumber="563">Etymology.</emphasis>
The new species is named after its
<typeStatus id="AC29DBE1FFD8B564FD1BFB5BFCAC3536" box="[734,782,1269,1293]" pageId="9" pageNumber="563">type</typeStatus>
locality – the
<taxonomicName id="B4921EC0FFD8B564FC6BFB5AFBA83537" box="[942,1034,1268,1292]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="568" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">Laroeha</taxonomicName>
River,
<taxonomicName id="B4921EC0FFD8B564FB9EFB5AFB613537" box="[1115,1219,1268,1292]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
,
<collectingCountry id="0B8525D3FFD8B564FB0AFB5AFA9D3537" box="[1231,1343,1268,1292]" name="Indonesia" pageId="9" pageNumber="563">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFD8B567FF0CFAB6FAE931F4" lastPageId="10" lastPageNumber="564" pageId="9" pageNumber="563" type="discussion">
<paragraph id="732D6543FFD8B564FF0CFAB6FD5B3777" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="563">
<emphasis id="41E6B951FFD8B564FF0CFAB6FE9F350B" bold="true" box="[201,317,1304,1328]" pageId="9" pageNumber="563">Remarks.</emphasis>
With its slender rostrum and unarmed distorostral dorsal margin,
<taxonomicName id="B4921EC0FFD8B564FBDAFAB6FB40350B" box="[1055,1250,1304,1328]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD8B564FBDAFAB6FB40350B" box="[1055,1250,1304,1328]" italics="true" pageId="9" pageNumber="563">Caridina laroeha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD8B564FB2CFAB7FA87350B" box="[1257,1317,1305,1328]" pageId="9" pageNumber="563" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="B4921EC0FFD8B564FF52FA93FEBE356F" box="[151,284,1341,1364]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FF52FA93FEBE356F" box="[151,284,1341,1364]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B4921EC0FFD8B564FE93FA93FE4B356F" box="[342,489,1340,1364]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD8B564FE93FA93FE4B356F" box="[342,489,1340,1364]" italics="true" pageId="9" pageNumber="563">C. mahalona</emphasis>
</taxonomicName>
.
<taxonomicName id="B4921EC0FFD8B564FE3CFA92FD1C356F" box="[505,702,1340,1364]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD8B564FE3CFA92FD1C356F" box="[505,702,1340,1364]" italics="true" pageId="9" pageNumber="563">Caridina laroeha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD8B564FD02FA93FCA7356F" box="[711,773,1341,1364]" pageId="9" pageNumber="563" rank="species">n. sp.</taxonomicNameLabel>
differs also from
<taxonomicName id="B4921EC0FFD8B564FC13FA93FBF9356F" box="[982,1115,1341,1364]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FC13FA93FBF9356F" box="[982,1115,1341,1364]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
by having a longer rostrum (reaching to or slightly overreaching distal end of scaphocerite vs. mostly reaching to or slightly overreaching third segment of antennular peduncle in
<taxonomicName id="B4921EC0FFD8B564FDE2FA2BFD0E35A7" box="[551,684,1413,1436]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FDE2FA2BFD0E35A7" box="[551,684,1413,1436]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
) with less postorbital teeth (2–3 vs.
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<taxonomicName id="B4921EC0FFD8B564FB65FA2BFA8635A7" box="[1184,1316,1413,1436]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FB65FA2BFA8635A7" box="[1184,1316,1413,1436]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
) but more ventral teeth (7–13 vs.
<quantity id="B46AC8A6FFD8B564FE6BFA06FE5835FB" box="[430,506,1448,1472]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.524" metricValueMin="0.762" pageId="9" pageNumber="563" unit="in" value="4.5" valueMax="6.0" valueMin="3.0">3–6 in</quantity>
<taxonomicName id="B4921EC0FFD8B564FDC3FA06FD2C35FB" box="[518,654,1448,1472]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FDC3FA06FD2C35FB" box="[518,654,1448,1472]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
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) and 0.25 distal dorsal margin unarmed (vs. less than
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<taxonomicName id="B4921EC0FFD8B564FA47FA06FF5B35DF" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FA47FA06FF5B35DF" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
). Penultimate segment of third maxilliped as long as or slightly longer than ultimate vs. much longer in
<taxonomicName id="B4921EC0FFD8B564FA47FA63FF5B3633" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FA47FA63FF5B3633" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
). The sixth abdominal segment is also longer (0.68–0.78 times carapace length vs. 0.53–0.63 times carapace length in
<taxonomicName id="B4921EC0FFD8B564FEB5F9BBFE573617" box="[368,501,1557,1580]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FEB5F9BBFE573617" box="[368,501,1557,1580]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
), the number of the spinule on uropodal diaeresis is lower (9–11 vs.
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<taxonomicName id="B4921EC0FFD8B564FA47F9BBFF5B366B" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FA47F9BBFF5B366B" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
), the number of the spinule on the dorsal margin of the telson is higher (4–6 pairs vs. 3–4 pairs in
<taxonomicName id="B4921EC0FFD8B564FA47F996FF5B364F" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FA47F996FF5B364F" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
), and the lateral spines on the distal end of the telson are longer than the inner spines (vs. as long as the inner spines with the second pair shorter in
<taxonomicName id="B4921EC0FFD8B564FD40F92EFCAB36A3" box="[645,777,1664,1688]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FD40F92EFCAB36A3" box="[645,777,1664,1688]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
). However, 2 out of 6 of the comparative specimens of
<taxonomicName id="B4921EC0FFD8B564FA47F92EFF5B3687" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FA47F92EFF5B3687" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
from Poniponiki (ZMB 29349) have relatively longer rostrum which are reaching to the end of the scaphocerite. Thus, these specimens resemble
<taxonomicName id="B4921EC0FFD8B564FD59F966FCB136DB" box="[668,787,1736,1760]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD8B564FD59F966FCB136DB" box="[668,787,1736,1760]" italics="true" pageId="9" pageNumber="563">C. laroeha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD8B564FCDCF966FCF736E4" box="[793,853,1736,1759]" pageId="9" pageNumber="563" rank="species">n. sp.</taxonomicNameLabel>
in shape and length of the rostrum. Other characters of these
<specimenCount id="6594AECAFFD8B564FF3FF942FE24373F" box="[250,390,1772,1796]" pageId="9" pageNumber="563" type="generic">2 specimens</specimenCount>
of
<taxonomicName id="B4921EC0FFD8B564FE75F943FD97373F" box="[432,565,1773,1796]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="opaensis">
<emphasis id="41E6B951FFD8B564FE75F943FD97373F" box="[432,565,1773,1796]" italics="true" pageId="9" pageNumber="563">C. opaensis</emphasis>
</taxonomicName>
from Poniponiki (e.g. number of postorbital teeth and spinules on uropodal diaeresis, relative length of segments of third maxilliped 6. abdominal segement and number and length of spines on telson) resembles typical specimens of
<taxonomicName id="B4921EC0FFD8B564FDB4F89BFD503777" box="[625,754,1844,1868]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="sulawesi">
<emphasis id="41E6B951FFD8B564FDB4F89BFD503777" box="[625,754,1844,1868]" italics="true" pageId="9" pageNumber="563">C. sulawesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="732D6543FFD8B567FF0CF8F6FAE931F4" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1436,151,535]" lastPageId="10" lastPageNumber="564" pageId="9" pageNumber="563">
<taxonomicName id="B4921EC0FFD8B564FF0CF8F6FE2E374B" box="[201,396,1880,1904]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFD8B564FF0CF8F6FE2E374B" box="[201,396,1880,1904]" italics="true" pageId="9" pageNumber="563">Caridina laroeha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFD8B564FE51F8F7FE73374B" box="[404,465,1881,1904]" pageId="9" pageNumber="563" rank="species">n. sp.</taxonomicNameLabel>
differs also from
<taxonomicName id="B4921EC0FFD8B564FD5BF8F6FC92374B" box="[670,816,1880,1904]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD8B564FD5BF8F6FC92374B" box="[670,816,1880,1904]" italics="true" pageId="9" pageNumber="563">C. mahalona</emphasis>
</taxonomicName>
by having a longer rostrum (0.82–1.20 times carapace length vs. 0.52–0.75 carapace length in
<taxonomicName id="B4921EC0FFD8B564FDADF8D3FD5E37AF" box="[616,764,1916,1940]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD8B564FDADF8D3FD5E37AF" box="[616,764,1916,1940]" italics="true" pageId="9" pageNumber="563">C. mahalona</emphasis>
</taxonomicName>
), less number of ventral rostral teeth (8–13 vs.
<quantity id="B46AC8A6FFD8B564FAEBF8D2FADA37AF" box="[1326,1400,1916,1940]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="17.78" metricValueMin="0.0" pageId="9" pageNumber="563" unit="in" value="3.5" valueMax="7.0" valueMin="0.0">0–7 in</quantity>
C.
<taxonomicName id="B4921EC0FFD8B564FF52F80EFEA53783" box="[151,263,1952,1976]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD8B564FF52F80EFEA53783" box="[151,263,1952,1976]" italics="true" pageId="9" pageNumber="563">mahalona</emphasis>
</taxonomicName>
), shorter stylocerite (0.83–0.88 as long as the first segment of antennular peduncle vs. 0.9–1.0 as long as the first segment of antennular peduncle in
<taxonomicName id="B4921EC0FFD8B564FD71F86BFCEF37E7" box="[692,845,1988,2012]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="563" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFD8B564FD71F86BFCEF37E7" box="[692,845,1988,2012]" italics="true" pageId="9" pageNumber="563">C. mahalona</emphasis>
</taxonomicName>
), relatively shorter penultimate segment of third maxilliped (as long as or slightly longer than ultimate vs. much longer penultimate in
<taxonomicName id="B4921EC0FFDBB567FBA9FF36FB5C308B" box="[1132,1278,152,176]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FBA9FF36FB5C308B" box="[1132,1278,152,176]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), epipod well developed on the second pereiopod but reduced on the third pereiopod (vs. absent from the third to the fifth pereiopods in
<taxonomicName id="B4921EC0FFDBB567FEF2FF4EFE6A30C3" box="[311,456,224,248]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FEF2FF4EFE6A30C3" box="[311,456,224,248]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), the carpus of the first and the second chelipeds are stouter (2.00–2.66 and 4.83–5.73 times as long as wide vs. 2.75–3.44 and 5.39–7.07 times as long as wide, respectivelyin
<taxonomicName id="B4921EC0FFDBB567FBBAFEABFAB23127" box="[1151,1296,260,284]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FBBAFEABFAB23127" box="[1151,1296,260,284]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), the carpus of the first pereiopod is relatively short (1.09–1.20 as long as merus vs. 1.33–1.44 as long as merus in
<taxonomicName id="B4921EC0FFDBB567FA47FE86FEA5315F" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FA47FE86FEA5315F" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), more spines on the dactylus of the third pereiopod (6–9 vs.
<quantity id="B46AC8A6FFDBB567FC08FEE2FBB7315F" box="[973,1045,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="564" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
<taxonomicName id="B4921EC0FFDBB567FBDBFEE3FB12315F" box="[1054,1200,332,356]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FBDBFEE3FB12315F" box="[1054,1200,332,356]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), more slender sixth abdominal segment (0.68–0.78 times as long as carapace vs. 0.54–0.61 times as long as carapace in
<taxonomicName id="B4921EC0FFDBB567FB3EFEDEFA2E31B3" box="[1275,1420,368,392]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FB3EFEDEFA2E31B3" box="[1275,1420,368,392]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
), less number of spinule on the uropodal diaeresis (9–12 vs.
<quantity id="B46AC8A6FFDBB567FCE3FE3AFC2A3197" box="[806,904,404,428]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="10" pageNumber="564" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">12–15 in</quantity>
<taxonomicName id="B4921EC0FFDBB567FC4BFE3BFBBD3197" box="[910,1055,404,428]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FC4BFE3BFBBD3197" box="[910,1055,404,428]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
) and longer disto-lateral spines of telson compare to the inner spines (vs. as long as the inner spines with second pair shorter in
<taxonomicName id="B4921EC0FFDBB567FB6FFE16FA9831EB" box="[1194,1338,440,464]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="mahalona">
<emphasis id="41E6B951FFDBB567FB6FFE16FA9831EB" box="[1194,1338,440,464]" italics="true" pageId="10" pageNumber="564">C. mahalona</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8836C8FFDBB567FF02FE75FD97322C" pageId="10" pageNumber="564" type="distribution">
<paragraph id="732D6543FFDBB567FF02FE75FD97322C" blockId="10.[151,1436,151,535]" pageId="10" pageNumber="564">
<emphasis id="41E6B951FFDBB567FF02FE75FE5331CF" bold="true" box="[199,497,475,500]" pageId="10" pageNumber="564">Distribution and habitat.</emphasis>
<taxonomicName id="B4921EC0FFDBB567FE32FE72FD1B31CF" box="[503,697,476,500]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="564" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">
<emphasis id="41E6B951FFDBB567FE32FE72FD1B31CF" box="[503,697,476,500]" italics="true" pageId="10" pageNumber="564">Caridina laroeha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD5042AFFDBB567FD05FE73FD5E31CF" box="[704,764,477,500]" pageId="10" pageNumber="564" rank="species">n. sp.</taxonomicNameLabel>
is only known from the
<typeStatus id="AC29DBE1FFDBB567FBCBFE73FB9C31CE" box="[1038,1086,477,501]" pageId="10" pageNumber="564">type</typeStatus>
locality—the
<taxonomicName id="B4921EC0FFDBB567FB1BFE72FA9831CF" box="[1246,1338,476,500]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="568" phylum="Arthropoda" rank="species" species="laroeha" status="sp. nov.">Laroeha</taxonomicName>
River in the western part of Central
<taxonomicName id="B4921EC0FFDBB567FE0DFE51FD93322C" box="[456,561,511,535]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="569" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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