470 lines
63 KiB
XML
470 lines
63 KiB
XML
<document ID-CLB-Dataset="34173" ID-DOI="http://dx.doi.org/10.3897/phytokeys.11.2563" ID-PMC="PMC3332035" ID-Pensoft-Pub="1314-2003-11-61" ID-Pensoft-UUID="FFF68F18FFF51E59717BA431FFE7FF8F" ID-PubMed="22577334" ID-Zenodo-Dep="576115" ModsDocID="1314-2003-11-61" checkinTime="1451252022863" checkinUser="pensoft" docAuthor="Kilian, Norbert, Djavadi, Seyyedeh Bahereh & Eskandari, Majid" docDate="2012" docId="FAE8D2A4CF3E0AF21FBBA97631A6F516" docLanguage="en" docName="PhytoKeys 11: 61-77" docOrigin="PhytoKeys 11" docPubDate="2012-04-18" docSource="http://dx.doi.org/10.3897/phytokeys.11.2563" docTitle="Lactuca denaensis N. Kilian & Djavadi 2012, sp. nov." docType="treatment" docVersion="8" id="FFF68F18FFF51E59717BA431FFE7FF8F" lastPageNumber="66" masterDocId="FFF68F18FFF51E59717BA431FFE7FF8F" masterDocTitle="Two new mountainous species of Lactuca (Cichorieae, Asteraceae) from Iran, one presenting a new, possibly myrmecochorous achene variant" masterLastPageNumber="77" masterPageNumber="61" pageNumber="62" updateTime="1720490830043" updateUser="ExternalLinkService">
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<mods:mods id="C3A7ABEAE79125BA731349F9A3AD0D3A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3359A679AA65E0613802451574D03058">
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<mods:title id="C78D9B3AA82EFAA524D212ABA014B4B6">Two new mountainous species of Lactuca (Cichorieae, Asteraceae) from Iran, one presenting a new, possibly myrmecochorous achene variant</mods:title>
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<mods:name id="9A20F85C74CC05C69E0EDE52A4788658" type="personal">
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<mods:role id="5B18FB227231E28BABEE92C968CFCBB4">
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<mods:roleTerm id="08748331A2C467E267A27F1E64B0F199">Author</mods:roleTerm>
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<mods:namePart id="D2622CA3D025313030495CF47EFE7A99">Kilian, Norbert</mods:namePart>
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<mods:affiliation id="B1B58A9470A21F7971151CCA091A6D2D">Botanic Garden and Botanical Museum Berlin-Dahlem, Dahlem Center of Plant Sciences, Free University Berlin, Koenigin-Luise-Str. 6 - 8, 14195 Berlin, Germany</mods:affiliation>
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<mods:nameIdentifier id="57BF0456D9159AC7B749C9835509A21C" type="email">n.kilian@bgbm.org</mods:nameIdentifier>
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<mods:name id="27DE478A94172B69341D08A1241422EA" type="personal">
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<mods:roleTerm id="5177545F45A136F4FF99F4E035CBF6F0">Author</mods:roleTerm>
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<mods:namePart id="B2B31DF6D30A1A6A98A0CD18D475F6E5">Djavadi, Seyyedeh Bahereh</mods:namePart>
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<mods:affiliation id="EFCC0441C46DBB68F0EA5DAC890540D0">Department of Botany, Iranian Research Institute of Plant Protection, P. O. Box 1454, Tehran 19395, Iran</mods:affiliation>
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</mods:name>
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<mods:name id="EFD7B274102A530DA48184C01AED6D7F" type="personal">
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<mods:roleTerm id="F08F794C086CE0309939DE4A5FA61B3A">Author</mods:roleTerm>
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<mods:namePart id="1594BE67206CAC248E4D40E968B0341F">Eskandari, Majid</mods:namePart>
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<mods:affiliation id="9F441784E2ADCF43464D6EA703FBCE47">Department of Botany, Iranian Research Institute of Plant Protection, P. O. Box 1454, Tehran 19395, Iran</mods:affiliation>
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<mods:typeOfResource id="865B38C416BB93A79C04421426896364">text</mods:typeOfResource>
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<mods:title id="85FEB3656AEBA5CB0F54A7CF70DB4182">PhytoKeys</mods:title>
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<mods:part id="78D2AFB3CB67A709E39C734598BF21A3">
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<mods:date id="5C209363154831AD0B226B68BC418853">2012</mods:date>
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<mods:number id="3E28E41532DD669BA9EC79D11239CE06">2012-04-18</mods:number>
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<mods:detail id="C7508A0628B2F64B528BB79F9EE6B3F1" type="volume">
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<mods:number id="BED252D06C91185896BF06032E07F1D6">11</mods:number>
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<mods:start id="4B8535E65D2289824B033C89CA8074E8">61</mods:start>
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<mods:end id="2CD866035CA24438B8AB460DD87075DD">77</mods:end>
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<mods:url id="61D216482D55C701A096C2CD0437B380">http://dx.doi.org/10.3897/phytokeys.11.2563</mods:url>
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<mods:classification id="7B530F5579671B49F596AD0A1EDE4EC0">journal article</mods:classification>
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<mods:identifier id="63280EB7C9F0E7C0EE904A9016DCCC2D" type="DOI">http://dx.doi.org/10.3897/phytokeys.11.2563</mods:identifier>
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<treatment id="FAE8D2A4CF3E0AF21FBBA97631A6F516" ID-GBIF-Taxon="152024217" LSID="urn:lsid:plazi:treatment:FAE8D2A4CF3E0AF21FBBA97631A6F516" httpUri="http://treatment.plazi.org/id/FAE8D2A4CF3E0AF21FBBA97631A6F516" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62">
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<subSubSection id="163EEA932CEDC1D9641B4EF11CCE7C4E" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="D3C7CCAF4A53A18C7852430F1EDDBD46" pageId="1" pageNumber="62">
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<taxonomicName id="6807F9FB5F1F69FB04A0980B97D2057D" ID-CoL="3RT3W" LSID="FAE8D2A4-CF3E-0AF2-1FBB-A97631A6F516" authority="N. Kilian & Djavadi" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lactuca denaensis" order="Asterales" pageId="1" pageNumber="62" phylum="Tracheophyta" rank="species" species="denaensis" status="sp. nov.">Lactuca denaensis N. Kilian & Djavadi</taxonomicName>
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<taxonomicNameLabel id="9AC5AE4DD28508258B697EE08B1AE44A" pageId="1" pageNumber="62">sp. nov.</taxonomicNameLabel>
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<figureCitation id="C1E9EAE9BE54780842723C808248B483" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="1" pageNumber="62">Fig. 1</figureCitation>
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<figureCitation id="09C8AD53E6D6359233949FD31C6ED891" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Achenes of the Lactuca rosularis group and Cicerbita polyclada - A, F-I Lactuca hazaranensis; complete achene (A); achene after loss of easily detachable beak, dorsal view (F); ventral view (G); longitudinal section of achene showing two segments, proximal large segment with embry, distal small segment solid with yellow tissue (H); embryo (I); B Lactuca rosularis C Lactuca denaensis D-E Cicerbita polyclada, complete achene (D) and longitudinal section of distal portion showing apical cavity (E). - A-D and F-H each to the same scale A, F-I from the holotype at IRAN B from Rechinger 55980 B, C from Kotschy 662 (B 100426936) D-E from Strauss 14091 (B 100312952)" httpUri="https://binary.pensoft.net/fig/10041" pageId="1" pageNumber="62">3C; Tuisl 1968: t. 2, fig. 12-13 = Tuisl 1977: t. 201, fig. 9-10 under Lactuca polyclada (achene, aspect and cross section)</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="1409D7A566B2670616F5A938910756E9" pageId="1" pageNumber="62" type="reference_group">
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<paragraph id="DF0C77DF30F51F8B46C078C16FB947F0" pageId="1" pageNumber="62">
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<taxonomicName id="E50B575BAFEB5B9473E4003BA775E9F0" authority="Kilian & Djavadi & Eskandari, 2012" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="1" pageNumber="62" phylum="Tracheophyta" rank="species" species="polyclada" status="sp. nov.">Lactuca polyclada</taxonomicName>
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sensu
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<bibRefCitation id="9055F0B05043BCE76D731863E4658037" author="Boissier, E" pageId="9" pageNumber="70" refId="B2" refString="Boissier, E, 1846. Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." title="Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." year="1846">Boissier (1846</bibRefCitation>
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: 10,
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<bibRefCitation id="27CBB4565C0C30B3DF05DC451E423363" author="Boissier, E" pageId="9" pageNumber="70" refId="B3" refString="Boissier, E, 1875. Flora orientalis 3. Basel & Geneve, Georg." title="Flora orientalis 3. Basel & Geneve, Georg." year="1875">1875</bibRefCitation>
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: 812) p.p. et
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<bibRefCitation id="CB27839FB740DB4662A8E0D802FA2E1B" author="Tuisl, G" journalOrPublisher="Annalen des Naturhistorischen Museums Wien" pageId="9" pageNumber="70" pagination="587 - 638" refId="B30" refString="Tuisl, G, 1968. Der Verwandtschaftskreis der Gattung Lactuca L. im iranischen Hochland. Vorarbeiten zur Flora Iranica Nr. 16. Annalen des Naturhistorischen Museums Wien 72: 587 - 638" title="Der Verwandtschaftskreis der Gattung Lactuca L. im iranischen Hochland. Vorarbeiten zur Flora Iranica Nr. 16." volume="72" year="1968">Tuisl (1968</bibRefCitation>
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: 606,
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<bibRefCitation id="D2ADF0C1C5E8BE3E8330BA58C3C59E9D" author="Tuisl, G" editor="Rechinger, KH" journalOrPublisher="Graz, Akademische Druck- und Verlagsanstalt" pageId="9" pageNumber="70" pagination="185 - 196" refId="B31" refString="Tuisl, G, 1977. Lactuca. In: Rechinger, KH, Ed., , Flora Iranica 122. Graz, Akademische Druck- und Verlagsanstalt: 185 - 196" title="Lactuca." volumeTitle=", Flora Iranica 122." year="1977">1977</bibRefCitation>
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: 190-191) p.p., non sensu lectotypi.
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</paragraph>
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</subSubSection>
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<subSubSection id="EA422200021C8F5253FA43FCAD0A1AF4" pageId="1" pageNumber="62" type="diagnosis">
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<paragraph id="8C0F2D488E5A1F02137A49ECDBB922B5" pageId="1" pageNumber="62">Diagnosis.</paragraph>
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<paragraph id="133360B95EF87FDBE61588B754FB1E95" pageId="1" pageNumber="62">
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Habitually similar to
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<taxonomicName id="04D58D11B8BD68C27668BE40FEB37232" authorityName="Boiss" authorityYear="1875" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="1" pageNumber="62" phylum="Tracheophyta" rank="species" species="rosularis">
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<emphasis id="7B2833ED6BB8D818172D7097BF564B3E" italics="true" pageId="1" pageNumber="62">Lactuca rosularis</emphasis>
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</taxonomicName>
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but clearly distinguished by the rosette leaves being undivided (instead of lyrately to irregularly pinnatifid to pinnatisect), the involucre being 10-12 mm (instead of 6-9(-10) mm) long, the achenes having a 4-5 mm (instead of 2.2-3 mm) long corpus, a 0.4-0.9(-2) mm (instead of (1.1-)3-5.3 mm) long beak and a 5-6 mm (instead of 2.5-3 mm) long pappus.
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</paragraph>
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</subSubSection>
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<subSubSection id="667A9C08EC5EDEA4577BC626FED4309C" pageId="1" pageNumber="62" type="holotype">
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<paragraph id="DFEACD2E95A472B87320424E0A31D164" pageId="1" pageNumber="62">Holotype.</paragraph>
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<paragraph id="88C53C863D66B8D967F68E31254F6CEA" pageId="1" pageNumber="62">
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[Iran, Kogiluyeh & Boyer Ahmad], in glareosis alpis
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<normalizedToken id="1CC352CBFC86C461E10B56B94945C5A6" originalValue="Kuh-Daëna">Kuh-Daena</normalizedToken>
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, fl. lutescens, 14 Jul 1842, Kotschy Pl. Pers. autral. 662 (G-BOIS G00330211, specimen annotated by
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<bibRefCitation id="72A8A14DDA583CD8E2657F4408FD1633" author="Beauverd, G" pageId="9" pageNumber="70" refId="B1" refString="Beauverd, G, 1910. Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99-145. http://biodiversitylibrary.org/page/5521285." title="Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99 - 145. http: // biodiversitylibrary. org / page / 5521285" year="1910">Beauverd in 1910</bibRefCitation>
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, see
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<bibRefCitation id="C579E156467B375C82D7741B80105A75" author="CHG," journalOrPublisher="Bibliotheca Botanica" pageId="9" pageNumber="70" refId="B7" refString="CHG, , 2012. Catalogue des herbiers de Geneve (CHG). Conservatoire & Jardin botaniques de la Ville de Geneve. Published at http://www.ville-ge.ch/musinfo/bd/cjb/chg." title="Catalogue des herbiers de Geneve (CHG). Conservatoire & Jardin botaniques de la Ville de Geneve. Published at http: // www. ville-ge. ch / musinfo / bd / cjb / chg" year="2012">CHG 2012</bibRefCitation>
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); selected isotypes: B 100426936, M 0030847 [p.p., two rosulate plants on the left], G [3 sheets], MO 6264530 [p.p., rosulate plants on the left and right], P 00750254 [p.p., left plant], P 00750251 [p.p., second bottom right rosulate plant], P 00750252 [p.p., first bottom right rosulate plant], P 00750253 [p.p., three rosulate plants at the bottom], WAG 0004075 [p.p., sterile leaf rosette on the left].
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</paragraph>
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</subSubSection>
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<subSubSection id="D0545AA4E2C1B50B71DAC74F1F893D43" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="description">
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<paragraph id="25B3790DABDA83F531F88E8A8A3E28D1" pageId="1" pageNumber="62">Description.</paragraph>
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<paragraph id="3DDD5D04A1751C45A75E7175B36EFF0B" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
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Perennial rosulate herb, with a woody subterranean caudex, rosette shoots often on vertical subterranean axes vested with cataphylls below the sometimes somewhat elongate rosette, acaulescent to usually very shortly caulescent and less than 5 cm tall (
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<figureCitation id="C6B72FB8877082987D9D6F53DDE79576" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="1" pageNumber="62">Fig. 1A</figureCitation>
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), rarely to c. 15 cm. Stem one per leaf rosette, usually not or little projecting above the leaf rosette. Rosette leaves (
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<figureCitation id="BC7A4028B6C5D39B0C1B9D9B678D6B1F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="1" pageNumber="62">Fig. 1A</figureCitation>
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) obovate to spatulate, tapering towards base, 2-7
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<normalizedToken id="3EA2CB19FB5FDF5A23D13D947397BFBE" originalValue="×">x</normalizedToken>
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(1-)1.5-3.2 cm, somewhat glaucous; base semiamplexicaul, margin
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<normalizedToken id="CD68BBE9B83B2363E63322F55613F081" originalValue="±">+/-</normalizedToken>
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densely dentate and denticulate, apex rounded to, more rarely, subacute. Synflorescence corymbosely paniculiform, condensed through conspicuously short axes, of c. 6-20 capitula (
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<figureCitation id="A2F42BEAEA1FC1BD9A0344D7CB7E54FD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="1" pageNumber="62">Fig. 1A, G</figureCitation>
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); peduncles 0.4-0.7 cm long. Capitula with c. 10-14 flowers. Involucre (
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<figureCitation id="BB5025B7B077CF206ACE4482A71A329A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="1" pageNumber="62">Fig. 1B-F</figureCitation>
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) narrowly cylindric at anthesis, 10-12 mm long, not
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<pageBreakToken id="DE8BE2FB7622770F4CA951D9F8AD79E9" pageId="2" pageNumber="63" start="start">elongating</pageBreakToken>
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during maturation; outer phyllaries imbricate, outermost
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<normalizedToken id="82788D38D8EAB4C04B66A56918D37563" originalValue="±">+/-</normalizedToken>
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narrowly ovate, c. 2 mm long, following ones gradually longer, lanceolate, the longest up to c. 1/2 as long as inner ones; inner phyllaries linear-lanceolate, 7-8,
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<normalizedToken id="B98609C2CE8E5BF5227C77005F0972E2" originalValue="±">+/-</normalizedToken>
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in one row, subequal in length, with
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<normalizedToken id="42BE8DCB71DB7F527701D5AC3346056E" originalValue="±">+/-</normalizedToken>
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narrow scarious margin. Receptacle flat to slightly convex, naked. Flowers with corolla yellow, ligule 6.7-7.5 mm, tube 5-5.5 mm long; anther tube without appendages 3.2-3.5 mm, basal appendages c. 0.4-0.6 mm, apical appendages 0.3-0.4 mm long; style arms 2.8-3.2 mm long. Achenes (
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<figureCitation id="B918D8FFE31FCBA47A5978AE40828E8C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lactuca denaensis - A habit, acaulescent form B-F capitula, all to the same scale, before flowering (B-D), with faded flowers (E), after flowering (F); G details of synflorescence, one capitulum at fruiting. - All from Remaudiere, 5252 E (IRAN 10625)" httpUri="https://binary.pensoft.net/fig/10039" pageId="2" pageNumber="63">Fig. 1G, 3C</figureCitation>
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) homomorphic, including beak 4.8-5.7 mm long; corpus 4-5 mm long, up to 1.1-1.3 mm in diam., slender-obovoid, compressed, apex contracted into a stout, easily detachable beak c. 0.4-0.9(-2) mm long; achene body apart from the two lateral ribs with 1 similarly strong median rib on either side, rarely dorsally with 2 equally strong ribs, secondary ribs missing; achene surface
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<normalizedToken id="F7BF3D2C7D9CB30E8B0B0A3C1A9F27F4" originalValue="±">+/-</normalizedToken>
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smooth, brown. Pappus simple, without an outer series of minute hairs, setae thin, white, 5-6 mm long, persistent. - Flowering and fruiting: July to September.
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</paragraph>
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<caption id="DEC98D618E1CA4408D96FD4BD140EF89" httpUri="https://binary.pensoft.net/fig/10039" pageId="2" pageNumber="63" start="Figure 1" startId="F1">
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<paragraph id="80543975A3A14571826440C5A7966606" pageId="2" pageNumber="63">
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<emphasis id="C06EE4C28057C954CF36176987741363" bold="true" pageId="2" pageNumber="63">Figure 1.</emphasis>
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<taxonomicName id="5554C2FFFB712861D0ECD0292F9EC1D0" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="DC4E506F3B3C64B321749804038AE344" italics="true" pageId="2" pageNumber="63">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<emphasis id="902F84C28AFFF59B624DC2C6DEEFDD41" bold="true" pageId="2" pageNumber="63">A</emphasis>
|
||
habit, acaulescent form
|
||
<emphasis id="686C1E100E7C2177AD75DD9BA5F05603" bold="true" pageId="2" pageNumber="63">B-F</emphasis>
|
||
capitula, all to the same scale, before flowering (
|
||
<emphasis id="4353FA84DE867F7B018DA286ACCBCF9D" bold="true" pageId="2" pageNumber="63">B-D</emphasis>
|
||
), with faded flowers (
|
||
<emphasis id="1B250EDE54895A02260B1DA6919CF742" bold="true" pageId="2" pageNumber="63">E</emphasis>
|
||
), after flowering (
|
||
<emphasis id="81B67AA8901BE19AF49267EB31ABC0E5" bold="true" pageId="2" pageNumber="63">F</emphasis>
|
||
);
|
||
<emphasis id="C59375D0E066E59CF8180109FB56B8AA" bold="true" pageId="2" pageNumber="63">G</emphasis>
|
||
details of synflorescence, one capitulum at fruiting. - All from
|
||
<normalizedToken id="7AD2B82AC95C4B80676114214AAEDB5D" originalValue="Remaudière">Remaudiere</normalizedToken>
|
||
, 5252E (IRAN 10625)
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="D928D9DFAB68D9B2F5D67EF63C8CCC06" lastPageId="5" lastPageNumber="66" pageId="2" pageNumber="63" type="notes">
|
||
<paragraph id="57923D756309450D84DEEC5ABBEC5B02" pageId="2" pageNumber="63">Notes.</paragraph>
|
||
<paragraph id="0715E839564BB977E15C329DCD06374E" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
|
||
<bibRefCitation id="8569DC61F3145994F8E9177CFA3F7430" author="Boissier, E" pageId="9" pageNumber="70" refId="B2" refString="Boissier, E, 1846. Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." title="Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." year="1846">Boissier (1846)</bibRefCitation>
|
||
described a new species,
|
||
<taxonomicName id="A8F64B40067F09926C3F7084599F892E" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="5E47D27F5B50C6EC4562A95551578F81" italics="true" pageId="2" pageNumber="63">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
, based on two collections (no. 603 and 662) made by T. Kotschy in the Zagros mountains, in the first half of July 1842. Coming from the village Dozdkurd (
|
||
<bibRefCitation id="6DB7CA3E86D0A494C0525F95BD4DF403" author="Edmondson, JR" journalOrPublisher="Willdenowia" pageId="9" pageNumber="70" pagination="579 - 588" publicationUrl="10.3372/wi.36.36154" refId="B8" refString="Edmondson, JR, Lack, HW, 2006. Karl Georg Theodor Kotschy's itinerary in southern Iran, 1841-42. Willdenowia 36: 579 - 588, 10.3372/wi.36.36154" title="Karl Georg Theodor Kotschy's itinerary in southern Iran, 1841 - 42." url="10.3372/wi.36.36154" volume="36" year="2006">Edmondson and Lack 2006</bibRefCitation>
|
||
), Kotschy collected the material on the upper slopes of the "Kuh Daena"[= Kuh e-Dena, c.
|
||
<geoCoordinate id="EBF9BF1D673B973B57D1F9BB122E332E" degrees="30" direction="north" minutes="56" orientation="latitude" precision="925" value="30.933332">30°56'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1B65A7950E286CAA54BBD54CC6F39C4A" degrees="51" direction="east" minutes="28" orientation="longitude" precision="925" value="51.466667">51°28'E</geoCoordinate>
|
||
; summit c. 4448 m, see
|
||
<bibRefCitation id="3159098AC7B05E7104FC854D7B1F0118" author="Wikipedia," journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="9" pageNumber="70" refId="B34" refString="Wikipedia, , 2012. Dena. http://en.wikipedia.org/wiki/Dena [accessed 20.2.2012]." title="Dena. http: // en. wikipedia. org / wiki / Dena [accessed 20.2.2012]" year="2012">Wikipedia (2012)</bibRefCitation>
|
||
, situated in the present-day province Kogiluyeh & Boyer Ahmad of Iran]. According to
|
||
<normalizedToken id="B2A6F5A93131DAAD46A055735376EC09" originalValue="Kotschy’s">Kotschy's</normalizedToken>
|
||
original labels present on the sheets in the Boissier herbarium in
|
||
<normalizedToken id="AB5D37F19974B59A8FC99014E273285F" originalValue="Genève">Geneve</normalizedToken>
|
||
(G-BOIS), the collection Kotschy 603 was collected on 10 July and Kotschy 662 on 14 July 1842. The entire material of
|
||
<normalizedToken id="2769DF8E0E48DC2192D56F6AC49BDD2F" originalValue="Kotschy’s">Kotschy's</normalizedToken>
|
||
gatherings from S Iran was revised by Boissier and subsequently edited and distributed by R. Hohenacker in his series of exsiccatae "Plantarum Persiae australis siccatarum species 440, collectae a Th. Kotschy, determinatae a Dre E. Boissier, editae a R. F. Hohenacker" (
|
||
<bibRefCitation id="332EDFDC6D23CC1AE0DEC532A7DA4C17" author="Triebel, D" journalOrPublisher="Canadian Journal of Botany" pageId="9" pageNumber="70" publicationUrl="http://indexs.botanischestaatssammlung.de" refId="B29" refString="Triebel, D, Scholz, P, 2001+. IndExs - Index of Exsiccatae. Botanische Staatssammlung Muenchen., http://indexs.botanischestaatssammlung.de" title="IndExs - Index of Exsiccatae. Botanische Staatssammlung Muenchen." url="http://indexs.botanischestaatssammlung.de" year="2001 +">Triebel and Scholz 2001+</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="55EE876D44CDF8D99E14E34D77210534" author="Edmondson, JR" journalOrPublisher="Willdenowia" pageId="9" pageNumber="70" pagination="579 - 588" publicationUrl="10.3372/wi.36.36154" refId="B8" refString="Edmondson, JR, Lack, HW, 2006. Karl Georg Theodor Kotschy's itinerary in southern Iran, 1841-42. Willdenowia 36: 579 - 588, 10.3372/wi.36.36154" title="Karl Georg Theodor Kotschy's itinerary in southern Iran, 1841 - 42." url="10.3372/wi.36.36154" volume="36" year="2006">Edmondson and Lack 2006</bibRefCitation>
|
||
). Duplicates are present today in many herbaria. The material of
|
||
<taxonomicName id="58E2F217D8009E1DC9F32DA1C7606E60" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="473266E72210C5459C1A7A79E641A18D" italics="true" pageId="2" pageNumber="63">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
was distributed as a single item under the united numbers "603. 662." and with a single collecting date cited as "14 Jul 1842" on the printed label. The syntypes in
|
||
<normalizedToken id="F995063F13CA120DB86AF44D067ADE14" originalValue="Boissier’s">Boissier's</normalizedToken>
|
||
herbarium as well as the duplicates distributed by Hohenacker in this series of exsiccatae as no. "603. 622." contain two morphologically distinct elements: (a) small leaf rosettes with usually very short, branched, slender to capillaceous flowering axes; (b) almost leafless, from base on divaricately and intricately branched, conspicuously inflated flowering axes. Apparently it has been taken as evident by Boissier and later workers that both elements represent different forms or stages of development of the same species. This assumption is backed by the leaves, which are fairly similar in colour, size, shape and denticulation of the margin in both elements, as well as by the existence of a plant with exceptionally well developed flowering shoots, approaching those of Kotschy 603, among the material of Kotschy 662 (on G00330211, the holotype sheet of
|
||
<taxonomicName id="8D22C3B29FCFC8090797E7E7B01F9CB4" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="6D8B6A12001D99BAE84B7EE7A43F67F9" italics="true" pageId="2" pageNumber="63">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="5181B8B2FC015B71F2A1F1F64E728E0C" author="Boissier, E" pageId="9" pageNumber="70" refId="B2" refString="Boissier, E, 1846. Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." title="Diagnoses plantarum orientalium novarum, Ser. 1, 7. Leipzig, Herrmann." year="1846">Boissier (1846</bibRefCitation>
|
||
: 10) expressed this hypothesis in the following way: "panicula corymbosa intra folia subsessili ... Post anthesin saepe panicula valde augetur, ramosissima fit semipedalis ramis elongatis intricatis dichotomis spongiose incrassatis
|
||
<pageBreakToken id="4DFB86F998BF02332DF1408F063E86E9" pageId="3" pageNumber="64" start="start">,</pageBreakToken>
|
||
sed haec forma monstrosa est, nam in ea nunquam achenia perfecta observavi." ["with a corymbose panicle subsessile among the leaves ... after anthesis panicle often strongly enlarged, very much branched, becomes half a foot long, with elongate, intricate, dichotomous, softly inflated branches, but this is a monstrous form, because I never have observed a perfect achene in it."].
|
||
<bibRefCitation id="D1C008E298C76132F50BDB491449CBB7" author="Tuisl, G" editor="Rechinger, KH" journalOrPublisher="Graz, Akademische Druck- und Verlagsanstalt" pageId="9" pageNumber="70" pagination="185 - 196" refId="B31" refString="Tuisl, G, 1977. Lactuca. In: Rechinger, KH, Ed., , Flora Iranica 122. Graz, Akademische Druck- und Verlagsanstalt: 185 - 196" title="Lactuca." volumeTitle=", Flora Iranica 122." year="1977">Tuisl (1977</bibRefCitation>
|
||
: 191) considered the two elements as different developmental stages: "caulis florifer abbreviatus, fructifer 10-30 cm longus" ["stems at anthesis very short, in fruit 10-30 cm long"].
|
||
</paragraph>
|
||
<paragraph id="E904A0796F4B2221266F8819307EDDF3" pageId="3" pageNumber="64">
|
||
<bibRefCitation id="4D1BE70B1C7A708A5FC7B1B3851DCA30" author="Beauverd, G" pageId="9" pageNumber="70" refId="B1" refString="Beauverd, G, 1910. Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99-145. http://biodiversitylibrary.org/page/5521285." title="Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99 - 145. http: // biodiversitylibrary. org / page / 5521285" year="1910">Beauverd (1910</bibRefCitation>
|
||
: 131-132), in contrast, came to the conclusion that the syntypes of
|
||
<taxonomicName id="EBDAC18C7EEC8105C3AF0DC8E5B1A695" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polycalda">
|
||
<emphasis id="07FD3189FB7B123EA3CB293F48AC9CC0" italics="true" pageId="3" pageNumber="64">Lactuca polycalda</emphasis>
|
||
</taxonomicName>
|
||
in the herbarium of Boissier (G-BOIS) represent two entirely different species and correctly distinguished them.
|
||
<bibRefCitation id="A3C3E9F917238CAD6E2447181DB230F8" author="Beauverd, G" pageId="9" pageNumber="70" refId="B1" refString="Beauverd, G, 1910. Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99-145. http://biodiversitylibrary.org/page/5521285." title="Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99 - 145. http: // biodiversitylibrary. org / page / 5521285" year="1910">Beauverd (1910</bibRefCitation>
|
||
: 131) consequently restricted the name
|
||
<taxonomicName id="C937DD6FBFC862E41D53662C178AA9E4" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="0E8485A6E3AA15CE36554CD51F0E9C21" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
to "K[otsch]y 603 solum! excl. No. 662 et descr. achen.," and formed the new combination
|
||
<taxonomicName id="B93AB17DB5242F265A86C57E3E139A59" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="64" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="4B308297B00D6A9228DC294C67CEE2F1" italics="true" pageId="3" pageNumber="64">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
(Boiss.) Beauverd for the taxon with divaricately branched inflated stems, smaller involucres, bluish flowers and a pappus with an outer row of minute hairs. Mature fruits of
|
||
<taxonomicName id="2C9EB96A054F8CC702274C8CE3CB9AC3" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="64" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="A25F87E58E75F0C4EA968DAD08C01061" italics="true" pageId="3" pageNumber="64">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
, missing in Kotschy 603, are known through a collection of T. Strauss (at B, see Appendix). They are illustrated here for the first time and show further differences between the two species, in particular with respect to the prominence of the ribs and shape and structure of the body apex (compare
|
||
<figureCitation id="D45273E2EFA2DA2F506726D8DDF985FC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lactuca hazaranensis - A habit, left plant of holotype sheet B-G capitula, all to the same scale, before flowering (B-C), at fruiting (D-E), after fruiting (F-G); H plant in natural habitat A-G from the holotype H plant in the natural habitat at the type locality; photograph by M. Eskandari, 10 Aug 2010." httpUri="https://binary.pensoft.net/fig/10040" pageId="3" pageNumber="64">Fig. 2C and D-E</figureCitation>
|
||
). For a summary of the differences see
|
||
<tableCitation id="C526D4AF378D344BBEFB333BB574983A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological differences between Lactuca hazaranensis and its relatives Lactuca rosularis and Lactuca denaensis, as well as between the latter and Cicerbita polyclada. Based on the material studied. - The investigated material is listed in the Appendix." httpUri="http://table.plazi.org/id/23D356992B6FF5B40DBAC23209CC6C17" pageId="3" pageNumber="64" tableUuid="23D356992B6FF5B40DBAC23209CC6C17">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="F481252106C49859F78BC8C02F925DB3" httpUri="https://binary.pensoft.net/fig/10040" pageId="3" pageNumber="64" start="Figure 2" startId="F2">
|
||
<paragraph id="FFE7A762DB37ADDA39EA7ADE7543BC01" pageId="3" pageNumber="64">
|
||
<emphasis id="F3FD13A7205C2445ED5F893CEAEF7478" bold="true" pageId="3" pageNumber="64">Figure 2.</emphasis>
|
||
<taxonomicName id="DC5AD77C3786C7F98356398DE61723DF" authorityName="Djavadi & N. Kilian" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="hazaranensis">
|
||
<emphasis id="E253F1E0DC8CE07A26D4FE77BAD20441" italics="true" pageId="3" pageNumber="64">Lactuca hazaranensis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<emphasis id="1A384BE67DF05EBAF9339BB4F058376A" bold="true" pageId="3" pageNumber="64">A</emphasis>
|
||
habit, left plant of holotype sheet
|
||
<emphasis id="3310A08966651767DB251DB855311C6C" bold="true" pageId="3" pageNumber="64">B-G</emphasis>
|
||
capitula, all to the same scale, before flowering (
|
||
<emphasis id="0AD349579AC5DD0BC69D3F80C6EB7A94" bold="true" pageId="3" pageNumber="64">B-C</emphasis>
|
||
), at fruiting (
|
||
<emphasis id="1F7145DDF6F22522D11A4C83CC1748DE" bold="true" pageId="3" pageNumber="64">D-E</emphasis>
|
||
), after fruiting (
|
||
<emphasis id="9ED9EC8130B05F9043179552AA0CDC7D" bold="true" pageId="3" pageNumber="64">F-G</emphasis>
|
||
);
|
||
<emphasis id="AB79509F92A6F36BAF7A59ABA46783E5" bold="true" pageId="3" pageNumber="64">H</emphasis>
|
||
plant in natural habitat
|
||
<emphasis id="792C0B93F2CE38A97E7261B6BBE7C5D2" bold="true" pageId="3" pageNumber="64">A-G</emphasis>
|
||
from the holotype
|
||
<emphasis id="78CD680612CD080B133EB05F4151D920" bold="true" pageId="3" pageNumber="64">H</emphasis>
|
||
plant in the natural habitat at the type locality; photograph by M. Eskandari, 10 Aug 2010.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9CE9CDFF26F88BE28C4A4BEAC336CFD2" pageId="3" pageNumber="64">
|
||
According to Art. 7.11 of the Vienna Code,
|
||
<normalizedToken id="648ED365FB76B23806B9023576B87F01" originalValue="Beauverd’s">Beauverd's</normalizedToken>
|
||
restriction of the name to Kotschy 603 does not, however, constitute a lectotypification, because he did not use "the term
|
||
<normalizedToken id="852C2BAEB791107A71554C00499F4E1F" originalValue="‘type’">'type'</normalizedToken>
|
||
(typus) or an equivalent" (
|
||
<bibRefCitation id="DE2BD5785706F589BD94D53C17C23CA9" author="McNeill, J" journalOrPublisher="Pollen and Spores" pageId="9" pageNumber="70" refId="B22" refString="McNeill, J, Barrie, FR, Burdet, HM, Demoulin, V, Hawksworth, DL, Marhold, K, Nicolson, DH, Prado, J, Silva, PC, Skog, JE, Wiersema, JH, Turland NJ (Eds &, Compilers), 2006. International Code of Botanical Nomenclature (Vienna Code), adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Regnum Vegetabile 146. http://www.ibot.sav.sk/icbn/main.htm." title="International Code of Botanical Nomenclature (Vienna Code), adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Regnum Vegetabile 146. http: // www. ibot. sav. sk / icbn / main. htm" year="2006">McNeill et al. 2006</bibRefCitation>
|
||
). Later authors apparently neglected
|
||
<normalizedToken id="CD30B5418782C412E64A2FFFFC134095" originalValue="Beauverd’s">Beauverd's</normalizedToken>
|
||
rectification and we are also not aware of any lectotypification of
|
||
<normalizedToken id="C39B65B7D187047696E3EE9F9D479A5D" originalValue="Boissier’s">Boissier's</normalizedToken>
|
||
name. In accordance with Art. 9.12 (
|
||
<bibRefCitation id="8A8C949CA9B79CAC6D4D897085F8D00C" author="McNeill, J" journalOrPublisher="Pollen and Spores" pageId="9" pageNumber="70" refId="B22" refString="McNeill, J, Barrie, FR, Burdet, HM, Demoulin, V, Hawksworth, DL, Marhold, K, Nicolson, DH, Prado, J, Silva, PC, Skog, JE, Wiersema, JH, Turland NJ (Eds &, Compilers), 2006. International Code of Botanical Nomenclature (Vienna Code), adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Regnum Vegetabile 146. http://www.ibot.sav.sk/icbn/main.htm." title="International Code of Botanical Nomenclature (Vienna Code), adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Regnum Vegetabile 146. http: // www. ibot. sav. sk / icbn / main. htm" year="2006">McNeill et al. 2006</bibRefCitation>
|
||
) and in this way maintaining the only unequivocal use of the name as established by
|
||
<bibRefCitation id="966E95DCF2E4D6B30B2854B85BF3863F" author="Beauverd, G" pageId="9" pageNumber="70" refId="B1" refString="Beauverd, G, 1910. Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99-145. http://biodiversitylibrary.org/page/5521285." title="Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99 - 145. http: // biodiversitylibrary. org / page / 5521285" year="1910">Beauverd (1910</bibRefCitation>
|
||
: 131) for the species placed by him in
|
||
<taxonomicName id="6A0CAD2BC3AEF9720A2D6300755FBCF7" class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="64" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="1B0C617509D385F112C4D09792198625" italics="true" pageId="3" pageNumber="64">Cicerbita</emphasis>
|
||
</taxonomicName>
|
||
, we formally lectotypify
|
||
<taxonomicName id="7C1FB12BBF4B3600682C724E9A21008B" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="A1FE144AEC893775955C36ECB4F998C1" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
Boiss. following his restriction:
|
||
<taxonomicName id="23EBC0608D49AF68B09744AC32A99D3C" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="64" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="1FBB10CF2B67D613F83AE0AFA2D5CE76" italics="true" pageId="3" pageNumber="64">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
(Boiss.) Beauverd in Bull. Soc. Bot.
|
||
<normalizedToken id="E1171332E3C983AFA4FFE9B3D8322A01" originalValue="Genève">Geneve</normalizedToken>
|
||
2: 131. 1910 ≡
|
||
<taxonomicName id="7BED5A35BB14FA5A951F3E32E6E21740" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="8C9323905CC7DBC7114FA6318F16AFD6" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
Boiss., Diagn. Pl. Orient., ser. 1, 7: 10. 1846. - Lectotype (designated here): [Iran, Kogiluyeh & Boyer Ahmad], in glareosis ad latera septentr. m.
|
||
<normalizedToken id="9BF25EF0EB4C25D3872472B0991F15E0" originalValue="Kuh-Daëna">Kuh-Daena</normalizedToken>
|
||
, flos caeruleus, 10 Jul 1842, Kotschy Pl. Pers. autral. 603 (G-BOIS G00330212 [2 sheets annotated by
|
||
<bibRefCitation id="85F3BA99B7F6948D6FEE142AB686FF8D" author="Beauverd, G" pageId="9" pageNumber="70" refId="B1" refString="Beauverd, G, 1910. Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99-145. http://biodiversitylibrary.org/page/5521285." title="Contribution a l'etude des Composees, 3. Le genre Cicerbita. - Bulletin de la Societe Botanique de Geneve, ser. 2, 2: 99 - 145. http: // biodiversitylibrary. org / page / 5521285" year="1910">Beauverd in 1910</bibRefCitation>
|
||
], see
|
||
<bibRefCitation id="0F6FCD579C46E75B38CEFCB675B73AD0" author="CHG," journalOrPublisher="Bibliotheca Botanica" pageId="9" pageNumber="70" refId="B7" refString="CHG, , 2012. Catalogue des herbiers de Geneve (CHG). Conservatoire & Jardin botaniques de la Ville de Geneve. Published at http://www.ville-ge.ch/musinfo/bd/cjb/chg." title="Catalogue des herbiers de Geneve (CHG). Conservatoire & Jardin botaniques de la Ville de Geneve. Published at http: // www. ville-ge. ch / musinfo / bd / cjb / chg" year="2012">CHG 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E69FC6C585F5D51D5AAC6C021C68B592" lastPageId="5" lastPageNumber="66" pageId="3" pageNumber="64">
|
||
The second species admixed by Boissier under the name
|
||
<taxonomicName id="12E7DAC88FE3086C3D80890EFB6D6A6D" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="4F99313E281D6D40B6E879EDB7FA7DC1" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
with a usually very short or almost missing flowering stem, yellow flowers, a pappus without an outer row of minute hairs and the achene anatomy as illustrated by
|
||
<bibRefCitation id="B90C1FF5651C71035E8A3D056FB43FB1" author="Tuisl, G" journalOrPublisher="Annalen des Naturhistorischen Museums Wien" pageId="9" pageNumber="70" pagination="587 - 638" refId="B30" refString="Tuisl, G, 1968. Der Verwandtschaftskreis der Gattung Lactuca L. im iranischen Hochland. Vorarbeiten zur Flora Iranica Nr. 16. Annalen des Naturhistorischen Museums Wien 72: 587 - 638" title="Der Verwandtschaftskreis der Gattung Lactuca L. im iranischen Hochland. Vorarbeiten zur Flora Iranica Nr. 16." volume="72" year="1968">Tuisl (1968</bibRefCitation>
|
||
: t. 2, fig. 12-13 = 1977: t. 201, fig. 9-10) was left by Beauverd without a legitimate name. We have named and described it therefore here as
|
||
<taxonomicName id="D304A50E33C867E85C1360ADC4D1B5D2" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="01E35CED3C11F10C2124CFC2AB9A5F94" italics="true" pageId="3" pageNumber="64">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
, the name being typified with the syntype "Kotschy 662" of
|
||
<taxonomicName id="70F216CC898B10F8C94F7299903D8F3A" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="4CB3E0844AADBB5F04C18BD6FE0991D3" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
Boiss. in
|
||
<normalizedToken id="597894B13350928A5489251210401730" originalValue="Boissier’s">Boissier's</normalizedToken>
|
||
herbarium. Since
|
||
<normalizedToken id="18AF8CD5AB816BAC5D645E7A38F78458" originalValue="Kotschy’s">Kotschy's</normalizedToken>
|
||
both collections no. 603 and 662 were combined by Hohenacker in his series of exsiccatae into a single unit of which each set usually contains both elements, isotypes are present in numerous herbaria. The corresponding exsiccatae sheets with the admixed material of
|
||
<taxonomicName id="DFA9CE846CC1C2D35D62716FD6539ED4" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="3" pageNumber="64" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="CCC2FC08631E3D9732DE62604F8D82D3" italics="true" pageId="3" pageNumber="64">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E701F52942DE0ACE35816E3511C417DD" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="920E30F3E2F98293EA88BCDFD342A6AC" italics="true" pageId="3" pageNumber="64">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
carry the printed label "Th. Kotschy, Pl. Pers. austr. Ed. R. F. Hohenacker. 1845 // 603. 662.
|
||
<taxonomicName id="C90AD91941C29A6C6F6096272FD129FC" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="3A73BA549FA3FC89F5CF5240D5F6A293" italics="true" pageId="3" pageNumber="64">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
<pageBreakToken id="F2FC57028B8FFC8A067C32CFFB82C993" pageId="4" pageNumber="65" start="start">/</pageBreakToken>
|
||
Boiss. n. sp. // In glareosis alpis
|
||
<normalizedToken id="0B034B60FCED9EEAA1225B915A4FF941" originalValue="Kuh-Daëna">Kuh-Daena</normalizedToken>
|
||
. D. 14. Jul. 1842. / Pl. lactescens." They are usually filed under
|
||
<taxonomicName id="AC35649E70294BD2F20CFC26B6FF8462" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A7B98216152CC02C8E75AA095EF309E2" italics="true" pageId="4" pageNumber="65">Lactuca</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4A81E0302D5C03F725F01A037EE09146" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="4" pageNumber="65" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="FA2278700951E8A1E4F227EB10E5B38E" italics="true" pageId="4" pageNumber="65">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
, respectively, or, sometimes erroneously under
|
||
<taxonomicName id="DB7A218A2FC402E0ED1639207DF14910" authorityName="Kirpicz" authorityYear="1964" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cephalorrhynchus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="4" pageNumber="65" phylum="Angiospermae" rank="species" species="polycladus">
|
||
<emphasis id="FA2FF46D4B9AE4EB7986AC5DD10DF318" italics="true" pageId="4" pageNumber="65">Cephalorrhynchus polycladus</emphasis>
|
||
</taxonomicName>
|
||
(Boiss.) Kirp. The latter name is not a further homotypic synonym of
|
||
<taxonomicName id="194E66570E79979A5FAC50F5C23C5626" authorityName="Kilian & Djavadi & Eskandari" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="33DBEADA1284F37B22E9501FD0528118" italics="true" pageId="4" pageNumber="65">Lactuca polyclada</emphasis>
|
||
</taxonomicName>
|
||
, but actually based on
|
||
<taxonomicName id="C52281DC818F44F81632E9712ACD8AA9" class="Magnoliopsida" family="Asteraceae" genus="Zollikoferia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="polyclada">
|
||
<emphasis id="9EFBABDCBDA2C82CD222073A474F3F9A" italics="true" pageId="4" pageNumber="65">Zollikoferia polyclada</emphasis>
|
||
</taxonomicName>
|
||
<pageBreakToken id="EC7B08E63EC510B74497469B0FEAE4AD" pageId="5" pageNumber="66" start="start">Boiss</pageBreakToken>
|
||
., which represents a different mountainous species distributed from E Iran and Afghanistan to Central Asia, also with intricately and divaricately branched stems. Habitually, the latter species can readily be distinguished by its indurate and never inflated stems and branches; also it is not present in the Zagros mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A17B46762226B713F8C681A89561C2AB" pageId="5" pageNumber="66" type="distribution">
|
||
<paragraph id="F02F36F15D4C327196E0751417549F80" pageId="5" pageNumber="66">Distribution and habitat.</paragraph>
|
||
<paragraph id="15A77DC1A5314890C3BA3C760CF1FF73" pageId="5" pageNumber="66">
|
||
As far as we know,
|
||
<taxonomicName id="C331847C81D1B4C51A6F3812FA2A28E2" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="5019E1E4A4903E65715CECDA896A851A" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the higher elevations, probably above 3000 m, of Kuh e-Dena in the Zagros mountains of SW Iran (
|
||
<figureCitation id="E24426260E7858BCC78E9273D676D19D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Lactuca hazaranensis (circles) and the related species Lactuca rosularis (squares) and Lactuca denaensis (rhomb, actucal position as indicated by arrow), as well as of Cicerbita polyclada (triangles). - Georeferenced map based on the known collections (see Appendix and supplemented by collection cited in Kirpicznikov 1964 and Tuisl 1977) and generated with DIVA-GIS (Hijmans 2011) using an adaptation of the SRTM 90 m digital elevation data (CGIAR-CSI 2004)." httpUri="https://binary.pensoft.net/fig/10042" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
). From the rare material with subterranean parts preserved (e.g. on the sheets MO 6264530 and P 00750252 with isotypes), showing a subterranean caudex producing several cm long shoots vested with cataphylls below the leaf rosettes, it can be concluded that the species is a scree plant.
|
||
</paragraph>
|
||
<paragraph id="9651DF1C345FD3AB416D359896BF21E0" pageId="5" pageNumber="66">
|
||
Additional specimen seen: Iran. Kogiluyeh & Boyer Ahmad: Dena [c.
|
||
<geoCoordinate id="C7D8CF037E2DA2F92A31DE6572EEA960" degrees="30" direction="north" minutes="56" orientation="latitude" precision="925" value="30.933332">30°56'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="81E36190BF06B7BBE256554E5DDF4D46" degrees="51" direction="east" minutes="28" orientation="longitude" precision="925" value="51.466667">51°28'E</geoCoordinate>
|
||
], 3300 m, Sep 1955,
|
||
<normalizedToken id="344538305C51D6FAD696C5A4B65446B3" originalValue="Remaudière">Remaudiere</normalizedToken>
|
||
5252E (IRAN 10625).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1A645C138F157CA9D8C8758624A51CE6" pageId="5" pageNumber="66" type="etymology">
|
||
<paragraph id="660F2BE7471C64A5B449B4A17C315337" pageId="5" pageNumber="66">Etymology.</paragraph>
|
||
<paragraph id="1B6898CEC95DED302FC1EE2FAA355D92" pageId="5" pageNumber="66">
|
||
<taxonomicName id="DD7DFE653EC0502B0AB2411D342ED45A" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="BF6A9A188FA173819E6B4C9F850D73C6" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
is named after the Kuh e-Dena massif of the Zagros mountains, where the only two collections known to us come from.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABB4099EF88611753D895D4E5D8274C6" pageId="5" pageNumber="66" type="morphological affinities and delimitation">
|
||
<paragraph id="73658B000E804346F438146FDE99414B" pageId="5" pageNumber="66">Morphological affinities and delimitation.</paragraph>
|
||
<paragraph id="825DE0A48217C407D9EDA3B38446475E" pageId="5" pageNumber="66">
|
||
Morphological comparison shows that
|
||
<taxonomicName id="C693C9BC41A0DE3F8537CB40814E53A6" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="FD11FCED0CCF770472A3B5212598CAE2" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="C6B5C4E60CF7BB631915A6F88055F5F8" authorityName="Boiss" authorityYear="1875" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="rosularis">
|
||
<emphasis id="D726346EB6B1C6AFBAAE27523F1435F0" italics="true" pageId="5" pageNumber="66">Lactuca rosularis</emphasis>
|
||
</taxonomicName>
|
||
. Both share the rosulate habit with a woody caudex, glaucous leaves and small yellow-flowered capitula but also the compressed, beaked achenes (compare
|
||
<figureCitation id="0492E13B19776514B87409777E4F2B9F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Achenes of the Lactuca rosularis group and Cicerbita polyclada - A, F-I Lactuca hazaranensis; complete achene (A); achene after loss of easily detachable beak, dorsal view (F); ventral view (G); longitudinal section of achene showing two segments, proximal large segment with embry, distal small segment solid with yellow tissue (H); embryo (I); B Lactuca rosularis C Lactuca denaensis D-E Cicerbita polyclada, complete achene (D) and longitudinal section of distal portion showing apical cavity (E). - A-D and F-H each to the same scale A, F-I from the holotype at IRAN B from Rechinger 55980 B, C from Kotschy 662 (B 100426936) D-E from Strauss 14091 (B 100312952)" httpUri="https://binary.pensoft.net/fig/10041" pageId="5" pageNumber="66">Fig. 3B and C</figureCitation>
|
||
), characterised by 4 (only exceptionally 5) similar and strongly prominent main ribs (two lateral and one median on either side, exceptionally dorsally 2) and a simple pappus without an outer series of minute hairs (see also
|
||
<tableCitation id="3240C6C86A3154FD90BC27F9CDBE38D9" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological differences between Lactuca hazaranensis and its relatives Lactuca rosularis and Lactuca denaensis, as well as between the latter and Cicerbita polyclada. Based on the material studied. - The investigated material is listed in the Appendix." httpUri="http://table.plazi.org/id/23D356992B6FF5B40DBAC23209CC6C17" pageId="5" pageNumber="66" tableUuid="23D356992B6FF5B40DBAC23209CC6C17">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="4F48DB1F147EB13DF626567F96DD1878" httpUri="https://binary.pensoft.net/fig/10041" pageId="5" pageNumber="66" start="Figure 3" startId="F3">
|
||
<paragraph id="5B1ED0B1DEEEFB1F87AFEF8302593A7D" pageId="5" pageNumber="66">
|
||
<emphasis id="3C0271C1EF21D53BB4371B0830737FA6" bold="true" pageId="5" pageNumber="66">Figure 3.</emphasis>
|
||
Achenes of the
|
||
<taxonomicName id="E187638758CD7C97DBC6B545BB4324FF" authorityName="Boiss" authorityYear="1875" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="rosularis">
|
||
<emphasis id="B870FF60A71FE6A85C16D2618EA6DCBA" italics="true" pageId="5" pageNumber="66">Lactuca rosularis</emphasis>
|
||
</taxonomicName>
|
||
group and
|
||
<taxonomicName id="8AA1239D716E59338A04F02C3801217F" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="5" pageNumber="66" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="CC692A903F8AB2249DDE9FF457BC56E1" italics="true" pageId="5" pageNumber="66">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<emphasis id="37F0374B27AFDC4FDDE84673D4E60FBE" bold="true" pageId="5" pageNumber="66">A, F-I</emphasis>
|
||
<taxonomicName id="4358A507D95BEB31FC0984CAC0EAB385" authorityName="Djavadi & N. Kilian" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="hazaranensis">
|
||
<emphasis id="0D9631205DFC39C267D0B35E8278E893" italics="true" pageId="5" pageNumber="66">Lactuca hazaranensis</emphasis>
|
||
</taxonomicName>
|
||
; complete achene (
|
||
<emphasis id="9AA43963753CDE6A16D1BA1570F4F598" bold="true" pageId="5" pageNumber="66">A</emphasis>
|
||
); achene after loss of easily detachable beak, dorsal view (
|
||
<emphasis id="CC916580EBCBDD397C8D21E42EC94827" bold="true" pageId="5" pageNumber="66">F</emphasis>
|
||
); ventral view (
|
||
<emphasis id="B7AC23642CC6E34B524E4705990E605C" bold="true" pageId="5" pageNumber="66">G</emphasis>
|
||
); longitudinal section of achene showing two segments, proximal large segment with embry, distal small segment solid with yellow tissue (
|
||
<emphasis id="16B72B64BC54EA9B99139D63D35FFB46" bold="true" pageId="5" pageNumber="66">H</emphasis>
|
||
); embryo (
|
||
<emphasis id="BDE30EE581A2A7E12E40D3352662E7C5" bold="true" pageId="5" pageNumber="66">I</emphasis>
|
||
);
|
||
<emphasis id="D821EF2DE34189F57427C5D0396B94C2" bold="true" pageId="5" pageNumber="66">B</emphasis>
|
||
<taxonomicName id="2FF99F2AC5BF3554A89D793E40F3DACE" authorityName="Boiss" authorityYear="1875" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="rosularis">
|
||
<emphasis id="62869C430D9210388DB65CC604281F7C" italics="true" pageId="5" pageNumber="66">Lactuca rosularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="138F11FF39F83CFB29713AAEDAA7ADB8" bold="true" pageId="5" pageNumber="66">C</emphasis>
|
||
<taxonomicName id="317334F941689747BFCA9A6EA19CC612" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="6C562FC8D606AC7EADB715618B1774A3" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9D68102528DB1A1C90A5C1AB33F18C4" bold="true" pageId="5" pageNumber="66">D-E</emphasis>
|
||
<taxonomicName id="95C4CE1295787D4B8729D17876AE13D8" authorityName="Beauverd" authorityYear="1910" baseAuthorityName="Boiss." class="Dicotyledoneae" family="Asteraceae" genus="Cicerbita" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="5" pageNumber="66" phylum="Angiospermae" rank="species" species="polyclada">
|
||
<emphasis id="6EE0E861F0D6CD6346D70CB5C87B4E79" italics="true" pageId="5" pageNumber="66">Cicerbita polyclada</emphasis>
|
||
</taxonomicName>
|
||
, complete achene (
|
||
<emphasis id="DF82EA5B59709D4239E0C5F935945B31" bold="true" pageId="5" pageNumber="66">D</emphasis>
|
||
) and longitudinal section of distal portion showing apical cavity (
|
||
<emphasis id="5BBC9C48451966B8F415D8282B807510" bold="true" pageId="5" pageNumber="66">E</emphasis>
|
||
). -
|
||
<emphasis id="7ECA4258E41689DD5B0678049D20EEE5" bold="true" pageId="5" pageNumber="66">A-D</emphasis>
|
||
and
|
||
<emphasis id="07D15E25F6A07D730C6B115E28036D2C" bold="true" pageId="5" pageNumber="66">F-H</emphasis>
|
||
each to the same scale
|
||
<emphasis id="37B736207776D85F823FE976F1302BDA" bold="true" pageId="5" pageNumber="66">A, F-I</emphasis>
|
||
from the holotype at IRAN
|
||
<emphasis id="26403F9713EA11532061C530FB7BAD0B" bold="true" pageId="5" pageNumber="66">B</emphasis>
|
||
from Rechinger 55980
|
||
<emphasis id="B6ACDA97A1D572265289B03BEB7D6322" bold="true" pageId="5" pageNumber="66">B, C</emphasis>
|
||
from Kotschy 662 (B 100426936)
|
||
<emphasis id="A9A0D07D157465BE217A58669944D0D8" bold="true" pageId="5" pageNumber="66">D-E</emphasis>
|
||
from Strauss 14091 (B 100312952)
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8E3B952DF76B8898EE6AB3F3F225AEF8" pageId="5" pageNumber="66" type="preliminary conservation status">
|
||
<paragraph id="E094824E61750D50140A14BBD75C24BF" pageId="5" pageNumber="66">Preliminary conservation status.</paragraph>
|
||
<paragraph id="78052306167817CB04B05D52D841051D" pageId="5" pageNumber="66">
|
||
<taxonomicName id="BA8FFADB01E8A503CE74976C3B65EE58" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="840A2E026118AAC61D77C899E1535044" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from the higher elevations of the Kuh e-Dena massif, which is part of a Protected Area, possibly also including the populations of the species. The species seems to be rare, since only two collections are known to us, but it has to be taken into account that it is rather inconspicuous.
|
||
<taxonomicName id="FD251E86C3ADD602D76507FF7F065002" authorityName="N. Kilian & Djavadi" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Lactuca" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="66" phylum="Tracheophyta" rank="species" species="denaensis">
|
||
<emphasis id="06F417209CDE144BE83D03408099B520" italics="true" pageId="5" pageNumber="66">Lactuca denaensis</emphasis>
|
||
</taxonomicName>
|
||
must currently be assessed as Data Deficient (
|
||
<bibRefCitation id="908D2111F122160D56BAF2665B328EBE" author="IUCN Species Survival Commision," journalOrPublisher="Planta" pageId="9" pageNumber="70" refId="B14" refString="IUCN Species Survival Commision, , 2001. IUCN Red List Categories: Version 3.1. IUCN, Gland and Cambridge." title="IUCN Red List Categories: Version 3.1. IUCN, Gland and Cambridge." year="2001">IUCN 2001</bibRefCitation>
|
||
) and an assessment of its populations in the field is strongly desirable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |