680 lines
56 KiB
XML
680 lines
56 KiB
XML
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<mods:titleInfo>
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<mods:title>A new semi-slug of the genus Microparmarion from Brunei, discovered, described and DNA-barcoded on citizen-science ' taxon expeditions' (Gastropoda, Stylommatophora, Ariophantidae)</mods:title>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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<mods:affiliation>Taxon Expeditions B. V., Leiden, Netherlands & Naturalis Biodiversity Center, Leiden, Netherlands & Institute Biology Leiden, Leiden University, Leiden, Netherlands & Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Berenyi, Simon</mods:namePart>
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<mods:affiliation>Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Taxon Expeditions, Leiden, Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ezzwan, Nurilya Syamimi Muhammad Nazirul</mods:namePart>
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<mods:affiliation>Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hamdani, Nur Izzah Amal Afiqah</mods:namePart>
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<mods:affiliation>Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wu, Harrison</mods:namePart>
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<mods:affiliation>University of British Columbia, Kelowna, Canada & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Antoni, Luca</mods:namePart>
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<mods:affiliation>Department of Biotechnology, University of Verona, Verona, Italy & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vincenzi, Leonardo</mods:namePart>
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<mods:affiliation>Department of Biotechnology, University of Verona, Verona, Italy & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>de Gier, Werner</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands & Taxon Expeditions B. V., Leiden, Netherlands & Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen, Netherlands</mods:affiliation>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>van Peursen, Anthonie D. P.</mods:namePart>
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<mods:affiliation>Taxon Expeditions B. V., Leiden, Netherlands & Naturalis Biodiversity Center, Leiden, Netherlands & Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei</mods:affiliation>
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</mods:name>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Njunjic, Iva</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5723-4068</mods:nameIdentifier>
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<mods:affiliation>Taxon Expeditions, Leiden, Netherlands & Naturalis Biodiversity Center, Leiden, Netherlands & Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei</mods:affiliation>
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<mods:nameIdentifier type="email">info@taxonexpeditions.com</mods:nameIdentifier>
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<mods:namePart>Delledonne, Massimo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7100-4581</mods:nameIdentifier>
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<mods:affiliation>Department of Biotechnology, University of Verona, Verona, Italy & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Slik, Ferry</mods:namePart>
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<mods:affiliation>Taxon Expeditions B. V., Leiden, Netherlands & Environmental and Life Sciences, Faculty of Science, Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Grafe, Ulmar</mods:namePart>
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<mods:affiliation>Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cicuzza, Daniele</mods:namePart>
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<mods:affiliation>Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei & Taxon Expeditions B. V., Leiden, Netherlands</mods:affiliation>
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<mods:title>Biodiversity Data Journal</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-10</mods:number>
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<mods:detail type="volume">
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|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:0904AEF2FF073B4F77AA17291F25BECB" country="Brunei" county="Occurrence" latitude="4.555" longitude="115.153" municipality="Menno Schilthuizen" name="Ulu-Ulu Resort Helipad" pageId="0" pageNumber="101579" stateProvince="Temburong District">Ulu-Ulu Resort Helipad</location>
|
||
; verbatimElevation:
|
||
<elevation pageId="0" pageNumber="101579">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
</elevation>
|
||
; decimalLatitude:
|
||
<geoCoordinate orientation="latitude" pageId="0" pageNumber="101579" value="4.555">4.555</geoCoordinate>
|
||
; decimalLongitude:
|
||
<geoCoordinate orientation="longitude" pageId="0" pageNumber="101579" value="115.153">115.153</geoCoordinate>
|
||
; georeferenceProtocol: GPS;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:7A68F978D1B72F8598499ACE35771811" country="Brunei" county="Occurrence" latitude="4.555" longitude="115.153" municipality="Menno Schilthuizen" name="Identification" stateProvince="Temburong District">Identification</location>
|
||
:
|
||
</emphasis>
|
||
identifiedBy:
|
||
<determinerName pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:311A21E213E3E7475D5A0227397BA694" country="Brunei" county="Occurrence" latitude="4.555" longitude="115.153" municipality="Menno Schilthuizen" name="Menno Schilthuizen" stateProvince="Temburong District">Menno Schilthuizen</location>
|
||
</determinerName>
|
||
; dateIdentified: 2022;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:866458108BEB0A91320637941BA5C7A4" country="Brunei" county="Occurrence" latitude="4.555" longitude="115.153" municipality="Menno Schilthuizen" name="Event" stateProvince="Temburong District">Event</location>
|
||
:
|
||
</emphasis>
|
||
eventDate:
|
||
<collectingDate pageId="0" pageNumber="101579" value="2022-10-14">14/10/2022</collectingDate>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Record Level:</emphasis>
|
||
language: en; basisOfRecord: PreservedSpecimen
|
||
<materialsCitation collectingDate="2022-10-16" collectorName="Iva Njunjic" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Iva Njunjic" pageId="0" pageNumber="101579" specimenCode="TxEx-BR 2202" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Paratype">
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Type status:</emphasis>
|
||
<materialsCitation collectingDate="2022-10-16" collectorName="Iva Njunjic" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Iva Njunjic" specimenCode="TxEx-BR 2202" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Paratype">
|
||
<typeStatus pageId="0" pageNumber="101579">Paratype</typeStatus>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<collectingCounty>Occurrence</collectingCounty>
|
||
:
|
||
</emphasis>
|
||
catalogNumber:
|
||
<specimenCode pageId="0" pageNumber="101579">TxEx-BR2202</specimenCode>
|
||
; recordedBy:
|
||
<collectorName pageId="0" pageNumber="101579">
|
||
<collectingMunicipality>
|
||
Iva
|
||
<normalizedToken originalValue="Njunjić">Njunjic</normalizedToken>
|
||
</collectingMunicipality>
|
||
</collectorName>
|
||
; individualCount:
|
||
<specimenCount pageId="0" pageNumber="101579" type="generic">1</specimenCount>
|
||
; lifeStage:
|
||
<specimenType pageId="0" pageNumber="101579">juvenile</specimenType>
|
||
; occurrenceID:
|
||
<uuid>E66016F3-86E1-5F46-AC2C-6C3D1535162D</uuid>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:961D3548EE9209186464208AC075C361" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Taxon" stateProvince="Temburong District">Taxon</location>
|
||
:
|
||
</emphasis>
|
||
scientificName:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:36072F0E10A48AAF20AA65F6134BBD72" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Microparmarion" stateProvince="Temburong District">Microparmarion</location>
|
||
sallehi;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:1694F01BD5CD4B3C100913FF68DFE663" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Location" stateProvince="Temburong District">Location</location>
|
||
:
|
||
</emphasis>
|
||
country:
|
||
<collectingCountry name="Brunei" pageId="0" pageNumber="101579">Brunei</collectingCountry>
|
||
; stateProvince:
|
||
<collectingRegion country="Brunei" name="Temburong">Temburong District</collectingRegion>
|
||
; locality:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:58F6CC339C9185152719A9DB123879F3" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Kuala Belalong Field Studies Centre, Ashton trail" pageId="0" pageNumber="101579" stateProvince="Temburong District">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:7C8A92E9A2454E86F03A403102DEA810" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Kuala Belalong Field Studies Centre" stateProvince="Temburong District">Kuala Belalong Field Studies Centre</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:19009F7A59A5F7BBFAC1AAF8CDF4A56C" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Ashton" stateProvince="Temburong District">Ashton</location>
|
||
trail
|
||
</location>
|
||
; verbatimElevation:
|
||
<elevation pageId="0" pageNumber="101579">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
</elevation>
|
||
; decimalLatitude:
|
||
<geoCoordinate orientation="latitude" pageId="0" pageNumber="101579" value="4.547">4.547</geoCoordinate>
|
||
; decimalLongitude:
|
||
<geoCoordinate orientation="longitude" pageId="0" pageNumber="101579" value="115.157">115.157</geoCoordinate>
|
||
; georeferenceProtocol: GPS;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:572F8C91E9356AD72FBAFF649E8CCE7E" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Identification" stateProvince="Temburong District">Identification</location>
|
||
:
|
||
</emphasis>
|
||
identifiedBy:
|
||
<determinerName pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:A0F2380178E715A661A9F48C397EAB52" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Menno Schilthuizen" stateProvince="Temburong District">Menno Schilthuizen</location>
|
||
</determinerName>
|
||
; dateIdentified: 2022;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:AD6D3D88D45CEBDF00A57947CFD8F997" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Event" stateProvince="Temburong District">Event</location>
|
||
:
|
||
</emphasis>
|
||
eventDate:
|
||
<collectingDate pageId="0" pageNumber="101579" value="2022-10-16">16/10/2022</collectingDate>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Record Level:</emphasis>
|
||
language: en; basisOfRecord: PreservedSpecimen
|
||
<materialsCitation collectingDate="2022-10-16" collectorName="Iva Njunjic" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Iva Njunjic" pageId="0" pageNumber="101579" specimenCode="TxEx-BR 2203" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Paratype">
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Type status:</emphasis>
|
||
<materialsCitation collectingDate="2022-10-16" collectorName="Iva Njunjic" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Iva Njunjic" specimenCode="TxEx-BR 2203" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Paratype">
|
||
<typeStatus pageId="0" pageNumber="101579">Paratype</typeStatus>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<collectingCounty>Occurrence</collectingCounty>
|
||
:
|
||
</emphasis>
|
||
catalogNumber:
|
||
<specimenCode pageId="0" pageNumber="101579">TxEx-BR2203</specimenCode>
|
||
; recordedBy:
|
||
<collectorName pageId="0" pageNumber="101579">
|
||
<collectingMunicipality>
|
||
Iva
|
||
<normalizedToken originalValue="Njunjić">Njunjic</normalizedToken>
|
||
</collectingMunicipality>
|
||
</collectorName>
|
||
; individualCount:
|
||
<specimenCount pageId="0" pageNumber="101579" type="generic">1</specimenCount>
|
||
; lifeStage:
|
||
<specimenType pageId="0" pageNumber="101579">juvenile</specimenType>
|
||
; occurrenceID:
|
||
<uuid>82288ED8-A035-581D-B1CC-3EC3838F2953</uuid>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:92C19D92126B13F5E43EE2B46E4890C7" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Taxon" stateProvince="Temburong District">Taxon</location>
|
||
:
|
||
</emphasis>
|
||
scientificName:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:C0951C56AC5914F6E051B15AF635692C" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Microparmarion" stateProvince="Temburong District">Microparmarion</location>
|
||
sallehi;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:93754EED812E9F26996685D1BE8DA982" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Location" stateProvince="Temburong District">Location</location>
|
||
:
|
||
</emphasis>
|
||
country:
|
||
<collectingCountry name="Brunei" pageId="0" pageNumber="101579">Brunei</collectingCountry>
|
||
; stateProvince:
|
||
<collectingRegion country="Brunei" name="Temburong">Temburong District</collectingRegion>
|
||
; locality:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:79BD5E23AA402DE732811825236C3573" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Kuala Belalong Field Studies Centre, Ashton trail" pageId="0" pageNumber="101579" stateProvince="Temburong District">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:EE552C4CC5CC46C96CC66B9D9199A54A" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Kuala Belalong Field Studies Centre" stateProvince="Temburong District">Kuala Belalong Field Studies Centre</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:618D76A42D766687F6D0ED7E87A7B2AC" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Ashton" stateProvince="Temburong District">Ashton</location>
|
||
trail
|
||
</location>
|
||
; verbatimElevation:
|
||
<elevation pageId="0" pageNumber="101579">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
</elevation>
|
||
; decimalLatitude:
|
||
<geoCoordinate orientation="latitude" pageId="0" pageNumber="101579" value="4.547">4.547</geoCoordinate>
|
||
; decimalLongitude:
|
||
<geoCoordinate orientation="longitude" pageId="0" pageNumber="101579" value="115.157">115.157</geoCoordinate>
|
||
; georeferenceProtocol: GPS;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:8E81AFCB8F7F98F6D3D110049E0C8F2E" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Identification" stateProvince="Temburong District">Identification</location>
|
||
:
|
||
</emphasis>
|
||
identifiedBy:
|
||
<determinerName pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:A83367B18DA177EBE84A4BE1AE4866A6" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Menno Schilthuizen" stateProvince="Temburong District">Menno Schilthuizen</location>
|
||
</determinerName>
|
||
; dateIdentified: 2022;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:2FC8B1D083AB4E6672AC6E5A1BD951B4" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Iva Njunjic" name="Event" stateProvince="Temburong District">Event</location>
|
||
:
|
||
</emphasis>
|
||
eventDate:
|
||
<collectingDate pageId="0" pageNumber="101579" value="2022-10-16">16/10/2022</collectingDate>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Record Level:</emphasis>
|
||
language: en; basisOfRecord: PreservedSpecimen
|
||
<materialsCitation collectingDate="2019-09-30" collectorName="Taxon Expeditions" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Taxon Expeditions" pageId="0" pageNumber="101579" specimenCode="TxEx-BR 1901" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Other material">
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Type status:</emphasis>
|
||
<materialsCitation collectingDate="2019-09-30" collectorName="Taxon Expeditions" country="Brunei" county="Occurrence" determinerName="Menno Schilthuizen" elevation="0" latitude="4.547" location="Taxon" longitude="115.157" municipality="Taxon Expeditions" specimenCode="TxEx-BR 1901" specimenCount="1" specimenCount-juvenile="1" stateProvince="Temburong District" typeStatus="Other material">
|
||
<typeStatus pageId="0" pageNumber="101579">Other material</typeStatus>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<collectingCounty>Occurrence</collectingCounty>
|
||
:
|
||
</emphasis>
|
||
catalogNumber:
|
||
<specimenCode pageId="0" pageNumber="101579">TxEx-BR1901</specimenCode>
|
||
; recordedBy:
|
||
<collectorName pageId="0" pageNumber="101579">
|
||
<collectingMunicipality>Taxon Expeditions</collectingMunicipality>
|
||
</collectorName>
|
||
; individualCount:
|
||
<specimenCount pageId="0" pageNumber="101579" type="generic">1</specimenCount>
|
||
; lifeStage:
|
||
<specimenType pageId="0" pageNumber="101579">juvenile</specimenType>
|
||
; occurrenceID:
|
||
<uuid>4B28FD4F-841C-5A38-A713-632C00FB1C1B</uuid>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:70C9950EE35D60080449996DC1A51EDD" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Taxon" stateProvince="Temburong District">Taxon</location>
|
||
:
|
||
</emphasis>
|
||
scientificName:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:0D4FBC0726DB790D8A62574C2628645D" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Microparmarion" stateProvince="Temburong District">Microparmarion</location>
|
||
sallehi;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:0FECA211F3DC06A58717A1717FE4E9C7" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Location" stateProvince="Temburong District">Location</location>
|
||
:
|
||
</emphasis>
|
||
country:
|
||
<collectingCountry name="Brunei" pageId="0" pageNumber="101579">Brunei</collectingCountry>
|
||
; stateProvince:
|
||
<collectingRegion country="Brunei" name="Temburong">Temburong District</collectingRegion>
|
||
; locality:
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:1E7F0D13B62B4BEB71F6FF0D225BA60F" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Kuala Belalong Field Studies Centre, Ashton trail" pageId="0" pageNumber="101579" stateProvince="Temburong District">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:36CDAC7399FA2DB2368B7A5A2C068EE3" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Kuala Belalong Field Studies Centre" stateProvince="Temburong District">Kuala Belalong Field Studies Centre</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:16C0B05118CF5B8B42E6E9F44A8AF5F9" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Ashton" stateProvince="Temburong District">Ashton</location>
|
||
trail
|
||
</location>
|
||
; verbatimElevation:
|
||
<elevation pageId="0" pageNumber="101579">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
</elevation>
|
||
; decimalLatitude:
|
||
<geoCoordinate orientation="latitude" pageId="0" pageNumber="101579" value="4.547">4.547</geoCoordinate>
|
||
; decimalLongitude:
|
||
<geoCoordinate orientation="longitude" pageId="0" pageNumber="101579" value="115.157">115.157</geoCoordinate>
|
||
; georeferenceProtocol: GPS;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:8DCFDE9A829AA69E7BA734C08D0F7A47" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Identification" stateProvince="Temburong District">Identification</location>
|
||
:
|
||
</emphasis>
|
||
identifiedBy:
|
||
<determinerName pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:4319A2A74EC9398FF42BD733435B4C58" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Menno Schilthuizen" stateProvince="Temburong District">Menno Schilthuizen</location>
|
||
</determinerName>
|
||
; dateIdentified: 2022;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">
|
||
<location LSID="urn:lsid:plazi:treatment:FAE460A4A4E25E4F8C741603B9095381:B63E02AE1A6BA2112291D1F7458B39E2" country="Brunei" county="Occurrence" latitude="4.547" longitude="115.157" municipality="Taxon Expeditions" name="Event" stateProvince="Temburong District">Event</location>
|
||
:
|
||
</emphasis>
|
||
eventDate:
|
||
<collectingDate pageId="0" pageNumber="101579" value="2019-09-30">30/09/2019</collectingDate>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="101579">Record Level:</emphasis>
|
||
language: en; basisOfRecord: PreservedSpecimen
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="description">
|
||
<paragraph pageId="0" pageNumber="101579">Description</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
Preserved specimens (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F8784720" captionText="Figure 1. Microparmarion sallehi, n. sp., photographs of preserved holotype specimen (UBDM. 7.00152), Kuala Belalong Field Studies Centre, Brunei Darussalam; a dorsal view; b left lateral view; c right lateral view." figureDoi="10.3897/BDJ.11.e101579.figure1" httpUri="https://binary.pensoft.net/fig/802820" pageId="0" pageNumber="101579">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F8784722" captionText="Figure 2. Microparmarion sallehi, n. sp., photographs of preserved paratype specimen (TxEx-BR 2203), Kuala Belalong Field Studies Centre, Brunei Darussalam; a dorsal view; b left lateral view; c right lateral view." figureDoi="10.3897/BDJ.11.e101579.figure2" httpUri="https://binary.pensoft.net/fig/802821" pageId="0" pageNumber="101579">2</figureCitation>
|
||
). Ethanol-preserved juveniles are 6-11 mm in length. On the dorsal and lateral sides excluding the foot and its margins, there is a blackish-brown mottled pattern on a pale white background. The foot is pale white with a smooth surface, with the lateral head margins of the foot having a blackish-brown mottled pattern; the rest of the margins remain pale white. Three distinct longitudinal dark brown bands run on the head, one starting posterior of each eye tentacle and the third dorsomedially in between these two, starting level with them. These three bands end at the level of the pneumostome. In between the rim of the shell and when the left and right mantle lobes are pulled back, there can be either blackish-brown mottled patterns or stripes that encircle the shell. An unpigmented peripheral ridge runs from the left frontal margin of the mantle, immediately behind the left eye tentacle, circles the shell, where, on the caudolateral side of the shell, it throws off a short side branch that leads dorsad to the mantle edge; the main branch of the keel continues and ends on the right at the pneumostome. The genital opening is visible on the right lateral side behind the head and appears as a circular pore. In the adult, 29 mm long, the colour patterns are similar to those in the juveniles, but the contrasts are much reduced and colours range from pale light brown to reddish-brown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
Living specimens (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F8784733" captionText="Figure 3. Microparmarion sallehi, n. sp., photographs of living individuals, Kuala Belalong Field Studies Centre, Brunei Darussalam; a holotype, UBDM. 7.00152, right lateral view; b holotype, UBDM. 7.00152, left lateral view; c paratype, TxEx-BR 2201, right dorsolateral view." figureDoi="10.3897/BDJ.11.e101579.figure3" httpUri="https://binary.pensoft.net/fig/802822" pageId="0" pageNumber="101579">3</figureCitation>
|
||
). Living specimens differ from preserved ones in size (in ethanol, the animals contract to 50% of their live extended length) and in colouration, as follows. The general colouration for the dorsal side and ventral side is blackish-brown and pale white, respectively. In juveniles, the foot is pale white with a smooth surface, with the lateral frontal margins of the foot having a blackish-brown mottled pattern and the rest of the margins remaining pale white. In adults, this blackish-brown mottled pattern can not only extend to the frontal margins of the foot, but also the caudal margins. The mantle carries a blackish-brown mottled pattern on a pale background, with a yellowish-white streak extending from the left mantle lobe to the mantle itself. In adults, the pattern on the mantle largely disappears and is replaced by an ochre colouration, with the streak being reduced or becoming non-existent. The left and right mantle lobes in both adults and juveniles meet along a curved slit. Anterior to the mantle in juveniles, two longitudinal black stripes on the head that start caudally at the base of the eye tentacles extend towards the anterior part of the mantle. In addition, a dorso-medial black line on the head in between the eye tentacles also extends caudally towards the mantle. In juveniles, all four tentacles may be reddish- or greyish-brown (though the oral tentacles can be pale as well), but in the adult, this is reduced to an ochre brown, with the oral tentacles becoming paler. The caudal horn, dorsal to the caudal end of the foot, is pointed caudally and slightly extends beyond the length of the foot. In the adult, the caudal horn can be pointed or curved downwards. In juveniles, the colouration of the tail is a greyish-brown, with a dark brown mottled pattern. In the adult, this high-contrast colouration is replaced with a general browner colouration, though some mottling remains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="shell">
|
||
<paragraph pageId="0" pageNumber="101579">Shell</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
Juvenile shells (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F8784736" captionText="Figure 4. Microparmarion sallehi, n. sp., dorsal views of shells; a pencil drawing of the shell of a paratype specimen (TxEx-BR 2201), Helipad, Ulu-Ulu Resort, Brunei Darussalam; b photograph of the shell of the holotype specimen (UBDM. 7.00152), Kuala Belalong Field Studies Centre, Brunei Darussalam." figureDoi="10.3897/BDJ.11.e101579.figure4" httpUri="https://binary.pensoft.net/fig/802823" pageId="0" pageNumber="101579">4</figureCitation>
|
||
a) are oval, 3.5-6.0 mm long and 2.0-4.0 mm wide, of a transparent yellowish-green colouration, slanting towards the left lateral side, which causes a bulge on the left side. The surface is smooth and developmental rings are visible. In lateral view, the shell rim is flattened while the shell smoothly curves into a round bump on top. Whorls are not discernible: the shell consists of a single, oval dish. The adult shell (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F8784736" captionText="Figure 4. Microparmarion sallehi, n. sp., dorsal views of shells; a pencil drawing of the shell of a paratype specimen (TxEx-BR 2201), Helipad, Ulu-Ulu Resort, Brunei Darussalam; b photograph of the shell of the holotype specimen (UBDM. 7.00152), Kuala Belalong Field Studies Centre, Brunei Darussalam." figureDoi="10.3897/BDJ.11.e101579.figure4" httpUri="https://binary.pensoft.net/fig/802823" pageId="0" pageNumber="101579">4</figureCitation>
|
||
b) is 11 mm long and 8 mm wide, fingernail-like and similarly displays several darker-coloured growth bands, but no traces of whorls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="genitalia">
|
||
<paragraph pageId="0" pageNumber="101579">Genitalia</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
The penis, at around 5 mm from the connection with the vagina, bends back upon itself in a 2-mm-long loop (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F8784749" captionText="Figure 5. Microparmarion sallehi, n. sp., preserved holotype specimen (UBDM. 7.00152), Kuala Belalong Field Studies Centre, Brunei Darussalam; a dissected genitalia; b detail of penis and epiphallus." figureDoi="10.3897/BDJ.11.e101579.figure5" httpUri="https://binary.pensoft.net/fig/802824" pageId="0" pageNumber="101579">5</figureCitation>
|
||
). The looped part is of the same width as the proximal part of the penis. Distally, the looped part narrows abruptly and is enveloped by a muscular extension of the penis; the epiphallus, 2.2 mm long, is visible as an initially relatively thick, but then narrowing, sinuous tube inside this muscular sheath. The penis retractor muscle appears to connect at the basis of the epiphallus. Terminally, the epiphallus gives way to the vas deferens. At 4 mm from the genital opening, the gametolytic sac emerges from the vagina. The dart sac is ca. 5 mm long and connects to the vagina at the opposite side from the penis. It is connected to the 10-mm-long, 2-mm-wide dart sac gland by a narrow looped stalk.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="dna barcode">
|
||
<paragraph pageId="0" pageNumber="101579">DNA barcode</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
The
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">COI</emphasis>
|
||
DNA barcode of the holotype (UBDM.7.00152; BOLD TXEX074-23) is as follows:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
5
|
||
<normalizedToken originalValue="’AACATTATATATAATTTTTGGAGTTTGATGTGGTATAGTAGGAACAGGCTTATCATTATTAATTCGATTAGAATTAGGTACAGCTGGTGTATTACTTGATGATCATTTTTTTAATGTAATTGTGACTGCACATGCATTTGTAATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTTGGAAATTGAATAGTTCCTTTATTAATTGGAGCTCCAGATATAAGTTTTCCACGAATAAATAATATAAGATTTTGATTATTACCTCCTTCATTTGTATTTTTAATTGTTTCTAGTATGGTTGAAGGTGGTGCAGGAACTGGTTGAACAGTTTATCCACCTTTAAGAGGACCAGTAGGACATGCAGGAGCCTCAGTAGATTTGGCTATTTTTTCTTTACATTTAGCAGGTATATCATCTATTTTAGGTGCTATTAATTTTATTACTACTATTTTTAACATACGGTCTTCTGGTATAACTATAGAACGTGTAAGACTATTTGTTTGATCTATTTTAGTAACTGTATTTTTATTACTATTATCTTTACCTGTACTTGCTGGGGCTATTACAATACTTTTAACTGATCGAAATTTTAATACTAGTTTTTTTGATCCTGCTGGAGGAGGAGATCCGATTTTATATCAACATTTATTT3’">'AACATTATATATAATTTTTGGAGTTTGATGTGGTATAGTAGGAACAGGCTTATCATTATTAATTCGATTAGAATTAGGTACAGCTGGTGTATTACTTGATGATCATTTTTTTAATGTAATTGTGACTGCACATGCATTTGTAATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTTGGAAATTGAATAGTTCCTTTATTAATTGGAGCTCCAGATATAAGTTTTCCACGAATAAATAATATAAGATTTTGATTATTACCTCCTTCATTTGTATTTTTAATTGTTTCTAGTATGGTTGAAGGTGGTGCAGGAACTGGTTGAACAGTTTATCCACCTTTAAGAGGACCAGTAGGACATGCAGGAGCCTCAGTAGATTTGGCTATTTTTTCTTTACATTTAGCAGGTATATCATCTATTTTAGGTGCTATTAATTTTATTACTACTATTTTTAACATACGGTCTTCTGGTATAACTATAGAACGTGTAAGACTATTTGTTTGATCTATTTTAGTAACTGTATTTTTATTACTATTATCTTTACCTGTACTTGCTGGGGCTATTACAATACTTTTAACTGATCGAAATTTTAATACTAGTTTTTTTGATCCTGCTGGAGGAGGAGATCCGATTTTATATCAACATTTATTT3'</normalizedToken>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="101579">Diagnosis</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
This new species of
|
||
<taxonomicName authorityName="Simroth" authorityYear="1893" class="Gastropoda" family="Ariophantidae" genus="Microparmarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparmarion" order="Stylommatophora" pageId="0" pageNumber="101579" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">Microparmarion</emphasis>
|
||
</taxonomicName>
|
||
stands out because of its shell, which, unlike other north Bornean species of the genus, has no discernible whorls. In that sense, it appears to be more closely related to
|
||
<taxonomicName authorityName="P. Fischer" authorityYear="1855" class="Gastropoda" family="Ariophantidae" genus="Parmarion" kingdom="Animalia" lsidName="Parmarion" order="Stylommatophora" pageId="0" pageNumber="101579" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">Parmarion</emphasis>
|
||
</taxonomicName>
|
||
, with which it also shares the clearly keeled mantle. However, the genital set-up, with the looped part of the penis, the vas deferens connected terminally to the epiphallus and the unstalked gametolytic sac clearly fits with the
|
||
<taxonomicName authorityName="Simroth" authorityYear="1893" class="Gastropoda" family="Ariophantidae" genus="Microparmarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparmarion" order="Stylommatophora" pageId="0" pageNumber="101579" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">Microparmarion</emphasis>
|
||
</taxonomicName>
|
||
genus definition (
|
||
<bibRefCitation author="Simroth, H." journalOrPublisher="E. J. Brill, Leiden" pageId="0" pageNumber="101579" refId="B8784682" refString="Simroth, H., 1893. Zoologische Ergebnisse einer Reise in Niederländisch Ost-Indien 3. E.J. Brill, Leiden" title="Zoologische Ergebnisse einer Reise in Niederla ̈ ndisch Ost-Indien 3." year="1893">Simroth 1893</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Vermeulen, J. J." journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="0" pageNumber="101579" refId="B8784712" refString="Vermeulen, J. J., Liew, T. S., 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Universiti Malaysia Sabah, Kota Kinabalu" title="Land snails and slugs of Sabah and Labuan (Malaysia)" year="2022">Vermeulen and Liew 2022</bibRefCitation>
|
||
). The looped part of the penis, which remains thick until the place where it is enveloped by the muscular sheath, places the species in 'group 2' of
|
||
<bibRefCitation author="Vermeulen, J. J." journalOrPublisher="Universiti Malaysia Sabah, Kota Kinabalu" pageId="0" pageNumber="101579" refId="B8784712" refString="Vermeulen, J. J., Liew, T. S., 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Universiti Malaysia Sabah, Kota Kinabalu" title="Land snails and slugs of Sabah and Labuan (Malaysia)" year="2022">Vermeulen and Liew (2022)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
Externally similar north Bornean species are
|
||
<taxonomicName lsidName="M. simrothi" pageId="0" pageNumber="101579" rank="species" species="simrothi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. simrothi</emphasis>
|
||
</taxonomicName>
|
||
Collinge & Godwin-Austen, 1895,
|
||
<taxonomicName lsidName="M. exquadratus" pageId="0" pageNumber="101579" rank="species" species="exquadratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. exquadratus</emphasis>
|
||
</taxonomicName>
|
||
Schilthuizen et al., 2019, and
|
||
<taxonomicName lsidName="M. convolutus" pageId="0" pageNumber="101579" rank="species" species="convolutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. convolutus</emphasis>
|
||
</taxonomicName>
|
||
Vermeulen & Liew, 2022. What sets
|
||
<taxonomicName lsidName="M. sallehi" pageId="0" pageNumber="101579" rank="species" species="sallehi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. sallehi</emphasis>
|
||
</taxonomicName>
|
||
apart from
|
||
<taxonomicName lsidName="M. simrothi" pageId="0" pageNumber="101579" rank="species" species="simrothi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. simrothi</emphasis>
|
||
</taxonomicName>
|
||
is that the new species lacks
|
||
<taxonomicName lsidName="M. simrothi" pageId="0" pageNumber="101579" rank="species" species="simrothi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. simrothi</emphasis>
|
||
</taxonomicName>
|
||
's papillose mantle surface and has partial mottled patterns on the foot margins, whereas in
|
||
<taxonomicName lsidName="M. simrothi" pageId="0" pageNumber="101579" rank="species" species="simrothi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. simrothi</emphasis>
|
||
</taxonomicName>
|
||
the foot margin has an uninterrupted mottled pattern. The mantle of the new species lacks the uneven, interrupted dark ring close to the periphery of the part of the mantle that covers the shell. In adult
|
||
<taxonomicName lsidName="M. convolutus" pageId="0" pageNumber="101579" rank="species" species="convolutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. convolutus</emphasis>
|
||
</taxonomicName>
|
||
, the mantle is smooth, while in the new species, it is covered in low nodules and ripples. The colouration in
|
||
<taxonomicName lsidName="M. convolutus" pageId="0" pageNumber="101579" rank="species" species="convolutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. convolutus</emphasis>
|
||
</taxonomicName>
|
||
is whitish, pale orange or pale pink with a slight greyish mottling on the sides and the tail, while the new species is a pale orange as an adult or pale white as a juvenile with a blackish-brown mottled pattern.
|
||
<taxonomicName lsidName="M. exquadratus" pageId="0" pageNumber="101579" rank="species" species="exquadratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. exquadratus</emphasis>
|
||
</taxonomicName>
|
||
, finally, is generally darker in colour: dull orange to pale red with dark brown patterning, while the new species is a pale orange as an adult or pale white as a juvenile with a blackish-brown mottled pattern.
|
||
<taxonomicName lsidName="M. sallehi" pageId="0" pageNumber="101579" rank="species" species="sallehi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. sallehi</emphasis>
|
||
</taxonomicName>
|
||
has either uninterrupted blackish-brown mottled patterns or horizontal stripes that encircle the shell in juveniles. In adults, these patterns or stripes do not appear.
|
||
<taxonomicName lsidName="M. exquadratus" pageId="0" pageNumber="101579" rank="species" species="exquadratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. exquadratus</emphasis>
|
||
</taxonomicName>
|
||
, on the other hand, has a mantle with an uneven, wide, locally interrupted dark ring surrounding the lumen otherwise coarsely spotted. In live
|
||
<taxonomicName lsidName="M. exquadratus" pageId="0" pageNumber="101579" rank="species" species="exquadratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. exquadratus</emphasis>
|
||
</taxonomicName>
|
||
, the surface of the mantle is covered in small, irregular pustules, whereas in
|
||
<taxonomicName lsidName="M. sallehi" pageId="0" pageNumber="101579" rank="species" species="sallehi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. sallehi</emphasis>
|
||
</taxonomicName>
|
||
, there are parallel ripples.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">
|
||
In the anatomy,
|
||
<taxonomicName lsidName="M. sallehi" pageId="0" pageNumber="101579" rank="species" species="sallehi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. sallehi</emphasis>
|
||
</taxonomicName>
|
||
n. sp. differs from all these three similar species by the long, strongly developed looped part of the penis, which is as thick and as long as the proximal part of the penis. In
|
||
<taxonomicName lsidName="M. simrothi" pageId="0" pageNumber="101579" rank="species" species="simrothi">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. simrothi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="M. exquadratus" pageId="0" pageNumber="101579" rank="species" species="exquadratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. exquadratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="M. convolutus" pageId="0" pageNumber="101579" rank="species" species="convolutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101579">M. convolutus</emphasis>
|
||
</taxonomicName>
|
||
, the looped part is always much thinner or differently shaped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="etymology">
|
||
<paragraph pageId="0" pageNumber="101579">Etymology</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">The species epithet is a masculine genitive and honours the supervisor of the Kuala Belalong Field Studies Centre, Mr. Md Salleh Abdullah Bat, who recently retired. The taxon expedition, during which the species was described, was the last group he hosted before retirement. The name won the winning number of votes out of seven options during a voting session in which all participants of the 2022 expedition took part.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101579" type="ecology">
|
||
<paragraph pageId="0" pageNumber="101579">Ecology</paragraph>
|
||
<paragraph pageId="0" pageNumber="101579">This species is generally crepuscular and nocturnal. It was seen at night and in the morning after rain, crawling underneath leaves of saplings and on vines, in both primary and secondary forest.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |