treatments-xml/data/FA/D0/2D/FAD02D57673C518ABABF6CB235AD9F39.xml

<document ID-CLB-Dataset="62259" ID-DOI="http://dx.doi.org/10.3897/subtbiol.43.76049" ID-GBIF-Dataset="0dfd9d7b-30ff-44d2-9a07-889035a9c2a4" ID-Pensoft-Pub="1314-2615-43-73" ID-Pensoft-UUID="5003A89E81C4599B9A2D3CC5E13924B8" ID-ZooBank="48C162B126CB4ED989E32F20BF612F90" ModsDocID="1314-2615-43-73" checkinTime="1654082229011" checkinUser="pensoft" docAuthor="Curcic, Srecko, Vesovic, Nikola, Kuraica, Milos, Bosco, Fabrizio, Curcic, Nina B. &amp; Vrbica, Maja" docDate="2022" docId="FAD02D57673C518ABABF6CB235AD9F39" docLanguage="en" docName="SubterBiol 43: 73-95" docOrigin="Subterranean Biology 43" docPubDate="2022-05-31" docSource="http://dx.doi.org/10.3897/subtbiol.43.76049" docTitle="Duvalius (Biharotrechus) semecensis subsp. semecensis semecensis Winkler 1926" docType="treatment" docVersion="6" id="5003A89E81C4599B9A2D3CC5E13924B8" lastPageNumber="73" masterDocId="5003A89E81C4599B9A2D3CC5E13924B8" masterDocTitle="A new subspecies of the genus Duvalius Delarouzee, 1859 (Coleoptera, Carabidae, Trechini) from western Serbia, with a key and an annotated catalogue of Serbian Biharotrechus and Duvalius s. str. taxa" masterLastPageNumber="95" masterPageNumber="73" pageNumber="73" updateTime="1732844599693" updateUser="ExternalLinkService">
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<mods:title id="E1A86E8764B50A9848BBCBBDED33A22E">A new subspecies of the genus Duvalius Delarouzee, 1859 (Coleoptera, Carabidae, Trechini) from western Serbia, with a key and an annotated catalogue of Serbian Biharotrechus and Duvalius s. str. taxa</mods:title>
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<mods:namePart id="F5F3A99DF02F90534F49E3EF7BDBA2AA">Curcic, Srecko</mods:namePart>
<mods:affiliation id="7E26C98286979923DE33AA5CAB8EAB13">Institute of Zoology, University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
<mods:nameIdentifier id="375A6C057DF548D6FF9EA34C62CA264C" type="email">srecko@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="0A6AEFD14E91CCFF191F34BBECDD5CF6">Vesovic, Nikola</mods:namePart>
<mods:affiliation id="BA6398290183CDE19C22F52827D30043">Institute of Zoology, University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="129FA133706C1D1E3BEF8E07021844D9">Kuraica, Milos</mods:namePart>
<mods:affiliation id="133BA9775399FEDF1DE7B5202B9A0F99">Institute of Zoology, University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="B9DC02B4542B60A61B53806B09E3C445">Bosco, Fabrizio</mods:namePart>
<mods:affiliation id="7DB6D805975DF30EF93E0A811261B68A">Natisone Valley Speleological Group, Via Antro, San Giovanni d'Antro, 33046 Pulfero, Italy</mods:affiliation>
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<mods:namePart id="6ED283E084C97065C324846BFB387C27">Curcic, Nina B.</mods:namePart>
<mods:affiliation id="F1E80A44563E3E10CDE7E7B9248109D5">Geographical Institute &quot; Jovan Cvijic &quot;, Serbian Academy of Sciences and Arts, Đure Jaksica 9, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="E640052D5564B11AB0A563A5864A0BA5">Vrbica, Maja</mods:namePart>
<mods:affiliation id="13B0A1EABE6ECFC5CAD960933809448B">Institute of Zoology, University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<subSection id="8BEF86564588B2393A3DA5B7AD6F8EDC" pageId="0" pageNumber="73" type="results">
<treatment id="FAD02D57673C518ABABF6CB235AD9F39" ID-GBIF-Taxon="196160431" LSID="urn:lsid:plazi:treatment:FAD02D57673C518ABABF6CB235AD9F39" httpUri="http://treatment.plazi.org/id/FAD02D57673C518ABABF6CB235AD9F39" lastPageNumber="73" pageId="0" pageNumber="73" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae" scope_tribe="Trechini">
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<paragraph id="EB7B6ED4C1FA3E16A9928B59830F360A" pageId="0" pageNumber="73">
<taxonomicName id="A3FEE3CD339E98FF3892B1B53BC63C05" ID-CoL="384ZN" LSID="FAD02D57-673C-518A-BABF-6CB235AD9F39" authority="semecensis Winkler, 1926" authorityName="semecensis Winkler" authorityYear="1926" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duvalius (Biharotrechus) semecensis subsp. semecensis" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="semecensis" subGenus="Biharotrechus" subSpecies="semecensis">Duvalius (Biharotrechus) semecensis semecensis Winkler, 1926</taxonomicName>
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<paragraph id="D945A0B97FABD8C2688C79191D345DB5" pageId="0" pageNumber="73">
<figureCitation id="1BE2F40B51BC9E8EDCD46268AB66F84A" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">Figs 10</figureCitation>
<figureCitation id="01510C464BC8E63265E91CF0042B2BAE" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">, 11</figureCitation>
</paragraph>
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<subSubSection id="06170DE164329F93DE5F5ACCF91131C2" pageId="0" pageNumber="73" type="type material">
<paragraph id="B245003725E1E4D3837D6C2BE7A908A3" pageId="0" pageNumber="73">Type material.</paragraph>
<paragraph id="121D38E09C0FFF0FADA327B44C2C4D1C" pageId="0" pageNumber="73">
<emphasis id="C2774485BCF9CDE0479CC5ABFF884C57" bold="true" pageId="0" pageNumber="73">
<emphasis id="B26350CFC3DD3C09AAF877EAD515884B" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
(
<emphasis id="0212F8E3E3DAACFE8F25052CD02C54C8" italics="true" pageId="0" pageNumber="73">by monotypy</emphasis>
)
</emphasis>
: male (NHMV) labeled as follows: 
<normalizedToken id="C0973A5D528FCA8D3B61F4B81A936050" originalValue="„Höhle">&quot;Hoehle</normalizedToken>
a. d. 
<normalizedToken id="7786801425A92738AB1F266C6AC82ACB" originalValue="Semeč">Semec</normalizedToken>
plan., Weirather&quot;, with no date provided (white label, handwritten) / &quot; 
<taxonomicName id="54065B03C7B96E73223BEE7C7B4785C4" authorityName="Winkler" authorityYear="1926" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duvalius semecensis" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="semecensis">
<emphasis id="AC27521DA470455712062E9644F94D44" italics="true" pageId="0" pageNumber="73">Duvalius semecensis</emphasis>
</taxonomicName>
Wnl.&quot; (white label, handwritten) / &quot;Holotypus 
<taxonomicName id="FF21428A3EE9A9F609808976E5709628" authorityName="Winkler" authorityYear="1926" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duvalius (Biharotrechus) semecensis" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="semecensis" subGenus="Biharotrechus">Duvalius (Biharotrechus) semecensis</taxonomicName>
Winkler, 1926 
<normalizedToken id="5C160D15C8107D76FCF3349C2BBC9006" originalValue="Ćurčić">Curcic</normalizedToken>
det. 2021&quot; (red label, printed) (Fig. 
<figureCitation id="612A025166B580AB4440DAB00D54E80A" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
).
</paragraph>
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<subSubSection id="9034C47E4A0F43F5FF3E5A1F5785310B" pageId="0" pageNumber="73" type="remarks">
<paragraph id="E7E89DFCFA7E158B44517B0FA79AF58B" pageId="0" pageNumber="73">Remarks.</paragraph>
<paragraph id="8C901492D3B83C7656F7E9B62962DB1A" pageId="0" pageNumber="73">
For the purpose of comparisons, we studied one male specimen of 
<taxonomicName id="29DC0EF4F452B6A63136B1A8F9330668" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
borrowed from the NHMV, labeled as 
<normalizedToken id="A2CD750B8EE93B634D34DE9FF97BA031" originalValue="„Höhle">&quot;Hoehle</normalizedToken>
a. d. 
<normalizedToken id="2542C5514AD3A0A5C3CD0366F4D0AB65" originalValue="Semeč">Semec</normalizedToken>
plan. Weirather / 
<taxonomicName id="0D58474CDCDC7B49BC3BF0A49851CAAC" authorityName="Winkler" authorityYear="1926" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Duvalius semecensis" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="semecensis">
<emphasis id="37C5DC3F208799D02130D8DCAC127160" italics="true" pageId="0" pageNumber="73">Duvalius semecensis</emphasis>
</taxonomicName>
Wnl.&quot; (Figs 
<figureCitation id="BAE2A63954706905FEF214F576147811" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10-13</figureCitation>
), but with no red type label added. The handwriting on both white labels appears to be that of Albert Winkler (Figs 
<figureCitation id="DEFF4BFEB27D619AD3466A552748FF8F" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">12</figureCitation>
, 
<figureCitation id="8D9A4C967478F4550D30F4871C228A8D" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">13</figureCitation>
). According to the available data (H. Schillhammer, pers. comm.), all red type labels in the collection of Albert Winkler that belong to the NHMV have been added subsequently. It seems that Winkler 
<normalizedToken id="41B203BC6B213AE2D8A70A664C513C8A" originalValue="didn’t">didn't</normalizedToken>
add red type labels for his type specimens. In the original description of 
<taxonomicName id="D627EAE44CA35F02A84FB8C83DD7D849" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
, it was indicated that Augusta Weirather collected one male specimen in an unnamed cave on Mt. 
<normalizedToken id="2E57DF99D58A7B27A24ACD5EA89BF1CD" originalValue="Sjemeć">Sjemec</normalizedToken>
in eastern Bosnia and Herzegovina (
<bibRefCitation id="4470F5EB80CC636C27FCD9849E6C9C9A" author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="258 - 266" refId="B27" refString="Winkler, A, 1926. Bestimmungstabelle der Duvalius -Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna). Koleopterologische Rundschau 12: 258 - 266" title="Bestimmungstabelle der Duvalius - Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna)." volume="12" year="1926">Winkler 1926</bibRefCitation>
). As the type series consists of only one male specimen with the same locality data, we designate the male specimen from the NHMV as the HT by monotypy of 
<taxonomicName id="19A29C2A64A95C68945052C004A47D6C" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
. According to 
<bibRefCitation id="D0ADD8D98B2DC375C233A63F3D0EA8CC" author="Pretner, E" journalOrPublisher="Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck" pageId="0" pageNumber="73" pagination="85 - 234" refId="B26" refString="Pretner, E, 2011. Die Verdienste von Leo Weirather um die Biospelaeologie, insbesondere Jugoslawiens, sein Hoehlenkataster und seine Sammelplaetze. Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck 97: 85 - 234" title="Die Verdienste von Leo Weirather um die Biospelaeologie, insbesondere Jugoslawiens, sein Hoehlenkataster und seine Sammelplaetze." volume="97" year="2011">Pretner (2011)</bibRefCitation>
, who decoded the list of the cave and above-ground collecting sites visited by Weirather, the right name of the type locality of this taxon is the Vrteljka Cave, which is situated in the village of 
<normalizedToken id="0257355E2F4AB3D3C56A3F786933F2E6" originalValue="Đipi">Đipi</normalizedToken>
(43°44'35&quot;N, 19°11'2&quot;E) on Mt. 
<normalizedToken id="908CB67819531D08EEDA93D800B1AA5C" originalValue="Sjemeć">Sjemec</normalizedToken>
, near the town of 
<normalizedToken id="118E28DAAB3EAE5A6FB5689CCD4E4D00" originalValue="Višegrad">Visegrad</normalizedToken>
, eastern Bosnia and Herzegovina. As we noted certain morphological intraspecific differences between the HT of 
<taxonomicName id="D8F136A04CBA37CFC24F1BE04F1CF90A" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
from the NHMV and 
<taxonomicName id="F03080DB29C82DB1CB961284ABA2A153" lsidName="D. (B.) semecensis subsp. tarensis" pageId="0" pageNumber="73" rank="subSpecies" species="semecensis" subGenus="B." subSpecies="tarensis">D. (B.) semecensis tarensis</taxonomicName>
ssp. nov., the need of establishing nominotypical subspecies [
<taxonomicName id="ECAFFC859BD5EBE96BBD13B71E42091E" authority="semecensis" lsidName="D. (B.) semecensis subsp. semecensis" pageId="0" pageNumber="73" rank="subSpecies" species="semecensis" subGenus="B." subSpecies="semecensis">D. (B.) semecensis semecensis</taxonomicName>
] of 
<taxonomicName id="2107C0D01744032071457055C5D3DD19" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
has arisen.
</paragraph>
<paragraph id="B1DB76F65CD9CAAE364524F5039FF4C9" pageId="0" pageNumber="73">
After reading both the original description of 
<taxonomicName id="1F529C2CC425DB0272CDF62B2AC46066" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
by 
<bibRefCitation id="FCF9A5CCE18D671AA056705F4867CEDE" author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="258 - 266" refId="B27" refString="Winkler, A, 1926. Bestimmungstabelle der Duvalius -Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna). Koleopterologische Rundschau 12: 258 - 266" title="Bestimmungstabelle der Duvalius - Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna)." volume="12" year="1926">Winkler (1926)</bibRefCitation>
and its subsequent description by 
<bibRefCitation id="3D50F6885F88951FDC7EC9646F216C22" author="Jeannel, R" journalOrPublisher="L'Abeille" pageId="0" pageNumber="73" pagination="1 - 808" refId="B18" refString="Jeannel, R, 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison). L'Abeille 35: 1 - 808" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison)." volume="35" year="1928">Jeannel (1928)</bibRefCitation>
and a careful examination of the HT of 
<taxonomicName id="576F6F675504FE89780407756512F544" lsidName="D. (B.) semecensis" pageId="0" pageNumber="73" rank="species" species="semecensis" subGenus="B.">D. (B.) semecensis</taxonomicName>
, we realized that some data on the morphology of the specimen listed in the previous literature do not correspond with the features of the observed male specimen. Namely, 
<bibRefCitation id="36C139996D5B5AA0EF6D1A4556C3E444" author="Jeannel, R" journalOrPublisher="L'Abeille" pageId="0" pageNumber="73" pagination="1 - 808" refId="B18" refString="Jeannel, R, 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison). L'Abeille 35: 1 - 808" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison)." volume="35" year="1928">Jeannel (1928)</bibRefCitation>
reported that the head of 
<taxonomicName id="22D4824CFDA45A90A3AF163A057E62FE" authority="semecensis" lsidName="D. (B.) semecensis subsp. semecensis" pageId="0" pageNumber="73" rank="subSpecies" species="semecensis" subGenus="B." subSpecies="semecensis">D. (B.) semecensis semecensis</taxonomicName>
is remarkably narrow, longer than wide, while 
<bibRefCitation id="E4E6319ACE9DBA59F11CF9C3308E4649" author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="258 - 266" refId="B27" refString="Winkler, A, 1926. Bestimmungstabelle der Duvalius -Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna). Koleopterologische Rundschau 12: 258 - 266" title="Bestimmungstabelle der Duvalius - Arten Jugoslaviens. Mit Neubeschreibungen (4. Beitrag zur Kenntnis der Blindkaeferfauna)." volume="12" year="1926">Winkler (1926)</bibRefCitation>
<normalizedToken id="7A1A902B321CC9A9A92FC7DAE2751C68" originalValue="didn’t">didn't</normalizedToken>
mention this in the description. The former author claimed that its pronotum is narrow, not wider than long, while the second author wrote that its pronotum is slightly wider than long. Both authors said that both discal setae are situated in fourth elytral intervals. Additionally, 
<normalizedToken id="9D349FC40448523A1DDFC178448F96A8" originalValue="Jeannel’s">Jeannel's</normalizedToken>
drawing of the specimen (
<bibRefCitation id="BD20D58743A6E9DCCCD111697FA0CD8F" author="Jeannel, R" journalOrPublisher="L'Abeille" pageId="0" pageNumber="73" pagination="1 - 808" refId="B18" refString="Jeannel, R, 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison). L'Abeille 35: 1 - 808" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison)." volume="35" year="1928">Jeannel 1928</bibRefCitation>
) is not perfect as it 
<normalizedToken id="93C965989B8AA88D43CB1F2B52F2D1CF" originalValue="doesn’t">doesn't</normalizedToken>
reflect its true shape (especially the head). Indeed, in the studied HT male, both head and pronotum are transverse, and the second discal seta is in the third stria (as was drawn by 
<bibRefCitation id="33FD4F79E46CEEAE88A3F57AD5FE8CEE" author="Jeannel, R" journalOrPublisher="L'Abeille" pageId="0" pageNumber="73" pagination="1 - 808" refId="B18" refString="Jeannel, R, 1928. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison). L'Abeille 35: 1 - 808" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (troisieme livraison)." volume="35" year="1928">Jeannel 1928</bibRefCitation>
) (Table 
<tableCitation id="0E50B9517DEB4D738C81708DB2EFB2C6" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Linear measurements and morphometric ratios of Duvalius (Biharotrechus) semecensis tarensis ssp. nov. and its closest conspecific relative D. (B.) semecensis semecensis." httpUri="http://table.plazi.org/id/DEDCD1CB1158ADF55C8EF83434D04611" pageId="0" pageNumber="73" tableUuid="DEDCD1CB1158ADF55C8EF83434D04611">1</tableCitation>
, Fig. 
<figureCitation id="56578899B24F6AA9E1219F23D9ACB70E" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
). For these reasons, we decided to redescribe the HT of 
<taxonomicName id="F60121464053E46F3AFDB4B0F8D7800F" authority="semecensis" lsidName="D. (B.) semecensis subsp. semecensis" pageId="0" pageNumber="73" rank="subSpecies" species="semecensis" subGenus="B." subSpecies="semecensis">D. (B.) semecensis semecensis</taxonomicName>
and to add here additional data on its morphology.
</paragraph>
</subSubSection>
<subSubSection id="C8DF74B13CD0855313D4463AC06993E1" pageId="0" pageNumber="73" type="redescription">
<paragraph id="07E448B9A51794662FD1C48E545F8254" pageId="0" pageNumber="73">Redescription.</paragraph>
<paragraph id="20EB3AB2830B3D61456B7EB5D2A666D2" pageId="0" pageNumber="73">Medium-sized. TL 4.17 mm, L 4.40 mm.</paragraph>
<paragraph id="E1BD49A44A2B9C67EF495D309A33AE22" pageId="0" pageNumber="73">
<emphasis id="C4C60B1F0411360D25426FA61EA5DAB5" bold="true" italics="true" pageId="0" pageNumber="73">Habitus</emphasis>
: Body elongate. Colour reddish-brown. Legs and palpi paler.
</paragraph>
<paragraph id="25C5E0C70BD6ED0ED3F53D4A91FD1C3C" pageId="0" pageNumber="73">
<emphasis id="C1C935958FED94D4D2BC92502E0FED0F" bold="true" italics="true" pageId="0" pageNumber="73">Integument</emphasis>
: Smooth, lustrous, head with a distinct isodiametric microsculpture, while microsculpture of both pronotum and elytra possesses both transverse and isodiametric meshes.
</paragraph>
<paragraph id="C294DA5539C1CF6FE4A79BBD985894F1" pageId="0" pageNumber="73">
<emphasis id="97F47811565A4C8FA7CA0B2C0CB4964E" bold="true" italics="true" pageId="0" pageNumber="73">Head</emphasis>
: Relatively large, around as long as 1/6 of TL, rounded (Fig. 
<figureCitation id="E9930FF419CF848E14257B3F9FE74A27" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
), narrower than pronotum, glabrous. Genae are rounded. Neck well-developed. HL 0.67 mm, HW 0.79 mm, HL/HW 0.85. Frontal furrows complete, deep, reaching neck constriction. Two pairs of supraorbital setae present, the anterior pair situated at the level of reduced eyes, while the posterior pair is near frontal furrows. Mandibles are thin, relatively long, sharply pointed, right one with a retinaculum. Labrum is deeply emarginate, with three pairs of setae. Clypeus is with two pairs of setae. Eyes are strongly reduced, oval, whitish, darkly bordered (Fig. 
<figureCitation id="319E11D66312A5FAB9F63878EB9AAC85" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">11</figureCitation>
), composed of 1-2 ommatidia. Antennae thin, long, haired, reaching almost half of elytral length when stretched backward. Antennomere 11 longest, followed by equally long antennomeres 1 and 3, antennomere 2 shortest (A2L/A2W 1.89), antennomeres 4 and 6-10 of similar length, antennomere 5 somewhat longer, A10L/A10W 2.60.
</paragraph>
<paragraph id="6A231FA46A5396F4CDD867D01CA504B8" pageId="0" pageNumber="73">
<emphasis id="714F584154B01225A084AD89D4127D18" bold="true" italics="true" pageId="0" pageNumber="73">Thorax</emphasis>
: Pronotum glabrous, transverse, cordiform (Fig. 
<figureCitation id="21DF44C9CA5D4C26B51B3D744BF7FE19" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
), slightly wider than long, with maximum width slightly before anterior third, at anterior margin wider than at pronotal base. PL 0.83 mm, PW 1.00 mm, PL/PW 0.83. Lateral margins rounded anteriorly, sinuated before hind angles. Anterior angles are rounded and obtuse. Posterior angles are sharp and pointed. Lateral furrows are relatively narrow and shallow, with two pairs of setae. Anterolateral pair of setae situated slightly before the anterior third of pronotal length, basolateral pair of setae before hind angles. Median furrow is well-developed, deep and long. Disc weakly convex.
</paragraph>
<paragraph id="79A3252763557C6095012E55AC567DB2" pageId="0" pageNumber="73">
<emphasis id="29C68E890037AE7292AEF4E61A2373AD" bold="true" italics="true" pageId="0" pageNumber="73">Elytra</emphasis>
: Elongate, sub-ovate, glabrous, with maximum width slightly after middle, apically rounded (Fig. 
<figureCitation id="CBFA905BFB82CE3EA17BFB847B76FD49" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
). EL 2.56 mm, EW 1.55 mm, EL/EW 1.65. Elytral base oblique. Shoulders are somewhat prominent, obtuse and sloped. Scutellum is small, sub-triangular, with one pair of scutellar setae. Elytral striae 1-5 well-developed, deep, outer striae reduced to rows of foveae. Punctuation on striae dense and deep. First discal seta is in fourth interval, second discal seta is in third stria, while apical seta is positioned at place where second and third striae coalesce, slightly below the level of second humeral seta. First discal seta slightly before the level of third humeral seta. Second discal seta after the middle of elytra. Intervals are slightly convex (inner intervals more than outer ones). Disc relatively convex. The umbilicate series consists of eight setae on each elytron (four humeral, two median and two apical), while the humeral group of umbilicate pores is aggregated.
</paragraph>
<paragraph id="521C770DFC1EEC909377B8DE6CE9AE9C" pageId="0" pageNumber="73">
<emphasis id="8229E4554E8229A7C90C3FB46BC3CBBF" bold="true" italics="true" pageId="0" pageNumber="73">Legs</emphasis>
: Elongate, thin, densely pubescent (Fig. 
<figureCitation id="190419CDBE84DDEFF196E123402E3DB8" captionStart="Figures 10–13" captionStartId="F5" captionText="Figures 10 - 13. Duvalius (Biharotrechus) semecensis semecensis from the Vrteljka Cave, village of Đipi, near the town of Visegrad, Mt. Sjemec, eastern Bosnia and Herzegovina 10 HT male, habitus, dorsal aspect 11 HT male, head, lateral aspect 12 locality data label of HT male 13 taxonomic name label of HT male. Scale bars: 2 mm (10); 0.5 mm (11)." figureDoi="10.3897/subtbiol.43.76049.figures10-13" httpUri="https://binary.pensoft.net/fig/690424" pageId="0" pageNumber="73">10</figureCitation>
), each protibia dorsally has a deep longitudinal fissure. The first two protarsomeres in the male are distinctly dilated and toothed at the internal margin. The first male protarsomere is longer than wide. Tarsal claws are long, thin, pointed apically.
</paragraph>
<paragraph id="565117C086DB4A57611014DDC90485A4" pageId="0" pageNumber="73">
<emphasis id="BBA32EDCC11BB6EAD1A59316D802438C" bold="true" italics="true" pageId="0" pageNumber="73">Abdomen</emphasis>
: Ventrites 4-6 and anal ventrite are glabrous, each with one pair of setae posteriorly. Male abdominal sternite IX (urite) missing, previously extracted from the male specimen.
</paragraph>
<paragraph id="DAFAB66805972888C51F6F7FE8F4AB7A" pageId="0" pageNumber="73">
<emphasis id="CDD211D5D274882686A6C86994B95465" bold="true" italics="true" pageId="0" pageNumber="73">Male genitalia</emphasis>
: Aedeagus missing, previously extracted from the male specimen.
</paragraph>
<paragraph id="EED7E432F80B8FBCAA86E77DF9C5C2BC" pageId="0" pageNumber="73">
<emphasis id="CA3E746EA829F63C041420936F009C6F" bold="true" italics="true" pageId="0" pageNumber="73">Female genitalia</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="499D7248E9582AA5D7D4F4D4D27ECF2C" pageId="0" pageNumber="73" type="distribution">
<paragraph id="0193979E576D9B943642F42C6F90FCBB" pageId="0" pageNumber="73">Geographic distribution.</paragraph>
<paragraph id="C810850D6AEC92680EDCBE35B49A3AA9" pageId="0" pageNumber="73">
So far known from its type locality (Vrteljka Cave) situated on Mt. 
<normalizedToken id="A0BA32844604D1A883EFCC26EF29B426" originalValue="Sjemeć">Sjemec</normalizedToken>
. Eric 
<normalizedToken id="EDA220BD85C866E90CF96A5BEBF43440" originalValue="Quéinnec">Queinnec</normalizedToken>
(pers. comm.) collected specimens of the same taxon in MSS in the surroundings of the type locality, at both sides of the Drina River, close to the town of 
<normalizedToken id="09BE3D41A60EC991406A374D71B04E45" originalValue="Višegrad">Visegrad</normalizedToken>
in eastern Bosnia and Herzegovina.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>