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<document id="6DB024FF4303A7C7F5988B43A80E6113" ID-CLB-Dataset="298662" ID-DOI="10.3897/zse.100.121512" ID-GBIF-Dataset="0b46f19d-c311-4c81-a69b-36b3fe37c5b0" ID-ZooBank="B0AF778B-E394-4C17-910D-FA2BDFB14A4F" ID-publisher-id="121512" URI-arpha="AF9BCD34-B3E8-5EBB-BDEF-F7DE25C0E7BD" URI-zoobank="http://zoobank.org/B0AF778B-E394-4C17-910D-FA2BDFB14A4F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1718792674186" checkinUser="pensoft" docAuthor="Tsukamoto, Sho &amp; Eguchi, Katsuyuki" docDate="2024" docId="FACE8FC162AE53B487E2AB55C607B251" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 821-840" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/121512/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dicellophilus pulcher" docType="treatment" docVersion="6" dtd-version="3.0" lastPageNumber="840" masterDocId="AF9BCD34B3E85EBBBDEFF7DE25C0E7BD" masterDocTitle="Integrative taxonomy of Dicellophilus Cook, 1896 (Chilopoda, Geophilomorpha, Mecistocephalidae) in Japan, with a description of a new species" masterLastPageNumber="840" masterPageNumber="821" pageNumber="821" updateTime="1718832442199" updateUser="ExternalLinkService">
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<mods:title id="A4FC42A609AF6AD9A0DE6E7E086EEB01">Integrative taxonomy of Dicellophilus Cook, 1896 (Chilopoda, Geophilomorpha, Mecistocephalidae) in Japan, with a description of a new species</mods:title>
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<mods:namePart id="B4CF67A8DCE0B6841A96E0591F58B4A5">Tsukamoto, Sho</mods:namePart>
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<mods:affiliation id="CE7265D7846151FE85FB91531B956311">Systematic Zoology Laboratory, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo 192 - 0397, Japan</mods:affiliation>
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<mods:namePart id="9F289576945850BF0F94E0F0373A2C23">Eguchi, Katsuyuki</mods:namePart>
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<mods:affiliation id="B3F1131C3B696C4D685C243A8566E420">Systematic Zoology Laboratory, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo 192 - 0397, Japan &amp; Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, 1 - 12 - 4 Sakamoto, Nagasaki 852 - 8523, Japan</mods:affiliation>
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<treatment id="FACE8FC162AE53B487E2AB55C607B251" ID-DOI="http://doi.org/10.5281/zenodo.12172784" ID-GBIF-Taxon="233233640" ID-Zenodo-Dep="12172784" ID-arpha="FACE8FC1-62AE-53B4-87E2-AB55C607B251" LSID="urn:lsid:plazi:treatment:FACE8FC162AE53B487E2AB55C607B251" httpUri="http://treatment.plazi.org/id/FACE8FC162AE53B487E2AB55C607B251">
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<taxonomicName id="D139AA09111854676D9500A204A496D5" authority="(Kishida, 1928)" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="BF939DC5238642FD5C7F198E80DDFDE8" italics="true">Dicellophilus pulcher</emphasis>
(
<bibRefCitation id="52EC128FB541631BC4481650EE6F6058" author="Kishida K" editor="Sengen Shrine" journalOrPublisher="Fuji. Kokinshoin, Tokyo" pagination="290-303" refId="B20" refString="Kishida K (1928) Ch. 6. Chilopoda. In: Sengen Shrine (Ed.) Animals of Mt. Fuji. Kokinshoin, Tokyo, 290 - 303. [in Japanese]" title="Ch. 6. Chilopoda." volumeTitle="Animals of Mt." year="1928">Kishida, 1928</bibRefCitation>
)
</taxonomicName>
</paragraph>
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<paragraph id="205BA1336CFFC46932212B6ECEF1D6FB">
<figureCitation id="BE693DC7BA82A20687B2A579B99127B5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicellophilus pulcher (TS 20210504-01). A. Cephalic plate, dorsal; B. Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure14" httpUri="https://binary.pensoft.net/fig/1073621">Figs 14</figureCitation>
,
<figureCitation id="967BE8F4FC217AA1325BC4AF822F9BCB" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicellophilus pulcher (TS 20210504-01). A. Forcipular segment, dorsal; B. Forcipular segment, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure15" httpUri="https://binary.pensoft.net/fig/1073622">15</figureCitation>
,
<figureCitation id="29895F65A407F915239435CB2AD7747D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicellophilus pulcher (TS 20210504-01). A. Ultimate leg-bearing segment and postpedal segment, dorsal; B. Ultimate leg-bearing segment and postpedal segment, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure16" httpUri="https://binary.pensoft.net/fig/1073623">16</figureCitation>
</paragraph>
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<paragraph id="1FE944233E0C6B0CF7B3D110B91DBA58">
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<taxonomicName id="C58F6276BF67E9CEC527697C282430E8" ID-CoL="72WLQ" authority="Kishida, 1928" authorityName="Kishida" authorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="2F0CC08616327C266C4C5327A53A4035" italics="true">Mecistocephalus pulcher</emphasis>
<bibRefCitation id="20520D4AABAA58A2167F08994D822C99" author="Kishida K" editor="Sengen Shrine" journalOrPublisher="Fuji. Kokinshoin, Tokyo" pagination="290-303" refId="B20" refString="Kishida K (1928) Ch. 6. Chilopoda. In: Sengen Shrine (Ed.) Animals of Mt. Fuji. Kokinshoin, Tokyo, 290 - 303. [in Japanese]" title="Ch. 6. Chilopoda." volumeTitle="Animals of Mt." year="1928">Kishida, 1928</bibRefCitation>
</taxonomicName>
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:
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<bibRefCitation id="2DD66BE665E688694F57FC7EEBE26BE2" author="Kishida" editor="Sengen Shrine" firstAuthor="Kishida" journalOrPublisher="Fuji. Kokinshoin, Tokyo" pagination="290-303" refId="B20" refString="Kishida K (1928) Ch. 6. Chilopoda. In: Sengen Shrine (Ed.) Animals of Mt. Fuji. Kokinshoin, Tokyo, 290303. [in Japanese]" title="Ch. 6. Chilopoda." volumeTitle="Animals of Mt." year="1928">Kishida 1928</bibRefCitation>
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, 300.
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</paragraph>
<paragraph id="49FDF2D97E678AF11AEB2FFB0664F552">
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<taxonomicName id="23EBFF0CB3DB2793C2D61EAAC99DDB03" ID-CoL="6CSRG" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="8F7293B51A31897DA6798B8DAA446CAB" italics="true">Dicellophilus latifrons</emphasis>
</taxonomicName>
:
<bibRefCitation id="7D6CD699176F1073AF7C344455989615" author="Takakuwa Y" issue="4" journalOrPublisher="Botany and Zoology" pagination="706-712" refId="B33" refString="Takakuwa Y (1934 a) The family Mecistocephalidae of Japan, I. Botany and Zoology 2 (4): 706712. [in Japanese]" title="The family Mecistocephalidae of Japan, I." volume="2">Takakuwa 1934 a</bibRefCitation>
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, 707;
<treatmentCitation id="1786211EF4A1D8DFACDE5FA5015A6A69">
<bibRefCitation id="258E58DBE47521F296DDA43698A5C6FA" author="Takakuwa Y" journalOrPublisher="Annotationes Zoologicae Japonenses" pagination="355-363" refId="B34" refString="Takakuwa Y (1934 b) Neue japanische Mecistocephalidae. Annotationes Zoologicae Japonenses 14: 355363." title="Neue japanische Mecistocephalidae." volume="14">Takakuwa 1934 b</bibRefCitation>
</treatmentCitation>
, 355;
<treatmentCitation id="6F749F4AB55593AD6761321C06E5F38A">
<bibRefCitation id="561D56282C9BB1A1905FCA45F058FF32" author="Takakuwa Y" issue="5" journalOrPublisher="Botany and Zoology" pagination="878-884" refId="B35" refString="Takakuwa Y (1934 c) The family Mecistocephalidae of Japan, II. Botany and Zoology 2 (5): 878884. [in Japanese]" title="The family Mecistocephalidae of Japan, II." volume="2">Takakuwa 1934 c</bibRefCitation>
</treatmentCitation>
, 878.
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</paragraph>
<paragraph id="49BAB5DE8A96D2253ADD37D1558142E2">
<treatmentCitationGroup id="859A4D00EE0EA274F20498237D3379F1">
<treatmentCitation id="8821F95FF3AB4FF2AFE08FE4A7B7092D">
<taxonomicName id="9A9CF110826F565E727B75D6434212CF" ID-CoL="35J7T" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="D394EBA4C7370A306C2225FAAEA66CE3" italics="true">Dicellophilus japonicus</emphasis>
</taxonomicName>
:
<bibRefCitation id="BEB43571D6ED96436B58C421CB56D117" author="Verhoeff" firstAuthor="Verhoeff" journalOrPublisher="Abteilung für Systematik" pagination="1-112" refId="B44" refString="Verhoeff KW (1934) Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher. Abteilung für Systematik 66: 1112." title="Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher." volume="66" year="1934">Verhoeff 1934</bibRefCitation>
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, 32.
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</paragraph>
<paragraph id="810940275F0991DDD997E0B4DC1FC20C">
<treatmentCitationGroup id="E92D988F3B9D9F40311B5ED73DFACDF6">
<treatmentCitation id="7AFC40AE9D883EBD99CAE0A9A48D9BD9">
<taxonomicName id="AB9CDFA827BDD57FAB33C41A2347CA60" ID-CoL="59SS5" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="6591DEC4C52D2384C59F1E601570821E" italics="true">Tygarrup monoporus</emphasis>
</taxonomicName>
:
<bibRefCitation id="D1A5FB0B3965EDDE2CF2BA67283EADBC" author="Shinohara" firstAuthor="Shinohara" issue="7" journalOrPublisher="Zoological Magazine" pagination="212-216" refId="B29" refString="Shinohara K (1961) Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda). Zoological Magazine 70 (7): 212216. [in Japanese with English Resume]" title="Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda)." volume="70" year="1961">Shinohara 1961</bibRefCitation>
</treatmentCitation>
, 212.
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</paragraph>
<paragraph id="1332A47A94E3D09783D0A92D1D32EFA1">
<treatmentCitationGroup id="60455166D9AC80D403F282DC5B13A6E1">
<treatmentCitation id="C7C57F170F100CF549B1AE27E60550A9">
<taxonomicName id="35EEB48DA5C6E64BCB3C3F9874B58D52" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="C897E9FC6A672E221A406461E1C733B4" italics="true">Dicellophilus pulcher</emphasis>
</taxonomicName>
:
<bibRefCitation id="2AF8496BAF4246741BA53150370B3C49" DOI="10.11646/zootaxa.1396.1.1" author="Uliana" etAl="et al." firstAuthor="Uliana" issue="1" journalOrPublisher="Zootaxa" pagination="1-84" refId="B43" refString="Uliana M, Bonato L, Minelli A (2007) The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha). Zootaxa 1396 (1): 184. https://doi.org/10.11646/zootaxa.1396.1.1" title="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)." volume="1396" year="2007">Uliana et al. 2007</bibRefCitation>
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, 27; Bonato et al. 2010, 525.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EOFBI" type="DI">
<paragraph id="1C3B80DD8FA467E3CFB90ADD2AEA021B">
<heading id="C247BCD1EB0E5E841B6D7583B9585FCD" reason="title">DI.</heading>
</paragraph>
<paragraph id="E51C5800995C03599CB52EAF1C9AA087">
<taxonomicName id="D605A1839EFE6933A904E05A91D70F77" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="74FAF1E60EE700F8FB73F5CA55D2B2C0" italics="true">Dicellophilus</emphasis>
sp.
</taxonomicName>
<ext-link id="0450152871699461196C9B03F48C1E2E" ext-link-type="uri" xlink_href="https://orcid.org/0000-0003-3020-8454" xlink_type="simple">0000-0003 - 3020 - 8454</ext-link>
_ 0068
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAGBI" type="material">
<paragraph id="311A060528064AF1AC1985F30AC65AF2">
<heading id="CA019AEEF334D102815E91D9C0598F5C" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="5FE8135EF87BD5C94F8A3056B1979417">
See Table
<tableCitation id="A3152C98F3533A45A5DBDB79D7F45B53" captionStart="Table 1" captionStartId="T1" captionText="Table 1. The list of specimens that were used in the phylogenetic analyses. Geographic coordinates enclosed by parentheses are secondary due to the lack of information in the labels." httpUri="http://table.plazi.org/id/9062C7A83B55F7C26FE5C8CDA5EB5F15" tableUuid="9062C7A83B55F7C26FE5C8CDA5EB5F15">1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKGBI" type="diagnosis">
<paragraph id="E3A8222C9747F7E3E5F4FE63B450C050">
<heading id="C0085C37064707B6762FB5D835307CE4" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="3416A96AE0FBB4573FE88232E55D0A54">
Mainly based on
<bibRefCitation id="E099F8FB3642525A3FDCF4ED914F6A3C" DOI="10.1111/j.1096-3642.2009.00557.x" author="Bonato L &amp; Dányi L &amp; Minelli A" issue="3" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="501-532" refId="B4" refString="Bonato L, Dányi L, Minelli A (2010 a) Morphology and phylogeny of Dicellophilus, a centipede genus with a highly disjunct distribution (Chilopoda: Mecistocephalidae). Zoological Journal of the Linnean Society 158 (3): 501532. https://doi.org/10.1111/j.1096-3642.2009.00557.x" title="Morphology and phylogeny of Dicellophilus, a centipede genus with a highly disjunct distribution (Chilopoda: Mecistocephalidae)." volume="158">Bonato et al. (2010 a</bibRefCitation>
),
<bibRefCitation id="679239D52419D5B1820F81B7871755F2" DOI="10.11646/zootaxa.1396.1.1" author="Uliana M &amp; Bonato L &amp; Minelli A" etAl="et al." firstAuthor="Uliana" issue="1" journalOrPublisher="Zootaxa" pagination="1-84" refId="B43" refString="Uliana M, Bonato L, Minelli A (2007) The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha). Zootaxa 1396 (1): 184. https://doi.org/10.11646/zootaxa.1396.1.1" title="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)." volume="1396" year="2007">Uliana et al. (2007)</bibRefCitation>
, and the present study. Trunk segments without dark patches; head 1.21.4 times as long as wide (Fig.
<figureCitation id="A3EE656B9381372BF6C48A8A65B49FD8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicellophilus pulcher (TS 20210504-01). A. Cephalic plate, dorsal; B. Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure14" httpUri="https://binary.pensoft.net/fig/1073621">14 A, B</figureCitation>
); lateral margin of cephalic plate abruptly converged posteriorly; clypeus with densely scattered setae (Fig.
<figureCitation id="B5D45A393A1EC9CDF2026B40512C6103" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicellophilus pulcher (TS 20210504-01). A. Cephalic plate, dorsal; B. Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure14" httpUri="https://binary.pensoft.net/fig/1073621">14 B</figureCitation>
); paraclypeal suture evidently converging posteriorly (Fig.
<figureCitation id="ADCDCABD415673FBEA7F31AA404757B3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicellophilus pulcher (TS 20210504-01). A. Cephalic plate, dorsal; B. Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure14" httpUri="https://binary.pensoft.net/fig/1073621">14 B</figureCitation>
); both ends of transverse suture convexed forward (Fig.
<figureCitation id="4FBD81F5CBB9A853180DF4EC3AE31F3F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicellophilus pulcher (TS 20210504-01). A. Cephalic plate, dorsal; B. Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure14" httpUri="https://binary.pensoft.net/fig/1073621">14 A</figureCitation>
); mandible with 57 lamellae; forcipular trochanteroprefemur almost as long as wide, with one small distal denticle (Fig.
<figureCitation id="D5FB364727320BC1274F0C23B75EF0C0" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicellophilus pulcher (TS 20210504-01). A. Forcipular segment, dorsal; B. Forcipular segment, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure15" httpUri="https://binary.pensoft.net/fig/1073622">15 A, B</figureCitation>
); forcipular tarsungulum with evident and variably spaced notches; metatergite subtrapezoidal (Fig.
<figureCitation id="E54CA4D94719F3A675395F852050D79A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicellophilus pulcher (TS 20210504-01). A. Ultimate leg-bearing segment and postpedal segment, dorsal; B. Ultimate leg-bearing segment and postpedal segment, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure16" httpUri="https://binary.pensoft.net/fig/1073623">16 A</figureCitation>
); metasternite trapezoidal, longer than wide (Fig.
<figureCitation id="103325EFA2729E71C3F65D9B81856D6A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicellophilus pulcher (TS 20210504-01). A. Ultimate leg-bearing segment and postpedal segment, dorsal; B. Ultimate leg-bearing segment and postpedal segment, ventral. Scale bars: 0.5 mm." figureDoi="10.3897/zse.100.121512.figure16" httpUri="https://binary.pensoft.net/fig/1073623">16 B</figureCitation>
); forty-one pairs of legs.
</paragraph>
<caption id="7FFE185B869858D5F488CC86E2515D45" ID-DOI="10.3897/zse.100.121512.figure14" ID-arpha="945CACD8-AA65-578F-8992-DE005124B95C" httpUri="https://binary.pensoft.net/fig/1073621" startId="F14">
<paragraph id="6BF7FCFC3925220A24F4DBF6029ADB60">
<label id="43F16098ABA4240AD0768CEA13C18C0B">Figure 14.</label>
</paragraph>
<paragraph id="7F633454903D43C8371BC897DF5CED97">
<taxonomicName id="862DC29A1BA126AEE7870540AE6041F7" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="AE3F1203F796ABA3D2C151AF37E00E75" italics="true">Dicellophilus pulcher</emphasis>
</taxonomicName>
(
<specimenCode id="CF4223BBD9E493DB59CD5BE7A376AA95">TS 20210504-01</specimenCode>
).
<emphasis id="50E09CAF157F8CEAEC2BC133874D2BB6" bold="true">A.</emphasis>
Cephalic plate, dorsal;
<emphasis id="EDC2FAACB2319FDD20C96F3CF5F0C054" bold="true">B.</emphasis>
Clypeus, and clypeal pleurite, ventral. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="8274C7BB6F04EC5D9FE79A6BDCA9622B" ID-DOI="10.3897/zse.100.121512.figure15" ID-arpha="3BE4AEAC-3446-5CA8-BEF0-4CCD09E41794" httpUri="https://binary.pensoft.net/fig/1073622" startId="F15">
<paragraph id="5DDBEBD8643EAB8A55D08AA3BA65CE51">
<label id="4E1BA6F4E37AFBAC545814BD68C95F2A">Figure 15.</label>
</paragraph>
<paragraph id="44C45E227F649224F3B1D519D51CDE19">
<taxonomicName id="4646A92901A735D3B0F840ACE0885F08" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="73EF23B0F66DEAE540B31BA4E2CA07E7" italics="true">Dicellophilus pulcher</emphasis>
</taxonomicName>
(
<specimenCode id="3D61F86D0B569DE4FBC7EEA0D6F8D564">TS 20210504-01</specimenCode>
).
<emphasis id="7B4CE5C8C16CC2489A63BEDFECC722F9" bold="true">A.</emphasis>
Forcipular segment, dorsal;
<emphasis id="8F94E43D15ABEF147C83C7E99223BB7B" bold="true">B.</emphasis>
Forcipular segment, ventral. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="492621AE5997AC1B22B5573F16DBC32F" ID-DOI="10.3897/zse.100.121512.figure16" ID-arpha="4F616965-F741-5640-8257-AF17CAA056E8" httpUri="https://binary.pensoft.net/fig/1073623" startId="F16">
<paragraph id="E1975FBF85D8A70448372AF181DD2922">
<label id="8AC369C429B827B953EAA57A940B1DA3">Figure 16.</label>
</paragraph>
<paragraph id="7EFF665C6C8849D6D1197E69BF389B1C">
<taxonomicName id="9F81CD05A3F67A4DAF7D5D3C10685335" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="069FF69BFB53E90CA1857173E95A95CF" italics="true">Dicellophilus pulcher</emphasis>
</taxonomicName>
(
<specimenCode id="676A2CB4B8B049426C9A2A6FD47F073E">TS 20210504-01</specimenCode>
).
<emphasis id="252086FF41878BD685CEBED9E7E24495" bold="true">A.</emphasis>
Ultimate leg-bearing segment and postpedal segment, dorsal;
<emphasis id="ABF2A0AAC76CC9B8264E04751CAFF195" bold="true">B.</emphasis>
Ultimate leg-bearing segment and postpedal segment, ventral. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIKBI" type="type locality">
<paragraph id="266DDBA4D44B2002C7904D08347C3831">
<heading id="6B4591DDCFD679686403AAB03550F57D" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="52838EC2CF939818DAA7EFD0A668C473">
The first section of the Subashiri trail of Mt. Fuji,
<collectingRegion id="C53D211ADC30F0525DECB3AD4E00C66D" country="Japan" name="Shizuoka">Shizuoka Pref.</collectingRegion>
,
<collectingCountry id="460258FC10EDC3526D3DABE801930F9E" name="Japan">Japan</collectingCountry>
(
<bibRefCitation id="2E0BDB1B71B29517A3E5CEBE6C4DC6A9" author="Kishida" editor="Sengen Shrine" firstAuthor="Kishida" journalOrPublisher="Fuji. Kokinshoin, Tokyo" pagination="290-303" refId="B20" refString="Kishida K (1928) Ch. 6. Chilopoda. In: Sengen Shrine (Ed.) Animals of Mt. Fuji. Kokinshoin, Tokyo, 290303. [in Japanese]" title="Ch. 6. Chilopoda." volumeTitle="Animals of Mt." year="1928">Kishida 1928</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESKBI" type="distribution">
<paragraph id="9133C92D05C9D770CF874EBB9469E9BF">
<heading id="8676D59EE7E04F00F4FAF2DE678C0786" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="6EFFED9144E2EEA55F14481FDD4258AA">
Honshu (
<collectingRegion id="8B69E1C532A63A134D69956EDD47C798" country="Japan" name="Fukushima">Fukushima Pref.</collectingRegion>
to
<collectingRegion id="6B8C77E1240E61113A5D0A5B1D2B2D80" country="Japan" name="Hyogo">Hyogo Pref.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXKBI" type="remarks">
<paragraph id="DBA1D417CA3E5CC3DDBE09B35E34DDAD">
<heading id="5E5BBCDFDE79C1BBC701906438D19F58" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="BB6B1EA9548DCB3842EDD1AFEB3A3FD7">
See remarks and the diagnosis of
<taxonomicName id="F759A3E4CB88C8743284F93CCC79C66F" authority="Tsukamoto &amp; Eguchi, 2024" authorityName="Tsukamoto &amp; Eguchi" authorityYear="2024" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="praetermissus" status="sp. nov.">
<emphasis id="2886119253C33A504CB38534496D901B" italics="true">D. praetermissus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="250A019631B301ACDD8930CF13AF12ED" rank="species">sp. nov.</taxonomicNameLabel>
for confirming how to distinguish
<taxonomicName id="07D82136BF7C8FCCD5D4295AA0FE7DE1" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="0BDC58F877746682A3F1D52E06806D5D" italics="true">D. pulcher</emphasis>
</taxonomicName>
from
<taxonomicName id="F5736FC12B5D51803F3C101F7B6AE6A9" authority="Tsukamoto &amp; Eguchi, 2024" authorityName="Tsukamoto &amp; Eguchi" authorityYear="2024" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="praetermissus" status="sp. nov.">
<emphasis id="60CC26D7EF94F0F2517FCDC5DB667270" italics="true">D. praetermissus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1B82441214C2904FAC6B9A2B39A8AF7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="52567B9572580E9BF13BFCDAE4A65DBC">
There are three junior synonyms under
<taxonomicName id="4ED98AB712EDA863826185FA55D97ABE" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="6A12A8AE2DF69DBA569007B1466F1E21" italics="true">D. pulcher</emphasis>
</taxonomicName>
, which were synonymized by previous authors based on morphological examination (
<bibRefCitation id="4FC795F255907F849A495B08AB27D1BE" author="Takakuwa" firstAuthor="Takakuwa" journalOrPublisher="Sanseido, Tokyo" refId="B36" refString="Takakuwa Y (1940) Fauna Nipponica 9 (8, 1). Geophilomorpha. Sanseido, Tokyo, 156 pp. [in Japanese]" title="Fauna Nipponica 9 (8, 1). Geophilomorpha." year="1940">Takakuwa 1940</bibRefCitation>
;
<bibRefCitation id="6F4B4FC94122A5C5E622CCB28028E028" author="Shinohara" firstAuthor="Shinohara" journalOrPublisher="Takakuwaia" pagination="5-6" refId="B30" refString="Shinohara K (1983) Revision on the scientific names of Japanese myriapods. V. On the Dicellophilus pulcher (Kishida). Takakuwaia 15: 56." title="Revision on the scientific names of Japanese myriapods. V. On the Dicellophilus pulcher (Kishida)." volume="15" year="1983">Shinohara 1983</bibRefCitation>
;
<bibRefCitation id="B5878A133D49B3BD0492BDA4B0E18B98" DOI="10.11646/zootaxa.1396.1.1" author="Uliana" etAl="et al." firstAuthor="Uliana" issue="1" journalOrPublisher="Zootaxa" pagination="1-84" refId="B43" refString="Uliana M, Bonato L, Minelli A (2007) The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha). Zootaxa 1396 (1): 184. https://doi.org/10.11646/zootaxa.1396.1.1" title="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)." volume="1396" year="2007">Uliana et al. 2007</bibRefCitation>
):
<taxonomicName id="36906D0D9634CA867408320609DF4D72" authority="Takakuwa, 1934" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="64882BD99566BB67964CF14ABF4A5F59" italics="true">D. latifrons</emphasis>
Takakuwa, 1934
</taxonomicName>
;
<taxonomicName id="8B2227EB995C0D5C912A6051A36BF2FC" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="03C171B97F4CF5636549D5AFE5ADD44A" italics="true">D. japonicus</emphasis>
<bibRefCitation id="F608A0538CFA54EFA943B6253019C63C" author="Verhoeff KW" journalOrPublisher="Abteilung für Systematik" pagination="1-112" refId="B44" refString="Verhoeff KW (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher. Abteilung fur Systematik 66: 1 - 112." title="Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher." volume="66" year="1934">Verhoeff, 1934</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E213EBFABFEC279E0D0F96B26FAD821F" authority="Shinohara, 1961" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="A424B476CBDB01C3014D7E817D82E220" italics="true">Tygarrup monoporus</emphasis>
<bibRefCitation id="A1C20A7B12B4D5ED90923C1CF400A985" author="Shinohara K" issue="7" journalOrPublisher="Zoological Magazine" pagination="212-216" refId="B29" refString="Shinohara K (1961) Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda). Zoological Magazine 70 (7): 212 - 216. [in Japanese with English Resume]" title="Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda)." volume="70" year="1961">Shinohara, 1961</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="741B1913CDA2E1C1B5D73F8025D6F3A6" authority="Takakuwa, 1934" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="C24C63206EAF0292CE04D09B024CB79F" italics="true">Dicellophilus latifrons</emphasis>
Takakuwa, 1934
</taxonomicName>
, which was described in a key to Japanese and Taiwanese species of
<taxonomicName id="E77FDDB8648B98DF2603BD1692638F5D" authorityName="Bollman" authorityYear="1893" class="Chilopoda" family="Mecistocephalidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Mecistocephalidae</taxonomicName>
by
<bibRefCitation id="1A62803DA426DE0C496882D1E9F2A396" author="Takakuwa Y" issue="4" journalOrPublisher="Botany and Zoology" pagination="706-712" refId="B33" refString="Takakuwa Y (1934 a) The family Mecistocephalidae of Japan, I. Botany and Zoology 2 (4): 706712. [in Japanese]" title="The family Mecistocephalidae of Japan, I." volume="2">Takakuwa (1934 a</bibRefCitation>
), was later described by
<bibRefCitation id="99963AB2D0ADB3A3BC784DFAC7350006" author="Takakuwa Y" journalOrPublisher="Annotationes Zoologicae Japonenses" pagination="355-363" refId="B34" refString="Takakuwa Y (1934 b) Neue japanische Mecistocephalidae. Annotationes Zoologicae Japonenses 14: 355363." title="Neue japanische Mecistocephalidae." volume="14">Takakuwa (1934 b</bibRefCitation>
,
<bibRefCitation id="7C63F89845906909D6F6F2B092C6AE9E" author="Takakuwa Y" issue="5" journalOrPublisher="Botany and Zoology" pagination="878-884" refId="B35" refString="Takakuwa Y (1934 c) The family Mecistocephalidae of Japan, II. Botany and Zoology 2 (5): 878884. [in Japanese]" title="The family Mecistocephalidae of Japan, II." volume="2">c</bibRefCitation>
) as a new species.
<bibRefCitation id="4B4978BF60E7F2AFA035C3CBCDC63475" author="Takakuwa Y" issue="4" journalOrPublisher="Botany and Zoology" pagination="706-712" refId="B33" refString="Takakuwa Y (1934 a) The family Mecistocephalidae of Japan, I. Botany and Zoology 2 (4): 706712. [in Japanese]" title="The family Mecistocephalidae of Japan, I." volume="2">Takakuwa (1934 a</bibRefCitation>
) did not designate a type locality for
<taxonomicName id="73F05B047B932D62BD3C62259BE92B6E" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="E60EC3943D660959562232E390449744" italics="true">D. latifrons</emphasis>
</taxonomicName>
, and
<bibRefCitation id="C8C7B3F7D3F0C186DD5EAA60613535C0" author="Takakuwa Y" journalOrPublisher="Annotationes Zoologicae Japonenses" pagination="355-363" refId="B34" refString="Takakuwa Y (1934 b) Neue japanische Mecistocephalidae. Annotationes Zoologicae Japonenses 14: 355363." title="Neue japanische Mecistocephalidae." volume="14">Takakuwa (1934 b</bibRefCitation>
,
<bibRefCitation id="3DD0A7DC066988A188220CDC3B552105" author="Takakuwa Y" issue="5" journalOrPublisher="Botany and Zoology" pagination="878-884" refId="B35" refString="Takakuwa Y (1934 c) The family Mecistocephalidae of Japan, II. Botany and Zoology 2 (5): 878884. [in Japanese]" title="The family Mecistocephalidae of Japan, II." volume="2">c</bibRefCitation>
) listed the localities: “ Kaibara (
<collectingRegion id="06DB6326D8CEE92E44FA9FEC48FFE7E9" country="Japan" name="Hyogo">Hyogo</collectingRegion>
) ” (= Tamba City,
<collectingRegion id="B751C1AB5926D35ED45EA1B248C38891" country="Japan" name="Hyogo">Hyogo Pref.</collectingRegion>
), “ Masudo (bei
<collectingRegion id="FC8D45FCC3955F4EE97CC1663427059F" country="Japan" name="Tokyo">Tokyo</collectingRegion>
) ” (possibly misread of Masuko-mura, currently in Akiruno-shi,
<collectingRegion id="D16B27189CCBA1D9A07A3389E934E31D" country="Japan" name="Tokyo">Tokyo Pref.</collectingRegion>
), “ Komono (Miye) ” (= Komono-cho,
<collectingRegion id="1632D9D881D08422AB09ED82CFD3730A" country="Japan" name="Mie">Mie Pref.</collectingRegion>
), “ Ikao (Gumma) ” (= Ikahocho, Shibukawa-shi,
<collectingRegion id="969AB1623F6A612848836013FA559AED" country="Japan" name="Gunma">Gunma Pref.</collectingRegion>
), “ Ōta (Gumma) ” (= Ota-shi,
<collectingRegion id="18A5B249FBD3603E1A79EAE25A8B66BE" country="Japan" name="Gunma">Gunma Pref.</collectingRegion>
), “ Odawara (
<collectingRegion id="2296BABBA7B14A95F2326FE5D8967FA2" country="Japan" name="Kanagawa">Kanagawa</collectingRegion>
) ” and “ Suwa (
<collectingRegion id="A3AE44544CD3939E6CE2A29E92FEF0F0" country="Japan" name="Nagano">Nagano</collectingRegion>
) ” (annotated by
<bibRefCitation id="075858787E857B2831B4C9AD38ACB69C" DOI="10.12782/specdiv.27.71" author="Jonishi and Nakano" firstAuthor="Jonishi" issue="1" journalOrPublisher="Species Diversity : An International Journal for Taxonomy, Systematics, Speciation, Biogeography, and Life History Research of Animals" pagination="71-81" refId="B17" refString="Jonishi T, Nakano T (2022) Correct Authorships, Synonymies, and Remarks on the Type Series of Fourteen Names of Centipedes Introduced by Yoshioki Takakuwa in 1934 and Mecistocephalus takakuwai (Chilopoda: Geophilomorpha and Scolopendromorpha). Species Diversity : An International Journal for Taxonomy, Systematics, Speciation, Biogeography, and Life History Research of Animals 27 (1): 7181. https://doi.org/10.12782/specdiv.27.71" title="Correct Authorships, Synonymies, and Remarks on the Type Series of Fourteen Names of Centipedes Introduced by Yoshioki Takakuwa in 1934 and Mecistocephalus takakuwai (Chilopoda: Geophilomorpha and Scolopendromorpha)." volume="27" year="2022">Jonishi and Nakano 2022</bibRefCitation>
). Considering the geographic distribution of
<taxonomicName id="45FD755C44CFCD78D8A846D99588C199" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="077D5AA9B842B6EFDD59FCD7A7B01AD2" italics="true">D. latifrons</emphasis>
</taxonomicName>
and
<taxonomicName id="3C60892B333A7E4F86CCCAC3B7A93D0A" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="AE56C117D40EB4A90177E1060F54015D" italics="true">D. pulcher</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40878AD92F12C4FA3F0C44F8AD5D6398" sensu="stricto">sensu stricto</taxonomicNameLabel>
,
<taxonomicName id="2C6A72658FCAE6A4CACCFB0B868D4531" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="43738C605571938BE394A01A58E38585" italics="true">D. latifrons</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="98FC7C1007B996E2D11680942674DDE4" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="1F9C5A13F51A4E18224D5FE0112A7BD8" italics="true">D. pulcher</emphasis>
</taxonomicName>
.
<taxonomicName id="15DECE76DC31E5D09E5643E4C6C30739" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="195718A593F208A70FFE61AED2D21EAD" italics="true">Dicellophilus japonicus</emphasis>
<bibRefCitation id="7B0BE506C3EB78985292F6F8FDE22E58" author="Verhoeff KW" journalOrPublisher="Abteilung für Systematik" pagination="1-112" refId="B44" refString="Verhoeff KW (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher. Abteilung fur Systematik 66: 1 - 112." title="Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher." volume="66" year="1934">Verhoeff, 1934</bibRefCitation>
</taxonomicName>
, was described based on a specimen from “
<collectingRegion id="9EAEC1CD2A650B8913B2ACE71825473A" country="Japan" name="Tokyo">Tokyo</collectingRegion>
” (
<bibRefCitation id="D73771402CD1DFC279D8C7B2D5DAE4FB" author="Verhoeff" firstAuthor="Verhoeff" journalOrPublisher="Abteilung für Systematik" pagination="1-112" refId="B44" refString="Verhoeff KW (1934) Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher. Abteilung für Systematik 66: 1112." title="Beiträge zur Systematik und Geographie der Chilopoden. Zoologische Jahrbücher." volume="66" year="1934">Verhoeff 1934</bibRefCitation>
) and later regarded as a junior synonym of
<taxonomicName id="C12FC78C2177B3A23BB2155681CBD5BA" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="09D86FF1005EDD4165DA8FFF8B944F6E" italics="true">D. latifrons</emphasis>
</taxonomicName>
based on the comparison of diagnostic characteristics (
<bibRefCitation id="40EF06E07906F82C188AB354EE517FBF" author="Takakuwa" firstAuthor="Takakuwa" journalOrPublisher="Sanseido, Tokyo" refId="B36" refString="Takakuwa Y (1940) Fauna Nipponica 9 (8, 1). Geophilomorpha. Sanseido, Tokyo, 156 pp. [in Japanese]" title="Fauna Nipponica 9 (8, 1). Geophilomorpha." year="1940">Takakuwa 1940</bibRefCitation>
;
<bibRefCitation id="9505C030EA272D9CD60DF20AF3A81933" author="Shinohara" firstAuthor="Shinohara" journalOrPublisher="Takakuwaia" pagination="5-6" refId="B30" refString="Shinohara K (1983) Revision on the scientific names of Japanese myriapods. V. On the Dicellophilus pulcher (Kishida). Takakuwaia 15: 56." title="Revision on the scientific names of Japanese myriapods. V. On the Dicellophilus pulcher (Kishida)." volume="15" year="1983">Shinohara 1983</bibRefCitation>
). Considering the geographic distribution of
<taxonomicName id="4EF371F8153B202805A7411373825157" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="092EA8C7DA23D5A01B94F4CE04E97528" italics="true">D. japonicus</emphasis>
</taxonomicName>
and
<taxonomicName id="96C703610F0E1DAA30B228F5396E442A" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="92AB5056A3C26703E73A8C3957A655E1" italics="true">D. pulcher</emphasis>
</taxonomicName>
and the phylogenetic analyses of the present study, including
<specimenCode id="95120821E28A4C0D2243BB98F569168F">TS 20181214-01</specimenCode>
,
<specimenCode id="67E7A8B5623B2F087522D3BA00CDFB85">TS 20191006-02</specimenCode>
,
<specimenCode id="F7788CDE1C936123220A07AD75CE8AC9">TS 20210401-03</specimenCode>
, and
<specimenCode id="2D41CF2BE4F85C2E4481732F049D5B5D">TS 20210819-01</specimenCode>
from
<collectingRegion id="A213FDB0C1AA1709A07B685A26AF134F" country="Japan" name="Tokyo">Tokyo Pref.</collectingRegion>
, it is not conflicting that
<taxonomicName id="3816D54576D730D2A526264E6E6F57D1" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B7F957A466CFC6372736B9F562825656" italics="true">D. japonicus</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="0B84C6E93DE82A72F87B9901E802941E" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="733CE00E966A53ADE3303024F2A4F235" italics="true">D. pulcher</emphasis>
</taxonomicName>
.
<taxonomicName id="B5FE6321230894EC27905EB7615E8884" authority="Shinohara, 1961" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="EF2B2F8358AA28DC9A3E9A0F515F8CE6" italics="true">Tygarrup monoporus</emphasis>
<bibRefCitation id="E96933E895612BB0FFF46C1103F474B8" author="Shinohara K" issue="7" journalOrPublisher="Zoological Magazine" pagination="212-216" refId="B29" refString="Shinohara K (1961) Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda). Zoological Magazine 70 (7): 212 - 216. [in Japanese with English Resume]" title="Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda)." volume="70" year="1961">Shinohara, 1961</bibRefCitation>
</taxonomicName>
, which is identical to the juvenile of
<taxonomicName id="228E8983B9E262A955A29D8DE59A1A0F" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="CC1F09E77C9987DFBBD8EE090230E59B" italics="true">D. pulcher</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="77886595924D44BD0730AD44436819BC" DOI="10.11646/zootaxa.1396.1.1" author="Uliana M &amp; Bonato L &amp; Minelli A" etAl="et al." firstAuthor="Uliana" issue="1" journalOrPublisher="Zootaxa" pagination="1-84" refId="B43" refString="Uliana M, Bonato L, Minelli A (2007) The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha). Zootaxa 1396 (1): 184. https://doi.org/10.11646/zootaxa.1396.1.1" title="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)." volume="1396" year="2007">Uliana et al. (2007)</bibRefCitation>
, was described based on the specimen from Manazuru-machi, Ashigarashimo-gun,
<collectingRegion id="72CF20C73A3171ED2AD8E74C3A7D85BB" country="Japan" name="Kanagawa">Kanagawa Pref.</collectingRegion>
(
<bibRefCitation id="00E96A585A4AA373F84C439D40377976" author="Shinohara" firstAuthor="Shinohara" issue="7" journalOrPublisher="Zoological Magazine" pagination="212-216" refId="B29" refString="Shinohara K (1961) Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda). Zoological Magazine 70 (7): 212216. [in Japanese with English Resume]" title="Taxonomical and morphological studies of Myriapoda VII. Two new species of Mecistocephalidae (Chilopoda)." volume="70" year="1961">Shinohara 1961</bibRefCitation>
). Considering the geographic distribution of
<taxonomicName id="EE5BAFA428ADE2292E2534D474B11211" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="A56CAA1884BA115B5F4A121B0DED9D62" italics="true">T. monoporus</emphasis>
</taxonomicName>
and
<taxonomicName id="D9696FCC2A884D64B764766ADE04DACA" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="8682E7ACAEC7C406A630C6B415B2A5EF" italics="true">D. pulcher</emphasis>
</taxonomicName>
and the phylogenetic analyses of the present study, including
<specimenCode id="C20A90A8A1B8182BBE59C35149F01DB2">TS 20210728-02</specimenCode>
, which was collected at a linear distance of approximately
<quantity id="B74564AC86E34B4381286FED194BA9D1" metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
from the type locality and could be identified as
<taxonomicName id="13FF7E4D8DF37C6C17A780B1B0F266B3" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="4196EFD66889A9BABC3F0D632CAE8A54" italics="true">T. monoporus</emphasis>
</taxonomicName>
,
<taxonomicName id="D8901F6D9EFF1A0A83EF3041E3C30768" authorityName="Shinohara" authorityYear="1961" class="Chilopoda" family="Mecistocephalidae" genus="Tygarrup" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="monoporus">
<emphasis id="A9C6C76985227C70DF32D2941CBCE453" italics="true">T. monoporus</emphasis>
</taxonomicName>
can also be regarded as a junior synonym of
<taxonomicName id="078DD95EF87018CD8C49D6B9564D3DAA" baseAuthorityName="Kishida" baseAuthorityYear="1928" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="812E02D501EC304E09806481EE91A82B" italics="true">D. pulcher</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>