212 lines
22 KiB
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212 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.79.57686" ID-GBIF-Dataset="054f2212-802b-42b3-bba6-e8bca2b22198" ID-Pensoft-Pub="1314-2607-79-77" ID-Pensoft-UUID="9771F5D9B2AA51558A0A56AC96840EC0" ID-Zenodo-Dep="4255453" ID-ZooBank="ED8D2D07C0384429B5FF1011D2B48907" ModsDocID="1314-2607-79-77" checkinTime="1604097865600" checkinUser="pensoft" docAuthor="Martins, Andre L. & Melo, Gabriel A. R." docDate="2020" docId="FAC40A926FFE5CE78F3CA94F5FC98304" docLanguage="en" docName="JourHymenoptRes 79: 77-88" docOrigin="Journal of Hymenoptera Research 79" docPubDate="2020-10-30" docSource="http://dx.doi.org/10.3897/jhr.79.57686" docTitle="Thaumatodryinus fuscescens Martins & Melo 2020, sp. nov." docType="treatment" docUuid="48DD7867-AB75-4443-9346-B7DF4A35C982" docUuidSource="ZooBank" docVersion="4" id="9771F5D9B2AA51558A0A56AC96840EC0" lastPageNumber="77" masterDocId="9771F5D9B2AA51558A0A56AC96840EC0" masterDocTitle="Revision of the fossil species of Thaumatodryinus Perkins from Dominican amber, with a new combination and description of a new species (Hymenoptera, Dryinidae)" masterLastPageNumber="88" masterPageNumber="77" pageNumber="77" updateTime="1678754720945" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the fossil species of Thaumatodryinus Perkins from Dominican amber, with a new combination and description of a new species (Hymenoptera, Dryinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martins, Andre L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4794-0644</mods:nameIdentifier>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Cx. postal 19020, 81531 - 980, Curitiba, PR, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Melo, Gabriel A. R.</mods:namePart>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Cx. postal 19020, 81531 - 980, Curitiba, PR, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">garmelo@ufpr.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-10-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>88</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.79.57686</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.79.57686</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-79-77</mods:identifier>
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<mods:identifier type="ZooBank">ED8D2D07C0384429B5FF1011D2B48907</mods:identifier>
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<mods:identifier type="Pensoft-UUID">9771F5D9B2AA51558A0A56AC96840EC0</mods:identifier>
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<treatment ID-GBIF-Taxon="168745941" LSID="urn:lsid:zoobank.org:act:48DD7867-AB75-4443-9346-B7DF4A35C982" httpUri="http://treatment.plazi.org/id/FAC40A926FFE5CE78F3CA94F5FC98304" lastPageNumber="77" pageId="0" pageNumber="77">
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<subSubSection pageId="0" pageNumber="77" type="nomenclature">
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<paragraph pageId="0" pageNumber="77">
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†
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<taxonomicName LSID="http://zoobank.org/48DD7867-AB75-4443-9346-B7DF4A35C982" authority="Martins & Melo, 2020" authorityName="Martins & Melo" authorityYear="2020" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus fuscescens" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="fuscescens" status="sp. nov.">Thaumatodryinus fuscescens</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="77">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="description">
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<paragraph pageId="0" pageNumber="77">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="diagnosis">
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<paragraph pageId="0" pageNumber="77">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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<taxonomicName authorityName="Martins & Melo" authorityYear="2020" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus fuscescens" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="fuscescens">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus fuscescens</emphasis>
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</taxonomicName>
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sp. nov. is characterized by the body predominantly testaceous, except head and part of pronotum and remainder of mesosoma black; eyes hemispherical, about 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide; maximum head width about 1.9
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<normalizedToken originalValue="×">x</normalizedToken>
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distance between inner margins of tegulae; frontal line present; mid portion of clypeus with anterior margin straight; antenna with 1st flagellomere as long as 2nd; POL 3.2
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<normalizedToken originalValue="×">x</normalizedToken>
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OL; OOL about 6
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<normalizedToken originalValue="×">x</normalizedToken>
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OPL; occipital carina complete; notauli percurrent, extending from anterior to posterior margins of mesoscutum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="description">
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<paragraph pageId="0" pageNumber="77">Description.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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Female holotype (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1A-F</figureCitation>
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). Approximate body length: 4.1 mm.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="77">Color</emphasis>
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. Head black, except mandible, clypeus and antenna testaceous (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1A-D</figureCitation>
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); pronotum black, except dorsal-posterior surface brown (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1B</figureCitation>
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); mesosoma black; legs testaceous, except pro- and metafemur with middle portion darkened (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1</figureCitation>
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); fore wing with one transverse darkened band; metasoma testaceous.
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<emphasis bold="true" pageId="0" pageNumber="77">
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<emphasis italics="true" pageId="0" pageNumber="77">Pubescence</emphasis>
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.
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</emphasis>
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Head with short setae (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1A-C</figureCitation>
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); antenna with setae; eye glabrous (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1C</figureCitation>
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); pronotum with fine and erect setae (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1B</figureCitation>
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); mesosoma with conspicuous erect setae (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1B</figureCitation>
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); propodeum with erect setae; metasoma with more conspicuous erect pilosity on last tergum.
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<emphasis bold="true" pageId="0" pageNumber="77">
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<taxonomicName genus="Integumental" lsidName="Integumental sculpture" pageId="0" pageNumber="77" rank="species" species="sculpture">
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<emphasis italics="true" pageId="0" pageNumber="77">Integumental sculpture</emphasis>
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</taxonomicName>
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.
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</emphasis>
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Head finely granulated (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1A-C</figureCitation>
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); frontal line in the form of a low carina and apparently complete, not visible near mid ocellus; malar sulcus well-marked; mandible with a marked groove at the base; pronotum granulate, except by rugosity on lateral portions of transverse impression; mesoscutum granulate; notauli complete; mesoscutellum rugulose, its anterior margin with a narrow row of small foveae, not with a groove; mesopleuron rugose; metapleuron reticulate-rugose; propodeum reticulate rugose.
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<emphasis bold="true" pageId="0" pageNumber="77">
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<emphasis italics="true" pageId="0" pageNumber="77">Structure and proportions</emphasis>
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.
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</emphasis>
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Vertex convex, except by ocellar triangle slightly elevated. Ocellar ratio: POL = 16; OL = 5; OOL = 25; OPL = 4; TL = 9. Antennomeres in following proportions: 35: 15: 45: 45: 60: 65: 55: 23: 27: 25. Eye hemispherical, about 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide; maximum head width about 1.9
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<normalizedToken originalValue="×">x</normalizedToken>
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distance between inner margins of tegulae; occipital carina complete; mid portion of clypeus with anterior margin straight; maxillary palpomeres elongated; malar space about as long as basal width of mandible; notaulus complete (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1B</figureCitation>
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). Head longer than pronotum (60:55); mesoscutum longer than mesoscutellum and metanotum shorter than propodeum. Fore wing with three cells closed by pigmented tubular veins; vein abscissa 3Rs&4Rs longer than 2r-rs (65:30). Dorsal surface of propodeum distinctly longer than posterior surface (50:45); mesopleuron with a groove in the central region. Articles of fore leg (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1E-F</figureCitation>
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) in the following proportions: 54 (coxa): 50 (trochanter): 63 (femur): 80 (tibia), 46: 8: 15: 34: 50 (tarsomeres 1-5), and 46 (enlarged claw). Segment 3 of protarsus with four long bristles and segment 4 with five long bristles. Chela (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Thaumatodryinus fuscescens sp. nov., female holotype. A habitus, dorsal view B right antenna, dorsal view C head and mesosoma, antero-ventral view D head and mesosoma, dorsal view E apical portion of flagellum, with details of flagellomeres 5 - 8 F-G Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (A), 0.5 mm (B), 0.5 mm (C-D), 0.3 mm (E-G)." figureDoi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77">1E-F</figureCitation>
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) and enlarged claw with two subapical teeth and one row of about 26 lamellae. Segment 5 of protarsus with two rows of at least 40 lamellae; distal apex with group of approximately seven long lamellae.
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</paragraph>
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<caption ID-Zenodo-Dep="4255455" doi="10.3897/jhr.79.57686.figure1" httpUri="https://binary.pensoft.net/fig/469490" pageId="0" pageNumber="77" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="77">
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<emphasis bold="true" pageId="0" pageNumber="77">Figure 1.</emphasis>
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†
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<taxonomicName authorityName="Martins & Melo" authorityYear="2020" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus fuscescens" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="fuscescens">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus fuscescens</emphasis>
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</taxonomicName>
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sp. nov., female holotype.
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<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="77">B</emphasis>
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right antenna, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="77">C</emphasis>
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head and mesosoma, antero-ventral view
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<emphasis bold="true" pageId="0" pageNumber="77">D</emphasis>
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head and mesosoma, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="77">E</emphasis>
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apical portion of flagellum, with details of flagellomeres 5-8
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<emphasis bold="true" pageId="0" pageNumber="77">F-G</emphasis>
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Apical portion of fore leg, with details of the chela. Scale bars: 2 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
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), 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">B</emphasis>
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), 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">C-D</emphasis>
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), 0.3 mm (
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<emphasis bold="true" pageId="0" pageNumber="77">E-G</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="remarks">
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<paragraph pageId="0" pageNumber="77">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="77">
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<taxonomicName authorityName="Martins & Melo" authorityYear="2020" class="Insecta" family="Dryinidae" genus="Thaumatodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thaumatodryinus fuscescens" order="Hymenoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="fuscescens">
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<emphasis italics="true" pageId="0" pageNumber="77">Thaumatodryinus fuscescens</emphasis>
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</taxonomicName>
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sp. nov. is more similar to the extant Neotropical fauna than to the other two species found in the Dominican amber. It has in common with some of the living species a convex vertex and slightly elevated ocellar triangle, straight lower clypeal margin, complete occipital carina, percurrent notauli and anterior margin of mesoscutellum with a narrow row of small foveae. Among the fossil species studied here,
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<taxonomicName lsidName="T. fuscescens" pageId="0" pageNumber="77" rank="species" species="fuscescens">
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<emphasis italics="true" pageId="0" pageNumber="77">T. fuscescens</emphasis>
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</taxonomicName>
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sp. nov. is more similar to
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<taxonomicName lsidName="T. miocenicus" pageId="0" pageNumber="77" rank="species" species="miocenicus">
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<emphasis italics="true" pageId="0" pageNumber="77">T. miocenicus</emphasis>
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</taxonomicName>
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and differs from
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<taxonomicName lsidName="T. priscus" pageId="0" pageNumber="77" rank="species" species="priscus">
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<emphasis italics="true" pageId="0" pageNumber="77">T. priscus</emphasis>
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</taxonomicName>
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by the head shape with eyes protuberant on the lateral side, frontal line present and ratio of OL short than TL. The new species differ from
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<taxonomicName lsidName="T. miocenicus" pageId="0" pageNumber="77" rank="species" species="miocenicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="77">T. miocenicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="T. priscus" pageId="0" pageNumber="77" rank="species" species="priscus">
|
||
<emphasis italics="true" pageId="0" pageNumber="77">T. priscus</emphasis>
|
||
</taxonomicName>
|
||
by the granulated integument of the head; clypeus with lower margin straight; 1st flagellomere as long as 2nd; hemispherical-shaped eyes; and percurrent notauli. The chela of
|
||
<taxonomicName lsidName="T. fuscescens" pageId="0" pageNumber="77" rank="species" species="fuscescens">
|
||
<emphasis italics="true" pageId="0" pageNumber="77">T. fuscescens</emphasis>
|
||
</taxonomicName>
|
||
is closed and for this reason it is not possible to determine the number of lamellae in the enlarged claw and in the 5th tarsomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="77" type="etimology">
|
||
<paragraph pageId="0" pageNumber="77">Etimology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="77">
|
||
The species is named in reference to its overall dark coloration, from the Latin
|
||
<emphasis italics="true" pageId="0" pageNumber="77">fuscus</emphasis>
|
||
, dark, dusky, and -
|
||
<emphasis italics="true" pageId="0" pageNumber="77">escens</emphasis>
|
||
, beginning of, becoming.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="77" type="type material">
|
||
<paragraph pageId="0" pageNumber="77">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="77">Holotype female, in amber from the Dominican Republic (piece DZUP Dom-018). According to the amber dealer, it is probably from La Toca or Los Cacaos mine, in Santiago de los Caballeros.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |