treatments-xml/data/FA/B0/50/FAB0508E0DEF54CABFF204D9D738AA58.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="FAB0508E0DEF54CABFF204D9D738AA58" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila bathyodon Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="29C5CA54-04BF-43EC-9B85-689E9F252E81" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:29C5CA54-04BF-43EC-9B85-689E9F252E81" httpUri="http://treatment.plazi.org/id/FAB0508E0DEF54CABFF204D9D738AA58" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/29C5CA54-04BF-43EC-9B85-689E9F252E81" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila bathyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bathyodon" status="sp. nov.">
Angustopila bathyodon 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 62" captionStartId="F62" captionText="Figure 62. Angustopila bathyodon Pall-Gergely &amp; Hunyadi, sp. nov. A, C-I holotype (HNHM 105295) B paratype. Apertural (A, B), lateral (C), apical (D) and ventral (E) sculpture on the protoconch (F), sides of the shell; aperture (G); ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure62" httpUri="https://binary.pensoft.net/fig/808871" pageId="0" pageNumber="1">Fig. 62</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Laos • 1 empty shell (H: 0.78 mm, D: 0.82 mm); Khammouane Province, 43.4 km from Nhommalath towards Nongchan (bridge on road #12), southern edge of Ban 
<normalizedToken originalValue="Khamhé">Khamhe</normalizedToken>
, right side of road (locality code 2019/108); 
<geoCoordinate degrees="17" direction="north" minutes="34.74" orientation="latitude" precision="9" value="17.579">17°34.74'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="27.52" orientation="longitude" precision="9" value="105.458664">105°27.52'E</geoCoordinate>
; 210 m a.s.l.; 30 Sep. 2019; A. Hunyadi leg.; HNHM 105295.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Laos • 34 shells; same data as for holotype; coll. HA • 3 shells; same data as for holotype; coll. JJV.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<collectingCountry name="Laos">Laos</collectingCountry>
• 11 j/b shells; same data as for holotype; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a conical to conical-globular shell, regularly developed parietal and palatal teeth that are situated inside the shell some distance from the peristome.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell small for the genus, slightly wider than high; off-white, translucent, conical to conical-globular; body whorl widest in standard apertural view; protoconch consists of 1.5 whorls with weak spiral striation preceding the first teleoconch whorl; teleoconch with very fine irregularly spaced, very weak radial growth lines crossed by much stronger, regularly spaced spiral striae (ca. 14-18 on body whorl from standard apertural view); on both ventral and dorsal surfaces of body whorl spiral lines far more dominant than radial lines; whorls 3.75-4, rounded or very slightly shouldered; aperture oblique to shell axis from lateral view; umbilicus wide, slightly excentric; aperture reniform, sinulus wide and weakly isolated due to the weak parietal and upper palatal teeth; peristome expanded, not reflected; parietal callus not protruding, not detached from penultimate whorl; parietal tooth rather low, short, situated inside some distance from parietal callus, perpendicular to parietal side; upper palatal tooth almost as high as parietal but wider, blunter, situated inside the shell some distance from peristome edge, at middle of palatal region, just opposite of parietal tooth.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure62" httpUri="https://binary.pensoft.net/fig/808871" pageId="0" pageNumber="1" start="Figure 62" startId="F62">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 62.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bathyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bathyodon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bathyodon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A, C-I</emphasis>
holotype (HNHM 105295) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
paratype. Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), sides of the shell; aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.67-0.78; D = 0.72-0.82, H/D*100 = 91.8-97.2 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4), RUD = 30.7-31.3 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila antidomedon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="antidomedon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila antidomedon</emphasis>
</taxonomicName>
sp. nov. has a higher spire, a more conical shell, and a lower situated parietal tooth. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila occidentalis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="occidentalis">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila occidentalis</emphasis>
</taxonomicName>
sp. nov. is much larger and has a lower situated parietal tooth. 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila pustulata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pustulata">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila pustulata</emphasis>
</taxonomicName>
sp. nov. has a concave-conical shell, it is also larger, has a weaker sculpture, and its palatal tooth is situated lower. See under 
<taxonomicName lsidName="A. tweediei" pageId="0" pageNumber="1" rank="species" species="tweediei">
<emphasis italics="true" pageId="0" pageNumber="1">A. tweediei</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet (
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bathyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bathyodon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bathyodon</emphasis>
</taxonomicName>
) refers to the position of the apertural barriers (
<emphasis italics="true" pageId="0" pageNumber="1">βαθύς</emphasis>
= deep, 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὀδούς">ὀδούς</normalizedToken>
</emphasis>
= tooth in Greek). Namely, they are situated more posteriorly than those of the most similar species 
<taxonomicName lsidName="A. tweediei" pageId="0" pageNumber="1" rank="species" species="tweediei">
<emphasis italics="true" pageId="0" pageNumber="1">A. tweediei</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila bathyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bathyodon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila bathyodon</emphasis>
</taxonomicName>
sp. nov. is known from the type locality only in Khammouane Province of Laos (Fig. 
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure55" httpUri="https://binary.pensoft.net/fig/808864" pageId="0" pageNumber="1">55</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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