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<document id="561D3DDA15A58B6D1E5CA1071BCB457D" ID-CLB-Dataset="306643" ID-DOI="10.11646/phytotaxa.664.3.5" ID-GBIF-Dataset="6777b37d-6dae-4211-8fe6-2e100734e29a" ID-ISSN="1179-3163" ID-Zenodo-Dep="14517360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734542859981" checkinUser="felipe" docAuthor="Nguyen, Cuong Huu, Dang, Ha Van, Nguyen, Yen Thi, Nguyen, Ly Van, Averyanov, Leonid V., Maisak, Tatiana V. &amp; Shvanova, Valeria V." docDate="2024" docId="FA663E377D6AFFB576C3F827FB559715" docLanguage="en" docName="phytotaxa.664.3.200-206.pdf" docOrigin="Phytotaxa 664 (3)" docSource="https://doi.org/10.11646/phytotaxa.664.3.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Dendrobium namdongense Aver. &amp; C. H. Nguyen 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="205" masterDocId="065F464F7D6BFFB0764BFFD3FF809453" masterDocTitle="Dendrobium namdongense (D. sect. Dendrobium), a new miniature orchid from limestone mountains of northern Vietnam" masterLastPageNumber="206" masterPageNumber="200" pageNumber="201" updateTime="1734543603947" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AF31994569B20BD9051507ECE157BEAD">Dendrobium namdongense (D. sect. Dendrobium), a new miniature orchid from limestone mountains of northern Vietnam</mods:title>
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<mods:namePart id="6D6EDCB0CE9B9FE1DC5E7B10BC037DA4">Nguyen, Cuong Huu</mods:namePart>
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<mods:namePart id="25D4FA1852AA9A633E2C651EF8FA84DF">Dang, Ha Van</mods:namePart>
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<mods:affiliation id="A52832F75BDF0EDC630295ACE1432999">Vietnam National University of Forestry, Xuan Mai, Chuong My, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="048EAA6A16A64EA92188B6FADFFC112D">Nguyen, Yen Thi</mods:namePart>
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<mods:affiliation id="F71D634213D25CBCF37ADC0B628A025E">Vietnam National University of Forestry, Xuan Mai, Chuong My, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="3544F701D60008AA22C9F3535B365669">Nguyen, Ly Van</mods:namePart>
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<mods:affiliation id="1B434131A3FC2C3CECCD62E7B0CD5D6B">Vietnam National University of Forestry, Xuan Mai, Chuong My, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="1A5761C28FC94B96C5312918E94B33DB">Averyanov, Leonid V.</mods:namePart>
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<mods:affiliation id="B69F469F7DE7315756906CA8E8E9E6A8">Komarov Botanical Institute Russian Academy of Sciences, Prof. Popov str., 2, St. Petersburg, 197022, Russia</mods:affiliation>
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<mods:namePart id="73062D965F87CA291D7BE517EE8AFFAD">Maisak, Tatiana V.</mods:namePart>
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<mods:affiliation id="E9E7C2D38D299FBC65C8163C02F4174E">Komarov Botanical Institute Russian Academy of Sciences, Prof. Popov str., 2, St. Petersburg, 197022, Russia</mods:affiliation>
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<mods:namePart id="0D645CF80313FCDC967ED16095C74839">Shvanova, Valeria V.</mods:namePart>
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<mods:affiliation id="F906AA71DFF66FB298E6808DFF51848A">Komarov Botanical Institute Russian Academy of Sciences, Prof. Popov str., 2, St. Petersburg, 197022, Russia</mods:affiliation>
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<mods:date id="C39A18F8ABA429E2231AA0DC0F8153A8">2024</mods:date>
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<treatment id="FA663E377D6AFFB576C3F827FB559715" ID-DOI="http://doi.org/10.5281/zenodo.14517358" ID-Zenodo-Dep="14517358" LSID="urn:lsid:plazi:treatment:FA663E377D6AFFB576C3F827FB559715" httpUri="http://treatment.plazi.org/id/FA663E377D6AFFB576C3F827FB559715" lastPageId="5" lastPageNumber="205" pageId="1" pageNumber="201">
<subSubSection id="3AD5DCAA7D6AFFB176C3F827FCF19C5D" box="[136,881,2036,2063]" pageId="1" pageNumber="201" type="nomenclature">
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<heading id="2938384D7D6AFFB176C3F827FCF19C5D" box="[136,881,2036,2063]" fontSize="11" level="2" pageId="1" pageNumber="201" reason="3">
<taxonomicName id="B5CFF4A27D6AFFB176C3F827FD379C5D" authority="Aver. &amp; C.H.Nguyen" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[136,695,2036,2063]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="201" phylum="Tracheophyta" rank="species" species="namdongense" status="sp. nov.">
<emphasis id="40BB53337D6AFFB176C3F827FE419C5C" bold="true" box="[136,449,2036,2063]" italics="true" pageId="1" pageNumber="201">Dendrobium namdongense</emphasis>
Aver. &amp; C.H.Nguyen
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,
<emphasis id="40BB53337D6AFFB17489F826FC979C5D" box="[706,791,2037,2062]" italics="true" pageId="1" pageNumber="201">
<taxonomicNameLabel id="5B88EE487D6AFFB17489F826FC979C5D" box="[706,791,2037,2062]" pageId="1" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="EAF493A47D6AFFB1756EF827FCE69C5D" box="[805,870,2036,2062]" captionStart="FIGURE 1" captionStartId="3.[136,229,1863,1885]" captionTargetBox="[243,1344,187,1838]" captionTargetId="figure-24@3.[243,1344,187,1838]" captionTargetPageId="3" captionText="FIGURE 1. Dendrobium namdongense. A. Flowering plant in natural habitat. B. Mature pseudobulbs with leaves. C. Flowering plant, side view. D–F. Intact flower, view from different sides. G. Lip and column in intact flower. H–K. Peduncle, pedicel, ovary and separated flower, view from different sides. M–O. Separated lip, view from different sides. P. Lip, side view. Q. Lip, sagittal section. R, S. Flattened lip, adaxial (R) and abaxial (S) side. T, U. Column apex, front view, with (T) and without anther cap (U). V. Anther cap with pollinaria, front view, view from below and side view. W. Anther cap, view from rear (with pollinaria) and from below (without pollinaria). X. Pollinaria, front and rear views. Y. Pollinia, side view. Photos by L. Averyanov and Cuong Huu Nguyen from living plants prior to the preparation of the type herbarium specimens AL3075. Photo correction and design by L. Averyanov and T. Maisak." figureDoi="http://doi.org/10.5281/zenodo.14517364" httpUri="https://zenodo.org/record/14517364/files/figure.png" pageId="1" pageNumber="201">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB276C3FF4CFC6995DE" pageId="2" pageNumber="202" type="materials_examined">
<paragraph id="72708F217D69FFB276C3FF4CFC6995DE" blockId="2.[136,1452,159,397]" pageId="2" pageNumber="202">
Type:—
<collectingCountry id="0AD8CFB17D69FFB2769DFF4CFEC894E7" box="[214,328,159,180]" name="Vietnam" pageId="2" pageNumber="202">VIETNAM</collectingCountry>
.
<collectingRegion id="B00B41C37D69FFB27718FF4CFDA194E6" box="[339,545,159,181]" country="Vietnam" name="Thanh Hoa" pageId="2" pageNumber="202">Thanh Hoa Province</collectingRegion>
: Quan Hoa District, Nam Dong Community, the territory of Nam Dong Conservation Area, primary broad-leaved evergreen and mixed (with
<taxonomicName id="B5CFF4A27D69FFB274D7FF17FCF2948B" box="[668,882,195,217]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="kwangtungensis">
<emphasis id="40BB53337D69FFB274D7FF17FCF2948B" box="[668,882,195,217]" italics="true" pageId="2" pageNumber="202">Pinus kwangtungensis</emphasis>
</taxonomicName>
) humid forest on very steep rocky slopes on solid, marble-like highly eroded limestone at elevation
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a.s.l. on steep rocky slope, clustering appressed epiphyte on mossy trees on rocky mountain top, flowers tepals white to light yellowish or pinkish with purple veins, lip dark purple, rare,
<date id="0671A9E17D69FFB27367FED8FA279573" box="[1324,1447,267,289]" pageId="2" pageNumber="202" value="2024-05-04">4 May 2024</date>
,
<emphasis id="40BB53337D69FFB276F6FEE3FCDA9517" box="[189,858,303,325]" italics="true" pageId="2" pageNumber="202">
Nguyen Huu Cuong, Nguyen Van Ly,
<collectionCode id="14DE17E47D69FFB27454FEE3FDAF9516" box="[543,559,304,325]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="202" type="Herbarium">L</collectionCode>
. Averyanov,
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. Maisak,
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3075
</emphasis>
(
<typeStatus id="AD7431837D69FFB2752DFEFCFC3C9516" box="[870,956,303,325]" pageId="2" pageNumber="202" type="holotype">holotype</typeStatus>
<collectionCode id="14DE17E47D69FFB2758BFEE3FC709516" box="[960,1008,304,325]" country="Viet Nam" httpUri="http://grbio.org/cool/merb-e6y6" name="Vietnam Forestry Herbarium" pageId="2" pageNumber="202" type="Herbarium">VNF</collectionCode>
<collectionCode id="14DE17E47D69FFB275BDFEFCFBA99516" box="[1014,1065,303,325]" pageId="2" pageNumber="202">NHC</collectionCode>
20240506001,
<typeStatus id="AD7431837D69FFB272FCFEFCFA869516" box="[1207,1286,303,325]" pageId="2" pageNumber="202" type="isotype">isotypes</typeStatus>
<collectionCode id="14DE17E47D69FFB27340FEE3FACB9516" box="[1291,1355,303,325]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="202">LE LE</collectionCode>
01276172 &amp;
<collectionCode id="14DE17E47D69FFB27697FE87FEA2953A" box="[220,290,339,361]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="202">LE LE</collectionCode>
01276518, photos of plants used for preparation of the type specimens
<collectionCode id="14DE17E47D69FFB2724CFE87FBCD953A" box="[1031,1101,339,361]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="202">LE LE</collectionCode>
01125079 https://en.herbariumle.ru/?t=occ&amp;id=226271,
<collectionCode id="14DE17E47D69FFB277CFFEABFE4595DE" box="[388,453,375,397]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="202">LE LE</collectionCode>
01255109 https://en.herbariumle.ru/?t=occ&amp;id=226270).
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB276C3FE6FFAE89586" box="[136,1384,443,470]" pageId="2" pageNumber="202" type="etymology">
<paragraph id="72708F217D69FFB276C3FE6FFAE89586" blockId="2.[136,1452,443,2090]" box="[136,1384,443,470]" pageId="2" pageNumber="202">
<emphasis id="40BB53337D69FFB276C3FE6FFEAE9585" bold="true" box="[136,302,444,470]" pageId="2" pageNumber="202">Etymology:—</emphasis>
The species epithet refers to the Nam Dong Conservation Area, where this plant was discovered.
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB276F6FE0CFCCB96DA" pageId="2" pageNumber="202" type="diagnosis">
<paragraph id="72708F217D69FFB276F6FE0CFCCB96DA" blockId="2.[136,1452,443,2090]" pageId="2" pageNumber="202">
<emphasis id="40BB53337D69FFB276F6FE0CFED695AA" bold="true" box="[189,342,479,505]" pageId="2" pageNumber="202">Diagnosis:—</emphasis>
The new species differs from most similar
<taxonomicName id="B5CFF4A27D69FFB27579FE0CFBD495A9" box="[818,1108,479,506]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D69FFB27579FE0CFBD495A9" box="[818,1108,479,506]" italics="true" pageId="2" pageNumber="202">Dendrobium fuscifaucium</emphasis>
</taxonomicName>
and
<taxonomicName id="B5CFF4A27D69FFB272C1FE33FAA195AA" box="[1162,1313,479,505]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="hekouense">
<emphasis id="40BB53337D69FFB272C1FE33FAA195AA" box="[1162,1313,479,505]" italics="true" pageId="2" pageNumber="202">D. hekouense</emphasis>
</taxonomicName>
in distinctly smaller pseudobulbs,
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tall; leaves,
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long with very short peduncle, less than
<quantity id="B53722C47D69FFB272E0FDD7FB6D964D" box="[1195,1261,516,542]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="202" unit="mm" value="1.0">1 mm</quantity>
long; sepals and petals with distinct purple veins; short lip,
<quantity id="B53722C47D69FFB2743BFDF4FC949611" box="[624,788,551,578]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="2" pageNumber="202" unit="mm" value="13.0" valueMax="14.5" valueMin="11.5">11.5–14.5 mm</quantity>
long, lip side lobes papillulate, obliquely semicircular, lip median lobe semicircular,
<quantity id="B53722C47D69FFB277F8FD98FDBB9635" box="[435,571,587,614]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="2" pageNumber="202" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 mm</quantity>
wide, lip disc flat or with two low insignificant keels; anther cap rectangular, semiglobular at apex, and time of flowering during April–May.
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB276F6FD40FF7092CE" pageId="2" pageNumber="202" type="description">
<paragraph id="72708F217D69FFB276F6FD40FF7092CE" blockId="2.[136,1452,443,2090]" pageId="2" pageNumber="202">
<emphasis id="40BB53337D69FFB276F6FD40FEED96FE" bold="true" box="[189,365,659,685]" pageId="2" pageNumber="202">Description:—</emphasis>
Miniature epiphytic glabrous herb with many distichous pseudobulbs densely clustering on thin, creeping plagiotropic rhizome.
<emphasis id="40BB53337D69FFB277A9FD6BFDF99681" bold="true" box="[482,633,696,722]" pageId="2" pageNumber="202">Pseudobulbs</emphasis>
brown to brown purple, 1–2-leaved, closely touching each other, narrowly ovate to broadly fusiform or ellipsoid, (5)6–8(9) mm tall, (2.5)3–4(4.5) mm in diameter, shortly stalked, consisting of 2(3) internodes, each (1.5)2–3(4) mm long, bladeless sheaths, pale green to scarious, with longitudinal nerves, soon weathered into papyraceous or fibrous remnants.
<emphasis id="40BB53337D69FFB274A0FCF7FCBD976D" bold="true" box="[747,829,804,830]" pageId="2" pageNumber="202">Leaves</emphasis>
subsessile; petiole sheathed, white, scarious,
<quantity id="B53722C47D69FFB27371FCF7FA2C976D" box="[1338,1452,804,830]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="202" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
long; leaf lamina narrowly ovate to broadly elliptic, dull green, often flushed with purple toward the apex and along the margin, coriaceous, conduplicate, (6)9–12(13) mm long, (3.5)4–5(5.5) mm wide, blunt to rounded at base and apex.
<emphasis id="40BB53337D69FFB27686FC5CFEF697FA" bold="true" box="[205,374,911,937]" pageId="2" pageNumber="202">Inflorescences</emphasis>
1–2 on individual pseudobulb, subterminal, arising from the apex of leafless pseudobulbs, 1- flowered; peduncle less than
<quantity id="B53722C47D69FFB27786FC67FD8E979D" box="[461,526,948,974]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="202" unit="mm" value="1.0">1 mm</quantity>
long, with 1 very small, insignificant, tubular sheath at base; floral bract triangular ovate, funnel-shaped, acute, (2.5)3–3.5(4) mm long, white to light gray, scarious; pedicel and ovary slightly curved, (4.5)5–6.5(7) mm long,
<quantity id="B53722C47D69FFB277D4FC2FFD939045" box="[415,531,1020,1046]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="202" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
in diameter, white to white tinged with purple.
<emphasis id="40BB53337D69FFB2720BFC2FFB1F9045" bold="true" box="[1088,1183,1020,1046]" pageId="2" pageNumber="202">Flowers</emphasis>
spreading horizontally, widely opening; sepals and petals white, sometimes with a light yellow or pink tint, with purple veins on both sides; the lip dark purple inside, white with a purple tint outside; side lobes white to light yellowish; the median lobe dark purple on both sides, disc and medial lobe adaxially with dense whitish hairs.
<emphasis id="40BB53337D69FFB275B6FBB4FB1E90D1" bold="true" box="[1021,1182,1127,1154]" pageId="2" pageNumber="202">Median sepal</emphasis>
oblong narrowly ovate, (6)6.5–7.5(8) mm long,
<quantity id="B53722C47D69FFB277EBFB58FD9590F5" box="[416,533,1163,1190]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="202" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
wide, obtuse, with 3 purple veins.
<emphasis id="40BB53337D69FFB275FAFB5FFBDC90F5" bold="true" box="[945,1116,1164,1190]" pageId="2" pageNumber="202">Lateral sepals</emphasis>
obliquely broadly triangular, (7)7.5–8(8.5) mm long, (10.5)11–12(12.5) mm wide at the base, acute, with 5 purple veins; mentum saccate, (9)10(11) mm long,
<quantity id="B53722C47D69FFB276B5FB00FEF390BD" box="[254,371,1235,1262]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="2" pageNumber="202" unit="mm" value="5.75" valueMax="6.0" valueMin="5.5">5.5–6 mm</quantity>
wide, slightly incurved, apically round, and slightly bilobed.
<emphasis id="40BB53337D69FFB2727DFB07FBFE90BD" bold="true" box="[1078,1150,1236,1262]" pageId="2" pageNumber="202">Petals</emphasis>
oblong narrowly obovate, (6)6.5–7(7.5) mm long, (3.2)3.5–4(4.2) mm wide, obtuse, with 1 violet vein.
<emphasis id="40BB53337D69FFB275B1FB24FBA49142" bold="true" box="[1018,1060,1271,1297]" pageId="2" pageNumber="202">Lip</emphasis>
trilobed, recurved, with erect side lobes, and strongly recurved median lobe, broadly obdeltoid (when flattened), (11.5)12–14(14.5) mm long and wide; side lobes erect, obliquely semicircular,
<quantity id="B53722C47D69FFB2741DFAECFD4B9109" box="[598,715,1343,1370]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="202" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 mm</quantity>
long,
<quantity id="B53722C47D69FFB2755DFAECFCF59109" box="[790,885,1343,1370]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="202" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
wide at the base, finely ciliate along the margin; median lobe, semicircular, entire or obscurely emarginate, (2.5)3(3.5) mm long, (4.5)5–6(6.5) mm wide, margin ciliate; disc and surface of median lobe adaxially densely pubescent with long hairs, often branching at apex, disc and base of median lobe almost flat or with two low, hardly visible longitudinal keels.
<emphasis id="40BB53337D69FFB2726DFA78FB069196" bold="true" box="[1062,1158,1451,1477]" pageId="2" pageNumber="202">Column</emphasis>
<quantity id="B53722C47D69FFB272C5FA78FA819195" box="[1166,1281,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="202" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
tall,
<quantity id="B53722C47D69FFB27372FA78FA2C9195" box="[1337,1452,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="202" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
wide, finely papillulate on sides, stigma concave, semicircular, about
<quantity id="B53722C47D69FFB275C2FA1CFC5E91B9" box="[905,990,1487,1514]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="202" unit="mm" value="1.5">1.5 mm</quantity>
in diameter; column foot
<quantity id="B53722C47D69FFB272B7FA1CFAED91B9" box="[1276,1389,1487,1514]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="2" pageNumber="202" unit="mm" value="9.25" valueMax="9.5" valueMin="9.0">9–9.5 mm</quantity>
long,
<quantity id="B53722C47D69FFB276C3FA20FF7B925D" box="[136,251,1523,1550]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="2" pageNumber="202" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
wide in broadest part, white proximally, dark purple in distal half, glabrous, almost round at apex.
<emphasis id="40BB53337D69FFB2731DFA20FF329261" bold="true" pageId="2" pageNumber="202">Anther cap</emphasis>
rectangular helmet-shaped, about
<quantity id="B53722C47D69FFB2747FF9CBFD0A9261" box="[564,650,1560,1586]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="202" unit="mm" value="1.8">1.8 mm</quantity>
tall,
<quantity id="B53722C47D69FFB27489F9CBFC839261" box="[706,771,1560,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="202" unit="mm" value="2.0">2 mm</quantity>
wide, at front view almost semiglobular at apex, light green, front margin white, finely papillulate; pollinia 4, bright yellow, narrowly conoid, slightly curved,
<quantity id="B53722C47D69FFB272A9F9EFFAEA9205" box="[1250,1386,1596,1622]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="202" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
long.
<emphasis id="40BB53337D69FFB276C3F98CFF73922A" bold="true" box="[136,243,1631,1657]" pageId="2" pageNumber="202">Capsules</emphasis>
(not ripe) dark green, narrowly obconoid, longitudinally ribbed,
<quantity id="B53722C47D69FFB2759FF9B3FBCE9229" box="[980,1102,1632,1658]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="202" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long,
<quantity id="B53722C47D69FFB272DDF98CFA9E9229" box="[1174,1310,1631,1658]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="2" pageNumber="202" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
in diameter, glabrous.
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB276F6F974FF4692B5" pageId="2" pageNumber="202" type="distribution">
<paragraph id="72708F217D69FFB276F6F974FF4692B5" blockId="2.[136,1452,443,2090]" pageId="2" pageNumber="202">
<emphasis id="40BB53337D69FFB276F6F974FEF59292" bold="true" box="[189,373,1703,1729]" pageId="2" pageNumber="202">Distribution:—</emphasis>
<collectingRegion id="B00B41C37D69FFB2773EF974FDE09292" box="[373,608,1703,1730]" country="Vietnam" name="Thanh Hoa" pageId="2" pageNumber="202">Thanh Hoa Province</collectingRegion>
, Quan Hoa District, endemic to limestone area of Nam Dong Conservation Area.
</paragraph>
</subSubSection>
<subSubSection id="3AD5DCAA7D69FFB576F6F923FB559715" lastPageId="5" lastPageNumber="205" pageId="2" pageNumber="202" type="biology_ecology">
<paragraph id="72708F217D69FFB276F6F923FD6C9C79" blockId="2.[136,1452,443,2090]" pageId="2" pageNumber="202">
<emphasis id="40BB53337D69FFB276F6F923FE43935A" bold="true" box="[189,451,1775,1802]" pageId="2" pageNumber="202">Ecology and habitat:</emphasis>
—Miniature creeping epiphyte with densely clustering appressed pseudobulbs; often observed on mossy trees in primary broad-leaved evergreen and mixed (with
<taxonomicName id="B5CFF4A27D69FFB275A4F8C7FB69937E" box="[1007,1257,1811,1837]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="kwangtungensis">
<emphasis id="40BB53337D69FFB275A4F8C7FB69937E" box="[1007,1257,1811,1837]" italics="true" pageId="2" pageNumber="202">Pinus kwangtungensis</emphasis>
</taxonomicName>
) humid forest on tops of remnant karstic mountains composed with solid, marble-like highly eroded limestone at elevations of
<quantity id="B53722C47D69FFB27377F8EBFF1E9325" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="2" pageNumber="202" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</quantity>
a.s.l. The climate in the area where
<taxonomicName id="B5CFF4A27D69FFB27461F888FCD79326" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[554,855,1883,1909]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D69FFB27461F888FCD79326" box="[554,855,1883,1909]" italics="true" pageId="2" pageNumber="202">Dendrobium namdongense</emphasis>
</taxonomicName>
was found is recognized as a tropical monsoon with a hot and humid summer, and dry and cool winter periods (Nguyen Khanh Van
<emphasis id="40BB53337D69FFB27596F853FB8D93CA" box="[989,1037,1919,1945]" italics="true" pageId="2" pageNumber="202">et al</emphasis>
. 2000). The summer rain season lasts from May until September, with daily usual temperatures between 23–38ºC (with a recorded maximum 41ºC). The dry winter season lasts from September to April, with usual temperatures between 26–23ºC (with a recorded minimum 6ºC). Annual rainfall averages nearly
<quantity id="B53722C47D69FFB2740BF838FD329C55" box="[576,690,2027,2054]" metricMagnitude="0" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="202" unit="mm" value="1050.0">1050 mm</quantity>
. Cloudy fogs and mists on top of limestone mountains harboring habitats of
<taxonomicName id="B5CFF4A27D69FFB2774FF7C3FE419C7A" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[260,449,2063,2089]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="202" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D69FFB2774FF7C3FE419C7A" box="[260,449,2063,2089]" italics="true" pageId="2" pageNumber="202">D. namdongense</emphasis>
</taxonomicName>
are very common all year.
</paragraph>
<caption id="26B0DFA97D68FFB376C3F894FB5A9C7B" ID-DOI="http://doi.org/10.5281/zenodo.14517364" ID-Zenodo-Dep="14517364" httpUri="https://zenodo.org/record/14517364/files/figure.png" pageId="3" pageNumber="203" startId="3.[136,229,1863,1885]" targetBox="[243,1344,187,1838]" targetPageId="3" targetType="figure">
<paragraph id="72708F217D68FFB376C3F894FB5A9C7B" blockId="3.[136,1452,1863,2088]" pageId="3" pageNumber="203">
<emphasis id="40BB53337D68FFB376C3F894FDB6930F" bold="true" box="[136,566,1863,1885]" pageId="3" pageNumber="203">
FIGURE 1.
<emphasis id="40BB53337D68FFB3774EF894FD98930E" bold="true" box="[261,536,1863,1885]" italics="true" pageId="3" pageNumber="203">
<taxonomicName id="B5CFF4A27D68FFB3774EF894FD95930E" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[261,533,1863,1885]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="203" phylum="Tracheophyta" rank="species" species="namdongense">Dendrobium namdongense</taxonomicName>
.
</emphasis>
A.
</emphasis>
Flowering plant in natural habitat.
<emphasis id="40BB53337D68FFB375C6F894FC23930F" bold="true" box="[909,931,1863,1884]" pageId="3" pageNumber="203">B.</emphasis>
Mature pseudobulbs with leaves.
<emphasis id="40BB53337D68FFB372A5F894FA86930E" bold="true" box="[1262,1286,1863,1885]" pageId="3" pageNumber="203">C.</emphasis>
Flowering plant, side view.
<emphasis id="40BB53337D68FFB376A6F8BAFE9D932D" bold="true" box="[237,285,1897,1918]" pageId="3" pageNumber="203">D–F.</emphasis>
Intact flower, view from different sides.
<emphasis id="40BB53337D68FFB374EFF8BBFD3D932D" bold="true" box="[676,701,1896,1918]" pageId="3" pageNumber="203">G.</emphasis>
Lip and column in intact flower.
<emphasis id="40BB53337D68FFB375B0F8BAFBB2932D" bold="true" box="[1019,1074,1897,1918]" pageId="3" pageNumber="203">H–K.</emphasis>
Peduncle, pedicel, ovary and separated flower, view from different sides.
<emphasis id="40BB53337D68FFB37787F859FD8793F3" bold="true" box="[460,519,1930,1952]" pageId="3" pageNumber="203">M–O.</emphasis>
Separated lip, view from different sides.
<emphasis id="40BB53337D68FFB375DBF859FC2293CC" bold="true" box="[912,930,1930,1951]" pageId="3" pageNumber="203">P.</emphasis>
Lip, side view.
<emphasis id="40BB53337D68FFB37273F859FBD193F3" bold="true" box="[1080,1105,1930,1952]" pageId="3" pageNumber="203">Q.</emphasis>
Lip, sagittal section.
<emphasis id="40BB53337D68FFB37356F859FACD93F3" bold="true" box="[1309,1357,1930,1952]" pageId="3" pageNumber="203">R, S.</emphasis>
Flattened lip, adaxial (R) and abaxial (S) side.
<emphasis id="40BB53337D68FFB377A5F878FDA09393" bold="true" box="[494,544,1963,1984]" pageId="3" pageNumber="203">T, U.</emphasis>
Column apex, front view, with (T) and without anther cap (U).
<emphasis id="40BB53337D68FFB372C6F878FB219393" bold="true" box="[1165,1185,1963,1984]" pageId="3" pageNumber="203">V.</emphasis>
Anther cap with pollinaria, front view, view from below and side view.
<emphasis id="40BB53337D68FFB37409F81EFDDD93B1" bold="true" box="[578,605,1997,2018]" pageId="3" pageNumber="203">W.</emphasis>
Anther cap, view from rear (with pollinaria) and from below (without pollinaria).
<emphasis id="40BB53337D68FFB373DFF81EFA2C93B1" bold="true" box="[1428,1452,1997,2018]" pageId="3" pageNumber="203">X.</emphasis>
Pollinaria, front and rear views.
<emphasis id="40BB53337D68FFB3778FF83DFE599C50" bold="true" box="[452,473,2030,2051]" pageId="3" pageNumber="203">Y.</emphasis>
Pollinia, side view. Photos by L. Averyanov and Cuong Huu Nguyen from living plants prior to the preparation of the type herbarium specimens
<emphasis id="40BB53337D68FFB37470F7C1FD079C74" box="[571,647,2066,2087]" italics="true" pageId="3" pageNumber="203">AL3075</emphasis>
. Photo correction and design by L. Averyanov and T. Maisak.
</paragraph>
</caption>
<paragraph id="72708F217D6FFFB476F6FF4DFC2C94EB" blockId="4.[136,1452,158,797]" box="[189,940,158,184]" pageId="4" pageNumber="204">
<emphasis id="40BB53337D6FFFB476F6FF4DFEC394EB" bold="true" box="[189,323,158,184]" pageId="4" pageNumber="204">Phenology:</emphasis>
—Flowers between April–May (onset of rainy season).
</paragraph>
<paragraph id="72708F217D6FFFB476F6FF11FE1A974F" blockId="4.[136,1452,158,797]" pageId="4" pageNumber="204">
<emphasis id="40BB53337D6FFFB476F6FF11FE89948F" bold="true" box="[189,265,194,220]" pageId="4" pageNumber="204">Notes:</emphasis>
—The new species described here belongs to the group of miniature species from
<taxonomicName id="B5CFF4A27D6FFFB4728DFF11FE959553" authority="section Dendrobium" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="section" section="Dendrobium">
<emphasis id="40BB53337D6FFFB4728DFF11FAD3948F" box="[1222,1363,194,220]" italics="true" pageId="4" pageNumber="204">Dendrobium</emphasis>
section
<emphasis id="40BB53337D6FFFB476C3FF35FE959553" box="[136,277,230,256]" italics="true" pageId="4" pageNumber="204">Dendrobium</emphasis>
</taxonomicName>
, including
<taxonomicName id="B5CFF4A27D6FFFB477D0FF34FCBE9552" authority="Souvann. &amp; Kumar" authorityName="Souvann. &amp; Kumar" box="[411,830,230,257]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D6FFFB477D0FF34FDD09553" box="[411,592,230,256]" italics="true" pageId="4" pageNumber="204">D. fuscifaucium</emphasis>
Souvann. &amp; Kumar
</taxonomicName>
in
<bibRefCitation id="165EF2D07D6FFFB47522FF35FA819553" author="Souvannakhoummane, K. &amp; Kumar, P. &amp; Phonepaseuth, P." box="[873,1281,230,256]" pageId="4" pageNumber="204" pagination="285 - 290" refId="ref5143" refString="Souvannakhoummane, K., Kumar, P., Phonepaseuth, P. (2022) Dendrobium fuscifaucium (Orchidaceae: Epidendroideae: Dendrobieae), a new Laotian species only known in cultivation. Phytotaxa 541 (3): 285 - 290. https: // doi. org / 10.11646 / phytotaxa. 541.3.8" type="journal article" year="2022">
Souvannakhoummane
<emphasis id="40BB53337D6FFFB47226FF34FB299553" box="[1133,1193,230,256]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
(2022)
</bibRefCitation>
,
<taxonomicName id="B5CFF4A27D6FFFB4735AFF34FEE69577" authority="Z. J. Liu &amp; L. J. Chen" authorityName="Z. J. Liu &amp; L. J. Chen" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="hekouense">
<emphasis id="40BB53337D6FFFB4735AFF34FA2C9553" box="[1297,1452,230,256]" italics="true" pageId="4" pageNumber="204">D. hekouense</emphasis>
Z.J.Liu &amp; L.J.Chen
</taxonomicName>
in
<bibRefCitation id="165EF2D07D6FFFB477C6FED9FDEC9577" author="Liu, Z. J. &amp; Chen, L. J." box="[397,620,266,292]" pageId="4" pageNumber="204" pagination="87 - 90" refId="ref4920" refString="Liu, Z. J. &amp; Chen, L. J. (2011) Dendrobium hekouense (Orchidaceae), a new species from Yunnan, China. Annales Botanici Fennici 48: 87 - 90. https: // www. jstor. org / stable / 23728247" type="journal article" year="2011">Liu &amp; Chen (2011)</bibRefCitation>
,
<taxonomicName id="B5CFF4A27D6FFFB47432FED8FC459576" authority="L. J. Chen &amp; W. H. Rao" authorityName="L. J. Chen &amp; W. H. Rao" box="[633,965,266,293]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="luoi">
<emphasis id="40BB53337D6FFFB47432FED8FD4A9577" box="[633,714,266,292]" italics="true" pageId="4" pageNumber="204">D. luoi</emphasis>
L.J.Chen &amp; W.H.Rao
</taxonomicName>
in
<bibRefCitation id="165EF2D07D6FFFB475A6FED8FB439577" author="Deng, X. X. &amp; Chen, Y. K. &amp; Rao, W. H. &amp; Chen, L. J." box="[1005,1219,266,293]" pageId="4" pageNumber="204" pagination="9 - 12" refId="ref4724" refString="Deng, X. X., Chen, Y. K., Rao, W. H. &amp; Chen, L. J. (2016) Dendrobium luoi, a new species of Orchidaceae from China. Plant Science Journal 34: 9 - 12. https: // doi. org / 10.11913 / PSJ. 2095 - 0837.2016.10009" type="journal article" year="2016">
Deng
<emphasis id="40BB53337D6FFFB47278FED8FBED9577" box="[1075,1133,266,292]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
(2016)
</bibRefCitation>
, and
<taxonomicName id="B5CFF4A27D6FFFB4734AFED8FE1A951B" authority="Hu, Long &amp; Jin (2008)" authorityName="G. W. Hu, C. L. Long &amp; X. H. Jin" authorityYear="2008" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="wangliangii">
<emphasis id="40BB53337D6FFFB4734AFED8FA2C9577" box="[1281,1452,266,292]" italics="true" pageId="4" pageNumber="204">D. wangliangii</emphasis>
<bibRefCitation id="165EF2D07D6FFFB476C3FEFCFE1A951B" author="Hu, G. W. &amp; Long, C. L. &amp; Jin, X. H." box="[136,410,302,329]" pageId="4" pageNumber="204" pagination="217 - 221" refId="ref4788" refString="Hu, G. W., Long, C. L. &amp; Jin, X. H. (2008) Dendrobium wangliangii (Orchidaceae), a new species belonging to section Dendrobium from Yunnan, China. Botanical Journal of the Linnean Society 157: 217 - 221. https: // doi. org / 10.1111 / j. 1095 - 8339.2008.00800. x" type="journal article" year="2008">Hu, Long &amp; Jin (2008)</bibRefCitation>
</taxonomicName>
. These species share remarkable characters such as an appressed, subtitle epiphytic habit, small stature of vegetative parts, short 1-flowered inflorescence, and comparatively larger flowers appearing on leafless pseudobulbs. Among these species,
<taxonomicName id="B5CFF4A27D6FFFB474C2FEA4FCC995C3" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[649,841,374,400]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D6FFFB474C2FEA4FCC995C3" box="[649,841,374,400]" italics="true" pageId="4" pageNumber="204">D. namdongense</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="B5CFF4A27D6FFFB472A4FEA4FA2495C3" box="[1263,1444,374,400]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D6FFFB472A4FEA4FA2495C3" box="[1263,1444,374,400]" italics="true" pageId="4" pageNumber="204">D. fuscifaucium</emphasis>
</taxonomicName>
, which originated supposedly from
<collectingCountry id="0AD8CFB17D6FFFB47442FE48FDBF95E6" box="[521,575,411,437]" name="Laos" pageId="4" pageNumber="204">Laos</collectingCountry>
(
<bibRefCitation id="165EF2D07D6FFFB47407FE49FC4595E7" author="Souvannakhoummane, K. &amp; Kumar, P. &amp; Phonepaseuth, P." box="[588,965,410,436]" pageId="4" pageNumber="204" pagination="285 - 290" refId="ref5143" refString="Souvannakhoummane, K., Kumar, P., Phonepaseuth, P. (2022) Dendrobium fuscifaucium (Orchidaceae: Epidendroideae: Dendrobieae), a new Laotian species only known in cultivation. Phytotaxa 541 (3): 285 - 290. https: // doi. org / 10.11646 / phytotaxa. 541.3.8" type="journal article" year="2022">
Souvannakhoummane
<emphasis id="40BB53337D6FFFB47506FE48FC0495E7" box="[845,900,410,436]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
2022
</bibRefCitation>
), and
<emphasis id="40BB53337D6FFFB4724EFE48FB2195E7" box="[1029,1185,410,436]" italics="true" pageId="4" pageNumber="204">
<taxonomicName id="B5CFF4A27D6FFFB4724EFE48FB1D95E7" box="[1029,1181,410,436]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="hekouense">D. hekouense</taxonomicName>
,
</emphasis>
recorded from southern
<collectingCountry id="0AD8CFB17D6FFFB476C3FE6DFF4B958B" box="[136,203,446,472]" name="China" pageId="4" pageNumber="204">China</collectingCountry>
and northern
<collectingCountry id="0AD8CFB17D6FFFB4772FFE6DFE43958B" box="[356,451,446,472]" name="Vietnam" pageId="4" pageNumber="204">Vietnam</collectingCountry>
(
<bibRefCitation id="165EF2D07D6FFFB4779BFE6DFD16958B" author="Liu, Z. J. &amp; Chen, L. J." box="[464,662,446,472]" pageId="4" pageNumber="204" pagination="87 - 90" refId="ref4920" refString="Liu, Z. J. &amp; Chen, L. J. (2011) Dendrobium hekouense (Orchidaceae), a new species from Yunnan, China. Annales Botanici Fennici 48: 87 - 90. https: // www. jstor. org / stable / 23728247" type="journal article" year="2011">Liu &amp; Chen 2011</bibRefCitation>
,
<bibRefCitation id="165EF2D07D6FFFB474EBFE6DFCD9958B" author="Zhou, X. &amp; Cheng, Z. &amp; Liu, Q. &amp; Zhang, J. &amp; Hu, A. &amp; Huang, M. &amp; Hu, C. &amp; Tian, H." box="[672,857,446,473]" pageId="4" pageNumber="204" pagination="1 - 148" refId="ref5265" refString="Zhou, X., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. &amp; Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. Phytotaxa 276 (1): 1 - 148. https: // doi. org / 10.11646 / phytotaxa. 276.1.1" type="journal article" year="2016">
Zhou
<emphasis id="40BB53337D6FFFB474AAFE6CFC91958B" box="[737,785,446,472]" italics="true" pageId="4" pageNumber="204">et al</emphasis>
. 2016
</bibRefCitation>
,
<bibRefCitation id="165EF2D07D6FFFB47529FE6DFBDA958B" author="Averyanov, L. V. &amp; Nguyen, V. C. &amp; Nguyen, H. T. &amp; Truong, B. V. &amp; Nguyen, P. T. &amp; Nguyen, S. K. &amp; Maisak, T. V. &amp; Nguyen, H. T. &amp; Bui, D. N. &amp; Chu, X. C." box="[866,1114,446,472]" pageId="4" pageNumber="204" pagination="195 - 219" refId="ref4452" refString="Averyanov, L. V., Nguyen, V. C., Nguyen, H. T., Truong, B. V., Nguyen, P. T., Nguyen, S. K., Maisak, T. V., Nguyen, H. T., Bui, D. N. &amp; Chu, X. C. (2018) New Orchids (Orchidaceae: Epidendroideae and Vandoideae) in the Flora of Vietnam. Taiwania 63 (3): 195 - 219. https: // doi. org / 10.6165 / tai. 2018.63.195" type="journal article" year="2018">
Averyanov
<emphasis id="40BB53337D6FFFB475A9FE6CFB99958B" box="[994,1049,446,472]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
2018
</bibRefCitation>
,
<bibRefCitation id="165EF2D07D6FFFB4722FFE6DFAA3958B" author="Cribb, P. &amp; Averyanov, L. &amp; Schuiteman, A. &amp; Want, T." box="[1124,1315,446,472]" pageId="4" pageNumber="204" pagination="82 - 88" refId="ref4676" refString="Cribb, P., Averyanov, L., Schuiteman, A. &amp; Want, T. (2021) 973. Dendrobium hekouense. Curtis's Botanical Magazine 38: 82 - 88. https: // doi. org / 10.1111 / curt. 12375" type="journal article" year="2021">
Cribb
<emphasis id="40BB53337D6FFFB472E1FE6CFB61958B" box="[1194,1249,446,472]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
2021
</bibRefCitation>
). Our newly described species can be differentiated from both these species by its distinctly smaller pseudobulbs,
<quantity id="B53722C47D6FFFB472BFFE31FAD195AE" box="[1268,1361,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="4" pageNumber="204" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
tall (vs.
<quantity id="B53722C47D6FFFB476C3FDD5FF739673" box="[136,243,518,544]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="4" pageNumber="204" unit="mm" value="11.0" valueMax="14.0" valueMin="8.0">8–14 mm</quantity>
) and leaves,
<quantity id="B53722C47D6FFFB477C9FDD5FE6C9672" box="[386,492,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="4" pageNumber="204" unit="mm" value="9.5" valueMax="13.0" valueMin="6.0">6–13 mm</quantity>
long (vs.
<quantity id="B53722C47D6FFFB47413FDD5FD549673" box="[600,724,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="5.2" metricValueMin="2.0" pageId="4" pageNumber="204" unit="mm" value="36.0" valueMax="52.0" valueMin="20.0">20–52 mm</quantity>
), very short peduncle, less than
<quantity id="B53722C47D6FFFB4727FFDD5FBF49672" box="[1076,1140,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="204" unit="mm" value="1.0">1 mm</quantity>
long (vs.
<quantity id="B53722C47D6FFFB472ABFDD5FAD59673" box="[1248,1365,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.5" metricValueMin="1.0" pageId="4" pageNumber="204" unit="mm" value="2.75" valueMax="4.5" valueMin="1.0">1–4.5 mm</quantity>
), sepals and petals with distinct purple veins (vs. purple veins absent, or hardly visible), median sepal oblong narrowly ovate (vs. median sepal ovate to broadly ovate), shorter lip,
<quantity id="B53722C47D6FFFB47492FD9DFCFA963A" box="[729,890,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="4" pageNumber="204" unit="mm" value="13.0" valueMax="14.5" valueMin="11.5">11.5–14.5 mm</quantity>
long (vs. lip
<quantity id="B53722C47D6FFFB47242FD9DFB05963B" box="[1033,1157,590,616]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="4" pageNumber="204" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20 mm</quantity>
), lip side lobes papillulate, obliquely semicircular (vs. side lobes fimbriate, obliquely triangular), lip median lobe semicircular,
<quantity id="B53722C47D6FFFB472ACFDA1FAEE96DE" box="[1255,1390,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="4" pageNumber="204" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 mm</quantity>
wide (vs. median lobe reniform,
<quantity id="B53722C47D6FFFB4778BFD45FDAD96E2" box="[448,557,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="204" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
wide), lip disc almost flat or with two low, hardly visible keels (vs. disk with two distinct prominent keels), hairs on the lip simple and branching (vs. hairs simple), and anther cap rectangular, semiglobular at apex (vs. anther cap conoid). The discriminative morphological characters of the mentioned species are presented in
<tableCitation id="3F4DBA9A7D6FFFB4770BFCD1FE15974F" box="[320,405,770,796]" captionStart="TABLE 3" captionStartId="4.[136,224,844,868]" captionText="TABLE 3. Morphological differences between Dendrobium namdongense, D. fuscifaucium, and D. hekouense." pageId="4" pageNumber="204">Table 3</tableCitation>
.
</paragraph>
<caption id="26B0DFA97D6FFFB476C3FC9FFAAA9737" box="[136,1322,844,868]" pageId="4" pageNumber="204" startId="4.[136,224,844,868]" targetType="table">
<paragraph id="72708F217D6FFFB476C3FC9FFAAA9737" blockId="4.[136,1322,844,868]" box="[136,1322,844,868]" pageId="4" pageNumber="204">
<emphasis id="40BB53337D6FFFB476C3FC9FFF7B9737" bold="true" box="[136,251,844,868]" pageId="4" pageNumber="204">TABLE 3.</emphasis>
Morphological differences between
<taxonomicName id="B5CFF4A27D6FFFB474C8FC9FFC239737" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[643,931,844,868]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D6FFFB474C8FC9FFC239737" box="[643,931,844,868]" italics="true" pageId="4" pageNumber="204">Dendrobium namdongense</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CFF4A27D6FFFB475FBFC9EFBD89737" box="[944,1112,844,868]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D6FFFB475FBFC9EFBD89737" box="[944,1112,844,868]" italics="true" pageId="4" pageNumber="204">D. fuscifaucium</emphasis>
</taxonomicName>
, and
<taxonomicName id="B5CFF4A27D6FFFB472D8FC9EFAA79737" box="[1171,1319,844,868]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="hekouense">D. hekouense</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="72708F217D6FFFB476D8FCA6FBFA9385" pageId="4" pageNumber="204">
<table id="00CF7D817D6F004F76D8FCA7FA0F9385" box="[147,1423,884,2006]" colsContinueIn="5.[147,1429,211,548]" gridcols="4" gridrows="17" pageId="4" pageNumber="204">
<tr id="CCFF8D637D6F004F76D8FCA7FA0F97D8" box="[147,1423,884,907]" gridrow="0" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FCA7FE3597D8" box="[147,437,884,907]" gridcol="0" gridrow="0" pageId="4" pageNumber="204">Characters</th>
<th id="8F2EE41F7D6F004F7780FCA7FD6897D8" box="[459,744,884,907]" gridcol="1" gridrow="0" pageId="4" pageNumber="204">
<taxonomicName id="B5CFF4A27D6FFFB47780FCA7FD5797D9" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[459,727,884,906]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D6FFFB47780FCA7FD5797D9" bold="true" box="[459,727,884,906]" italics="true" pageId="4" pageNumber="204">Dendrobium namdongense</emphasis>
</taxonomicName>
</th>
<th id="8F2EE41F7D6F004F7549FCA7FB9897D8" box="[770,1048,884,907]" gridcol="2" gridrow="0" pageId="4" pageNumber="204">
<taxonomicName id="B5CFF4A27D6FFFB47549FCA7FB8997D9" box="[770,1033,884,906]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D6FFFB47549FCA7FB8997D9" bold="true" box="[770,1033,884,906]" italics="true" pageId="4" pageNumber="204">Dendrobium fuscifaucium</emphasis>
</taxonomicName>
</th>
<th id="8F2EE41F7D6F004F7271FCA7FA0F97D8" box="[1082,1423,884,907]" gridcol="3" gridrow="0" pageId="4" pageNumber="204">
<taxonomicName id="B5CFF4A27D6FFFB47271FCA7FAA697D9" authority="Pseudobulb" authorityName="Pseudobulb" box="[1082,1318,884,906]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="204" phylum="Tracheophyta" rank="species" species="hekouense">
<emphasis id="40BB53337D6FFFB47271FCA7FAA697D9" bold="true" box="[1082,1318,884,906]" italics="true" pageId="4" pageNumber="204">Dendrobium hekouense</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="CCFF8D637D6F004F76D8FC4FFA0F9783" box="[147,1423,924,976]" gridrow="1" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FC4FFE359783" box="[147,437,924,976]" gridcol="0" gridrow="1" pageId="4" pageNumber="204">Pseudobulb shape</th>
<td id="8F2EE41F7D6F004F7780FC4FFD689783" box="[459,744,924,976]" gridcol="1" gridrow="1" pageId="4" pageNumber="204">narrowly ovate to broadly fusiform or ellipsoid</td>
<td id="8F2EE41F7D6F004F7549FC4FFB989783" box="[770,1048,924,976]" gridcol="2" gridrow="1" pageId="4" pageNumber="204">fusiform, narrowly ellipsoid</td>
<td id="8F2EE41F7D6F004F7271FC4FFA0F9783" box="[1082,1423,924,976]" gridcol="3" gridrow="1" pageId="4" pageNumber="204">sub-ellipsoid</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FC32FA0F9047" box="[147,1423,993,1044]" gridrow="2" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FC32FE359047" box="[147,437,993,1044]" gridcol="0" gridrow="2" pageId="4" pageNumber="204">Pseudobulb size</th>
<td id="8F2EE41F7D6F004F7780FC32FD689047" box="[459,744,993,1044]" gridcol="1" gridrow="2" pageId="4" pageNumber="204">5–9 mm tall, 2.5–4.5 mm in diameter</td>
<td id="8F2EE41F7D6F004F7549FC32FB989047" box="[770,1048,993,1044]" gridcol="2" gridrow="2" pageId="4" pageNumber="204">8–15 mm tall, 2–5 mm in diameter</td>
<td id="8F2EE41F7D6F004F7271FC32FA0F9047" box="[1082,1423,993,1044]" gridcol="3" gridrow="2" pageId="4" pageNumber="204">8–14 mm tall, 4–6 mm in diameter</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FBF5FA0F9025" box="[147,1423,1062,1142]" gridrow="3" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FBF5FE359025" box="[147,437,1062,1142]" gridcol="0" gridrow="3" pageId="4" pageNumber="204">Leaf lamina shape and texture</th>
<td id="8F2EE41F7D6F004F7780FBF5FD689025" box="[459,744,1062,1142]" gridcol="1" gridrow="3" pageId="4" pageNumber="204">narrowly ovate to broadly elliptic, coriaceous, blunt to rounded at apex</td>
<td id="8F2EE41F7D6F004F7549FBF5FB989025" box="[770,1048,1062,1142]" gridcol="2" gridrow="3" pageId="4" pageNumber="204">narrowly elliptic to oblong- elliptic, herbaceous, acute to round at apex</td>
<td id="8F2EE41F7D6F004F7271FBF5FA0F9025" box="[1082,1423,1062,1142]" gridcol="3" gridrow="3" pageId="4" pageNumber="204">Narrowly elliptic to oblong elliptic, coriaceous, acute</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FB5BFA0F90CD" box="[147,1423,1160,1182]" gridrow="4" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FB5BFE3590CD" box="[147,437,1160,1182]" gridcol="0" gridrow="4" pageId="4" pageNumber="204">Leaf lamina size</th>
<td id="8F2EE41F7D6F004F7780FB5BFD6890CD" box="[459,744,1160,1182]" gridcol="1" gridrow="4" pageId="4" pageNumber="204">6–13 × 3.5–5.5 mm</td>
<td id="8F2EE41F7D6F004F7549FB5BFB9890CD" box="[770,1048,1160,1182]" gridcol="2" gridrow="4" pageId="4" pageNumber="204">20–52 × 3.5–8 mm</td>
<td id="8F2EE41F7D6F004F7271FB5BFA0F90CD" box="[1082,1423,1160,1182]" gridcol="3" gridrow="4" pageId="4" pageNumber="204">about 20 × 3.5–6 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FB7CFA0F90B1" box="[147,1423,1199,1250]" gridrow="5" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FB7CFE3590B1" box="[147,437,1199,1250]" gridcol="0" gridrow="5" pageId="4" pageNumber="204">The number of inflorescences on individual pseudobulb</th>
<td id="8F2EE41F7D6F004F7780FB7CFD6890B1" box="[459,744,1199,1250]" gridcol="1" gridrow="5" pageId="4" pageNumber="204">1–2</td>
<td id="8F2EE41F7D6F004F7549FB7CFB9890B1" box="[770,1048,1199,1250]" gridcol="2" gridrow="5" pageId="4" pageNumber="204">1</td>
<td id="8F2EE41F7D6F004F7271FB7CFA0F90B1" box="[1082,1423,1199,1250]" gridcol="3" gridrow="5" pageId="4" pageNumber="204">1</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FB27FA0F9159" box="[147,1423,1268,1290]" gridrow="6" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FB27FE359159" box="[147,437,1268,1290]" gridcol="0" gridrow="6" pageId="4" pageNumber="204">Peduncle length</th>
<td id="8F2EE41F7D6F004F7780FB27FD689159" box="[459,744,1268,1290]" gridcol="1" gridrow="6" pageId="4" pageNumber="204">less than 1 mm long</td>
<td id="8F2EE41F7D6F004F7549FB27FB989159" box="[770,1048,1268,1290]" gridcol="2" gridrow="6" pageId="4" pageNumber="204">1–2 mm</td>
<td id="8F2EE41F7D6F004F7271FB27FA0F9159" box="[1082,1423,1268,1290]" gridcol="3" gridrow="6" pageId="4" pageNumber="204">4–4.5 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FAC8FA0F913F" box="[147,1423,1307,1388]" gridrow="7" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FAC8FE35913F" box="[147,437,1307,1388]" gridcol="0" gridrow="7" pageId="4" pageNumber="204">Sepals and petals color</th>
<td id="8F2EE41F7D6F004F7780FAC8FD68913F" box="[459,744,1307,1388]" gridcol="1" gridrow="7" pageId="4" pageNumber="204">White with distinct purple nerves, rarely with light yellow or pink tint</td>
<td id="8F2EE41F7D6F004F7549FAC8FB98913F" box="[770,1048,1307,1388]" gridcol="2" gridrow="7" pageId="4" pageNumber="204">Uniform pink to purple</td>
<td id="8F2EE41F7D6F004F7271FAC8FA0F913F" box="[1082,1423,1307,1388]" gridcol="3" gridrow="7" pageId="4" pageNumber="204">White to light yellowish green, sometimes flushed with purple, or with indistinct purple veins</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FAAEFA0F91E3" box="[147,1423,1405,1456]" gridrow="8" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FAAEFE3591E3" box="[147,437,1405,1456]" gridcol="0" gridrow="8" pageId="4" pageNumber="204">Median sepal shape and size</th>
<td id="8F2EE41F7D6F004F7780FAAEFD6891E3" box="[459,744,1405,1456]" gridcol="1" gridrow="8" pageId="4" pageNumber="204">oblong narrowly ovate, 6–8 × 4–4.5 mm</td>
<td id="8F2EE41F7D6F004F7549FAAEFB9891E3" box="[770,1048,1405,1456]" gridcol="2" gridrow="8" pageId="4" pageNumber="204">ovate, 6–6.5 × 5–5.5 mm</td>
<td id="8F2EE41F7D6F004F7271FAAEFA0F91E3" box="[1082,1423,1405,1456]" gridcol="3" gridrow="8" pageId="4" pageNumber="204">broadly ovate, 7–7.5 × 5.5–6 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FA12FA0F918B" box="[147,1423,1473,1496]" gridrow="9" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FA12FE35918B" box="[147,437,1473,1496]" gridcol="0" gridrow="9" pageId="4" pageNumber="204">Lateral sepal size</th>
<td id="8F2EE41F7D6F004F7780FA12FD68918B" box="[459,744,1473,1496]" gridcol="1" gridrow="9" pageId="4" pageNumber="204">7–8.5 × 10.5–12.5 mm</td>
<td id="8F2EE41F7D6F004F7549FA12FB98918B" box="[770,1048,1473,1496]" gridcol="2" gridrow="9" pageId="4" pageNumber="204">6.5–7.0 × 13–15 mm</td>
<td id="8F2EE41F7D6F004F7271FA12FA0F918B" box="[1082,1423,1473,1496]" gridcol="3" gridrow="9" pageId="4" pageNumber="204">11–12 × 13–15 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8FA3AFA0F924F" box="[147,1423,1513,1564]" gridrow="10" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8FA3AFE35924F" box="[147,437,1513,1564]" gridcol="0" gridrow="10" pageId="4" pageNumber="204">Petal shape and size</th>
<td id="8F2EE41F7D6F004F7780FA3AFD68924F" box="[459,744,1513,1564]" gridcol="1" gridrow="10" pageId="4" pageNumber="204">oblong narrowly obovate, 6–7.5 × 3.2–4.2 mm</td>
<td id="8F2EE41F7D6F004F7549FA3AFB98924F" box="[770,1048,1513,1564]" gridcol="2" gridrow="10" pageId="4" pageNumber="204">elliptic-ovate, 4.5–6 × 3.5–4 mm</td>
<td id="8F2EE41F7D6F004F7271FA3AFA0F924F" box="[1082,1423,1513,1564]" gridcol="3" gridrow="10" pageId="4" pageNumber="204">elliptic-obovate 7.5–8 × 4–4.5 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F9FEFA0F9217" box="[147,1423,1581,1604]" gridrow="11" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F9FEFE359217" box="[147,437,1581,1604]" gridcol="0" gridrow="11" pageId="4" pageNumber="204">Mentum size and shape</th>
<td id="8F2EE41F7D6F004F7780F9FEFD689217" box="[459,744,1581,1604]" gridcol="1" gridrow="11" pageId="4" pageNumber="204">9–11 × 5.5–6 mm, saccate</td>
<td id="8F2EE41F7D6F004F7549F9FEFB989217" box="[770,1048,1581,1604]" gridcol="2" gridrow="11" pageId="4" pageNumber="204">9–10 × 5.5–6 mm, saccate</td>
<td id="8F2EE41F7D6F004F7271F9FEFA0F9217" box="[1082,1423,1581,1604]" gridcol="3" gridrow="11" pageId="4" pageNumber="204">12–13 × 6–8 mm, cylindrical</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F986FA0F9238" box="[147,1423,1621,1643]" gridrow="12" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F986FE359238" box="[147,437,1621,1643]" gridcol="0" gridrow="12" pageId="4" pageNumber="204">Flattened lip shape and size</th>
<td id="8F2EE41F7D6F004F7780F986FD689238" box="[459,744,1621,1643]" gridcol="1" gridrow="12" pageId="4" pageNumber="204">11.5–14.5 mm long and wide</td>
<td id="8F2EE41F7D6F004F7549F986FB989238" box="[770,1048,1621,1643]" gridcol="2" gridrow="12" pageId="4" pageNumber="204">14–16 mm long and wide</td>
<td id="8F2EE41F7D6F004F7271F986FA0F9238" box="[1082,1423,1621,1643]" gridcol="3" gridrow="12" pageId="4" pageNumber="204">18–22 mm long and wide,</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F9AFFA0F929E" box="[147,1423,1660,1741]" gridrow="13" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F9AFFE35929E" box="[147,437,1660,1741]" gridcol="0" gridrow="13" pageId="4" pageNumber="204">Lip side lobes</th>
<td id="8F2EE41F7D6F004F7780F9AFFD68929E" box="[459,744,1660,1741]" gridcol="1" gridrow="13" pageId="4" pageNumber="204">obliquely semicircular, 3.5–4 × 6–7 mm, margin finely ciliate or papillulate</td>
<td id="8F2EE41F7D6F004F7549F9AFFB98929E" box="[770,1048,1660,1741]" gridcol="2" gridrow="13" pageId="4" pageNumber="204">obliquely triangular, 3.5–4 × 8–9 mm, margin fimbriate</td>
<td id="8F2EE41F7D6F004F7271F9AFFA0F929E" box="[1082,1423,1660,1741]" gridcol="3" gridrow="13" pageId="4" pageNumber="204">long-subtriangular, 5–5.5 × 13-14 mm, margin fimbriate</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F90DFA0F937C" box="[147,1423,1758,1839]" gridrow="14" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F90DFE35937C" box="[147,437,1758,1839]" gridcol="0" gridrow="14" pageId="4" pageNumber="204">Lip median lobe</th>
<td id="8F2EE41F7D6F004F7780F90DFD68937C" box="[459,744,1758,1839]" gridcol="1" gridrow="14" pageId="4" pageNumber="204">semicircular, entire or obscurely emarginate, 2.5– 3.5 × 4.5–6.5 mm</td>
<td id="8F2EE41F7D6F004F7549F90DFB98937C" box="[770,1048,1758,1839]" gridcol="2" gridrow="14" pageId="4" pageNumber="204">reniform, distinctly emarginate, 2.8–3 × 9.5–10 mm</td>
<td id="8F2EE41F7D6F004F7271F90DFA0F937C" box="[1082,1423,1758,1839]" gridcol="3" gridrow="14" pageId="4" pageNumber="204">reniform, distinctly emarginate, 3.5–5.5 × 7–7.5 mm</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F893FA0F93FD" box="[147,1423,1856,1966]" gridrow="15" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F893FE3593FD" box="[147,437,1856,1966]" gridcol="0" gridrow="15" pageId="4" pageNumber="204">Lip ornamentation</th>
<td id="8F2EE41F7D6F004F7780F893FD6893FD" box="[459,744,1856,1966]" gridcol="1" gridrow="15" pageId="4" pageNumber="204">long pubescent only on disc and median lobe, almost flat, or with two low hardly visible longitudinal keels</td>
<td id="8F2EE41F7D6F004F7549F893FB9893FD" box="[770,1048,1856,1966]" gridcol="2" gridrow="15" pageId="4" pageNumber="204">long pubescent on almost all adaxial surface, with two distinct lamellar longitudinal keels</td>
<td id="8F2EE41F7D6F004F7271F893FA0F93FD" box="[1082,1423,1856,1966]" gridcol="3" gridrow="15" pageId="4" pageNumber="204">finely pubescent on almost all adaxial surface, with two distinct convex longitudinal keels</td>
</tr>
<tr id="CCFF8D637D6F004F76D8F813FA0F9385" box="[147,1423,1984,2006]" gridrow="16" pageId="4" pageNumber="204">
<th id="8F2EE41F7D6F004F76D8F813FE359385" box="[147,437,1984,2006]" gridcol="0" gridrow="16" pageId="4" pageNumber="204">Hairs of lip indumentum</th>
<td id="8F2EE41F7D6F004F7780F813FD689385" box="[459,744,1984,2006]" gridcol="1" gridrow="16" pageId="4" pageNumber="204">simple and branching</td>
<td id="8F2EE41F7D6F004F7549F813FB989385" box="[770,1048,1984,2006]" gridcol="2" gridrow="16" pageId="4" pageNumber="204">simple</td>
<td id="8F2EE41F7D6F004F7271F813FA0F9385" box="[1082,1423,1984,2006]" gridcol="3" gridrow="16" pageId="4" pageNumber="204">simple</td>
</tr>
</table>
</paragraph>
<paragraph id="72708F217D6FFFB47232F83BFA2093AE" blockId="4.[1145,1440,2024,2045]" box="[1145,1440,2024,2045]" pageId="4" pageNumber="204">
<tableNote id="8F298EAF7D6FFFB47232F83BFA2093AE" box="[1145,1440,2024,2045]" pageId="4" pageNumber="204" targetBox="[147,1423,884,2006]" targetPageId="4">......continued on the next page</tableNote>
</paragraph>
<caption id="26B0DFA97D6EFFB576D8FF79FE0E9491" box="[147,398,170,194]" isContinuationCaption="true" pageId="5" pageNumber="205" startId="5.[147,235,170,194]" targetType="table">
<paragraph id="72708F217D6EFFB576D8FF79FE0E9491" blockId="5.[147,398,170,194]" box="[147,398,170,194]" pageId="5" pageNumber="205">
<emphasis id="40BB53337D6EFFB576D8FF79FE869491" bold="true" box="[147,262,170,194]" pageId="5" pageNumber="205">TABLE 3.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="72708F217D6EFFB576D8FF07FA899677" pageId="5" pageNumber="205">
<table id="00CF7D817D6E004F76D8FF00FA159677" box="[147,1429,211,548]" colsContinueFrom="4.[147,1423,884,2006]" gridcols="4" gridrows="6" pageId="5" pageNumber="205">
<tr id="CCFF8D637D6E004F76D8FF00FA1594B9" box="[147,1429,211,234]" gridrow="0" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FF00FE0094B9" box="[147,384,211,234]" gridcol="0" gridrow="0" pageId="5" pageNumber="205">Characters</th>
<th id="8F2EE41F7D6E004F7780FF00FD5794B9" box="[459,727,211,234]" gridcol="1" gridrow="0" pageId="5" pageNumber="205">
<taxonomicName id="B5CFF4A27D6EFFB57780FF00FD5794BA" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[459,727,211,233]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="205" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D6EFFB57780FF00FD5794BA" bold="true" box="[459,727,211,233]" italics="true" pageId="5" pageNumber="205">Dendrobium namdongense</emphasis>
</taxonomicName>
</th>
<th id="8F2EE41F7D6E004F7549FF00FB8994B9" box="[770,1033,211,234]" gridcol="2" gridrow="0" pageId="5" pageNumber="205">
<taxonomicName id="B5CFF4A27D6EFFB57549FF00FB8994BA" box="[770,1033,211,233]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="205" phylum="Tracheophyta" rank="species" species="fuscifaucium">
<emphasis id="40BB53337D6EFFB57549FF00FB8994BA" bold="true" box="[770,1033,211,233]" italics="true" pageId="5" pageNumber="205">Dendrobium fuscifaucium</emphasis>
</taxonomicName>
</th>
<th id="8F2EE41F7D6E004F7271FF00FA1594B9" box="[1082,1429,211,234]" gridcol="3" gridrow="0" pageId="5" pageNumber="205">
<taxonomicName id="B5CFF4A27D6EFFB57271FF00FAA694BA" authority="Column" authorityName="Column" box="[1082,1318,211,233]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="205" phylum="Tracheophyta" rank="species" species="hekouense">
<emphasis id="40BB53337D6EFFB57271FF00FAA694BA" bold="true" box="[1082,1318,211,233]" italics="true" pageId="5" pageNumber="205">Dendrobium hekouense</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="CCFF8D637D6E004F76D8FF28FA15957C" box="[147,1429,251,303]" gridrow="1" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FF28FE00957C" box="[147,384,251,303]" gridcol="0" gridrow="1" pageId="5" pageNumber="205">Column size and surface</th>
<td id="8F2EE41F7D6E004F7780FF28FD57957C" box="[459,727,251,303]" gridcol="1" gridrow="1" pageId="5" pageNumber="205">2.5–3 × 3–3.5 mm, finely papillulate on sides</td>
<td id="8F2EE41F7D6E004F7549FF28FB89957C" box="[770,1033,251,303]" gridcol="2" gridrow="1" pageId="5" pageNumber="205">2–2.4 × 2–2.5 mm, puberulent on sides</td>
<td id="8F2EE41F7D6E004F7271FF28FA15957C" box="[1082,1429,251,303]" gridcol="3" gridrow="1" pageId="5" pageNumber="205">about 3 mm tall and wide, smooth</td>
</tr>
<tr id="CCFF8D637D6E004F76D8FE93FA159520" box="[147,1429,320,371]" gridrow="2" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FE93FE009520" box="[147,384,320,371]" gridcol="0" gridrow="2" pageId="5" pageNumber="205">Column foot</th>
<td id="8F2EE41F7D6E004F7780FE93FD579520" box="[459,727,320,371]" gridcol="1" gridrow="2" pageId="5" pageNumber="205">9–9.5 × 5–5.5 mm wide in broadest part</td>
<td id="8F2EE41F7D6E004F7549FE93FB899520" box="[770,1033,320,371]" gridcol="2" gridrow="2" pageId="5" pageNumber="205">7–8 × 5–5.5 mm wide in broadest part</td>
<td id="8F2EE41F7D6E004F7271FE93FA159520" box="[1082,1429,320,371]" gridcol="3" gridrow="2" pageId="5" pageNumber="205">13–14 × 6–7.5 mm wide in broadest part</td>
</tr>
<tr id="CCFF8D637D6E004F76D8FE56FA159586" box="[147,1429,389,469]" gridrow="3" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FE56FE009586" box="[147,384,389,469]" gridcol="0" gridrow="3" pageId="5" pageNumber="205">Anther cap</th>
<td id="8F2EE41F7D6E004F7780FE56FD579586" box="[459,727,389,469]" gridcol="1" gridrow="3" pageId="5" pageNumber="205">rectangular, semiglobular at apex, light green, front margin white</td>
<td id="8F2EE41F7D6E004F7549FE56FB899586" box="[770,1033,389,469]" gridcol="2" gridrow="3" pageId="5" pageNumber="205">conoid, light green, front margin white</td>
<td id="8F2EE41F7D6E004F7271FE56FA159586" box="[1082,1429,389,469]" gridcol="3" gridrow="3" pageId="5" pageNumber="205">conoid, gray green, front margin purple</td>
</tr>
<tr id="CCFF8D637D6E004F76D8FE34FA1595AE" box="[147,1429,487,509]" gridrow="4" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FE34FE0095AE" box="[147,384,487,509]" gridcol="0" gridrow="4" pageId="5" pageNumber="205">Flowering period</th>
<td id="8F2EE41F7D6E004F7780FE34FD5795AE" box="[459,727,487,509]" gridcol="1" gridrow="4" pageId="5" pageNumber="205">April–May</td>
<td id="8F2EE41F7D6E004F7549FE34FB8995AE" box="[770,1033,487,509]" gridcol="2" gridrow="4" pageId="5" pageNumber="205">July</td>
<td id="8F2EE41F7D6E004F7271FE34FA1595AE" box="[1082,1429,487,509]" gridcol="3" gridrow="4" pageId="5" pageNumber="205">August–September</td>
</tr>
<tr id="CCFF8D637D6E004F76D8FDDDFA159677" box="[147,1429,526,548]" gridrow="5" pageId="5" pageNumber="205">
<th id="8F2EE41F7D6E004F76D8FDDDFE009677" box="[147,384,526,548]" gridcol="0" gridrow="5" pageId="5" pageNumber="205">Distribution</th>
<td id="8F2EE41F7D6E004F7780FDDDFD579677" box="[459,727,526,548]" gridcol="1" gridrow="5" pageId="5" pageNumber="205">N Vietnam</td>
<td id="8F2EE41F7D6E004F7549FDDDFB899677" box="[770,1033,526,548]" gridcol="2" gridrow="5" pageId="5" pageNumber="205">central Laos?</td>
<td id="8F2EE41F7D6E004F7271FDDDFA159677" box="[1082,1429,526,548]" gridcol="3" gridrow="5" pageId="5" pageNumber="205">SE China, N Vietnam</td>
</tr>
</table>
</paragraph>
<paragraph id="72708F217D6EFFB576F6FD87FB559715" blockId="5.[136,1452,596,838]" pageId="5" pageNumber="205">
<emphasis id="40BB53337D6EFFB576F6FD87FEE7963D" bold="true" box="[189,359,596,622]" pageId="5" pageNumber="205">Conservation:</emphasis>
—According to the available data,
<taxonomicName id="B5CFF4A27D6EFFB574BCFD87FBA7963D" authorityName="Aver. &amp; C. H. Nguyen" authorityYear="2024" box="[759,1063,596,622]" class="Liliopsida" family="Orchidaceae" genus="Dendrobium" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="205" phylum="Tracheophyta" rank="species" species="namdongense">
<emphasis id="40BB53337D6EFFB574BCFD87FBA7963D" box="[759,1063,596,622]" italics="true" pageId="5" pageNumber="205">Dendrobium namdongense</emphasis>
</taxonomicName>
is an extremely rare orchid. Until now, only one tiny wild population with an extent of occurrence (EOO) and an area of occupancy (AOO) less than
<quantity id="B53722C47D6EFFB573CAFDABFF2C96EB" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="205" unit="km" value="100.0">100 km</quantity>
² and
<quantity id="B53722C47D6EFFB576A6FD4FFEB596EB" box="[237,309,668,696]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="205" unit="km" value="10.0">10 km</quantity>
² respectively, has been confirmed. According to our field observation, the population likely includes less than 50 colonies, almost all of which are of flowering maturity. The species is speculated to have ornamental value and thus highly desired in the unofficial wild plant market. It is possible that the species can quickly turn extinct in the wild due to overcollection due to poaching and illegal trade. According to the IUCN Red List (2024) criteria, we establish its status as Critically Endangered, CR A3(d); B1ab(i–v)+2ab(i-v); C1a(i–ii)+2(i–ii); D1.
</paragraph>
</subSubSection>
</treatment>
</document>
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