139 lines
19 KiB
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139 lines
19 KiB
XML
<document id="8881FFB183319D442DC54916B97BD94F" ID-DOI="10.5281/zenodo.274221" ID-GBIF-Dataset="10609322-6a33-4d2f-ad00-f0d99c62a792" ID-ISSN="1175-5326" ID-Zenodo-Dep="274221" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459836727268" checkinUser="plazi" docAuthor="Woodley, Norman E." docDate="2008" docId="FA60FE397755311893A96871FE9DF775" docLanguage="en" docName="zt01746p045.pdf" docOrigin="Zootaxa 1746" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Kerteszmyia Woodley, 2008, gen. nov." docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="0659864177553119933E6F56FFDBF069" masterDocTitle="Kerteszmyia, a new genus of Pachygastrinae from the Neotropical Region (Diptera: Stratiomyidae)" masterLastPageNumber="45" masterPageNumber="39" pageNumber="39" updateTime="1698223771803" updateUser="plazi">
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<mods:titleInfo id="9C3C638486A71D2468A14834B717112A">
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<mods:title id="0CEC2C60773E5819BA4C8CB1AA8175F3">Kerteszmyia, a new genus of Pachygastrinae from the Neotropical Region (Diptera: Stratiomyidae)</mods:title>
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<mods:name id="868EDEBA451434FC3DCDDDA89C4942F9" type="personal">
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<mods:roleTerm id="7E8E077A14ED552B5E65942DFF6BF2DF">Author</mods:roleTerm>
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<mods:namePart id="BF935148BE14E762F1AEE20B8FA1ADA0">Woodley, Norman E.</mods:namePart>
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<mods:typeOfResource id="92347FC796F880D54943C1B236D3A5A0">text</mods:typeOfResource>
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<mods:title id="29D74F9DC2CA169C51644C098555B7F6">Zootaxa</mods:title>
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<mods:part id="86C7A357B82D4A9FBF783251DDB3BA4F">
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<mods:date id="F243FE02DA44CB686092B0725A9F5202">2008</mods:date>
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<mods:detail id="CD56568C46D5E3233B17768F22B7BF36" type="volume">
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<mods:number id="288587FFE433BECB249060DBC923999A">1746</mods:number>
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<mods:start id="84C588DC287A9820059FB6AA93527689">39</mods:start>
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<mods:end id="0A5A3E8334E25842AD931A20D7B8823E">45</mods:end>
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<mods:classification id="728B3CA09F0F5D2D9C35D2BA3EE3D2DD">journal article</mods:classification>
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<mods:identifier id="013CE7FFBEB0313F380A521427A0B75A" type="DOI">10.5281/zenodo.274221</mods:identifier>
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<mods:identifier id="229AFC4F87CCA12C5234AF2478623D0B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="FA60FE397755311893A96871FE9DF775" ID-DOI="http://doi.org/10.5281/zenodo.6230243" ID-GBIF-Taxon="119359158" ID-Zenodo-Dep="6230243" LSID="urn:lsid:plazi:treatment:FA60FE397755311893A96871FE9DF775" httpUri="http://treatment.plazi.org/id/FA60FE397755311893A96871FE9DF775" lastPageId="1" lastPageNumber="40" pageId="0" pageNumber="39">
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<subSubSection id="3AD31CA47755311993A96871FE45F728" box="[151,414,1831,1857]" pageId="0" pageNumber="39" type="nomenclature">
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<paragraph id="72764F2F7755311993A96871FE45F728" blockId="0.[151,414,1831,1857]" box="[151,414,1831,1857]" pageId="0" pageNumber="39">
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<heading id="293EF8437755311993A96871FE45F728" bold="true" box="[151,414,1831,1857]" fontSize="11" level="1" pageId="0" pageNumber="39" reason="1">
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<emphasis id="40BD933D7755311993A96871FE45F728" bold="true" box="[151,414,1831,1857]" pageId="0" pageNumber="39">
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<taxonomicName id="B5C934AC7755311993A96871FEF1F728" ID-CoL="583G" box="[151,298,1831,1857]" class="Insecta" family="Stratiomyidae" genus="Kerteszmyia" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="40BD933D7755311993A96871FEF1F728" bold="true" box="[151,298,1831,1857]" italics="true" pageId="0" pageNumber="39">Kerteszmyia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5B8E2E4677553119920F6871FE45F728" box="[305,414,1831,1857]" pageId="0" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3AD31CA47755311993A96824FC1DF7E5" box="[151,966,1906,1932]" pageId="0" pageNumber="39" type="materials_examined">
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<paragraph id="72764F2F7755311993A96824FC1DF7E5" blockId="0.[151,1436,1906,2012]" box="[151,966,1906,1932]" pageId="0" pageNumber="39">
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<typeStatus id="AD72F18D7755311993A96824FF0AF7E5" box="[151,209,1906,1932]" pageId="0" pageNumber="39">Type</typeStatus>
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species,
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<taxonomicName id="B5C934AC7755311992026825FD99F7E5" box="[316,578,1907,1932]" class="Insecta" family="Stratiomyidae" genus="Kerteszmyia" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="ecuadora" status="sp. nov.">
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<emphasis id="40BD933D7755311992026825FD99F7E5" box="[316,578,1907,1932]" italics="true" pageId="0" pageNumber="39">Kerteszmyia ecuadora</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5B8E2E467755311991746824FD7CF7E5" box="[586,679,1906,1932]" pageId="0" pageNumber="39" rank="species">
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<emphasis id="40BD933D7755311991746824FD7CF7E5" bold="true" box="[586,679,1906,1932]" pageId="0" pageNumber="39">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="3AD31CA47755311893F868CCFC5BF143" lastPageId="1" lastPageNumber="40" pageId="0" pageNumber="39" type="diagnosis">
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<paragraph id="72764F2F7755311893F868CCFC5BF143" blockId="0.[151,1436,1906,2012]" lastBlockId="1.[151,1437,152,1820]" lastPageId="1" lastPageNumber="40" pageId="0" pageNumber="39">
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<emphasis id="40BD933D7755311993F868CCFE9DF7DD" bold="true" box="[198,326,1946,1972]" pageId="0" pageNumber="39">Diagnosis.</emphasis>
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This distinctive genus can be easily separated from other Neotropical pachygastrines that have four scutellar spines by the characteristic antennal flagellum that is kidney-shaped, distinctly higher than long, with a subapical arista-like style that is inserted on the dorsal one-third of the flagellar complex (
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<figureCitation id="EAF253AA7754311896256FCEFA55F0DB" box="[1307,1422,152,178]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Figs. 1–2</figureCitation>
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). All other genera with scutellar spines have the antennal flagellar complex as long as high or much longer. Other characters useful in recognition are the densely hairy eyes (
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<figureCitation id="EAF253AA7754311890966FBEFBC1F16B" box="[936,1050,232,258]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Figs. 1–4</figureCitation>
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), the head distinctly higher than long (
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<figureCitation id="EAF253AA7754311893E56E46FE97F143" box="[219,332,272,298]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Figs. 1–2</figureCitation>
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), and the short abdomen that is wider than long.
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</paragraph>
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</subSubSection>
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<subSubSection id="3AD31CA47754311893F86E61FEC7F425" pageId="1" pageNumber="40" type="description">
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<paragraph id="72764F2F7754311893F86E61FB64F2FB" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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<emphasis id="40BD933D7754311893F86E61FE84F138" bold="true" box="[198,351,311,337]" pageId="1" pageNumber="40">Description.</emphasis>
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<emphasis id="40BD933D77543118925B6E6EFE71F138" box="[357,426,312,337]" italics="true" pageId="1" pageNumber="40">Male.</emphasis>
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Head (
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<figureCitation id="EAF253AA7754311892C06E6EFDB7F13B" box="[510,620,312,338]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Figs. 1, 3</figureCitation>
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) with eye very large, occupying most of anterior and lateral head surface, eyes narrowly separated; upper frons reduced to a very narrow strip that is 0.06 width of head at narrowest point (
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<figureCitation id="EAF253AA7754311892306EDEFE8CF1CB" box="[270,343,392,418]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
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); lower frons gradually diverging toward oral margin; face reduced to an extremely narrow strip below antennae, antennae thus inserted very close to oral margin. Ocellar tubercle slightly prominent. Gena only slightly visible in lateral view, very narrow, about width of two ommatidia; occiput not visible in lateral view. Eye very densely haired, ommatidia uniform in size. Antenna short, scape slightly longer than pedicel, the latter very slightly produced on inner side; flagellum bilaterally flattened, kidney-shaped, about 2.5 times higher than long, last flagellomere an arista-form style that is inserted on the upper third of flagellar complex. Palpus two-segmented, first segment shorter than second, second segment ovoid.
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</paragraph>
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<paragraph id="72764F2F7754311893F86DF6FAB4F333" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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Thorax with scutum convex; scutellum forming angle with scutum (not in same plane), rounded posteriorly with distinct margin, armed with 4 marginal spines, the lateral spines about two-thirds length of medial spines and more slender; subscutellum slightly developed, largely bare, with only very sparse scattered tomentum. Legs unremarkable, without significant modifications. Wing (
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<figureCitation id="EAF253AA7754311890C86C4EFBE6F35B" box="[1014,1085,792,818]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1941,1965]" captionTargetBox="[361,1225,186,1914]" captionTargetId="figure@3.[361,1225,186,1914]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Kerteszmyia ecuadora, morphological features. 1, male head, lateral view. 2, female head, lateral view. 3, male head anterodorsal view. 4, female head, anterodorsal view. 5, female scutum, dorsal view. 6, female abdomen, posterior view. 7, female right wing, dorsal view." httpUri="https://zenodo.org/record/274222/files/figure.png" pageId="1" pageNumber="40">Fig. 7</figureCitation>
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) with microtrichia present on darkened areas, and scattered on posterior portion; R2+3 originating well beyond crossvein r-m; R4 present.
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</paragraph>
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<paragraph id="72764F2F7754311893F86C3CFD3CF3C5" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">Abdomen short and convex, wider than thorax, approximately 1.7 times wider than long, posterior half of tergite 4 and all of 5 nearly vertical in lateral view.</paragraph>
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<paragraph id="72764F2F7754311893F86CEDFCC5F395" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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<emphasis id="40BD933D7754311893F86CEDFEFDF3BD" box="[198,294,955,980]" italics="true" pageId="1" pageNumber="40">Female.</emphasis>
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Differs from male as follows: Head with eye slightly smaller than in male, upper frons wider, gradually diverging ventrally, 0.14–0.18 width of head.
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</paragraph>
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<paragraph id="72764F2F7754311893F86B5CFEC7F425" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">Abdomen with cercus two-segmented, segments subequal in length and diameter, second segment elongate-ovoid.</paragraph>
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</subSubSection>
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<subSubSection id="3AD31CA47754311893F86B0CFC2DF4AD" pageId="1" pageNumber="40" type="etymology">
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<paragraph id="72764F2F7754311893F86B0CFC2DF4AD" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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<emphasis id="40BD933D7754311893F86B0CFE88F41D" bold="true" box="[198,339,1114,1140]" pageId="1" pageNumber="40">Etymology.</emphasis>
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This genus is named after Kalman Kertész (
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<date id="067769EF7754311890546B0CFC2FF41D" box="[874,1012,1114,1140]" pageId="1" pageNumber="40" value="1867" valueMax="1922">1867–1922</date>
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), a Hungarian dipterist who was an excellent taxonomist working on
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<taxonomicName id="B5C934AC7754311891176BD4FD16F4F5" box="[553,717,1154,1180]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="family">Stratiomyidae</taxonomicName>
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among other families. He published several comprehensive and important papers on
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<taxonomicName id="B5C934AC7754311892846BFCFDB5F4AD" box="[442,622,1194,1220]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Pachygastrinae">Pachygastrinae</taxonomicName>
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, including a key to world genera.
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</paragraph>
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</subSubSection>
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<subSubSection id="3AD31CA47754311893F86B84FE9DF775" pageId="1" pageNumber="40" type="discussion">
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<paragraph id="72764F2F7754311893F86B84FA40F50D" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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<emphasis id="40BD933D7754311893F86B84FE9BF485" bold="true" box="[198,320,1234,1260]" pageId="1" pageNumber="40">Remarks.</emphasis>
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<taxonomicName id="B5C934AC7754311892766B85FE02F485" box="[328,473,1235,1260]" class="Insecta" family="Stratiomyidae" genus="Kerteszmyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="40BD933D7754311892766B85FE02F485" box="[328,473,1235,1260]" italics="true" pageId="1" pageNumber="40">Kerteszmyia</emphasis>
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</taxonomicName>
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has a general appearance that is different from most other Neotropical genera of pachygastrines with scutellar spines due to its smaller size; less elongate thorax; and short, wide abdomen with strongly declivitous apical segments. To the naked eye it is similar to genera such as
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<taxonomicName id="B5C934AC7754311896366A75FF2AF50D" authority="Kertesz" authorityName="Kertesz" class="Insecta" family="Stratiomyidae" genus="Popanomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="40BD933D7754311896366A75FA47F555" box="[1288,1436,1315,1340]" italics="true" pageId="1" pageNumber="40">Popanomyia</emphasis>
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Kertész
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</taxonomicName>
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and similar medium-sized black taxa, but these lack scutellar spines and do not have darkened wings.
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</paragraph>
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<paragraph id="72764F2F7754311893F86A24FC1EF63D" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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No phylogenetic work has been done on the large, species-rich subfamily
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<taxonomicName id="B5C934AC77543118976B6A24FADDF5E5" box="[1109,1286,1394,1420]" class="Insecta" family="Stratiomyidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Pachygastrinae">Pachygastrinae</taxonomicName>
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on a global scale. The character state that unites the genera discussed here, having two or more scutellar spines, is plesiomorphic relative to many genera in the subfamily that have no scutellar spines (see
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<bibRefCitation id="165832DE77543118974A6A94FAFFF5B5" author="Woodley" box="[1140,1316,1474,1500]" pageId="1" pageNumber="45" refString="Woodley, N. E. (2001) A world catalog of Stratiomyidae (Insecta: Diptera). Myia, 11, (8) + 1 - 475." type="journal article" year="2001">Woodley 2001</bibRefCitation>
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: 5–6 for a general discussion of this character for the family). Thus, it is possible that the species with scutellar spines form a paraphyletic or polyphyletic assemblage within the pachygastrines depending on how those without spines evolved and how many times loss of scutellar spines occurred.
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</paragraph>
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<paragraph id="72764F2F7754311893F86934FE9DF775" blockId="1.[151,1437,152,1820]" pageId="1" pageNumber="40">
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Due to lack of phylogenetic work, it is not possible to discuss the relationship of
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<taxonomicName id="B5C934AC77543118974E6935FADAF615" box="[1136,1281,1635,1660]" class="Insecta" family="Stratiomyidae" genus="Kerteszmyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="40BD933D77543118974E6935FADAF615" box="[1136,1281,1635,1660]" italics="true" pageId="1" pageNumber="40">Kerteszmyia</emphasis>
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</taxonomicName>
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to other genera with scutellar spines. However, some features that the genus exhibits, such as the short, tall head and small compact body form are similar to those found in some genera that lack scutellar spines. This possibly indicates that
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<taxonomicName id="B5C934AC775431189231698DFE7BF69D" box="[271,416,1755,1780]" class="Insecta" family="Stratiomyidae" genus="Kerteszmyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="40BD933D775431189231698DFE7BF69D" box="[271,416,1755,1780]" italics="true" pageId="1" pageNumber="40">Kerteszmyia</emphasis>
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</taxonomicName>
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is more derived among the taxa with scutellar spines, but only a more rigorous study can verify this.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |