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<document id="CD6E068C040511191CBCC9A138046123" ID-CLB-Dataset="8048" ID-DOI="10.11646/zootaxa.4877.1.4" ID-GBIF-Dataset="a4126ee5-4b17-474c-b96a-fa4662e0c653" ID-ISSN="1175-5326" ID-Zenodo-Dep="4423887" ID-ZooBank="7F5AF885-F6F2-4A65-B0A7-CBE943787943" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610026657991" checkinUser="plazi" docAuthor="Gaviria-Ortiz, Fabian Guillermo, Dolibaina, Diego Rodrigo, Carneiro, Eduardo, Warren, Andrew D., Mielke, Olaf Hermann Hendrik &amp; Casagrande, Mirna Martins" docDate="2020" docId="FA5DEF439F65A07CFF497830FCE41490" docLanguage="en" docName="zootaxa.4877.1.4.pdf" docOrigin="Zootaxa 4877 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sodalia spangla Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande 2020, comb. nov." docType="treatment" docVersion="9" lastPageNumber="122" masterDocId="0664973B9F74A068FFDE7E04FF961336" masterDocTitle="A review of the Neotropical skipper genus Sodalia Evans, 1955 (Lepidoptera Hesperiidae: Hesperiinae) with the description of a new species" masterLastPageNumber="124" masterPageNumber="102" pageNumber="119" updateTime="1698923397729" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="209BFF76EFABAB727DA30E37B79099F5">A review of the Neotropical skipper genus Sodalia Evans, 1955 (Lepidoptera Hesperiidae: Hesperiinae) with the description of a new species</mods:title>
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<mods:namePart id="58EC13F9470FFF9AD4B8A6CB22A9791F">Gaviria-Ortiz, Fabian Guillermo</mods:namePart>
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<mods:namePart id="D44B2CDF42D3DC401DFF6E13140C624A">Carneiro, Eduardo</mods:namePart>
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<mods:affiliation id="31F594F5B3AD5D18D2162D54D8A4639B">eduardo. carneiro @ ufpr. br; https: // orcid. org / 0000 - 0002 - 4796 - 0020</mods:affiliation>
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<mods:affiliation id="D3B746D7AE98C9EA2CBD364FCFE8CC10">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, U. S. A. awhesp @ gmail. com; https: // orcid. org / 0000 - 0002 - 6730 - 1938</mods:affiliation>
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<mods:namePart id="BA859460A0196E8A8AC233B7602BC34D">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<mods:affiliation id="D555391442CE240368A6EC7B89B36CBB">omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606</mods:affiliation>
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<treatment id="FA5DEF439F65A07CFF497830FCE41490" ID-DOI="http://doi.org/10.5281/zenodo.4560505" ID-GBIF-Taxon="179907942" ID-Zenodo-Dep="4560505" LSID="urn:lsid:plazi:treatment:FA5DEF439F65A07CFF497830FCE41490" httpUri="http://treatment.plazi.org/id/FA5DEF439F65A07CFF497830FCE41490" lastPageId="20" lastPageNumber="122" pageId="17" pageNumber="119">
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<paragraph id="724B5E559F65A079FF497830FD011579" blockId="17.[151,663,1588,1652]" box="[151,663,1588,1615]" pageId="17" pageNumber="119">
<heading id="2903E9399F65A079FF497830FD011579" bold="true" box="[151,663,1588,1615]" fontSize="11" level="1" pageId="17" pageNumber="119" reason="1">
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<taxonomicName id="B5F425D69F65A079FF497830FD9F1578" authority="(Evans, 1955)" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[151,521,1588,1615]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="119" phylum="Arthropoda" rank="species" species="spangla" status="comb. nov.">
<emphasis id="408082479F65A079FF497830FECF1579" bold="true" box="[151,345,1588,1615]" italics="true" pageId="17" pageNumber="119">Sodalia spangla</emphasis>
(
<bibRefCitation id="166523A49F65A079FEB67830FD971578" author="Evans, W. H." box="[360,513,1588,1614]" pageId="17" pageNumber="119" refId="ref24541" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
)
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<taxonomicNameLabel id="5BB33F3C9F65A079FDCE7831FD011579" box="[528,663,1589,1615]" pageId="17" pageNumber="119" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="3AEE0DDE9F65A079FF49785DFE701542" box="[151,486,1625,1652]" pageId="17" pageNumber="119" type="description">
<paragraph id="724B5E559F65A079FF49785DFE701542" blockId="17.[151,663,1588,1652]" box="[151,486,1625,1652]" pageId="17" pageNumber="119">
<figureCitation id="EACF42D09F65A079FF49785DFE811545" box="[151,279,1625,1651]" captionStart="FIGURES 118" captionStartId="13.[151,264,1337,1361]" captionTargetBox="[157,1433,184,1312]" captionTargetId="figure-26@13.[151,1436,181,1313]" captionTargetPageId="13" captionText="FIGURES 118. Habitus of species of Sodalia, males (left) and females (right). 14. S. sodalis, 12. Male from Santo Antônio do Tauá, Pará, Brazil (DZ 35.537), 34. Female from Cacaulândia, Rôndonia, Brazil (OM 21.494). 58. S. argyrospila, 56. Male from Antonina, Paraná, Brazil (OM 14.223), 78. Female from Pontal do Paraná, Paraná, Brazil (OM 25.358). 912. S. coler, 910. Male from Pontal do Paraná, Paraná, Brazil (DZ 35.842), 1112. Female from Seara, Santa Catarina, Brazil (DZ 35.696). 1316. S. petiti sp. nov., 1314. Male holotype from Alluriquin, Ecuador (OM 52.619). 1516. Female allotype from Alluriguin, Ecuador (DZ 31.487). 1718. S. spangla comb. nov., male from Oso Perdido, Amazonas, Peru (OM 83.945). Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4423891" httpUri="https://zenodo.org/record/4423891/files/figure.png" pageId="17" pageNumber="119">Figs 1718</figureCitation>
,
<figureCitation id="EACF42D09F65A079FEFC785DFED61545" box="[290,320,1625,1651]" captionStart="FIGURES 1923" captionStartId="14.[151,264,1165,1189]" captionTargetBox="[152,1436,452,1136]" captionTargetId="figure-140@14.[151,1436,452,1141]" captionTargetPageId="14" captionText="FIGURES 1923. Head of species of Sodalia in antero-dorsal view. 19. S. sodalis, male from Santo Antônio do Tauá, Pará, Brazil (DZ 31.331). 20. S. argyrospila, male from Antonina, Paraná, Brazil (OM 14.223). 21. S. coler, male from Agrolândia, Santa Catarina, Brazil (DZ 35.352). 22. S. petiti sp. nov., male holotype. 23. S. spangla comb. nov., male from Oso Perdido, Amazonas, Peru (OM 83.945). Arrows indicate areas of frons with iridescent bluish-green. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4423893" httpUri="https://zenodo.org/record/4423893/files/figure.png" pageId="17" pageNumber="119">23</figureCitation>
,
<figureCitation id="EACF42D09F65A079FE92785DFEFC1545" box="[332,362,1625,1651]" captionStart="FIGURES 2428" captionStartId="14.[151,264,1895,1919]" captionTargetBox="[152,1436,1327,1871]" captionTargetId="figure-224@14.[151,1436,1321,1874]" captionTargetPageId="14" captionText="FIGURES 2428. Head and thorax of species of Sodalia in dorsal view. 24. S. sodalis, male from Yavita, Amazonas, Venezuela (OM 39.809). 25. S. argyrospila, male from Joinville, Santa Catarina, Brazil (DZ 35.654). 26. S. coler, male from Pontal do Paraná, Paraná, Brazil (DZ 35.842). 27. S. petiti sp. nov., male holotype. 28. S. spangla comb. nov., male from Oso Perdido, Amazonas, Peru (OM 83.945). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4423895" httpUri="https://zenodo.org/record/4423895/files/figure.png" pageId="17" pageNumber="119">28</figureCitation>
,
<figureCitation id="EACF42D09F65A079FEA8785DFE021545" box="[374,404,1625,1651]" captionStart="FIGURES 2933" captionStartId="15.[151,264,1877,1901]" captionTargetBox="[152,1436,1317,1853]" captionTargetId="figure-528@15.[151,1436,1317,1853]" captionTargetPageId="15" captionText="FIGURES 2933. Forewing brands of male Sodalia. 29. S. sodalis, from Yavita, Amazonas, Venezuela (OM 39.809). 30. S. argyrospila, from Joinville, Santa Catarina, Brazil (DZ 35.654). 31. S. coler, from Pontal do Paraná, Paraná, Brazil (DZ 35.842). 32. S. petiti sp. nov., paratype from Tinalândia, Alluriquin, Santo Domingo de los Tsáchilas, Ecuador (OM 52.619). 33. S. spangla comb. nov., from Oso Perdido, Amazonas, Peru (OM 83.945). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4423897" httpUri="https://zenodo.org/record/4423897/files/figure.png" pageId="17" pageNumber="119">33</figureCitation>
,
<figureCitation id="EACF42D09F65A079FE7E785DFE281545" box="[416,446,1625,1651]" captionStart="FIGURES 3638" captionStartId="17.[151,264,1374,1398]" captionTargetBox="[151,1436,181,1350]" captionTargetId="figure-26@17.[151,1436,181,1350]" captionTargetPageId="17" captionText="FIGURES 3638. Male genitalia of Sodalia. 36. S. coler, from Pelotas, Rio Grande do Sul, Brazil (DZ 396). 37. S. petiti sp. nov., paratype from Santo Domingo de los Colorados, Santo Domingo de los Tsáchilas, Ecuador (OM 33.708). 38. S. spangla comb. nov., from Oso Perdido, Amazonas, Peru (OM 83.645). Structures and views as described in the legend of figures 3435. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4423903" httpUri="https://zenodo.org/record/4423903/files/figure.png" pageId="17" pageNumber="119">38</figureCitation>
,
<figureCitation id="EACF42D09F65A079FE14785DFE701542" box="[458,486,1625,1652]" captionStart="FIGURE 45" captionStartId="21.[151,250,1120,1144]" captionTargetBox="[157,1429,185,1090]" captionTargetId="figure-26@21.[151,1436,181,1096]" captionTargetPageId="21" captionText="FIGURE 45. Geographic distribution of Sodalia petiti sp. nov. and Sodalia splanga comb. nov." figureDoi="http://doi.org/10.5281/zenodo.4423913" httpUri="https://zenodo.org/record/4423913/files/figure.png" pageId="17" pageNumber="119">45</figureCitation>
</paragraph>
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<paragraph id="724B5E559F65A079FF497899FB551427" blockId="17.[151,1437,1693,1840]" pageId="17" pageNumber="119">
<taxonomicName id="B5F425D69F65A079FF497899FE7D1583" ID-CoL="43S9T" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[151,491,1693,1717]" class="Insecta" family="Hesperiidae" genus="Mnasitheus" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="119" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F65A079FF497899FEF51583" box="[151,355,1693,1717]" italics="true" pageId="17" pageNumber="119">Mnasitheus spangla</emphasis>
<bibRefCitation id="166523A49F65A079FEB4789AFE7D1583" author="Evans, W. H." box="[362,491,1693,1717]" pageId="17" pageNumber="119" refId="ref24541" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
.
<emphasis id="408082479F65A079FE2B7899FD541583" bold="true" box="[501,706,1693,1717]" pageId="17" pageNumber="119">Cat. Amer. Hesp. 4</emphasis>
, p. 146, pl. 63 (male gen.); [holo]
<typeStatus id="AD4FE0F79F65A079FBC47899FBDE1583" box="[1050,1096,1693,1717]" pageId="17" pageNumber="119">type</typeStatus>
male,
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, [
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,]
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,
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<collectorName id="DF013B839F65A079FE1878B8FDD015E5" box="[454,582,1724,1747]" pageId="17" pageNumber="119">F. M. Brown</collectorName>
leg.; BM(NH).—
<collectorName id="DF013B839F65A079FD2778B8FCDD15E2" box="[761,843,1724,1748]" pageId="17" pageNumber="119">Bridges</collectorName>
, 1983.
<emphasis id="408082479F65A079FC4D78B8FB8F15E2" bold="true" box="[915,1049,1724,1748]" pageId="17" pageNumber="119">
<collectorName id="DF013B839F65A079FC4D78B8FB9115E2" box="[915,1031,1724,1748]" pageId="17" pageNumber="119">Lep. Hesp.</collectorName>
1
</emphasis>
, p. 111;
<emphasis id="408082479F65A079FBB178B8FBEA15E2" bold="true" box="[1135,1148,1724,1748]" pageId="17" pageNumber="119">2</emphasis>
, p. 21.—
<collectorName id="DF013B839F65A079FB0978B8FABC15E2" box="[1239,1322,1724,1748]" pageId="17" pageNumber="119">Bridges</collectorName>
, 1988.
<collectorName id="DF013B839F65A079FAAD78B8FE9415C4" pageId="17" pageNumber="119">Cat. Hesp.</collectorName>
1, p. 176; 2, p. 34.—
<collectorName id="DF013B839F65A079FE3E78DFFDB115C5" box="[480,551,1755,1779]" pageId="17" pageNumber="119">Lamas</collectorName>
, 2003.
<emphasis id="408082479F65A079FDAD78DEFCF415C4" bold="true" box="[627,866,1754,1779]" pageId="17" pageNumber="119">
<collectorName id="DF013B839F65A079FDAD78DEFCF415C4" box="[627,866,1754,1779]" pageId="17" pageNumber="119">Marip. Machu Picchu</collectorName>
</emphasis>
, p. 190.—
<collectorName id="DF013B839F65A079FC1178DFFB8D15C5" box="[975,1051,1755,1779]" pageId="17" pageNumber="119">Mielke</collectorName>
, 2004.
<collectorName id="DF013B839F65A079FBB978DFFB7915C5" box="[1127,1263,1755,1779]" pageId="17" pageNumber="119">Hesperioidea</collectorName>
, p. 72,
<emphasis id="408082479F65A079FAE278DFFAC715C5" bold="true" box="[1340,1361,1755,1779]" italics="true" pageId="17" pageNumber="119">in</emphasis>
<collectorName id="DF013B839F65A079FA8778DFFA0B15C4" box="[1369,1437,1755,1778]" pageId="17" pageNumber="119">Lamas</collectorName>
(ed.).
<emphasis id="408082479F65A079FEDD78FDFC951427" bold="true" box="[259,771,1785,1809]" pageId="17" pageNumber="119">
<collectorName id="DF013B839F65A079FEDD78FDFEFA1427" box="[259,364,1785,1809]" pageId="17" pageNumber="119">Checklist</collectorName>
:
<collectorName id="DF013B839F65A079FEA978FDFE311427" box="[375,423,1785,1809]" pageId="17" pageNumber="119">Part</collectorName>
4A,
<collectorName id="DF013B839F65A079FE0778FDFC951427" box="[473,771,1785,1809]" pageId="17" pageNumber="119">Hesperioidea-Papilionoidea</collectorName>
</emphasis>
,
<emphasis id="408082479F65A079FCCE78FEFCB21427" box="[784,804,1786,1809]" italics="true" pageId="17" pageNumber="119">in</emphasis>
<collectorName id="DF013B839F65A079FCF478FEFC171427" box="[810,897,1786,1809]" pageId="17" pageNumber="119">Heppner</collectorName>
(ed.).
<emphasis id="408082479F65A079FC1D78FDFB2B1427" bold="true" box="[963,1213,1785,1809]" pageId="17" pageNumber="119">
<collectorName id="DF013B839F65A079FC1D78FDFB0E1427" box="[963,1176,1785,1809]" pageId="17" pageNumber="119">Atlas Neotrop. Lep.</collectorName>
5A
</emphasis>
.
</paragraph>
<paragraph id="724B5E559F65A079FF49791CFD491406" blockId="17.[151,1437,1693,1840]" box="[151,735,1816,1840]" pageId="17" pageNumber="119">
(no genus)
<taxonomicName id="B5F425D69F65A079FED7791CFECF1406" ID-CoL="43S9T" authorityName="Evans" authorityYear="1955" box="[265,345,1816,1840]" class="Insecta" family="Hesperiidae" genus="Mnasitheus" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="119" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F65A079FED7791CFECF1406" box="[265,345,1816,1840]" italics="true" pageId="17" pageNumber="119">spangla</emphasis>
</taxonomicName>
; Beattie, 1976.
<emphasis id="408082479F65A079FE25791CFD131406" bold="true" box="[507,645,1816,1840]" pageId="17" pageNumber="119">Rhop. Direct</emphasis>
., p. 262.
</paragraph>
</subSubSection>
<subSubSection id="3AEE0DDE9F65A079FF49795AFD2B14AA" pageId="17" pageNumber="119" type="diagnosis">
<paragraph id="724B5E559F65A079FF49795AFD2B14AA" blockId="17.[151,1436,1886,2021]" pageId="17" pageNumber="119">
<emphasis id="408082479F65A079FF49795AFE84144E" bold="true" box="[151,274,1886,1912]" pageId="17" pageNumber="119">Diagnosis.</emphasis>
<taxonomicName id="B5F425D69F65A079FEC4795AFE44144E" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[282,466,1886,1912]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="119" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F65A079FEC4795AFE44144E" box="[282,466,1886,1912]" italics="true" pageId="17" pageNumber="119">Sodalia spangla</emphasis>
</taxonomicName>
is the smallest species of
<taxonomicName id="B5F425D69F65A079FCDF795AFCC1144E" authorityName="Evans" authorityYear="1955" box="[769,855,1886,1912]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="408082479F65A079FCDF795AFCC1144E" box="[769,855,1886,1912]" italics="true" pageId="17" pageNumber="119">Sodalia</emphasis>
</taxonomicName>
and the only one with the dorsal and ventral wing surfaces completely speckled with yellow scales.
</paragraph>
</subSubSection>
<subSubSection id="3AEE0DDE9F65A07BFF1979A2FCEF11FB" lastPageId="19" lastPageNumber="121" pageId="17" pageNumber="119" type="description">
<paragraph id="724B5E559F65A079FF1979A2FA0A14D3" blockId="17.[151,1436,1886,2021]" pageId="17" pageNumber="119">
<emphasis id="408082479F65A079FF1979A2FEFB14F6" bold="true" box="[199,365,1958,1984]" pageId="17" pageNumber="119">Redescription</emphasis>
.
<emphasis id="408082479F65A079FEA479A2FE2114F6" box="[378,439,1958,1984]" italics="true" pageId="17" pageNumber="119">Head</emphasis>
: Mostly brown mixed with yellow, frons with a patch of short, compact, iridescent bluishgreen scales. Labial palpus mostly dark brown and yellow. Antenna brown, ventrally yellow at base of segments;
</paragraph>
<paragraph id="724B5E559F66A07AFF497E93FD8013E3" blockId="18.[151,1436,151,249]" pageId="18" pageNumber="120">club brown, with yellow ring at base; apiculus shorter than club, ventrally with a yellow line; nudum of 11 segments, dark brown, restricted to apiculus.</paragraph>
<paragraph id="724B5E559F66A07AFF197EDBFC4E13CF" blockId="18.[151,1436,151,249]" box="[199,984,223,249]" pageId="18" pageNumber="120">
<emphasis id="408082479F66A07AFF197EDBFE8E13CF" box="[199,280,223,249]" italics="true" pageId="18" pageNumber="120">Thorax</emphasis>
: Entirely brown and yellow including patagia, tegula and legs.
</paragraph>
<caption id="268B0EDD9F66A07AFF497914FE62148E" ID-DOI="http://doi.org/10.5281/zenodo.4423905" ID-Zenodo-Dep="4423905" httpUri="https://zenodo.org/record/4423905/files/figure.png" pageId="18" pageNumber="120" startId="18.[151,264,1808,1832]" targetBox="[151,1435,278,1783]" targetPageId="18">
<paragraph id="724B5E559F66A07AFF497914FE62148E" blockId="18.[151,1437,1808,1977]" pageId="18" pageNumber="120">
<emphasis id="408082479F66A07AFF497914FEC1141E" bold="true" box="[151,343,1808,1832]" pageId="18" pageNumber="120">FIGURES 3942.</emphasis>
Female genitalia of
<taxonomicName id="B5F425D69F66A07AFDEE7914FDEB141E" authorityName="Evans" authorityYear="1955" box="[560,637,1808,1832]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="408082479F66A07AFDEE7914FDEB141E" box="[560,637,1808,1832]" italics="true" pageId="18" pageNumber="120">Sodalia</emphasis>
</taxonomicName>
. 39.
<taxonomicName id="B5F425D69F66A07AFD6B7915FC8E141E" baseAuthorityName="Butler" baseAuthorityYear="1877" box="[693,792,1808,1832]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="120" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="408082479F66A07AFD6B7915FC8E141E" box="[693,792,1808,1832]" italics="true" pageId="18" pageNumber="120">S. sodalis</emphasis>
</taxonomicName>
, from Tingo Maria, Huánuco, Peru (DZ 31.675). 40.
<taxonomicName id="B5F425D69F66A07AFA957915FF48147A" baseAuthorityName="Mabille" baseAuthorityYear="1876" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="120" phylum="Arthropoda" rank="species" species="argyrospila">
<emphasis id="408082479F66A07AFA957915FF48147A" italics="true" pageId="18" pageNumber="120">S. argyrospila</emphasis>
</taxonomicName>
, from Paranaguá, Paraná, Brazil (OM 15.992). 41.
<taxonomicName id="B5F425D69F66A07AFD3B7931FCA4147A" baseAuthorityName="Schaus" baseAuthorityYear="1902" box="[741,818,1844,1868]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="120" phylum="Arthropoda" rank="species" species="coler">
<emphasis id="408082479F66A07AFD3B7931FCA4147A" box="[741,818,1844,1868]" italics="true" pageId="18" pageNumber="120">S. coler</emphasis>
</taxonomicName>
, from Seara, Santa Catarina, Brazil (DZ 35.306). 42.
<taxonomicName id="B5F425D69F66A07AFA917931FA0A147A" authority="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande, 2020" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" box="[1359,1436,1845,1868]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="120" phylum="Arthropoda" rank="species" species="petiti" status="sp. nov.">
<emphasis id="408082479F66A07AFA917931FA0A147A" box="[1359,1436,1845,1868]" italics="true" pageId="18" pageNumber="120">S. petiti</emphasis>
</taxonomicName>
<emphasis id="408082479F66A07AFF49795DFF7F1447" bold="true" box="[151,233,1881,1905]" pageId="18" pageNumber="120">
<taxonomicNameLabel id="5BB33F3C9F66A07AFF49795DFF7F1447" box="[151,233,1881,1905]" pageId="18" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype from La Palma, Alluriquin, Santo Domingo de los Tsáchilas, Ecuador (DZ 35.375). Top to bottom: lateral view of tergum VIII, posterior apophysis, stergima, papilla analis and part of the ductus bursae; ventral view of the sterigma and bursa copulatrix. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="724B5E559F67A07BFF197E93FE4B1277" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF197E93FE9D1386" box="[199,267,151,176]" italics="true" pageId="19" pageNumber="121">Wings</emphasis>
: Forewing length
<quantity id="B50CF3B09F67A07BFE037E93FDDA1387" box="[477,588,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="19" pageNumber="121" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
. DFW and DHW dark brown speckled with yellow; DFW with two brands over and under CuA
<subScript id="EE705C109F67A07BFEA37EC9FE1313ED" attach="left" box="[381,389,205,219]" fontSize="6" pageId="19" pageNumber="121">2</subScript>
, covered by dark brown scales, the former longer than in all other species of the genus, with base only slightly wider than tip, bordering CuA, the latter brand narrower and shorter than former, bordering CuA
<subScript id="EE705C109F67A07BFA537EF5FA0313C9" attach="left" box="[1421,1429,241,255]" fontSize="6" pageId="19" pageNumber="121">2</subScript>
. VFW dark brown, yellow speckling denser than on DFW; VHW as VFW but yellow speckling denser. Fringe of both wings yellowish brown.
</paragraph>
<paragraph id="724B5E559F67A07BFF197F4FFABA1253" blockId="19.[151,1437,151,933]" box="[199,1324,331,357]" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF197F4FFEA71253" box="[199,305,331,357]" italics="true" pageId="19" pageNumber="121">Abdomen</emphasis>
: Dorsally dark brown, densely speckled with yellow scales; ventrally as dorsally but paler.
</paragraph>
<paragraph id="724B5E559F67A07BFF197F6BFCEF11FB" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF197F6BFEFD12BF" box="[199,363,367,393]" italics="true" pageId="19" pageNumber="121">Male genitalia</emphasis>
: Tegumen more than two times wider than long in dorsal view, proximal margin with a short indentation. Fenestra semicircular, almost reaching distal margin of uncus in dorsal view. Saccus slightly longer than tegumen. Uncus more than twice as long as tegumen; distal margin of base with a wide, shallow indentation; arms parallel to tegumen, about six times longer than base, narrowed distally to a sharp ventrally bent tip. Gnathos arms about 2/3 length of uncus, markedly narrowed distally to an upturned tip. Valva 2.5 longer than wide, base narrow, widening distally; ampulla 2/3 width of valva, wider distally, distal margin rounded; harpe longer than ampulla, with a sharp, serrated and upcurved distal projection, slightly curved anteriorly. Fultura inferior the smallest of the genus, lateral sides dorsally projected, base with an anterior and another, short, ventral rounded projection. Aedeagus about 90% as long as valva, straight in dorsal and slightly concave in lateral view; distal opening restricted to tip; aedeagus distal margin with a short median dorsal indentation; no cornutus.
</paragraph>
</subSubSection>
<subSubSection id="3AEE0DDE9F67A07BFF197CD3FBF3106B" pageId="19" pageNumber="121" type="materials_examined">
<paragraph id="724B5E559F67A07BFF197CD3FBF3106B" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="121">
<materialsCitation id="C29C54089F67A07BFF197CD3FBB31023" ID-GBIF-Occurrence="3042108371" collectionCode="NHMUK" location="Rio Blanco" pageId="19" pageNumber="121" specimenCount="2" typeStatus="holotype">
<emphasis id="408082479F67A07BFF197CD3FEE211C6" bold="true" box="[199,372,726,753]" pageId="19" pageNumber="121">
<typeStatus id="AD4FE0F79F67A07BFF197CD3FE9411C7" box="[199,258,727,753]" pageId="19" pageNumber="121">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="B5F425D69F67A07BFEA47CD3FD7C11C7" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[378,746,726,753]" class="Insecta" family="Hesperiidae" genus="Mnasitheus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="121" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F67A07BFEA47CD3FDCC11C7" box="[378,602,727,753]" italics="true" pageId="19" pageNumber="121">Mnasitheus spangla</emphasis>
<bibRefCitation id="166523A49F67A07BFDBE7CD3FD7C11C7" author="Evans, W. H." box="[608,746,726,753]" pageId="19" pageNumber="121" refId="ref24541" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
was described based on the [holo]
<typeStatus id="AD4FE0F79F67A07BFBBB7CD2FB0E11C7" box="[1125,1176,726,753]" pageId="19" pageNumber="121">type</typeStatus>
male from
<location id="772B088E9F67A07BFAC97CD3FA0E11C7" LSID="urn:lsid:plazi:treatment:FA5DEF439F65A07CFF497830FCE41490:772B088E9F67A07BFAC97CD3FA0E11C7" box="[1303,1432,727,753]" name="Rio Blanco" pageId="19" pageNumber="121">Rio Blanco</location>
, Ecuador. This specimen is deposited at the
<collectionCode id="14E5C6909F67A07BFDA97CFFFD771023" box="[631,737,763,789]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="121" type="Museum">NHMUK</collectionCode>
bearing the following labels
</materialsCitation>
: /
<materialsCitation id="C29C54089F67A07BFBE27CFFFD2D106B" ID-GBIF-Occurrence="3042108313" collectingDate="1949-05" collectionCode="PHOTO" country="Ecuador" elevation="1700" location="Rio Blanco" pageId="19" pageNumber="121" specimenCount="1" typeStatus="holotype">
<typeStatus id="AD4FE0F79F67A07BFBE27CFFFBE21023" box="[1084,1140,763,789]" pageId="19" pageNumber="121">Type</typeStatus>
/ 432 /
<collectingCountry id="0AE31EC59F67A07BFB167CFFFAC21023" box="[1224,1364,763,789]" name="Ecuador" pageId="19" pageNumber="121">ECUADOR</collectingCountry>
:
<collectorName id="DF013B839F67A07BFA837CFFFE16100F" pageId="19" pageNumber="121">Cord. Oriental E. Foothills</collectorName>
[, Morona Santinado,]
<location id="772B088E9F67A07BFD547D1BFC9C100F" LSID="urn:lsid:plazi:treatment:FA5DEF439F65A07CFF497830FCE41490:772B088E9F67A07BFD547D1BFC9C100F" box="[650,778,799,825]" country="Ecuador" name="Rio Blanco" pageId="19" pageNumber="121">Rio Blanco</location>
<quantity id="B50CF3B09F67A07BFCCC7D1BFCFA100F" box="[786,876,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="121" unit="m" value="1700.0">
<elevation id="F9D9B9669F67A07BFCCC7D1BFCFA100F" box="[786,876,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="121" unit="m" value="1700.0">1700 m</elevation>
</quantity>
.
<date id="064A78959F67A07BFCA67D1BFC7A100F" box="[888,1004,799,825]" pageId="19" pageNumber="121" value="1949-05">
<collectingDate id="160E817D9F67A07BFCA67D1BFC7A100F" box="[888,1004,799,825]" pageId="19" pageNumber="121" value="1949-05">May 1949</collectingDate>
</date>
<collectorName id="DF013B839F67A07BFC2A7D1BFB13100F" box="[1012,1157,799,825]" pageId="19" pageNumber="121">F. M. Brown</collectorName>
/
<collectorName id="DF013B839F67A07BFB407D1BFA84100F" box="[1182,1298,799,825]" pageId="19" pageNumber="121">Brit. Mus.</collectorName>
1950-369. / accidentally damaged after description /
<collectionCode id="14E5C6909F67A07BFD807D47FD2D106B" box="[606,699,835,861]" pageId="19" pageNumber="121">PHOTO</collectionCode>
</materialsCitation>
AA /
<collectionCode id="14E5C6909F67A07BFD237D47FCC2106B" box="[765,852,835,861]" pageId="19" pageNumber="121">BMNH</collectionCode>
(E)# 806018 /
<collectionCode id="14E5C6909F67A07BFC2D7D47FBF7106B" box="[1011,1121,835,861]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="121" type="Museum">NHMUK</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="3AEE0DDE9F67A07BFF197D62FE961093" pageId="19" pageNumber="121" type="distribution">
<paragraph id="724B5E559F67A07BFF197D62FE961093" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF197D62FEF710B6" bold="true" box="[199,353,870,896]" pageId="19" pageNumber="121">Distribution.</emphasis>
This species is distributed at mid to high elevations of the eastern Andes of
<collectingCountry id="0AE31EC59F67A07BFB0F7D63FAB810B7" box="[1233,1326,871,897]" name="Ecuador" pageId="19" pageNumber="121">Ecuador</collectingCountry>
and
<collectingCountry id="0AE31EC59F67A07BFAB77D63FA0A10B7" box="[1385,1436,871,897]" name="Peru" pageId="19" pageNumber="121">Peru</collectingCountry>
(
<figureCitation id="EACF42D09F67A07BFF7E7D8FFF621093" box="[160,244,906,933]" captionStart="FIGURE 45" captionStartId="21.[151,250,1120,1144]" captionTargetBox="[157,1429,185,1090]" captionTargetId="figure-26@21.[151,1436,181,1096]" captionTargetPageId="21" captionText="FIGURE 45. Geographic distribution of Sodalia petiti sp. nov. and Sodalia splanga comb. nov." figureDoi="http://doi.org/10.5281/zenodo.4423913" httpUri="https://zenodo.org/record/4423913/files/figure.png" pageId="19" pageNumber="121">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="268B0EDD9F67A07BFF497963FC021449" ID-DOI="http://doi.org/10.5281/zenodo.4423907" ID-Zenodo-Dep="4423907" box="[151,916,1895,1919]" httpUri="https://zenodo.org/record/4423907/files/figure.png" pageId="19" pageNumber="121" startId="19.[151,250,1895,1919]" targetBox="[168,1422,964,1855]" targetPageId="19">
<paragraph id="724B5E559F67A07BFF497963FC021449" blockId="19.[151,916,1895,1919]" box="[151,916,1895,1919]" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF497963FEB61449" bold="true" box="[151,288,1895,1919]" pageId="19" pageNumber="121">FIGURE 43.</emphasis>
Geographic distribution of
<taxonomicName id="B5F425D69F67A07BFDE77963FD431449" baseAuthorityName="Butler" baseAuthorityYear="1877" box="[569,725,1895,1919]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="121" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="408082479F67A07BFDE77963FD431449" box="[569,725,1895,1919]" italics="true" pageId="19" pageNumber="121">Sodalia sodalis</emphasis>
</taxonomicName>
and
<taxonomicName id="B5F425D69F67A07BFCD87963FC181449" baseAuthorityName="Schaus" baseAuthorityYear="1902" box="[774,910,1895,1919]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="121" phylum="Arthropoda" rank="species" species="coler">
<emphasis id="408082479F67A07BFCD87963FC181449" box="[774,910,1895,1919]" italics="true" pageId="19" pageNumber="121">Sodalia coler</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="3AEE0DDE9F67A07CFF1979A9FCE41490" lastPageId="20" lastPageNumber="122" pageId="19" pageNumber="121" type="discussion">
<paragraph id="724B5E559F67A07CFF1979A9FDA212BF" blockId="19.[151,1436,1965,2028]" lastBlockId="20.[151,1437,151,393]" lastPageId="20" lastPageNumber="122" pageId="19" pageNumber="121">
<emphasis id="408082479F67A07BFF1979A9FED914F1" bold="true" box="[199,335,1965,1991]" pageId="19" pageNumber="121">Comments.</emphasis>
<taxonomicName id="B5F425D69F67A07BFE8879A9FD9D14F1" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[342,523,1965,1991]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="121" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F67A07BFE8879A9FD9D14F1" box="[342,523,1965,1991]" italics="true" pageId="19" pageNumber="121">Sodalia spangla</emphasis>
</taxonomicName>
is the smallest species of the genus and the only one with wings densely speckled with yellow scales, an atypical pattern among skippers. This color pattern is also found, albeit to a lesser extent, in the Andean endemic
<taxonomicName id="B5F425D69F60A07CFE5E7E93FE431386" authority="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande, 2020" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" box="[384,469,151,177]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="petiti" status="sp. nov.">
<emphasis id="408082479F60A07CFE5E7E93FE431386" box="[384,469,151,177]" italics="true" pageId="20" pageNumber="122">S. petiti</emphasis>
</taxonomicName>
<emphasis id="408082479F60A07CFE057E93FDA21387" bold="true" box="[475,564,151,177]" pageId="20" pageNumber="122">
<taxonomicNameLabel id="5BB33F3C9F60A07CFE057E93FDA21387" box="[475,564,151,177]" pageId="20" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The range of
<taxonomicName id="B5F425D69F60A07CFD0A7E93FCDE1387" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[724,840,151,177]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F60A07CFD0A7E93FCDE1387" box="[724,840,151,177]" italics="true" pageId="20" pageNumber="122">S. spangla</emphasis>
</taxonomicName>
extends along a narrow zone of elfin forest in the Andes of
<collectingCountry id="0AE31EC59F60A07CFF3E7EBFFEA913E3" box="[224,319,187,213]" name="Ecuador" pageId="20" pageNumber="122">Ecuador</collectingCountry>
,
<collectingCountry id="0AE31EC59F60A07CFE977EBFFEEA13E3" box="[329,380,187,213]" name="Peru" pageId="20" pageNumber="122">Peru</collectingCountry>
and possibly
<collectingCountry id="0AE31EC59F60A07CFDCB7EBFFD1513E3" box="[533,643,187,213]" name="Colombia" pageId="20" pageNumber="122">Colombia</collectingCountry>
(which needs further confirmation), where it is believed to be endemic, in areas at least between
<quantity id="B50CF3B09F60A07CFE727EDBFDD113CF" box="[428,583,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="20" pageNumber="122" unit="m" value="2600.0" valueMax="3000.0" valueMin="2200.0">22003000 m</quantity>
elevation, although two specimens are known from the
<typeStatus id="AD4FE0F79F60A07CFB667EDBFB7E13CF" box="[1208,1256,223,249]" pageId="20" pageNumber="122">type</typeStatus>
locality at
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. At the MGCL there are seven males from
<collectingCountry id="0AE31EC59F60A07CFD467F07FD63122B" box="[664,757,259,285]" name="Ecuador" pageId="20" pageNumber="122">Ecuador</collectingCountry>
labeled by Lafebre from the NW slope of Volcan
<collectingRegion id="B03090B79F60A07CFAF07F07FA0A122B" box="[1326,1436,259,285]" country="Ecuador" name="Pichincha" pageId="20" pageNumber="122">Pichincha</collectingRegion>
(
<collectingRegion id="B03090B79F60A07CFF417F23FEED1274" box="[159,379,295,322]" country="Ecuador" name="Pichincha" pageId="20" pageNumber="122">Pichincha Province</collectingRegion>
) at
<quantity id="B50CF3B09F60A07CFE7B7F22FE6A1277" box="[421,508,294,321]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="122" unit="m" value="3500.0">3500 m</quantity>
, and a male labeled from the Cordillera Cotacachi (
<collectingRegion id="B03090B79F60A07CFBE37F23FA801274" box="[1085,1302,295,322]" country="Ecuador" name="Imbabura" pageId="20" pageNumber="122">Imbabura Province</collectingRegion>
) at
<quantity id="B50CF3B09F60A07CFA9E7F22FA0E1277" box="[1344,1432,294,321]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="20" pageNumber="122" unit="m" value="3750.0">3750 m</quantity>
. Given that various Lafebre specimens from
<collectingCountry id="0AE31EC59F60A07CFD527F4FFD7F1253" box="[652,745,331,357]" name="Ecuador" pageId="20" pageNumber="122">Ecuador</collectingCountry>
are known to be mislabeled, the occurrence of
<taxonomicName id="B5F425D69F60A07CFAD97F4FFAEB1253" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1287,1405,331,357]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F60A07CFAD97F4FFAEB1253" box="[1287,1405,331,357]" italics="true" pageId="20" pageNumber="122">S. spangla</emphasis>
</taxonomicName>
as high as
<quantity id="B50CF3B09F60A07CFF307F6AFED512BF" box="[238,323,366,393]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="20" pageNumber="122" unit="m" value="3750.0">3750 m</quantity>
requires verification.
</paragraph>
<caption id="268B0EDD9F60A07CFF497B48FC911652" ID-DOI="http://doi.org/10.5281/zenodo.4423911" ID-Zenodo-Dep="4423911" box="[151,775,1356,1381]" httpUri="https://zenodo.org/record/4423911/files/figure.png" pageId="20" pageNumber="122" startId="20.[151,250,1356,1380]" targetBox="[158,1430,424,1327]" targetPageId="20">
<paragraph id="724B5E559F60A07CFF497B48FC911652" blockId="20.[151,775,1356,1381]" box="[151,775,1356,1381]" pageId="20" pageNumber="122">
<emphasis id="408082479F60A07CFF497B48FEB61652" bold="true" box="[151,288,1356,1380]" pageId="20" pageNumber="122">FIGURE 44.</emphasis>
Geographic distribution of
<taxonomicName id="B5F425D69F60A07CFDE77B49FC971653" baseAuthorityName="Mabille" baseAuthorityYear="1876" box="[569,769,1357,1381]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="argyrospila">
<emphasis id="408082479F60A07CFDE77B49FC971653" box="[569,769,1357,1381]" italics="true" pageId="20" pageNumber="122">Sodalia argyrospila</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="724B5E559F60A07CFF197B97FD0D159F" blockId="20.[151,1437,1427,1958]" pageId="20" pageNumber="122">
This species was originally placed by
<bibRefCitation id="166523A49F60A07CFDAE7B97FC931698" author="Evans, W. H." box="[624,773,1427,1454]" pageId="20" pageNumber="122" refId="ref24541" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
in the genus
<taxonomicName id="B5F425D69F60A07CFC477B97FB8C169B" authorityName="Godman" authorityYear="1900" box="[921,1050,1427,1453]" class="Insecta" family="Hesperiidae" genus="Mnasitheus" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="408082479F60A07CFC477B97FB8C169B" box="[921,1050,1427,1453]" italics="true" pageId="20" pageNumber="122">Mnasitheus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="B5F425D69F60A07CFB457B97FA86169B" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1179,1296,1427,1453]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F60A07CFB457B97FA86169B" box="[1179,1296,1427,1453]" italics="true" pageId="20" pageNumber="122">S. spangla</emphasis>
</taxonomicName>
differs from most of the species included in that genus by the presence of a transverse brand over CuA
<subScript id="EE705C109F60A07CFB5F7BCDFB1F16E1" attach="left" box="[1153,1161,1481,1495]" fontSize="6" pageId="20" pageNumber="122">2</subScript>
, the broadly divided uncus, and coecum of the aedeagus of similar thickness as the rest of the structure, with the opening of the ejaculatory bulb almost as long as half the length of the aedeagus. Most of the species included in
<taxonomicName id="B5F425D69F60A07CFBB47BFBFB7D152F" authorityName="Godman" authorityYear="1900" box="[1130,1259,1535,1561]" class="Insecta" family="Hesperiidae" genus="Mnasitheus" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="408082479F60A07CFBB47BFBFB7D152F" box="[1130,1259,1535,1561]" italics="true" pageId="20" pageNumber="122">Mnasitheus</emphasis>
</taxonomicName>
have a longitudinal or sagitated brand over CuA
<subScript id="EE705C109F60A07CFDC77831FDB71575" attach="left" box="[537,545,1589,1603]" fontSize="6" pageId="20" pageNumber="122">2</subScript>
, a slightly divided uncus, and the coecum of the aedeagus is distinctly thinner than the rest of the structure. Therefore, the characters above support the combination of
<taxonomicName id="B5F425D69F60A07CFB527843FAD51557" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1164,1347,1607,1633]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F60A07CFB527843FAD51557" box="[1164,1347,1607,1633]" italics="true" pageId="20" pageNumber="122">Sodalia spangla</emphasis>
</taxonomicName>
(
<emphasis id="408082479F60A07CFA8A7843FF5E15B3" bold="true" pageId="20" pageNumber="122">comb. nov.</emphasis>
). Although the female of
<taxonomicName id="B5F425D69F60A07CFE32786FFDF415B3" authorityName="Gaviria-Ortiz &amp; Dolibaina &amp; Carneiro &amp; Warren &amp; Mielke &amp; Casagrande" authorityYear="2020" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[492,610,1643,1669]" class="Insecta" family="Hesperiidae" genus="Sodalia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="122" phylum="Arthropoda" rank="species" species="spangla">
<emphasis id="408082479F60A07CFE32786FFDF415B3" box="[492,610,1643,1669]" italics="true" pageId="20" pageNumber="122">S. spangla</emphasis>
</taxonomicName>
remains unknown, it is expected that the morphology of its genitalia will reinforce the new combination proposed here.
</paragraph>
<paragraph id="724B5E559F60A07CFF1978B7FCE41490" blockId="20.[151,1437,1427,1958]" pageId="20" pageNumber="122">
<emphasis id="408082479F60A07CFF1978B7FE2015FB" bold="true" box="[199,438,1715,1741]" pageId="20" pageNumber="122">
<collectingRegion id="B03090B79F60A07CFF1978B7FED615FB" box="[199,320,1715,1741]" pageId="20" pageNumber="122">Examined</collectingRegion>
material.
</emphasis>
<materialsCitation id="C29C54089F60A07CFE6178B7FEA415C7" ID-GBIF-Occurrence="3042108718" collectingDate="1971-11" collectionCode="R, MCGL" collectorName="de Lafebre" country="Ecuador" elevation="3750" location="Imbabura" pageId="20" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Imbabura">
<collectingCountry id="0AE31EC59F60A07CFE6178B7FDD015FB" box="[447,582,1715,1741]" name="Ecuador" pageId="20" pageNumber="122">ECUADOR</collectingCountry>
:
<emphasis id="408082479F60A07CFD8778B7FD5115FB" box="[601,711,1715,1741]" italics="true" pageId="20" pageNumber="122">
<collectingRegion id="B03090B79F60A07CFD8778B7FD5115FB" box="[601,711,1715,1741]" country="Ecuador" name="Imbabura" pageId="20" pageNumber="122">Imbabura</collectingRegion>
</emphasis>
<emphasis id="408082479F60A07CFD1978B7FC7715FB" bold="true" box="[711,993,1715,1741]" pageId="20" pageNumber="122">—Cordillera Cotacachi</emphasis>
,
<quantity id="B50CF3B09F60A07CFC2F78B7FBDC15FB" box="[1009,1098,1715,1742]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="20" pageNumber="122" unit="m" value="3750.0">
<elevation id="F9D9B9669F60A07CFC2F78B7FBDC15FB" box="[1009,1098,1715,1742]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="20" pageNumber="122" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="064A78959F60A07CFB8678B7FB2115FB" box="[1112,1207,1715,1741]" pageId="20" pageNumber="122" value="1971-11">
<collectingDate id="160E817D9F60A07CFB8678B7FB2115FB" box="[1112,1207,1715,1741]" pageId="20" pageNumber="122" value="1971-11">XI.1971</collectingDate>
</date>
,
<collectionCode id="14E5C6909F60A07CFB1B78B7FB4D15FB" box="[1221,1243,1715,1741]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="122">R</collectionCode>
.
<collectorName id="DF013B839F60A07CFB3678B7FAF515FB" box="[1256,1379,1715,1741]" pageId="20" pageNumber="122">de Lafebre</collectorName>
leg.,
<specimenCount id="64F295DC9F60A07CFF4978D3FF5415C7" box="[151,194,1751,1777]" pageId="20" pageNumber="122" type="male">1 ♂</specimenCount>
(
<collectionCode id="14E5C6909F60A07CFF0C78D3FEBF15C7" box="[210,297,1751,1777]" pageId="20" pageNumber="122">MCGL</collectionCode>
)
</materialsCitation>
.
<emphasis id="408082479F60A07CFEE378D3FE5115C7" box="[317,455,1751,1777]" italics="true" pageId="20" pageNumber="122">Pichincha—</emphasis>
old road from Quito to coast,
<materialsCitation id="C29C54089F60A07CFCC478D3FD9A1423" ID-GBIF-Occurrence="3042108711" collectingDate="2005-10-15" collectionCode="MCGL" collectorName="J. D. Turner" country="Ecuador" elevation="2835" latitude="-0.30094445" longLatPrecision="1" longitude="-78.648605" pageId="20" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Cordillera Cotacachi">
<quantity id="B50CF3B09F60A07CFCC478D3FC2E15C7" box="[794,952,1751,1778]" metricMagnitude="3" metricUnit="m" metricValue="2.835" metricValueMax="2.957" metricValueMin="2.713" pageId="20" pageNumber="122" unit="m" value="2835.0" valueMax="2957.0" valueMin="2713.0">
<elevation id="F9D9B9669F60A07CFCC478D3FC2E15C7" box="[794,952,1751,1778]" metricMagnitude="3" metricUnit="m" metricValue="2.835" metricValueMax="2.957" metricValueMin="2.713" pageId="20" pageNumber="122" unit="m" value="2835.0" valueMax="2957.0" valueMin="2713.0">27132957 m</elevation>
</quantity>
,
<geoCoordinate id="17C038929F60A07CFC1A78D3FBF415C7" box="[964,1122,1751,1777]" degrees="00" direction="south" minutes="18" orientation="latitude" pageId="20" pageNumber="122" precision="1" seconds="03.4" value="-0.30094445">00°1803.4S</geoCoordinate>
<geoCoordinate id="17C038929F60A07CFBB478D3FA8515C4" box="[1130,1299,1751,1778]" degrees="78" direction="west" minutes="38" orientation="longitude" pageId="20" pageNumber="122" precision="1" seconds="55.0" value="-78.648605">78°3855.0W</geoCoordinate>
,
<date id="064A78959F60A07CFAC178D3FA0E15C4" box="[1311,1432,1751,1778]" pageId="20" pageNumber="122" value="2005-10-15">
<collectingDate id="160E817D9F60A07CFAC178D3FA0E15C4" box="[1311,1432,1751,1778]" pageId="20" pageNumber="122" value="2005-10-15">15.X.2005</collectingDate>
</date>
,
<collectorName id="DF013B839F60A07CFF4978FFFEB61423" box="[151,288,1787,1813]" pageId="20" pageNumber="122">J. D. Turner</collectorName>
leg.,
<specimenCount id="64F295DC9F60A07CFEBC78FFFE181423" box="[354,398,1787,1813]" pageId="20" pageNumber="122" type="male">1 ♂</specimenCount>
(
<collectionCode id="14E5C6909F60A07CFE4178FFFE6F1423" box="[415,505,1787,1813]" pageId="20" pageNumber="122">MCGL</collectionCode>
*)
</materialsCitation>
.
<materialsCitation id="C29C54089F60A07CFDC678FFFC3F1420" ID-GBIF-Occurrence="3042108639" box="[536,937,1787,1814]" collectingDate="2005-10-15" collectionCode="NW" collectorName="Slope &amp; Vol." country="Ecuador" elevation="2835" isTailEnumeration="true" latitude="-0.30094445" longLatPrecision="1" longitude="-78.648605" pageId="20" pageNumber="122" specimenCount="1" stateProvince="Pichincha">
<emphasis id="408082479F60A07CFDC678FFFDF61423" bold="true" box="[536,608,1787,1813]" pageId="20" pageNumber="122">Niebli</emphasis>
(
<collectionCode id="14E5C6909F60A07CFDAE78FFFD361423" box="[624,672,1787,1813]" pageId="20" pageNumber="122">NW</collectionCode>
<collectorName id="DF013B839F60A07CFD7778FFFD7A1423" box="[681,748,1787,1813]" pageId="20" pageNumber="122">Slope</collectorName>
,
<collectorName id="DF013B839F60A07CFD2678FFFCB01423" box="[760,806,1787,1813]" pageId="20" pageNumber="122">Vol.</collectorName>
<collectingRegion id="B03090B79F60A07CFCF078FFFC371420" box="[814,929,1787,1814]" country="Ecuador" name="Pichincha" pageId="20" pageNumber="122">Pichincha</collectingRegion>
)
</materialsCitation>
,
<materialsCitation id="C29C54089F60A07CFC6B78FFFE8D140F" ID-GBIF-Occurrence="3042108439" collectingDate="1971-08-01" collectionCode="R, MCGL" collectorName="de Lafebre" country="Ecuador" elevation="3500" pageId="20" pageNumber="122" specimenCount="7" specimenCount-male="7" stateProvince="Niebli">
<quantity id="B50CF3B09F60A07CFC6B78FFFB981423" box="[949,1038,1787,1814]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="122" unit="m" value="3500.0">
<elevation id="F9D9B9669F60A07CFC6B78FFFB981423" box="[949,1038,1787,1814]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="122" unit="m" value="3500.0">3500 m</elevation>
</quantity>
,
<date id="064A78959F60A07CFBC278FFFB341423" box="[1052,1186,1787,1813]" pageId="20" pageNumber="122" value="1971-08-01">
<collectingDate id="160E817D9F60A07CFBC278FFFB341423" box="[1052,1186,1787,1813]" pageId="20" pageNumber="122" value="1971-08-01">1.VIII.1971</collectingDate>
</date>
,
<collectionCode id="14E5C6909F60A07CFB6E78FFFB501423" box="[1200,1222,1787,1813]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="122">R</collectionCode>
.
<collectorName id="DF013B839F60A07CFB0C78FFFADA1423" box="[1234,1356,1787,1813]" pageId="20" pageNumber="122">de Lafebre</collectorName>
leg.,
<specimenCount id="64F295DC9F60A07CFA5078FFFF3A140F" pageId="20" pageNumber="122" type="male">7 ♂</specimenCount>
(
<collectionCode id="14E5C6909F60A07CFF62791BFE85140F" box="[188,275,1823,1849]" pageId="20" pageNumber="122">MCGL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C29C54089F60A07CFEF8791BFE37140F" ID-GBIF-Occurrence="3042108592" box="[294,417,1823,1849]" country="Ecuador" location="Tugurahua" pageId="20" pageNumber="122" specimenCount="1" stateProvince="Niebli">
<emphasis id="408082479F60A07CFEF8791BFE37140F" box="[294,417,1823,1849]" italics="true" pageId="20" pageNumber="122">
<location id="772B088E9F60A07CFEF8791BFE37140F" LSID="urn:lsid:plazi:treatment:FA5DEF439F65A07CFF497830FCE41490:772B088E9F60A07CFEF8791BFE37140F" box="[294,417,1823,1849]" country="Ecuador" name="Tugurahua" pageId="20" pageNumber="122" stateProvince="Niebli">Tugurahua</location>
</emphasis>
</materialsCitation>
<materialsCitation id="C29C54089F60A07CFE63791BFB22140F" ID-GBIF-Occurrence="3042108351" box="[445,1204,1823,1849]" collectingDate="1971-08" collectionCode="R, MGCL" collectorName="de Lafebre" country="Ecuador" elevation="1700" location="Rio Blanco" pageId="20" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Rio Blanco">
<emphasis id="408082479F60A07CFE63791BFDD7140F" bold="true" box="[445,577,1823,1849]" pageId="20" pageNumber="122">
<collectingRegion id="B03090B79F60A07CFE63791BFDD7140F" box="[445,577,1823,1849]" pageId="20" pageNumber="122">Rio Blanco</collectingRegion>
</emphasis>
,
<quantity id="B50CF3B09F60A07CFD8E791BFD3E140F" box="[592,680,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="122" unit="m" value="1700.0">
<elevation id="F9D9B9669F60A07CFD8E791BFD3E140F" box="[592,680,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="20" pageNumber="122" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="064A78959F60A07CFD6A791BFCB0140F" box="[692,806,1823,1849]" pageId="20" pageNumber="122" value="1971-08">
<collectingDate id="160E817D9F60A07CFD6A791BFCB0140F" box="[692,806,1823,1849]" pageId="20" pageNumber="122" value="1971-08">VIII.1971</collectingDate>
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,
<collectionCode id="14E5C6909F60A07CFCEC791BFCDE140F" box="[818,840,1823,1849]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="122">R</collectionCode>
.
<collectorName id="DF013B839F60A07CFC8D791BFC5B140F" box="[851,973,1823,1849]" pageId="20" pageNumber="122">de Lafebre</collectorName>
leg.,
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(
<collectionCode id="14E5C6909F60A07CFB99791BFB37140F" box="[1095,1185,1823,1849]" pageId="20" pageNumber="122">MGCL</collectionCode>
*)
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.
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<emphasis id="408082479F60A07CFB617924FB64140F" box="[1215,1266,1824,1849]" italics="true" pageId="20" pageNumber="122">
<collectingRegion id="B03090B79F60A07CFB617924FB64140F" box="[1215,1266,1824,1849]" country="Ecuador" name="Loja" pageId="20" pageNumber="122">Loja</collectingRegion>
</emphasis>
<emphasis id="408082479F60A07CFAD1791BFEA9146B" bold="true" pageId="20" pageNumber="122">Podocarpus National Park</emphasis>
(
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),
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,
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,
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<collectingDate id="160E817D9F60A07CFBEA7947FB201468" box="[1076,1206,1859,1886]" pageId="20" pageNumber="122" value="2009-11-15">15.XI.2009</collectingDate>
</date>
, C.
<collectionCode id="14E5C6909F60A07CFB017947FB65146B" box="[1247,1267,1859,1885]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="122" type="Museum">V</collectionCode>
.
<collectorName id="DF013B839F60A07CFB227947FAF1146B" box="[1276,1383,1859,1885]" pageId="20" pageNumber="122">Covell Jr.</collectorName>
leg.,
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(
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)
</materialsCitation>
. PERU—
<emphasis id="408082479F60A07CFE7F7963FDBB14B7" box="[417,557,1895,1921]" italics="true" pageId="20" pageNumber="122">Amazonas—</emphasis>
<materialsCitation id="C29C54089F60A07CFDF37963FF681490" ID-GBIF-Occurrence="3042108673" collectingDate="2016-11" collectionCode="OM" collectorName="J. Pintado" country="Ecuador" elevation="2200" latitude="-5.7166667" location="Oso Perdido" longLatPrecision="1305" longitude="-77.833336" pageId="20" pageNumber="122" specimenCode="OM 83.645" specimenCount="1" specimenCount-male="1" stateProvince="Oso Perdido">
<emphasis id="408082479F60A07CFDF37963FD5514B7" bold="true" box="[557,707,1895,1921]" pageId="20" pageNumber="122">
<collectingRegion id="B03090B79F60A07CFDF37963FD5514B7" box="[557,707,1895,1921]" pageId="20" pageNumber="122">Oso Perdido</collectingRegion>
</emphasis>
,
<quantity id="B50CF3B09F60A07CFD0C7963FCBC14B7" box="[722,810,1895,1921]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="122" unit="m" value="2200.0">
<elevation id="F9D9B9669F60A07CFD0C7963FCBC14B7" box="[722,810,1895,1921]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="122" unit="m" value="2200.0">2200 m</elevation>
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,
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,
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,
<date id="064A78959F60A07CFBC87963FBE314B7" box="[1046,1141,1895,1921]" pageId="20" pageNumber="122" value="2016-11">
<collectingDate id="160E817D9F60A07CFBC87963FBE314B7" box="[1046,1141,1895,1921]" pageId="20" pageNumber="122" value="2016-11">XI.2016</collectingDate>
</date>
,
<collectorName id="DF013B839F60A07CFB5C7963FB6414B7" box="[1154,1266,1895,1921]" pageId="20" pageNumber="122">J. Pintado</collectorName>
leg.,
<specimenCount id="64F295DC9F60A07CFAEC7963FAC814B7" box="[1330,1374,1895,1921]" pageId="20" pageNumber="122" type="male">1 ♂</specimenCount>
(
<specimenCode id="2252F62E9F60A07CFAB07963FF7C1490" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="20" pageNumber="122">OM 83.645</specimenCode>
*)
</materialsCitation>
,
<materialsCitation id="C29C54089F60A07CFED4798FFCF81490" ID-GBIF-Occurrence="3042108309" box="[266,878,1931,1958]" collectingDate="2017-04" collectionCode="OM" collectorName="J. Pintado" country="Ecuador" pageId="20" pageNumber="122" specimenCode="OM 83.945, OM 83.985" specimenCount="2" specimenCount-male="2" stateProvince="Oso Perdido">
<date id="064A78959F60A07CFED4798FFEF31493" box="[266,357,1931,1957]" pageId="20" pageNumber="122" value="2017-04">
<collectingDate id="160E817D9F60A07CFED4798FFEF31493" box="[266,357,1931,1957]" pageId="20" pageNumber="122" value="2017-04">IV.2017</collectingDate>
</date>
,
<collectorName id="DF013B839F60A07CFEAE798FFE491493" box="[368,479,1931,1957]" pageId="20" pageNumber="122">J. Pintado</collectorName>
leg.
<specimenCount id="64F295DC9F60A07CFDC8798FFDD61493" box="[534,576,1931,1957]" pageId="20" pageNumber="122" type="male">2 ♂</specimenCount>
(
<specimenCode id="2252F62E9F60A07CFD91798FFD431490" box="[591,725,1931,1958]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="20" pageNumber="122">OM 83.945</specimenCode>
,
<specimenCode id="2252F62E9F60A07CFD3E798FFCF01490" box="[736,870,1931,1958]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="20" pageNumber="122">OM 83.985</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>