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<document id="1C2D90A98DCDB4E94098A610924DC57E" ID-CLB-Dataset="32404" ID-DOI="10.11646/zootaxa.4306.1.7" ID-GBIF-Dataset="c8239bdd-76a0-481f-893c-e7969f4a6e2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="843151" ID-ZooBank="9447E681-4C29-44D4-BB59-0AC2FCC204EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502783969055" checkinUser="plazi" docAuthor="Xue, Guo-Xi, Li, Xiao-Juan &amp; Li, Ding-Kai" docDate="2017" docId="FA5787D6FF8E861C12D1FE63D518FE7F" docLanguage="en" docName="zootaaxa.4306.1.7.pdf" docOrigin="Zootaxa 4306 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coladenia simianensis Xue &amp; Li &amp; Li 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="124" masterDocId="066EFFAEFF8F86181246FFF2D725FF8B" masterDocTitle="Description of Coladenia simianensis sp. nov. (Lepidoptera: Hesperiidae) from China, with an annotated checklist of the genus Coladenia Moore, 1881" masterLastPageNumber="129" masterPageNumber="121" pageNumber="122" updateTime="1698466593472" updateUser="ExternalLinkService">
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<mods:title id="1926C981E15D9A9A28B2E1520203A21D">Description of Coladenia simianensis sp. nov. (Lepidoptera: Hesperiidae) from China, with an annotated checklist of the genus Coladenia Moore, 1881</mods:title>
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<mods:namePart id="A23DF4E1DA1A7696A17AE5CF10066298">Xue, Guo-Xi</mods:namePart>
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<mods:namePart id="61901F044FBFF1184EDD06FCACEFEC17">Li, Xiao-Juan</mods:namePart>
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<mods:namePart id="54252445354ED5B6DC0417C14742A7D5">Li, Ding-Kai</mods:namePart>
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<mods:date id="A63EFAA5ABB8B587D53F2816E88970C8">2017</mods:date>
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<treatment id="FA5787D6FF8E861C12D1FE63D518FE7F" ID-DOI="http://doi.org/10.5281/zenodo.6032362" ID-GBIF-Taxon="132650262" ID-Zenodo-Dep="6032362" LSID="urn:lsid:plazi:treatment:FA5787D6FF8E861C12D1FE63D518FE7F" httpUri="http://treatment.plazi.org/id/FA5787D6FF8E861C12D1FE63D518FE7F" lastPageId="4" lastPageNumber="124" pageId="1" pageNumber="122">
<subSubSection id="3AE4654BFF8E861912D1FE63D6A7FE47" pageId="1" pageNumber="122" type="nomenclature">
<paragraph id="724136C0FF8E861912D1FE63D562FE20" blockId="1.[151,583,401,461]" box="[151,583,401,427]" pageId="1" pageNumber="122">
<heading id="290981ACFF8E861912D1FE63D562FE20" bold="true" box="[151,583,401,427]" fontSize="11" level="1" pageId="1" pageNumber="122" reason="1">
<emphasis id="408AEAD2FF8E861912D1FE63D562FE20" bold="true" box="[151,583,401,427]" pageId="1" pageNumber="122">
<taxonomicName id="B5FE4D43FF8E861912D1FE63D6C7FE20" authority="Xue" authorityName="Xue &amp; Li &amp; Li" authorityYear="2017" box="[151,482,401,427]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="simianensis" status="sp. nov.">
<emphasis id="408AEAD2FF8E861912D1FE63D682FE20" bold="true" box="[151,423,401,427]" italics="true" pageId="1" pageNumber="122">Coladenia simianensis</emphasis>
Xue
</taxonomicName>
<taxonomicNameLabel id="5BB957A9FF8E861913AFFE63D562FE20" box="[489,583,401,427]" pageId="1" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="724136C0FF8E861912D1FE46D6A7FE47" blockId="1.[151,583,401,461]" box="[151,386,436,461]" pageId="1" pageNumber="122">
(
<figureCitation id="EAC52A45FF8E861912D9FE46D62FFE47" box="[159,266,436,461]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[184,1374,205,1916]" captionTargetId="figure@2.[175,1387,195,1924]" captionTargetPageId="2" captionText="FIGURES 1 4. Adults of Coladenia simianensis. a. dorsal view; b. ventral view. 1. male, holotype; 2 3. male, paratypes; 4. female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/843153/files/figure.png" pageId="1" pageNumber="122">Figs. 14</figureCitation>
,
<figureCitation id="EAC52A45FF8E86191350FE46D674FE46" box="[278,337,436,461]" captionStart="FIGURES 5 17" captionStartId="3.[151,264,951,974]" captionTargetBox="[163,1424,209,918]" captionTargetId="figure@3.[151,1436,193,930]" captionTargetPageId="3" captionText="FIGURES 5 17. Male genitalia of Coladenia simianensis. 5. genital capsule, lateral view, with the left valva and aedeagus removed; 6. ditto, oblique view; 7. tegumen and uncus, dorsal view; 8. ditto, ventral view; 9. gnathos, oblique view; 10. uncus and gnathos, posterior view; 11. left valva, inner side; 12. ditto, dorsal view; 13. ditto, outer side; 14. aedeagus, lateral view; 15. ditto, dorsal view; 16. juxta, lateral view; 17. juxta. Scale bar = 1 mm." httpUri="https://zenodo.org/record/843155/files/figure.png" pageId="1" pageNumber="122">517</figureCitation>
,
<figureCitation id="EAC52A45FF8E8619131BFE46D65CFE47" box="[349,377,436,460]" captionStart="FIGURE 18" captionStartId="3.[151,250,1629,1651]" captionTargetBox="[175,1419,1115,1588]" captionTargetId="figure@3.[151,1436,1115,1607]" captionTargetPageId="3" captionText="FIGURE 18. Female genitalia of Coladenia simianensis, ventral view (bursa copulatrix broken). Scale bar = 1 mm." httpUri="https://zenodo.org/record/843157/files/figure.png" pageId="1" pageNumber="122">18</figureCitation>
)
</paragraph>
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<subSubSection id="3AE4654BFF8E861912D1FE0ED689FD0B" pageId="1" pageNumber="122" type="materials_examined">
<paragraph id="724136C0FF8E861912D1FE0ED7E4FDB3" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
<materialsCitation id="C2963C9DFF8E861912D1FE0ED7E4FDB3" ID-GBIF-Occurrence="1586033289" collectingDate="2015-07-06" collectorName="G. X. Xue" country="China" elevation="1100" location="Simian Mountain" municipality="Jiangjin" pageId="1" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="holotype">
<emphasis id="408AEAD2FF8E861912D1FE0ED663FD9E" bold="true" box="[151,326,508,533]" pageId="1" pageNumber="122">
<typeStatus id="AD458862FF8E861912D1FE0ED7F7FD9E" box="[151,210,508,533]" pageId="1" pageNumber="122">Type</typeStatus>
material.
</emphasis>
<typeStatus id="AD458862FF8E86191308FE0ED6C7FD9F" box="[334,482,508,532]" pageId="1" pageNumber="122" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="64F8FD49FF8E861913ACFE0FD527FD9F" box="[490,514,509,532]" pageId="1" pageNumber="122" type="male"></specimenCount>
:
<location id="7721601BFF8E86191049FE0ED5F2FD9E" LSID="urn:lsid:plazi:treatment:FA5787D6FF8E861C12D1FE63D518FE7F:7721601BFF8E86191049FE0ED5F2FD9E" box="[527,727,508,533]" country="China" municipality="Jiangjin" name="Simian Mountain" pageId="1" pageNumber="122" stateProvince="Chongqing">Simian Mountain</location>
,
<collectingMunicipality id="9225ACBAFF8E861910A5FE0ED41AFD9E" box="[739,831,508,533]" pageId="1" pageNumber="122">Jiangjin</collectingMunicipality>
,
<collectingRegion id="B03AF822FF8E8619110DFE0ED4EFFD9E" box="[843,970,508,533]" country="China" name="Chongqing" pageId="1" pageNumber="122">Chongqing</collectingRegion>
,
<collectingCountry id="0AE97650FF8E86191191FE0ED338FD9E" box="[983,1053,508,533]" name="China" pageId="1" pageNumber="122">China</collectingCountry>
,
<elevation id="F9D3D1F3FF8E8619166CFE0ED3A4FD9F" box="[1066,1153,508,532]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="122" unit="m" value="1100.0">
<quantity id="B5069B25FF8E8619166CFE0ED3A4FD9F" box="[1066,1153,508,532]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="122" unit="m" value="1100.0">1100 m</quantity>
</elevation>
,
<collectingDate id="1604E9E8FF8E861916C8FE0ED238FD9E" box="[1166,1309,508,533]" pageId="1" pageNumber="122" value="2015-07-06">
<date id="06401000FF8E861916C8FE0ED238FD9E" box="[1166,1309,508,533]" pageId="1" pageNumber="122" value="2015-07-06">6. VII. 2015</date>
</collectingDate>
,
<collectorName id="DF0B5316FF8E8619176FFE0ED2B9FD9F" box="[1321,1436,508,532]" pageId="1" pageNumber="122">G. X. Xue</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="724136C0FF8E86191281FDB6D689FD0B" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
<materialsCitation id="C2963C9DFF8E86191281FDB6D58EFDD6" ID-GBIF-Occurrence="1586033279" box="[199,683,580,605]" collectingDate="2015-07-06" collectorName="G. X. Xue" country="China" elevation="1100" location="Simian Mountain" municipality="Jiangjin" pageId="1" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="paratype">
<typeStatus id="AD458862FF8E86191281FDB6D643FDD7" box="[199,358,580,604]" pageId="1" pageNumber="122" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="64F8FD49FF8E86191337FDB6D6B8FDD7" box="[369,413,580,604]" pageId="1" pageNumber="122" type="male">1 ♂</specimenCount>
, same data as Holotype
</materialsCitation>
;
<materialsCitation id="C2963C9DFF8E861910F1FDB6D4C8FDD6" ID-GBIF-Occurrence="1586033284" box="[695,1005,580,605]" collectingDate="2015-07-06" collectorName="S. Y. Huang" country="China" elevation="1100" location="Simian Mountain" municipality="Jiangjin" pageId="1" pageNumber="122" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="paratype">
<specimenCount id="64F8FD49FF8E861910F1FDB6D5C6FDD7" box="[695,739,580,604]" pageId="1" pageNumber="122" type="male">1 ♂</specimenCount>
,
<emphasis id="408AEAD2FF8E861910B6FDB6D406FDD7" box="[752,803,580,604]" italics="true" pageId="1" pageNumber="122">ditto</emphasis>
,
<collectorName id="DF0B5316FF8E86191177FDB6D49CFDD6" box="[817,953,580,605]" pageId="1" pageNumber="122">S. Y. Huang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="C2963C9DFF8E861911BFFDB6D689FD0B" ID-GBIF-Occurrence="1586033270" collectingDate="2011-08" collectorName="J. Y. Qiu" country="China" elevation="1100" location="Simian Mountain" municipality="Jiangjin" pageId="1" pageNumber="122" specimenCount="1" specimenCount-female="1" stateProvince="Chongqing" typeStatus="paratype">
<specimenCount id="64F8FD49FF8E861911BFFDB6D306FDD7" box="[1017,1059,580,604]" pageId="1" pageNumber="122" type="female">1 ♀</specimenCount>
, same locality as Holotype, early
<collectingDate id="1604E9E8FF8E861912D1FD9AD62BFD0B" box="[151,270,616,640]" pageId="1" pageNumber="122" value="2011-08">
<date id="06401000FF8E861912D1FD9AD62BFD0B" box="[151,270,616,640]" pageId="1" pageNumber="122" value="2011-08">VIII. 2011</date>
</collectingDate>
,
<collectorName id="DF0B5316FF8E8619135CFD9AD65EFD0B" box="[282,379,615,640]" pageId="1" pageNumber="122">J. Y. Qiu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AE4654BFF8E86191281FD7ED3ADF891" pageId="1" pageNumber="122" type="description">
<paragraph id="724136C0FF8E86191281FD7ED5F9FA97" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
<emphasis id="408AEAD2FF8E86191281FD7ED67CFD2E" bold="true" box="[199,345,652,677]" pageId="1" pageNumber="122">Description.</emphasis>
Male. (
<figureCitation id="EAC52A45FF8E861913F3FD7ED504FD2F" box="[437,545,652,677]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[184,1374,205,1916]" captionTargetId="figure@2.[175,1387,195,1924]" captionTargetPageId="2" captionText="FIGURES 1 4. Adults of Coladenia simianensis. a. dorsal view; b. ventral view. 1. male, holotype; 2 3. male, paratypes; 4. female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/843153/files/figure.png" pageId="1" pageNumber="122">Figs. 13</figureCitation>
) Antennae:
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in length, dorsal side dark brown, ventral side checkered with gray; apiculus sharply pointed, nudum
<quantity id="B5069B25FF8E861910C2FD42D5C3FD43" box="[644,742,687,712]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="1" pageNumber="122" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">1314 in</quantity>
number. Labial palpi: ventral side of second segment covered with yellowish white scales, mixed with sparse black hairs; third segment porrect, thicker than shaft of antennae, dark brown, with a blunt point. Thorax and abdomen covered with brown scales and hairs on both dorsal and ventral sides; hind tibiae with a pale yellowish hair pencil. Forewing:
<quantity id="B5069B25FF8E861911F9FCEED335FCBF" box="[959,1040,796,820]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="122" unit="mm" value="20.0">20 mm</quantity>
in length, dorsal side dark brown, basal half clothed with brown scales, spots white, hyaline; subapical spots in spaces R3M2 arranged in S-shape, the one in R3 and R
<quantity id="B5069B25FF8E86191302FC87D64DFCF4" box="[324,360,870,899]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="122" unit="in" value="5.0">5 in</quantity>
same size, bigger than those in R4, M1 and M2; spot in space Sc shorter than half length of widest portion of cell spot; cell spot located in middle of wing base and wing apex, rectangular, with outer lower angle protruding; spot in space M3 bigger than apical spots, closer to the one in Cu1 than to cell spot; spot in space Cu1 nearly equidistant from spot in M3 and cell spot, much bigger than the former but smaller than the latter, overlapping with outer lower angle of cell spot; under spot in Cu1 there are two separated tiny dots in space Cu2, of which the lower one moved inwards and almost invisible; ventral side of forewing dark brown, wing base covered with sparse brown scales and hairs, spots as in dorsal side except for two pale patches beyond tiny dots in space Cu2. Hindwing: dorsal side brown, basal area covered with hairs, marginal area darker; discal area with a series of black spots from space Sc+R1 to Cu2, of which the two in spaces M1M2 longer, and the two in Cu2 blurred; cell with a very indistinct black discoidal spot, sometimes completely invisible; ventral side dark brown, overlaid with brown scales, slightly greenish near dorsum; spots as in dorsal side. Cilia dark brown, with yellowish-white at apex of forewing and before end of vein M1 of hindwing.
</paragraph>
<paragraph id="724136C0FF8E86191281FAD8D241F9B4" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
Male genitalia (
<figureCitation id="EAC52A45FF8E8619133EFAD8D6D1FAC8" box="[376,500,1322,1347]" captionStart="FIGURES 5 17" captionStartId="3.[151,264,951,974]" captionTargetBox="[163,1424,209,918]" captionTargetId="figure@3.[151,1436,193,930]" captionTargetPageId="3" captionText="FIGURES 5 17. Male genitalia of Coladenia simianensis. 5. genital capsule, lateral view, with the left valva and aedeagus removed; 6. ditto, oblique view; 7. tegumen and uncus, dorsal view; 8. ditto, ventral view; 9. gnathos, oblique view; 10. uncus and gnathos, posterior view; 11. left valva, inner side; 12. ditto, dorsal view; 13. ditto, outer side; 14. aedeagus, lateral view; 15. ditto, dorsal view; 16. juxta, lateral view; 17. juxta. Scale bar = 1 mm." httpUri="https://zenodo.org/record/843155/files/figure.png" pageId="1" pageNumber="122">Figs. 517</figureCitation>
): Tegumen humpy in lateral view, conspicuously higher than uncus. In dorsal view, basal half of uncus tapered, with a small triangular process on each side; distal half finger-like, with a round tip. In lateral view, uncus nearly 2/3 times as long as tegumen, falcate, with a pointed tip. Gnathos arm-like in lateral view, ventral side connected and very broad, as wide as base of uncus, densely covered with tiny teeth. Saccus very short. Valva elongate, with its distal portion bifid into two strongly curved and sharply pointed long branches, of which the dorsal one thin and smooth, evenly tapered, and the distal one much more robust, with its apical area outwardly serrated. Aedeagus a little longer than ventral margin of valva; in lateral view, middle of dorsal margin concave before a serrated ridge, ventral margin sinuolate; coecum short, slender and curved upwards. Juxta cordiform.
</paragraph>
<paragraph id="724136C0FF8E86191281F9B8D253F90C" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
Female (
<figureCitation id="EAC52A45FF8E8619136CF9B8D655F9E8" box="[298,368,1610,1635]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1971,1994]" captionTargetBox="[184,1374,205,1916]" captionTargetId="figure@2.[175,1387,195,1924]" captionTargetPageId="2" captionText="FIGURES 1 4. Adults of Coladenia simianensis. a. dorsal view; b. ventral view. 1. male, holotype; 2 3. male, paratypes; 4. female, paratype. Scale bar = 1 cm." httpUri="https://zenodo.org/record/843153/files/figure.png" pageId="1" pageNumber="122">Fig. 4</figureCitation>
): Forewing
<quantity id="B5069B25FF8E861913BCF9B9D56EF9E9" box="[506,587,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="122" unit="mm" value="21.0">21 mm</quantity>
in length; wing pattern similar to male, but spot in space Sc half length of widest portion of cell spot; spot in space Cu1 obviously longer than male; end of abdomen with a gray anal tuft.
</paragraph>
<paragraph id="724136C0FF8E86191281F967D3ADF891" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
Female genitalia (
<figureCitation id="EAC52A45FF8E861913C9F967D6C4F925" box="[399,481,1685,1710]" captionStart="FIGURE 18" captionStartId="3.[151,250,1629,1651]" captionTargetBox="[175,1419,1115,1588]" captionTargetId="figure@3.[151,1436,1115,1607]" captionTargetPageId="3" captionText="FIGURE 18. Female genitalia of Coladenia simianensis, ventral view (bursa copulatrix broken). Scale bar = 1 mm." httpUri="https://zenodo.org/record/843157/files/figure.png" pageId="1" pageNumber="122">Fig. 18</figureCitation>
): Papillae anales reniform, covered with short hairs and thorns. Lamella postvaginalis very wide, middle portion of distal edge shallowly concaved and higher than shoulders on each side. Lamella antevaginalis shovel-like, basal portion tapered, medial concavity of distal edge shallow V-shaped with a round bottom. Ductus bursae twisted; bursa copulatrix bursiform, membranous, without signum.
</paragraph>
</subSubSection>
<subSubSection id="3AE4654BFF8E86191281F8D7D4E3F80D" pageId="1" pageNumber="122" type="distribution">
<paragraph id="724136C0FF8E86191281F8D7D5D4F8B5" blockId="1.[151,1437,508,1962]" box="[199,753,1829,1854]" pageId="1" pageNumber="122">
<emphasis id="408AEAD2FF8E86191281F8D7D645F8B5" bold="true" box="[199,352,1829,1854]" pageId="1" pageNumber="122">Distribution.</emphasis>
Only known from the
<typeStatus id="AD458862FF8E86191026F8D4D5B5F8B5" box="[608,656,1830,1854]" pageId="1" pageNumber="122">type</typeStatus>
locality.
</paragraph>
<paragraph id="724136C0FF8E86191281F8BBD4E3F80D" blockId="1.[151,1437,508,1962]" pageId="1" pageNumber="122">
<emphasis id="408AEAD2FF8E86191281F8BBD66FF8E9" bold="true" box="[199,330,1865,1890]" pageId="1" pageNumber="122">Bionomics.</emphasis>
The adult flies rapidly along forest paths on sunny days (
<figureCitation id="EAC52A45FF8E861911A5F8BBD312F8EA" box="[995,1079,1865,1890]" captionStart="FIGURE 19" captionStartId="4.[151,250,1559,1581]" captionTargetBox="[151,1436,571,1538]" captionTargetId="figure@4.[151,1436,571,1538]" captionTargetPageId="4" captionText="FIGURE 19. Habitat of Coladenia simianensis. Photo by Guo-xi Xue." httpUri="https://zenodo.org/record/843159/files/figure.png" pageId="1" pageNumber="122">Fig. 19</figureCitation>
), and it is very alert. The three male specimens in the
<typeStatus id="AD458862FF8E861913D0F89CD6E3F80D" box="[406,454,1902,1926]" pageId="1" pageNumber="122">type</typeStatus>
series were captured after seven days of rain.
</paragraph>
</subSubSection>
<subSubSection id="3AE4654BFF8E86191281F863D451F821" box="[199,884,1937,1962]" pageId="1" pageNumber="122" type="etymology">
<paragraph id="724136C0FF8E86191281F863D451F821" blockId="1.[151,1437,508,1962]" box="[199,884,1937,1962]" pageId="1" pageNumber="122">
<emphasis id="408AEAD2FF8E86191281F863D66BF821" bold="true" box="[199,334,1937,1962]" pageId="1" pageNumber="122">Etymology.</emphasis>
The new species is named after the
<typeStatus id="AD458862FF8E861910A5F860D436F821" box="[739,787,1938,1962]" pageId="1" pageNumber="122">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="26816648FF8D861A12D1F841D6D8F863" httpUri="https://zenodo.org/record/843153/files/figure.png" pageId="2" pageNumber="123" targetBox="[184,1374,205,1916]" targetPageId="2">
<paragraph id="724136C0FF8D861A12D1F841D6D8F863" blockId="2.[151,1436,1971,2024]" pageId="2" pageNumber="123">
<emphasis id="408AEAD2FF8D861A12D1F841D610F843" bold="true" box="[151,309,1971,1994]" pageId="2" pageNumber="123">FIGURES 14</emphasis>
. Adults of
<taxonomicName id="B5FE4D43FF8D861A13EDF841D5B7F842" authorityName="Xue &amp; Li &amp; Li" authorityYear="2017" box="[427,658,1971,1993]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="simianensis">
<emphasis id="408AEAD2FF8D861A13EDF841D5B7F842" box="[427,658,1971,1993]" italics="true" pageId="2" pageNumber="123">Coladenia simianensis</emphasis>
</taxonomicName>
. a. dorsal view; b. ventral view. 1. male, holotype; 23. male, paratypes; 4. female, paratype. Scale bar = 1 cm.
</paragraph>
</caption>
<caption id="26816648FF8C861B12D1FC45D478FBA1" httpUri="https://zenodo.org/record/843155/files/figure.png" pageId="3" pageNumber="124" targetBox="[163,1424,209,918]" targetPageId="3">
<paragraph id="724136C0FF8C861B12D1FC45D478FBA1" blockId="3.[151,1436,951,1066]" pageId="3" pageNumber="124">
<emphasis id="408AEAD2FF8C861B12D1FC45D667FC46" bold="true" box="[151,322,951,974]" pageId="3" pageNumber="124">FIGURES 517</emphasis>
. Male genitalia of
<taxonomicName id="B5FE4D43FF8C861B104DFC4AD5D6FC45" authorityName="Xue &amp; Li &amp; Li" authorityYear="2017" box="[523,755,952,974]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="simianensis">
<emphasis id="408AEAD2FF8C861B104DFC4AD5D6FC45" box="[523,755,952,974]" italics="true" pageId="3" pageNumber="124">Coladenia simianensis</emphasis>
</taxonomicName>
. 5. genital capsule, lateral view, with the left valva and aedeagus removed; 6.
<emphasis id="408AEAD2FF8C861B1351FC24D660FC67" box="[279,325,982,1004]" italics="true" pageId="3" pageNumber="124">ditto</emphasis>
, oblique view; 7. tegumen and uncus, dorsal view; 8.
<emphasis id="408AEAD2FF8C861B112AFC24D4BFFC67" box="[876,922,982,1004]" italics="true" pageId="3" pageNumber="124">ditto</emphasis>
, ventral view; 9. gnathos, oblique view; 10. uncus and gnathos, posterior view; 11. left valva, inner side; 12.
<emphasis id="408AEAD2FF8C861B1098FC07D429FB80" box="[734,780,1013,1035]" italics="true" pageId="3" pageNumber="124">ditto</emphasis>
, dorsal view; 13.
<emphasis id="408AEAD2FF8C861B1186FC07D4CBFB80" box="[960,1006,1013,1035]" italics="true" pageId="3" pageNumber="124">ditto</emphasis>
, outer side; 14. aedeagus, lateral view; 15.
<emphasis id="408AEAD2FF8C861B12D1FBE6D7E0FBA1" box="[151,197,1044,1066]" italics="true" pageId="3" pageNumber="124">ditto</emphasis>
, dorsal view; 16. juxta, lateral view; 17. juxta. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="26816648FF8C861B12D1F9AFD20BF9F9" box="[151,1326,1628,1651]" httpUri="https://zenodo.org/record/843157/files/figure.png" pageId="3" pageNumber="124" targetBox="[175,1419,1115,1588]" targetPageId="3">
<paragraph id="724136C0FF8C861B12D1F9AFD20BF9F9" blockId="3.[151,1326,1628,1651]" box="[151,1326,1628,1651]" pageId="3" pageNumber="124">
<emphasis id="408AEAD2FF8C861B12D1F9AFD63CF9F9" bold="true" box="[151,281,1629,1651]" pageId="3" pageNumber="124">FIGURE 18</emphasis>
. Female genitalia of
<taxonomicName id="B5FE4D43FF8C861B13A9F9AED5F0F9F8" authorityName="Xue &amp; Li &amp; Li" authorityYear="2017" box="[495,725,1628,1651]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="simianensis">
<emphasis id="408AEAD2FF8C861B13A9F9AED5F0F9F8" box="[495,725,1628,1651]" italics="true" pageId="3" pageNumber="124">Coladenia simianensis</emphasis>
</taxonomicName>
, ventral view (bursa copulatrix broken). Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="3AE4654BFF8C861C1281F968D518FE7F" lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124" type="diagnosis">
<paragraph id="724136C0FF8C861B1281F968D64BF8C8" blockId="3.[151,1436,1690,2010]" pageId="3" pageNumber="124">
<emphasis id="408AEAD2FF8C861B1281F968D664F938" bold="true" box="[199,321,1690,1715]" pageId="3" pageNumber="124">Diagnosis.</emphasis>
Judged by appearance, the new species is similar to
<taxonomicName id="B5FE4D43FF8C861B11DEF968D2B9F939" authority="Elwes &amp; Edwards, 1897" authorityName="Elwes &amp; Edwards" authorityYear="1897" box="[920,1436,1690,1715]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="agnioides">
<emphasis id="408AEAD2FF8C861B11DEF968D3A7F939" box="[920,1154,1690,1714]" italics="true" pageId="3" pageNumber="124">Coladenia agnioides</emphasis>
Elwes &amp; Edwards, 1897
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8C861B1280F94DD58BF95C" authority="de Jong &amp; Treadaway, 1992" authorityName="de Jong &amp; Treadaway" authorityYear="1992" box="[198,686,1726,1751]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
<emphasis id="408AEAD2FF8C861B1280F94DD64BF95D" box="[198,366,1726,1750]" italics="true" pageId="3" pageNumber="124">C. agni sundae</emphasis>
de
<bibRefCitation id="166F4B31FF8C861B13D0F94DD58BF95C" author="Jong" box="[406,686,1726,1751]" pageId="3" pageNumber="124" refString="de Jong, R. &amp; Treadaway, C. G. (1992) Revisional notes on Coladenia Moore, 1881 (Lepidoptera: Hesperiidae). Zoologische Mededelingen, 66 (15), 283 - 293." type="journal article" year="1992">
Jong &amp; Treadaway,
<quantity id="B5069B25FF8C861B1030F94DD58BF95C" box="[630,686,1727,1751]" metricMagnitude="1" metricUnit="m" metricValue="5.05968" pageId="3" pageNumber="124" unit="in" value="1992.0">1992</quantity>
</bibRefCitation>
</taxonomicName>
in having narrow and separated discal spots on the forewing, but it can be immediately distinguished from the latter two by the remarkable long and curved branches of the valva (for figures of the male genitalia of
<taxonomicName id="B5FE4D43FF8C861B1047F8F5D5B5F895" authorityName="Elwes &amp; Edwards" authorityYear="1897" box="[513,656,1798,1822]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="agnioides">
<emphasis id="408AEAD2FF8C861B1047F8F5D5B5F895" box="[513,656,1798,1822]" italics="true" pageId="3" pageNumber="124">C. agnioides</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8C861B108CF8F5D450F895" authorityName="de Jong &amp; Treadaway" authorityYear="1992" box="[714,885,1798,1822]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
<emphasis id="408AEAD2FF8C861B108CF8F5D450F895" box="[714,885,1798,1822]" italics="true" pageId="3" pageNumber="124">C. agni sundae</emphasis>
</taxonomicName>
, see
<bibRefCitation id="166F4B31FF8C861B11F4F8F4D342F894" author="Devyatkin" box="[946,1127,1798,1823]" pageId="3" pageNumber="124" refString="Devyatkin, A. L. (2002) Hesperiidae of Vietnam, 10. A new species of Coladenia Moore, [1881] (Lepidoptera, Hesperiidae). Atalanta, 33 (1 / 2), 123 - 125." type="journal article" year="2002">Devyatkin 2002</bibRefCitation>
and de
<bibRefCitation id="166F4B31FF8C861B1683F8F5D7EAF8C9" author="Jong" pageId="3" pageNumber="124" refString="de Jong, R. &amp; Treadaway, C. G. (1992) Revisional notes on Coladenia Moore, 1881 (Lepidoptera: Hesperiidae). Zoologische Mededelingen, 66 (15), 283 - 293." type="journal article" year="1992">Jong &amp; Treadaway 1992</bibRefCitation>
respectively).
</paragraph>
<paragraph id="724136C0FF8C861C1281F8BDD518FE7F" blockId="3.[151,1436,1690,2010]" lastBlockId="4.[151,1436,151,501]" lastPageId="4" lastPageNumber="125" pageId="3" pageNumber="124">
Structures of the male genitalia of
<taxonomicName id="B5FE4D43FF8C861B100BF8BCD469F8ED" authorityName="Xue &amp; Li &amp; Li" authorityYear="2017" box="[589,844,1870,1894]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="simianensis">
<emphasis id="408AEAD2FF8C861B100BF8BCD469F8ED" box="[589,844,1870,1894]" italics="true" pageId="3" pageNumber="124">Coladenia simianensis</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="B5FE4D43FF8C861B1617F8BDD252F8EC" authority="Devyatkin, 2002" authorityName="Devyatkin" authorityYear="2002" box="[1105,1399,1870,1895]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8C861B1617F8BDD397F8ED" box="[1105,1202,1871,1894]" italics="true" pageId="3" pageNumber="124">C. tanya</emphasis>
<bibRefCitation id="166F4B31FF8C861B16FEF8BCD252F8EC" author="Devyatkin" box="[1208,1399,1870,1895]" pageId="3" pageNumber="124" refString="Devyatkin, A. L. (2002) Hesperiidae of Vietnam, 10. A new species of Coladenia Moore, [1881] (Lepidoptera, Hesperiidae). Atalanta, 33 (1 / 2), 123 - 125." type="journal article" year="2002">Devyatkin, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B5FE4D43FF8C861B17C4F8BDD6A8F801" authority="Huang, 2003" authorityName=", Huang" authorityYear="2003" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8C861B17C4F8BDD7D1F801" italics="true" pageId="3" pageNumber="124">C. uemurai</emphasis>
<bibRefCitation id="166F4B31FF8C861B12BAF881D6A8F801" author="Huang" box="[252,397,1906,1931]" pageId="3" pageNumber="124" refString="Huang, H. (2003) A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongjiang, China with descriptions of new species, new subspecies and revisional notes. Neue Entomologische Nachrichten, 55, 3 - 114." type="journal article" year="2003">Huang, 2003</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8C861B1382F880D437F800" authority="Huang &amp; Li, 2006" authorityName="Huang &amp; Li" authorityYear="2006" box="[452,786,1906,1931]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="motuoa">
<emphasis id="408AEAD2FF8C861B1382F880D51CF801" box="[452,569,1906,1930]" italics="true" pageId="3" pageNumber="124">C. motuoa</emphasis>
<bibRefCitation id="166F4B31FF8C861B1006F881D437F800" author="Huang" box="[576,786,1906,1931]" pageId="3" pageNumber="124" refString="Huang, H. &amp; Li, L. Z. (2006) A new subspecies of Coladenia uemurai Huang, 2003 from SE Tibet (Lepidoptera, Lycaenidae) Atalanta, 37 (3 / 4), 322 - 324." type="journal article" year="2006">Huang &amp; Li, 2006</bibRefCitation>
</taxonomicName>
. However, the new taxon differs from these species in the following characters (based on the male): discal spots on the forewing are separated, spots in spaces Sc, Cu1 and the cell are narrow (in
<taxonomicName id="B5FE4D43FF8C861B132AF84FD6EFF85E" authorityName="Devyatkin" authorityYear="2002" box="[364,458,1981,2005]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8C861B132AF84FD6EFF85E" box="[364,458,1981,2005]" italics="true" pageId="3" pageNumber="124">C. tanya</emphasis>
</taxonomicName>
,
<taxonomicName id="B5FE4D43FF8C861B139EF84FD573F85E" authorityName=", Huang" authorityYear="2003" box="[472,598,1981,2005]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8C861B139EF84FD573F85E" box="[472,598,1981,2005]" italics="true" pageId="3" pageNumber="124">C. uemurai</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8C861B10CAF84FD424F85E" authorityName="Huang &amp; Li" authorityYear="2006" box="[652,769,1981,2005]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="motuoa">
<emphasis id="408AEAD2FF8C861B10CAF84FD424F85E" box="[652,769,1981,2005]" italics="true" pageId="3" pageNumber="124">C. motuoa</emphasis>
</taxonomicName>
, the discal spots are contiguous, the spots in spaces Sc, Cu1 and the cell are conspicuously wider); the uncus is nearly 2/3 times the length of the tegumen (it is less than half of the tegumen in
<taxonomicName id="B5FE4D43FF8B861C130DFF4ED689FF5F" authorityName="Devyatkin" authorityYear="2002" box="[331,428,188,212]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8B861C130DFF4ED689FF5F" box="[331,428,188,212]" italics="true" pageId="4" pageNumber="125">C. tanya</emphasis>
</taxonomicName>
); the dorsal branch of the valva is long and strongly curved (it is markedly shorter and erect in
<taxonomicName id="B5FE4D43FF8B861C12B4FF12D677FF73" authorityName="Devyatkin" authorityYear="2002" box="[242,338,224,248]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8B861C12B4FF12D677FF73" box="[242,338,224,248]" italics="true" pageId="4" pageNumber="125">C. tanya</emphasis>
</taxonomicName>
, and relatively shorter and not curved that much in
<taxonomicName id="B5FE4D43FF8B861C11DAFF12D33EFF7C" authorityName=", Huang" authorityYear="2003" box="[924,1051,224,247]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8B861C11DAFF12D33EFF7C" box="[924,1051,224,247]" italics="true" pageId="4" pageNumber="125">C. uemurai</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8B861C1615FF12D3EDFF73" authorityName="Huang &amp; Li" authorityYear="2006" box="[1107,1224,224,248]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="motuoa">
<emphasis id="408AEAD2FF8B861C1615FF12D3EDFF73" box="[1107,1224,224,248]" italics="true" pageId="4" pageNumber="125">C. motuoa</emphasis>
</taxonomicName>
); the distal branch of the valva is long and strongly curved, with its apical area outwardly serrated (in
<taxonomicName id="B5FE4D43FF8B861C161BFEF6D3F1FE97" authorityName="Huang &amp; Li" authorityYear="2006" box="[1117,1236,260,284]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="motuoa">
<emphasis id="408AEAD2FF8B861C161BFEF6D3F1FE97" box="[1117,1236,260,284]" italics="true" pageId="4" pageNumber="125">C. motuoa</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8B861C1749FEF6D255FE97" authorityName="Devyatkin" authorityYear="2002" box="[1295,1392,260,284]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8B861C1749FEF6D255FE97" box="[1295,1392,260,284]" italics="true" pageId="4" pageNumber="125">C. tanya</emphasis>
</taxonomicName>
the distal branch is much shorter, erect and smooth); the distal edge of the valva is smooth (in
<taxonomicName id="B5FE4D43FF8B861C16C9FEDAD3CBFECB" authorityName="Devyatkin" authorityYear="2002" box="[1167,1262,296,320]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8B861C16C9FEDAD3CBFECB" box="[1167,1262,296,320]" italics="true" pageId="4" pageNumber="125">C. tanya</emphasis>
</taxonomicName>
it is serrated to the base of the erect branch); the coecum of the aedeagus is curved 45 degrees upwards (it is curved nearly 90 degrees in
<taxonomicName id="B5FE4D43FF8B861C1349FE82D6A9FE0C" authorityName=", Huang" authorityYear="2003" box="[271,396,368,391]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8B861C1349FE82D6A9FE0C" box="[271,396,368,391]" italics="true" pageId="4" pageNumber="125">C. uemurai</emphasis>
</taxonomicName>
); in lateral view, the ventral margin in opposite of the serrate ridge of the aedeagus is slightly concaved (it is more or less convex in
<taxonomicName id="B5FE4D43FF8B861C100AFE66D5EFFE27" authorityName=", Huang" authorityYear="2003" box="[588,714,404,428]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8B861C100AFE66D5EFFE27" box="[588,714,404,428]" italics="true" pageId="4" pageNumber="125">C. uemurai</emphasis>
</taxonomicName>
). (For figures of the male genitalia of
<taxonomicName id="B5FE4D43FF8B861C1638FE66D3FBFE27" authorityName="Devyatkin" authorityYear="2002" box="[1150,1246,404,428]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="tanya">
<emphasis id="408AEAD2FF8B861C1638FE66D3FBFE27" box="[1150,1246,404,428]" italics="true" pageId="4" pageNumber="125">C. tanya</emphasis>
</taxonomicName>
,
<taxonomicName id="B5FE4D43FF8B861C16ABFE66D24EFE27" authorityName=", Huang" authorityYear="2003" box="[1261,1387,404,428]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="uemurai">
<emphasis id="408AEAD2FF8B861C16ABFE66D24EFE27" box="[1261,1387,404,428]" italics="true" pageId="4" pageNumber="125">C. uemurai</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FE4D43FF8B861C12D1FE4AD629FE5B" authorityName="Huang &amp; Li" authorityYear="2006" box="[151,268,440,464]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="motuoa">
<emphasis id="408AEAD2FF8B861C12D1FE4AD629FE5B" box="[151,268,440,464]" italics="true" pageId="4" pageNumber="125">C. motuoa</emphasis>
</taxonomicName>
, see
<bibRefCitation id="166F4B31FF8B861C1303FE45D6D8FE5B" author="Devyatkin" box="[325,509,439,464]" pageId="4" pageNumber="125" refString="Devyatkin, A. L. (2002) Hesperiidae of Vietnam, 10. A new species of Coladenia Moore, [1881] (Lepidoptera, Hesperiidae). Atalanta, 33 (1 / 2), 123 - 125." type="journal article" year="2002">Devyatkin 2002</bibRefCitation>
,
<bibRefCitation id="166F4B31FF8B861C104EFE4AD5B0FE5B" author="Huang" box="[520,661,440,464]" pageId="4" pageNumber="125" refString="Huang, H. (2003) A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongjiang, China with descriptions of new species, new subspecies and revisional notes. Neue Entomologische Nachrichten, 55, 3 - 114." type="journal article" year="2003">Huang 2003</bibRefCitation>
,
<bibRefCitation id="166F4B31FF8B861C10E7FE4AD455FE5B" author="Huang" box="[673,880,439,464]" pageId="4" pageNumber="125" refString="Huang, H. &amp; Li, L. Z. (2006) A new subspecies of Coladenia uemurai Huang, 2003 from SE Tibet (Lepidoptera, Lycaenidae) Atalanta, 37 (3 / 4), 322 - 324." type="journal article" year="2006">Huang &amp; Li, 2006</bibRefCitation>
and
<bibRefCitation id="166F4B31FF8B861C11E1FE4AD358FE5B" author="Huang" box="[935,1149,439,464]" pageId="4" pageNumber="125" refString="Huang, H. &amp; Wu, Z. J. (2016) Report on some little known butterflies from SE Tibet and NW Yunnan. Atalanta, 47 (1 / 2), 174 - 178." type="journal article" year="2016">Huang &amp; Wu 2016</bibRefCitation>
respectively; distribution of these species are shown in fig. 20.)
</paragraph>
</subSubSection>
</treatment>
</document>