523 lines
46 KiB
XML
523 lines
46 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.81.13106" ID-PMC="PMC5523872" ID-Pensoft-Pub="1314-2003-81-79" ID-Pensoft-UUID="8A4FFF88B20CFF87FFA5FFC2FFA8FF88" ID-PubMed="28785166" ID-Zenodo-Dep="816476" ModsDocID="1314-2003-81-79" checkinTime="1502745056618" checkinUser="pensoft" docAuthor="Legler, Ben S. & Dillenberger, Markus S." docDate="2017" docId="FA55A2560F62553FB327EC5DB788E97A" docLanguage="en" docName="PhytoKeys 81: 79-102" docOrigin="PhytoKeys 81" docPubDate="2017-06-15" docSource="http://dx.doi.org/10.3897/phytokeys.81.13106" docTitle="Sabulina sororia B. S. Legler 2017, sp. nov." docType="treatment" docVersion="3" id="8A4FFF88B20CFF87FFA5FFC2FFA8FF88" lastPageNumber="92" masterDocId="8A4FFF88B20CFF87FFA5FFC2FFA8FF88" masterDocTitle="Two new species of Sabulina (Caryophyllaceae) from Washington State, U. S. A." masterLastPageNumber="102" masterPageNumber="79" pageNumber="86" updateTime="1668141985303" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of Sabulina (Caryophyllaceae) from Washington State, U. S. A.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Legler, Ben S.</mods:namePart>
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<mods:affiliation>WTU Herbarium, Burke Museum, Box 355325, University of Washington, Seattle, WA 98195 - 5325, U. S. A.</mods:affiliation>
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<mods:nameIdentifier type="email">blegler@u.washington.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dillenberger, Markus S.</mods:namePart>
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<mods:affiliation>Department of Botany and Plant Pathology, Oregon State University, 2082 Cordley Hall, Corvallis, OR 97331, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-06-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>79</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.81.13106</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.81.13106</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-81-79</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8A4FFF88B20CFF87FFA5FFC2FFA8FF88</mods:identifier>
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<mods:identifier type="Zenodo-Dep">816476</mods:identifier>
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||
</mods:mods>
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<subSection lastPageId="13" lastPageNumber="92" pageId="7" pageNumber="86" type="results">
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<treatment ID-GBIF-Taxon="182226062" LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A" httpUri="http://treatment.plazi.org/id/FA55A2560F62553FB327EC5DB788E97A" lastPageId="13" lastPageNumber="92" pageId="7" pageNumber="86">
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<subSubSection pageId="7" pageNumber="86" type="nomenclature">
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<paragraph pageId="7" pageNumber="86">
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<taxonomicName LSID="FA55A256-0F62-553F-B327-EC5DB788E97A" authority="B. S. Legler" authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="7" pageNumber="86" phylum="Tracheophyta" rank="species" species="sororia" status="sp. nov.">Sabulina sororia B.S.Legler</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="86">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Line drawings of Sabulina basaltica and Sabulina sororia. A-E. Sabulina basaltica. A Habit B Dried leaf with 3 veins C Cymose, bracteate inflorescence with two flowers D Capsule with dried, 3 - veined sepals (with sepals pushed outwards and withered petals removed to reveal capsule) E Seed F-I Sabulina sororia F Habit G Dried leaf with 1 vein H Flower I Capsule with dried, 3 - veined sepals (withered petals removed)." figureDoi="10.3897/phytokeys.81.13106.figure2" httpUri="https://binary.pensoft.net/fig/140638" pageId="7" pageNumber="86">Figs 2F -I</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sabulina sororia. A Plant forming a loose mat (Legler 14263) B Excavated plant with loosely sprawling stems (Legler 14268) C Fresh leaves and persisting, 1 - veined, dead leaves (Legler 14263) D Excavated plant (Legler 14263) E-F Cymose inflorescences (Legler 14263) G Dried, 1 - veined leaves (Legler 14263) H Dead leaves with only midvein persisting (Legler 14263) I Ovate-lanceolate, purple-tinged sepals (Legler 14263) J Sepals and petals, showing shapes and relative lengths (Legler 14263) K-L Flowers with different combinations of stamen and style lengths (Legler 14263) M Dried flower with dehisced capsule longer than sepals (Legler 14263) N Seeds (Legler 14263). Black scale bars are 1 mm." figureDoi="10.3897/phytokeys.81.13106.figure4" httpUri="https://binary.pensoft.net/fig/140640" pageId="7" pageNumber="86">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="87" type="materials_examined">
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<paragraph pageId="8" pageNumber="87">
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<pageBreakToken pageId="8" pageNumber="87" start="start">
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<typeStatus>Type</typeStatus>
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</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="8" pageNumber="87">
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<materialsCitation collectingDate="2016-08-06" collectorName="B. S. Legler" country="U. S. A." county="Whatcom Co." elevation="1414" latitude="48.70713" location="Sisters Divide" longLatPrecision="1" longitude="-121.93409" municipality="Mt. Baker-Snoqualmie National Forest" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<collectingCountry name="United States of America">U.S.A.</collectingCountry>
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<collectingRegion country="United States of America" name="Washington">Washington</collectingRegion>
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,
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<collectingCounty>Whatcom Co.</collectingCounty>
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:
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<collectingMunicipality>Mt. Baker-Snoqualmie National Forest</collectingMunicipality>
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, on west side of ridge along
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:BA714D4E9DCC669AE3FBD44F6AF3D8CA" country="U. S. A." county="Whatcom Co." latitude="48.70713" longLatPrecision="1" longitude="-121.93409" municipality="Mt. Baker-Snoqualmie National Forest" name="Sisters Divide" stateProvince="Washington">Sisters Divide</location>
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0.45 air km southeast of outlet of
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:88748367FCE0DFA9E23D9C2F2E69B682" country="U. S. A." county="Whatcom Co." latitude="48.70713" longLatPrecision="1" longitude="-121.93409" municipality="Mt. Baker-Snoqualmie National Forest" name="Lake Wiseman" stateProvince="Washington">Lake Wiseman</location>
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,
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:495471151A77FD49AC1F77B194E01411" country="U. S. A." county="Whatcom Co." latitude="48.70713" longLatPrecision="1" longitude="-121.93409" municipality="Mt. Baker-Snoqualmie National Forest" name="Twin Sisters Range" stateProvince="Washington">Twin Sisters Range</location>
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,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.414" unit="m" value="1414.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.414" unit="m" value="1414.0">1414 m</elevation>
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</quantity>
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,
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<geoCoordinate degrees="48.707131" direction="north" orientation="latitude" precision="1" value="48.70713">48.707131°N</geoCoordinate>
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,
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<geoCoordinate degrees="121.934086" direction="west" orientation="longitude" precision="1" value="-121.93409">121.934086°W</geoCoordinate>
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,
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<collectingDate value="2016-08-06">6 Aug 2016</collectingDate>
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,
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<emphasis italics="true" pageId="8" pageNumber="87">
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<collectorName>B.S. Legler</collectorName>
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14263
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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: WTU!; isotypes: MICH!, MO!, NY!, UBC!)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="87" type="diagnosis">
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<paragraph pageId="8" pageNumber="87">Diagnosis.</paragraph>
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<paragraph pageId="8" pageNumber="87">
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Differs from all other glabrous, perennial
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<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina" order="Caryophyllales" pageId="8" pageNumber="87" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="8" pageNumber="87">Sabulina</emphasis>
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</taxonomicName>
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species in North America by the combination of 1-veined dried leaves, flowers partly in 2-3-flowered cymes, sepals mostly <2.5 mm long, petals conspicuously longer than the sepals, capsules 1.8-2.6 mm long and mostly> sepals, and reddish-black seeds 0.6-0.8 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="89" pageId="8" pageNumber="87" type="description">
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<paragraph pageId="8" pageNumber="87">Description.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="89" pageId="8" pageNumber="87">
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<emphasis italics="true" pageId="8" pageNumber="87">Plants</emphasis>
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perennial, forming loose to dense mats 2-20 cm in diameter, glabrous throughout.
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<emphasis italics="true" pageId="8" pageNumber="87">Taproot</emphasis>
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slender to slightly thickened, 1-3 mm diameter near summit.
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<emphasis italics="true" pageId="8" pageNumber="87">Stems</emphasis>
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numerous, radially spreading from the taproot, prolifically branching; older stems decumbent to ascending, 1-10 cm, brown to tan; new shoots arising from axillary fascicles on previous
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<normalizedToken originalValue="year’s">year's</normalizedToken>
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stems, ascending to erect, 1-4 cm, internodes of flowering shoots 0.3-2(-3) times as long as leaves, deep green or purplish.
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<emphasis italics="true" pageId="8" pageNumber="87">Leaves</emphasis>
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slightly to strongly overlapping or well-spaced, connate proximally to form a tight, scarious sheath; blade 1.2-3.5(-5)
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<normalizedToken originalValue="×">x</normalizedToken>
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0.4-0.7 mm, ascending to spreading-ascending, straight to slightly incurved or slightly recurved, green to deep green, often maroon-tinged, shiny, subulate, rounded abaxially, nearly flat adaxially, veins not visible in life, margins rounded, not scarious, smooth, apex obtuse to rounded, usually maroon; axillary fascicles of leaves usually present; previous
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<normalizedToken originalValue="year’s">year's</normalizedToken>
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leaves loosely marcescent on
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<pageBreakToken pageId="9" pageNumber="88" start="start">older</pageBreakToken>
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stems, with only the midvein visible and persisting (no lateral veins).
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<emphasis italics="true" pageId="9" pageNumber="88">Inflorescences</emphasis>
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terminal, 2-3-flowered, open cymes, usually mixed with solitary terminal flowers; bracts 0.7-1.6 mm, lanceolate to ovate-lanceolate, incurved, green or maroon with scarious margins, rounded abaxially, flat to concave adaxially, apex obtuse to bluntly acute.
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<emphasis italics="true" pageId="9" pageNumber="88">Pedicels</emphasis>
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(1-)2-8(-15) mm, glabrous.
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<emphasis italics="true" pageId="9" pageNumber="88">Flowers</emphasis>
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perfect or functionally male or functionally female, most plants functionally monoecious to nearly dioecious.
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<emphasis italics="true" pageId="9" pageNumber="88">Hypanthium</emphasis>
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obscure, disc-shaped.
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<emphasis italics="true" pageId="9" pageNumber="88">Sepals</emphasis>
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spreading-ascending at anthesis, deep green, often lightly maroon-tinged, glabrous, broadly ovate to ovate-lanceolate, (1.4-)1.7-2.5(-3)
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<normalizedToken originalValue="×">x</normalizedToken>
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0.6-1.1(-1.3) mm, 1.5-2.5(-3.5) times as long as wide, scarious margins ca. 0.05-0.15 mm wide, base cupped, apex green to maroon, acute, outer surface convex, smooth to very weakly 3-veined at anthesis, becoming 3-veined in fruit or when dried.
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<emphasis italics="true" pageId="9" pageNumber="88">Petals</emphasis>
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white, spreading, broadly oblong to obovate, 3.2-4(-5.2)
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<normalizedToken originalValue="×">x</normalizedToken>
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1.2-2(-2.6) mm, 1.3-2(-2.5) times as long as sepals, base gradually tapered to a short, greenish-yellow claw, apex rounded to weakly truncate.
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<emphasis italics="true" pageId="9" pageNumber="88">Nectaries</emphasis>
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5, at base of outer stamens, greenish-yellow, ca. 0.3-0.4 mm, truncate, alternate with the petals.
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<emphasis italics="true" pageId="9" pageNumber="88">Stamens</emphasis>
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10, in 2 series of
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<pageBreakToken pageId="10" pageNumber="89" start="start">5</pageBreakToken>
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, either all fertile or all abortive; filaments subulate, whitish-green; anthers orbiculate, pale yellow; fertile stamens with filaments 1.5-2.8 mm and anthers (0.3-)0.4-0.5 mm; abortive stamens with filaments 0.2-0.6 mm and anthers 0.1-0.3 mm.
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<emphasis italics="true" pageId="10" pageNumber="89">Ovary</emphasis>
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superior; placentation shortly free-central; ovules usually 12 per ovary.
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<emphasis italics="true" pageId="10" pageNumber="89">Styles</emphasis>
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3, distinct, erect to ascending; functionally male flowers with styles 0.6-0.9 mm and stigmas scarcely developed; functionally female flowers with styles 1.1-2.1 mm and stigmas linear, glandular-puberulent adaxially.
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<emphasis italics="true" pageId="10" pageNumber="89">Capsules</emphasis>
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light green to greenish-tan (valve margins tan), on stipe ca. 0.1-0.2 mm, ovoid-conical, 1.8-2.6
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<normalizedToken originalValue="×">x</normalizedToken>
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1.1-1.8 mm slightly longer than (rarely slightly shorter than) and mostly enclosed by the appressed sepals and withering-persistent petals, dehiscing in upper half by 3 valves, these becoming incurved on margins and slightly recurved at tip.
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<emphasis italics="true" pageId="10" pageNumber="89">Seeds</emphasis>
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apparently 8 per capsule, 0.7-0.8 mm, reddish-black, obliquely reniform with radicle prolonged into a curved bump, somewhat compressed, surfaces sculpted with low bumps at> 10
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="89" type="materials_examined">
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<paragraph pageId="10" pageNumber="89">Additional specimens examined.</paragraph>
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<paragraph pageId="10" pageNumber="89">
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<materialsCitation collectingDate="1939-08-11" collectingDateMax="2016-08-07" collectingDateMin="1939-08-11" collectorName="W. C. Muenscher, A. R. Kruckeberg, R. J. Taylor, B. S. Legler" country="U. S. A." county="Whatcom Co." elevation="1494" latitude="48.704998" location="Head of Orsina Creek" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" specimenCount="1" stateProvince="Washington">
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||
<emphasis bold="true" pageId="10" pageNumber="89">
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<collectingCountry name="United States of America">U.S.A.</collectingCountry>
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<collectingRegion country="United States of America" name="Washington">Washington</collectingRegion>
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,
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<collectingCounty>Whatcom Co.</collectingCounty>
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</emphasis>
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:
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<collectingCounty>Twin Sisters Range</collectingCounty>
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,
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<collectingDate value="1939-08-11">11 Aug 1939</collectingDate>
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,
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<emphasis italics="true" pageId="10" pageNumber="89">
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<collectorName>W.C. Muenscher</collectorName>
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10281
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</emphasis>
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||
(WTU);
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<collectingMunicipality>Twin Sisters Range</collectingMunicipality>
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,
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<collectingDate value="1939-08-12">12 Aug 1939</collectingDate>
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,
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||
<emphasis italics="true" pageId="10" pageNumber="89">
|
||
<collectorName>W.C. Muenscher</collectorName>
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||
10306
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||
</emphasis>
|
||
(WTU);
|
||
<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:C53D7F42E0809ADCBDFDF3100AF0568D" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Head of Orsina Creek" stateProvince="Washington">Head of Orsina Creek</location>
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||
, at west base of
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:88A4AF9C6AEA77DD0DBD83CB91D675EE" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Twin Sisters Mountain" stateProvince="Washington">Twin Sisters Mountain</location>
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||
,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.49352" unit="ft" value="4900.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.49352" unit="ft" value="4900.0">4900 ft</elevation>
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</quantity>
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, T37N R6E S11,
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<collectingDate value="1961-07-12">12 Jul 1961</collectingDate>
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,
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<emphasis italics="true" pageId="10" pageNumber="89">
|
||
<collectorName>A.R. Kruckeberg</collectorName>
|
||
5225
|
||
</emphasis>
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||
(WTU);
|
||
<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:AFB168BD7D5FFDB7BE12F3B1EA55CB79" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Northwest" stateProvince="Washington">Northwest</location>
|
||
slope of
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||
<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:82A1FF21160CBD5CDEF10B1452878CFE" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Twin Sisters" stateProvince="Washington">Twin Sisters</location>
|
||
, ca.
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||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" unit="ft" value="6200.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" unit="ft" value="6200.0">6200 ft</elevation>
|
||
</quantity>
|
||
,
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||
<collectingDate value="1968-07-28">28 Jul 1968</collectingDate>
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||
,
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||
<emphasis italics="true" pageId="10" pageNumber="89">
|
||
<collectorName>R.J. Taylor</collectorName>
|
||
2158
|
||
</emphasis>
|
||
(WWB);
|
||
<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:6BC04746ED4392BBDA9DC0007705B1C8" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Crest" stateProvince="Washington">Crest</location>
|
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of ridge along
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:0403088E2738441979ABDC387216DD4D" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Sisters Divide" stateProvince="Washington">Sisters Divide</location>
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0.7 air km southeast of outlet of
|
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:184B0DC02B2A8D77D16CFCD74915B2A8" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Lake Wiseman" stateProvince="Washington">Lake Wiseman</location>
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,
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<location LSID="urn:lsid:plazi:treatment:FA55A2560F62553FB327EC5DB788E97A:53CBDFB0F24C505744428975601DA996" country="U. S. A." county="Whatcom Co." latitude="48.704998" longLatPrecision="1" longitude="-121.93141" municipality="Twin Sisters Range" name="Twin Sisters Range" stateProvince="Washington">Twin Sisters Range</location>
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,
|
||
<geoCoordinate degrees="48.704998" direction="north" orientation="latitude" precision="1" value="48.704998">48.704998°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="121.931408" direction="west" orientation="longitude" precision="1" value="-121.93141">121.931408°W</geoCoordinate>
|
||
;
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.508" unit="m" value="1508.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.508" unit="m" value="1508.0">1508 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="2016-08-07">7 Aug 2016</collectingDate>
|
||
,
|
||
<emphasis italics="true" pageId="10" pageNumber="89">
|
||
<collectorName>B.S. Legler</collectorName>
|
||
14268
|
||
</emphasis>
|
||
(ID, US, WTU)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="89" type="etymology">
|
||
<paragraph pageId="10" pageNumber="89">Etymology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="89">
|
||
The epithet
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="sororia" order="Caryophyllales" pageId="10" pageNumber="89" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="10" pageNumber="89">sororia</emphasis>
|
||
</taxonomicName>
|
||
is from the Latin word
|
||
<emphasis italics="true" pageId="10" pageNumber="89">sororis</emphasis>
|
||
, sister, in reference to the Twin Sisters Range.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="89" type="vernacular name">
|
||
<paragraph pageId="10" pageNumber="89">Vernacular name.</paragraph>
|
||
<paragraph pageId="10" pageNumber="89">Twin Sisters sandwort.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="13" lastPageNumber="92" pageId="10" pageNumber="89" type="distribution">
|
||
<paragraph pageId="10" pageNumber="89">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="89">
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="10" pageNumber="89" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="10" pageNumber="89">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
is known only from the Twin Sisters Range on the western flank of the Cascade Mountains in Whatcom County, Washington, U.S.A. (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Sabulina basaltica and S. sororia. A-B Reference maps of western North America and Washington State, indicating locations of inset maps C Known locations for Sabulina basaltica (open black circles) within the northeastern Olympic Mountains; gray shading indicates the predicted extent of potential habitat based on exposures of oceanic basalt rocks at subalpine to alpine elevations D Known locations for Sabulina sororia (open black squares) within the Twin Sisters Range; gray shading indicates the predicted extent of potential habitat based on exposures of dunite rock away from large snowfields at subalpine to alpine elevations." figureDoi="10.3897/phytokeys.81.13106.figure6" httpUri="https://binary.pensoft.net/fig/140642" pageId="10" pageNumber="89">6D</figureCitation>
|
||
). The Twin Sisters Range consists of a large body of relatively unaltered dunite rock aproximately 16 km long by 6.5 km wide (
|
||
<bibRefCitation author="Tabor, RW" journalOrPublisher="Systematic Biology" pageId="20" pageNumber="99" refId="B34" refString="Tabor, RW, Haugerud, RA, Hildreth, W, Brown, EH, 2003. Geologic Map of the Mount Baker 30- by 60-minute quadrangle, Washington. U.S. Geological Survey Investigation Series I-2660." title="Geologic Map of the Mount Baker 30 - by 60 - minute quadrangle, Washington. U. S. Geological Survey Investigation Series I- 2660." year="2003">Tabor et al. 2003</bibRefCitation>
|
||
) oriented in a northwest to southeast direction, with a maximum elevation of 2135 meters and sustained ridgeline elevations above 1500 meters. The dunite rock likely formed in the
|
||
<normalizedToken originalValue="earth’s">earth's</normalizedToken>
|
||
mantle and was subsequently uplifted along a series of nearly vertical thrust faults (
|
||
<bibRefCitation DOI="https://doi.org/10.2475/ajs.261.6.549" author="Ragan, DM" journalOrPublisher="American Journal of Science" pageId="20" pageNumber="99" pagination="549 - 565" refId="B30" refString="Ragan, DM, 1963. Emplacement of the Twin Sisters Dunite. American Journal of Science 261 (6): 549 - 565, DOI: https://doi.org/10.2475/ajs.261.6.549" title="Emplacement of the Twin Sisters Dunite." url="https://doi.org/10.2475/ajs.261.6.549" volume="261" year="1963">Ragan 1963</bibRefCitation>
|
||
); it is a dense, crystalline, ultramafic rock composed mostly of olivine with lesser amounts of chromite and pyroxenite, rich in magnesium, iron, chromium and nickel (
|
||
<bibRefCitation DOI="https://doi.org/10.2475/ajs.261.6.549" author="Ragan, DM" journalOrPublisher="American Journal of Science" pageId="20" pageNumber="99" pagination="549 - 565" refId="B30" refString="Ragan, DM, 1963. Emplacement of the Twin Sisters Dunite. American Journal of Science 261 (6): 549 - 565, DOI: https://doi.org/10.2475/ajs.261.6.549" title="Emplacement of the Twin Sisters Dunite." url="https://doi.org/10.2475/ajs.261.6.549" volume="261" year="1963">Ragan 1963</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1130/0016-7606(1978)89<1459:TSDPAM>2.0.CO;2" author="Onyeagocha, AC" journalOrPublisher="Geological Society of America Bulletin" pageId="20" pageNumber="99" pagination="1459 - 1474" refId="B23" refString="Onyeagocha, AC, 1978. Twin Sisters dunite: Petrology and mineral chemistry. Geological Society of America Bulletin 89 (10): 1459 - 1474, DOI: https://doi.org/10.1130/0016-7606(1978)89<1459:TSDPAM>2.0.CO;2" title="Twin Sisters dunite: Petrology and mineral chemistry." url="https://doi.org/10.1130/0016-7606(1978)89<1459:TSDPAM>2.0.CO;2" volume="89" year="1978">Onyeagocha 1978</bibRefCitation>
|
||
). The rock weathers to a distinctive light reddish-brown color with a coarse-grained surface. Ultramafic rocks display a pronounced effect on the overlying vegetation (
|
||
<bibRefCitation author="Kruckeberg, AR" journalOrPublisher="University of Washington Press, Seattle, Washington" pageId="19" pageNumber="98" refId="B20" refString="Kruckeberg, AR, 2002. Geology and Plant Life: The Effects of Landforms and Rock Types on Plants. University of Washington Press, Seattle, Washington" title="Geology and Plant Life: The Effects of Landforms and Rock Types on Plants." year="2002">Kruckeberg 2002</bibRefCitation>
|
||
, and references therein), and the Twin Sisters dunite is no exception with its depressed treeline and sparse vegetation cover above treeline.
|
||
</paragraph>
|
||
<paragraph lastPageId="11" lastPageNumber="90" pageId="10" pageNumber="89">
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="10" pageNumber="89" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="10" pageNumber="89">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
is apparently restricted to rocky or gravelly, sparsely vegetated, subalpine and alpine slopes. Documented elevations range from 1490 to 1890 meters. Habitat information from older herbarium specimens is sparse, indicating only a "west-facing alpine ridgeline" (
|
||
<emphasis italics="true" pageId="10" pageNumber="89">Grable 5023</emphasis>
|
||
), "moist, gravelly, serpentine soil on an alpine slope" (
|
||
<emphasis italics="true" pageId="10" pageNumber="89">Taylor 2158</emphasis>
|
||
), "along streambank" (
|
||
<emphasis italics="true" pageId="10" pageNumber="89">Muenscher 10281</emphasis>
|
||
), and "olivine in massive
|
||
<pageBreakToken pageId="11" pageNumber="90" start="start">fell-fields</pageBreakToken>
|
||
and talus, with krummholz lodgepole pine and subalpine fir in snow-melt basin" (
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Kruckeberg 5225</emphasis>
|
||
). At the two sites visited by B. Legler in August 2016,
|
||
<taxonomicName lsidName="S. sororia" pageId="11" pageNumber="90" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">S. sororia</emphasis>
|
||
</taxonomicName>
|
||
was observed growing most frequently in mesic, coarse, gravelly and rocky soil derived from dunite on erosional surfaces with slopes ranging from flat to ca. 30° (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Representative habitats. A-C Sabulina basaltica habitat A Basalt slope near the type locality (Legler 14177) B-C Plants in crevices of basalt rock faces (Legler 14177, Legler 14183) D-F Sabulina sororia habitat D Reddish-colored dunite slope at the type locality (Legler 14263) E-F Plants among dunite rock and gravel (Legler 14263, Legler 14268)." figureDoi="10.3897/phytokeys.81.13106.figure5" httpUri="https://binary.pensoft.net/fig/140641" pageId="11" pageNumber="90">5D-F</figureCitation>
|
||
). A few plants were found in exposed crevices of stable dunite rock outcrops along a narrow ridgeline with slopes of ca. 45-60°. The species apparently avoids areas with late-lying snow.
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="11" pageNumber="90" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
occurs as scattered individuals, forming a minor component of the sparse, low vegetation cover. Total vegetation cover of all plant species at these two sites is estimated at 5-20%. Average precipitation for the higher elevations of the Twin Sisters Range is estimated at ca. 180-190 cm per year, with about 30% of the total precipitation falling during May-September (
|
||
<bibRefCitation author="PRISM" journalOrPublisher="Quaternary Research" pageId="20" pageNumber="99" publicationUrl="http://prism.oregonstate.edu" refId="B27" refString="PRISM, 2017. PRISM Climate Group, Oregon State University. http://prism.oregonstate.edu" title="PRISM Climate Group, Oregon State University." url="http://prism.oregonstate.edu" year="2017">PRISM 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph lastPageId="12" lastPageNumber="91" pageId="11" pageNumber="90">
|
||
Directly associated species consist of scattered tufts or mats of
|
||
<taxonomicName class="Monocotyledoneae" family="Cyperaceae" genus="Carex" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carex spectabilis" order="Cyperales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="spectabilis">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Carex spectabilis</emphasis>
|
||
</taxonomicName>
|
||
Dewey,
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" genus="Cassiope" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cassiope mertensiana" order="Ericales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="mertensiana">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Cassiope mertensiana</emphasis>
|
||
</taxonomicName>
|
||
(Bong.) G. Don,
|
||
<taxonomicName class="Eudicots" family="Caryophyllaceae" genus="Cerastium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cerastium arvense subsp. strictum" order="Caryophyllales" pageId="11" pageNumber="90" phylum="Magnoliophyta" rank="subSpecies" species="arvense" subSpecies="strictum">Cerastium arvense L. subsp. strictum</taxonomicName>
|
||
Gaudin,
|
||
<taxonomicName class="Filicopsida" family="Pteridaceae" genus="Cryptogramma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cryptogramma acrostichoides" order="Polypodiales" pageId="11" pageNumber="90" phylum="Pteridophyta" rank="species" species="acrostichoides">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Cryptogramma acrostichoides</emphasis>
|
||
</taxonomicName>
|
||
R. Br.,
|
||
<taxonomicName authorityName="Vasey" authorityYear="1883" class="Monocotyledoneae" family="Poaceae" genus="Danthonia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Danthonia intermedia" order="Graminales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="intermedia">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Danthonia intermedia</emphasis>
|
||
</taxonomicName>
|
||
Vasey,
|
||
<taxonomicName authorityName="Greene" authorityYear="1891" class="Magnoliopsida" family="Asteraceae" genus="Erigeron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Erigeron aureus" order="Asterales" pageId="11" pageNumber="90" phylum="Tracheophyta" rank="species" species="aureus">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Erigeron aureus</emphasis>
|
||
</taxonomicName>
|
||
Greene,
|
||
<taxonomicName authorityName="Underw" authorityYear="1900" class="Bryopsida" family="Dryopteridaceae" genus="Polystichum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Polystichum lemmonii" order="Polypodiales" pageId="11" pageNumber="90" phylum="Bryophyta" rank="species" species="lemmonii">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Polystichum lemmonii</emphasis>
|
||
</taxonomicName>
|
||
Underw.,
|
||
<taxonomicName authorityName="Dillenb & Kadereit" authorityYear="2014" baseAuthorityName="Wahlenb." class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina rubella" order="Caryophyllales" pageId="11" pageNumber="90" phylum="Tracheophyta" rank="species" species="rubella">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Sabulina rubella</emphasis>
|
||
</taxonomicName>
|
||
(Wahlenb.) Dillenb. & Kadereit,
|
||
<taxonomicName class="Dicotyledoneae" family="Saxifragaceae" genus="Saxifraga" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Saxifraga cespitosa" order="Rosales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="cespitosa">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Saxifraga cespitosa</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Sibbaldia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sibbaldia procumbens" order="Rosales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="procumbens">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Sibbaldia procumbens</emphasis>
|
||
</taxonomicName>
|
||
L.,
|
||
<taxonomicName class="Eudicots" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Silene acaulis" order="Caryophyllales" pageId="11" pageNumber="90" phylum="Magnoliophyta" rank="species" species="acaulis">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Silene acaulis</emphasis>
|
||
</taxonomicName>
|
||
(L.) Jacq., and
|
||
<taxonomicName class="Dicotyledoneae" family="Brassicaceae" genus="Smelowskia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Smelowskia ovalis" order="Papaverales" pageId="11" pageNumber="90" phylum="Angiospermae" rank="species" species="ovalis">
|
||
<emphasis italics="true" pageId="11" pageNumber="90">Smelowskia ovalis</emphasis>
|
||
</taxonomicName>
|
||
Rydb. Trees and taller shrubs are absent from these sites, though
|
||
<pageBreakToken pageId="12" pageNumber="91" start="start">adjacent</pageBreakToken>
|
||
ridgelines and slopes hold patches of
|
||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Abies lasiocarpa" order="Pinales" pageId="12" pageNumber="91" phylum="Coniferophyta" rank="species" species="lasiocarpa">
|
||
<emphasis italics="true" pageId="12" pageNumber="91">Abies lasiocarpa</emphasis>
|
||
</taxonomicName>
|
||
(Hook.) Nutt.,
|
||
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Callitropsis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Callitropsis nootkatensis" order="Cupressales" pageId="12" pageNumber="91" phylum="Tracheophyta" rank="species" species="nootkatensis">
|
||
<emphasis italics="true" pageId="12" pageNumber="91">Callitropsis nootkatensis</emphasis>
|
||
</taxonomicName>
|
||
(D. Don) D.P. Little,
|
||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pinus contorta subsp. var. var. latifolia" order="Pinales" pageId="12" pageNumber="91" phylum="Coniferophyta" rank="variety" species="contorta" subSpecies="var." variety="latifolia">Pinus contorta Douglas ex Loudon var. latifolia</taxonomicName>
|
||
Engelm.,
|
||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Tsuga" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tsuga mertensiana" order="Pinales" pageId="12" pageNumber="91" phylum="Coniferophyta" rank="species" species="mertensiana">
|
||
<emphasis italics="true" pageId="12" pageNumber="91">Tsuga mertensiana</emphasis>
|
||
</taxonomicName>
|
||
(Bong.)
|
||
<normalizedToken originalValue="Carrière">Carriere</normalizedToken>
|
||
,
|
||
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juniperus communis subsp. var. var. kelleyi" order="Pinales" pageId="12" pageNumber="91" phylum="Coniferophyta" rank="variety" species="communis" subSpecies="var." variety="kelleyi">Juniperus communis L. var. kelleyi</taxonomicName>
|
||
R.P. Adams, and
|
||
<taxonomicName authorityName="D.Don" authorityYear="1834" class="Dicotyledoneae" family="Ericaceae" genus="Phyllodoce" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Phyllodoce empetriformis" order="Ericales" pageId="12" pageNumber="91" phylum="Angiospermae" rank="species" species="empetriformis">
|
||
<emphasis italics="true" pageId="12" pageNumber="91">Phyllodoce empetriformis</emphasis>
|
||
</taxonomicName>
|
||
(Sm.) D. Don. Crustose lichens are sparse, and bryophytes nearly absent.
|
||
</paragraph>
|
||
<paragraph lastPageId="13" lastPageNumber="92" pageId="12" pageNumber="91">
|
||
The southern terminus of the Twin Sisters Range extends slightly into adjacent Skagit County, and 6 km farther to the southeast of this are two smaller dunite bodies
|
||
<pageBreakToken pageId="13" pageNumber="92" start="start">exposed</pageBreakToken>
|
||
at slightly lower elevation (
|
||
<bibRefCitation author="Tabor, RW" journalOrPublisher="Systematic Biology" pageId="20" pageNumber="99" refId="B34" refString="Tabor, RW, Haugerud, RA, Hildreth, W, Brown, EH, 2003. Geologic Map of the Mount Baker 30- by 60-minute quadrangle, Washington. U.S. Geological Survey Investigation Series I-2660." title="Geologic Map of the Mount Baker 30 - by 60 - minute quadrangle, Washington. U. S. Geological Survey Investigation Series I- 2660." year="2003">Tabor et al. 2003</bibRefCitation>
|
||
). An examination of aerial imagery suggests marginally suitable habitat for
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
may occur in these areas, though no surveys have been conducted to determine its presence. It seems unlikely that
|
||
<taxonomicName lsidName="S. sororia" pageId="13" pageNumber="92" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">S. sororia</emphasis>
|
||
</taxonomicName>
|
||
will be found elsewhere in the Cascades Mountains or over non-dunite rocks; however, small, subalpine exposures of ultramaphic rocks in Skagit and Snohomish counties may warrant investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="13" pageNumber="92" type="phenology">
|
||
<paragraph pageId="13" pageNumber="92">Phenology.</paragraph>
|
||
<paragraph pageId="13" pageNumber="92">
|
||
Specimens indicate the flowering period for
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
extends from mid July to mid August, and fruiting period from early to mid August. The full ranges of flowering and fruiting periods likely vary based on timing of snowmelt and site exposure.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="13" pageNumber="92" type="conservation status">
|
||
<paragraph pageId="13" pageNumber="92">Conservation status.</paragraph>
|
||
<paragraph pageId="13" pageNumber="92">
|
||
Although apparently restricted to the Twin Sisters Range,
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
may occur in suitable microsites throughout the upper elevations of the range within an extent of occurrence estimated at ca. 16 km2. The total number of plants cannot be estimated due to inadequate sampling across the range, possibly preventing assignment of a formal conservation status at this time. The Twin Sisters Range lies almost fully within the Mt. Baker-Snoqualmie National Forest, and the entire northeastern slope of the range lies within the Mt. Baker Wilderness. No roads or trails penetrate the range, resource extraction is absent from the higher elevations, and very few people visit each year due to difficulty of access. Direct anthropogenic impacts are therefore assumed to be very minimal.
|
||
</paragraph>
|
||
<caption doi="10.3897/phytokeys.81.13106.figure4" httpUri="https://binary.pensoft.net/fig/140640" pageId="13" pageNumber="92" start="Figure 4" startId="F4">
|
||
<paragraph pageId="13" pageNumber="92">
|
||
<emphasis bold="true" pageId="13" pageNumber="92">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="13" pageNumber="92">A</emphasis>
|
||
Plant forming a loose mat (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">B</emphasis>
|
||
Excavated plant with loosely sprawling stems (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14268</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">C</emphasis>
|
||
Fresh leaves and persisting, 1-veined, dead leaves (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">D</emphasis>
|
||
Excavated plant (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">E-F</emphasis>
|
||
Cymose inflorescences (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">G</emphasis>
|
||
Dried, 1-veined leaves (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">H</emphasis>
|
||
Dead leaves with only midvein persisting (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">I</emphasis>
|
||
Ovate-lanceolate, purple-tinged sepals (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">J</emphasis>
|
||
Sepals and petals, showing shapes and relative lengths (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">K-L</emphasis>
|
||
Flowers with different combinations of stamen and style lengths (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">M</emphasis>
|
||
Dried flower with dehisced capsule longer than sepals (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">N</emphasis>
|
||
Seeds (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
). Black scale bars are 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/phytokeys.81.13106.figure5" httpUri="https://binary.pensoft.net/fig/140641" pageId="13" pageNumber="92" start="Figure 5" startId="F5">
|
||
<paragraph pageId="13" pageNumber="92">
|
||
<emphasis bold="true" pageId="13" pageNumber="92">Figure 5.</emphasis>
|
||
Representative habitats.
|
||
<emphasis bold="true" pageId="13" pageNumber="92">A-C</emphasis>
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina basaltica" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="basaltica">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina basaltica</emphasis>
|
||
</taxonomicName>
|
||
habitat
|
||
<emphasis bold="true" pageId="13" pageNumber="92">A</emphasis>
|
||
Basalt slope near the type locality (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14177</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">B-C</emphasis>
|
||
Plants in crevices of basalt rock faces (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14177</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14183</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">D-F</emphasis>
|
||
<taxonomicName authorityName="B. S. Legler" authorityYear="2017" class="Magnoliopsida" family="Caryophyllaceae" genus="Sabulina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sabulina sororia" order="Caryophyllales" pageId="13" pageNumber="92" phylum="Tracheophyta" rank="species" species="sororia">
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Sabulina sororia</emphasis>
|
||
</taxonomicName>
|
||
habitat
|
||
<emphasis bold="true" pageId="13" pageNumber="92">D</emphasis>
|
||
Reddish-colored dunite slope at the type locality (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="13" pageNumber="92">E-F</emphasis>
|
||
Plants among dunite rock and gravel (
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14263</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="13" pageNumber="92">Legler 14268</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |