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<mods:title id="BA70CA5EAA8A0039E62992984ED374AF">On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae)</mods:title>
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<mods:affiliation id="E8B4E6D1D8E223A9C6371A599DC588C1">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. &amp; Department of Biology and Ecology; the Research Centre for Environmental “ Monitoring ”, Toraighyrov University, Lomova str. 64, KZ- 140008, Pavlodar, Kazakhstan. &amp; Scientific Center “ Bioarchaeology ”, Institute of Archaeological Research, Margulan University, Olzhabay Batyr str. 60, KZ- 140002, Pavlodar, Kazakhstan. &amp; sandibicumar @ gmail. com; https: // orcid. org / 0000 - 0003 - 2196 - 280 X</mods:affiliation>
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Genus
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<emphasis id="4089EAA4FFD5A627FF4D4DC3FEA332FA" bold="true" box="[238,354,655,682]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
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<taxonomicName id="B5FD4D35FFD5A627FF344D98FE7232BC" ID-CoL="92CJF" authority="Staudinger, 1882" authorityName="Staudinger" authorityYear="1882" box="[151,435,724,748]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FF344D98FF3932BC" box="[151,248,724,748]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
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,
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, 43: 39
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(
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species:
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<emphasis id="4089EAA4FFD5A627FBAC4D98FB7A32BC" box="[1039,1211,724,748]" italics="true" pageId="2" pageNumber="411">Isochlora viridis</emphasis>
<bibRefCitation id="166C4B47FFD5A627FB674D98FAB432BC" author="Staudinger, O." box="[1220,1397,724,748]" pageId="2" pageNumber="411" pagination="35 - 78" refId="ref8501" refString="Staudinger, O. (1882) Beitrag zur Lepidopterenfauna Central-Asiens. Stettiner entomologische Zeitung, 43, 35 - 78. [in German]" type="journal article" year="1882">Staudinger, 1882</bibRefCitation>
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, by monotypy).
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=
<taxonomicName id="B5FD4D35FFD5A627FF0F4C5DFE6A3379" ID-CoL="929XZ" authority="Alpheraky, 1892" authorityName="Alpheraky" authorityYear="1892" box="[172,427,785,809]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FF0F4C5DFF3A3378" box="[172,251,785,808]" italics="true" pageId="2" pageNumber="411">Grumia</emphasis>
<bibRefCitation id="166C4B47FFD5A627FEA34C5DFE6A3379" author="Alpheraky, S." box="[256,427,785,809]" pageId="2" pageNumber="411" pagination="444 - 459" refId="ref7412" refString="Alpheraky, S. (1892) Lepidoptera nova a Gr. Grum-Grshimailo in Asia Centrali novissime lecta. Horae Societatis entomologicae Rossicae, 26 (3 - 4), 444 - 459. [in Latin]" type="journal article" year="1892">Alphéraky, 1892</bibRefCitation>
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,
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Horae Societatis entomologicae vossicae, variis semonibus in
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usitatis editae
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, 26 (34): 453
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(
<typeStatus id="AD468814FFD5A627FF6D4C7CFEC33318" box="[206,258,816,840]" pageId="2" pageNumber="411">Type</typeStatus>
species:
<taxonomicName id="B5FD4D35FFD5A627FEFC4C7CFD543318" ID-CoL="3HF84" authority="Alpheraky, 1892" authorityName="Alpheraky" authorityYear="1892" box="[351,661,816,840]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD5A627FEFC4C7CFE243318" box="[351,485,816,840]" italics="true" pageId="2" pageNumber="411">Grumia flora</emphasis>
<bibRefCitation id="166C4B47FFD5A627FE494C7CFD543318" author="Alpheraky, S." box="[490,661,816,840]" pageId="2" pageNumber="411" pagination="444 - 459" refId="ref7412" refString="Alpheraky, S. (1892) Lepidoptera nova a Gr. Grum-Grshimailo in Asia Centrali novissime lecta. Horae Societatis entomologicae Rossicae, 26 (3 - 4), 444 - 459. [in Latin]" type="journal article" year="1892">Alphéraky, 1892</bibRefCitation>
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, by monotypy),
<emphasis id="4089EAA4FFD5A627FC984C7CFC523318" bold="true" box="[827,915,816,840]" pageId="2" pageNumber="411">syn. rev.</emphasis>
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<paragraph id="724236B6FFD5A627FF344C03FC2E3336" blockId="2.[151,1437,724,901]" box="[151,1007,846,870]" pageId="2" pageNumber="411">
=
<taxonomicName id="B5FD4D35FFD5A627FF0D4C02FE023336" ID-CoL="92548" authority="Staudinger, 1900" authorityName="Staudinger" authorityYear="1900" box="[174,451,846,870]" class="Insecta" family="Noctuidae" genus="Chamyla" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FF0D4C02FEC83336" box="[174,265,846,870]" italics="true" pageId="2" pageNumber="411">Chamyla</emphasis>
<bibRefCitation id="166C4B47FFD5A627FEB04C02FE023336" author="Staudinger, O." box="[275,451,846,870]" pageId="2" pageNumber="411" pagination="331 - 351" refId="ref8555" refString="Staudinger, O. (1900) Ueber Lepidoptereren aus dem ostlichsten Thian Schan-Gebiet. Deutsche entomologische Zeitschrift Iris, 12 (2), 331 - 351. [in German]" type="journal article" year="1900">Staudinger, 1900</bibRefCitation>
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,
<emphasis id="4089EAA4FFD5A627FE734C02FCB23336" box="[464,883,846,870]" italics="true" pageId="2" pageNumber="411">Deutsche entomologische Zeitschrift Iris</emphasis>
, 12(2): 343
</paragraph>
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<paragraph id="724236B6FFD5A627FC5B4C02FE3B33D5" blockId="2.[151,1437,724,901]" pageId="2" pageNumber="411">
(
<typeStatus id="AD468814FFD5A627FC5C4C02FBF23336" box="[1023,1075,846,870]" pageId="2" pageNumber="411">Type</typeStatus>
species:
<taxonomicName id="B5FD4D35FFD5A627FB364C02FF3D33D5" ID-CoL="9695L" authority="Staudinger, 1900" authorityName="Staudinger" authorityYear="1900" class="Insecta" family="Noctuidae" genus="Chamyla" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="idia">
<emphasis id="4089EAA4FFD5A627FB364C02FAE13336" box="[1173,1312,846,870]" italics="true" pageId="2" pageNumber="411">Chamyla idia</emphasis>
<bibRefCitation id="166C4B47FFD5A627FA8A4C02FF3D33D5" author="Staudinger, O." pageId="2" pageNumber="411" pagination="331 - 351" refId="ref8555" refString="Staudinger, O. (1900) Ueber Lepidoptereren aus dem ostlichsten Thian Schan-Gebiet. Deutsche entomologische Zeitschrift Iris, 12 (2), 331 - 351. [in German]" type="journal article" year="1900">Staudinger, 1900</bibRefCitation>
</taxonomicName>
, by monotypy),
<emphasis id="4089EAA4FFD5A627FE014C21FE3B33D5" bold="true" box="[418,506,877,901]" pageId="2" pageNumber="411">syn. rev.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3AE7653DFFD5A627FF344CFFFEA8340D" pageId="2" pageNumber="411" type="reference_group">
<paragraph id="724236B6FFD5A627FF344CFFFEA8340D" blockId="2.[151,1436,947,1117]" pageId="2" pageNumber="411">
<emphasis id="4089EAA4FFD5A627FF344CFFFF1B339D" bold="true" box="[151,218,947,973]" pageId="2" pageNumber="411">Notes</emphasis>
.
<emphasis id="4089EAA4FFD5A627FF4B4CFFFEC9339D" bold="true" box="[232,264,947,973]" pageId="2" pageNumber="411">(1)</emphasis>
The
<typeStatus id="AD468814FFD5A627FEE34CF8FEB1339E" box="[320,368,948,974]" pageId="2" pageNumber="411">type</typeStatus>
species of
<taxonomicName id="B5FD4D35FFD5A627FE4F4CFFFD90339D" ID-CoL="92548" authorityName="Staudinger" authorityYear="1900" box="[492,593,947,973]" class="Insecta" family="Noctuidae" genus="Chamyla" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FE4F4CFFFD90339D" box="[492,593,947,973]" italics="true" pageId="2" pageNumber="411">Chamyla</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD5A627FDC34CFFFD13339D" box="[608,722,947,973]" captionStart="FIGURES 912" captionStartId="5.[151,264,1763,1787]" captionTargetBox="[196,1404,948,1728]" captionTargetId="figure-303@5.[184,1404,930,1740]" captionTargetPageId="5" captionText="FIGURES 912. Isochlora arctomys: adults. Depositories of the specimens: 911 in ZISP; 12 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152645" httpUri="https://zenodo.org/record/10152645/files/figure.png" pageId="2" pageNumber="411">Figs 912</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FD7E4CFFFCE4339E" box="[733,805,947,974]" captionStart="FIGURES 2427" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 2427. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="2" pageNumber="411">24, 25</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FC934CFFFC8C339D" box="[816,845,947,973]" captionStart="FIGURES 2832" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 2832. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="2" pageNumber="411">29</figureCitation>
) and
<taxonomicName id="B5FD4D35FFD5A627FC284CF8FC23339E" ID-CoL="929XZ" authorityName="Alpheraky" authorityYear="1892" box="[907,994,948,974]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FC284CF8FC23339E" box="[907,994,948,974]" italics="true" pageId="2" pageNumber="411">Grumia</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD5A627FC534CFFFBB0339E" box="[1008,1137,947,974]" captionStart="FIGURES 1319" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 1319. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="2" pageNumber="411">Figs 1517</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FBDF4CFFFB5B339D" box="[1148,1178,947,973]" captionStart="FIGURES 2427" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 2427. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="2" pageNumber="411">27</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FB054CFFFB02339D" box="[1190,1219,947,973]" captionStart="FIGURES 2832" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 2832. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="2" pageNumber="411">32</figureCitation>
) display no significant morphological differences from the typical
<taxonomicName id="B5FD4D35FFD5A627FD134C9BFCDA33A1" ID-CoL="92CJF" authorityName="Staudinger" authorityYear="1882" box="[688,795,983,1009]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FD134C9BFCDA33A1" box="[688,795,983,1009]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD5A627FC894C9BFC4C33A1" box="[810,909,983,1009]" captionStart="FIGURES 18" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[163,1423,182,1902]" captionTargetId="figure-18@3.[158,1430,179,1918]" captionTargetPageId="3" captionText="FIGURES 18. Isochlora spp.: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 46 and 8 in ZISP; 7 in CAV." figureDoi="http://doi.org/10.5281/zenodo.10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="2" pageNumber="411">Figs 1, 2</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FC3B4C9BFC2133A1" box="[920,992,983,1009]" captionStart="FIGURES 2023" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 2023. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="2" pageNumber="411">20, 21</figureCitation>
,
<figureCitation id="EAC62A33FFD5A627FC484C9BFBC933A1" box="[1003,1032,983,1009]" captionStart="FIGURES 2832" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 2832. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="2" pageNumber="411">30</figureCitation>
) and therefore cannot be considered as subgenera of the latter. For this reason, we follow the wide concept of the genus suggested by
<bibRefCitation id="166C4B47FFD5A627FB724CB7FA5D3445" author="Lafontaine, J. D." box="[1233,1436,1019,1045]" pageId="2" pageNumber="411" pagination="1 - 348" refId="ref8218" refString="Lafontaine, J. D. (1998) Noctuoidea: Noctuidae (Part), Noctuinae, part. In: Dominik, R. B. et al. (Eds.), The Moths of America North of Mexico. Fasc. 27.3. The Wedge Entomological Research Foundation, Washington, pp. 1 - 348." type="book chapter" year="1998">Lafontaine (1998)</bibRefCitation>
and consider
<taxonomicName id="B5FD4D35FFD5A627FE8D4B6CFE44346A" ID-CoL="929XZ" authorityName="Alpheraky" authorityYear="1892" box="[302,389,1056,1082]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FE8D4B6CFE44346A" box="[302,389,1056,1082]" italics="true" pageId="2" pageNumber="411">Grumia</emphasis>
</taxonomicName>
and
<taxonomicName id="B5FD4D35FFD5A627FE1D4B53FDE23469" ID-CoL="92548" authorityName="Staudinger" authorityYear="1900" box="[446,547,1055,1081]" class="Insecta" family="Noctuidae" genus="Chamyla" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FE1D4B53FDE23469" box="[446,547,1055,1081]" italics="true" pageId="2" pageNumber="411">Chamyla</emphasis>
</taxonomicName>
as junior synonyms of
<taxonomicName id="B5FD4D35FFD5A627FC8D4B53FC583469" ID-CoL="92CJF" authorityName="Staudinger" authorityYear="1882" box="[814,921,1055,1081]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="4089EAA4FFD5A627FC8D4B53FC583469" box="[814,921,1055,1081]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
</taxonomicName>
.
<emphasis id="4089EAA4FFD5A627FC0B4B53FC083469" bold="true" box="[936,969,1055,1081]" pageId="2" pageNumber="411">(2)</emphasis>
The genus was characterised in detail by
<bibRefCitation id="166C4B47FFD5A627FF344B0FFEA7340D" author="Lafontaine, J. D." box="[151,358,1091,1117]" pageId="2" pageNumber="411" pagination="1 - 348" refId="ref8218" refString="Lafontaine, J. D. (1998) Noctuoidea: Noctuidae (Part), Noctuinae, part. In: Dominik, R. B. et al. (Eds.), The Moths of America North of Mexico. Fasc. 27.3. The Wedge Entomological Research Foundation, Washington, pp. 1 - 348." type="book chapter" year="1998">Lafontaine (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>