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<document id="CE16EE0AD72F368480A43D75196357EB" ID-DOI="10.11646/zootaxa.5374.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10152641" ID-ZooBank="75A1CC8A-D4EB-4457-86D0-01A89891C535" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700265469172" checkinUser="plazi" docAuthor="Volynkin, Anton V., Titov, Sergey V., Matov, Alexey Yu., Tóth, Balázs, Saldaitis, Aidas, Rakhimov, Ruslan D. &amp; Egorov, Petr V." docDate="2023" docId="FA5487A0FFD1A62DFF344CE2FAAF330D" docLanguage="en" docName="zootaxa.5374.3.5.pdf" docOrigin="Zootaxa 5374 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5374.3.5/52296" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isochlora kozlovi Volynkin, Titov, Matov &amp; Saldaitis 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="417" masterDocId="066DFFD8FFD7A625FFA34F4CFFC13050" masterDocTitle="On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae)" masterLastPageNumber="423" masterPageNumber="409" pageNumber="415" updateTime="1700265878127" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="05C2BDC5F0DEBF418B0FDD291E17AF07">On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae)</mods:title>
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<mods:affiliation id="BDC1B4C94E5FCE8D72D444DA641028F9">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. &amp; Department of Biology and Ecology; the Research Centre for Environmental “ Monitoring ”, Toraighyrov University, Lomova str. 64, KZ- 140008, Pavlodar, Kazakhstan. &amp; Scientific Center “ Bioarchaeology ”, Institute of Archaeological Research, Margulan University, Olzhabay Batyr str. 60, KZ- 140002, Pavlodar, Kazakhstan. &amp; sandibicumar @ gmail. com; https: // orcid. org / 0000 - 0003 - 2196 - 280 X</mods:affiliation>
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<mods:namePart id="7F75BC741AF83BEB51BF384A80E60D9A">Matov, Alexey Yu.</mods:namePart>
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<mods:affiliation id="1FFC516D65F881C3E8EB09D473C844C4">Zoological Institute, Russian Academy of Sciences, Universitetskaya Embankment 1, St. - Petersburg, RF- 199034 Russia. alexey. matov @ zin. ru; https: // orcid. org / 0000 - 0002 - 6066 - 6440</mods:affiliation>
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<mods:affiliation id="6858DEB9D6E79E1511E888224B0A9E60">Department of Zoology, Hungarian Natural History Museum, Baross utca, 13, H- 1088 Budapest, Hungary. toth. balazs @ nhmus. hu; https: // orcid. org / 0000 - 0002 - 6089 - 1218</mods:affiliation>
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<mods:affiliation id="ED8994B96A6957C0D43DAFEAF60DD59F">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. &amp; frontosa 88 @ mail. ru; https: // orcid. org / 0000 - 0001 - 5650 - 7239</mods:affiliation>
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<mods:affiliation id="71819975F2486D6B70BE59518A811881">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. &amp; ent. helios _ 2908 @ mail. ru; https: // orcid. org / 0000 - 0001 - 7686 - 6845</mods:affiliation>
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<mods:date id="D405966F619DB417B9D80A030645C0EC">2023</mods:date>
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<taxonomicName id="B5FD4D35FFD1A623FF344CE2FCE33398" authority="Volynkin, Titov, Matov &amp; Saldaitis" authorityName="Volynkin, Titov, Matov &amp; Saldaitis" authorityYear="2023" box="[151,802,942,969]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. nov.">
<emphasis id="4089EAA4FFD1A623FF344CE2FEA63399" bold="true" box="[151,359,942,969]" italics="true" pageId="6" pageNumber="415">Isochlora kozlovi</emphasis>
Volynkin, Titov, Matov &amp; Saldaitis
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,
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(
<figureCitation id="EAC62A33FFD1A623FF3C4C9FFE8B33BD" box="[159,330,979,1005]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Figs 13, 14, 18</figureCitation>
,
<figureCitation id="EAC62A33FFD1A623FEF54C9FFEB533BD" box="[342,372,979,1005]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 24–27. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="6" pageNumber="415">26</figureCitation>
,
<figureCitation id="EAC62A33FFD1A623FE234C9FFE5C33BD" box="[384,413,979,1005]" captionStart="FIGURES 28–32" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 28–32. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="6" pageNumber="415">31</figureCitation>
)
</paragraph>
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<emphasis id="4089EAA4FFD1A623FF344B57FE803464" bold="true" box="[151,321,1050,1077]" pageId="6" pageNumber="415">Type material</emphasis>
.
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<typeStatus id="AD468814FFD1A623FEF14B57FE7E3465" box="[338,447,1051,1077]" pageId="6" pageNumber="415" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="EAC62A33FFD1A623FE734B57FDEB3465" box="[464,554,1051,1077]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Figs 13</figureCitation>
,
<figureCitation id="EAC62A33FFD1A623FD9A4B57FD983465" box="[569,601,1051,1077]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 24–27. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="6" pageNumber="415">26</figureCitation>
): male, “
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<collectingDate id="1607E99EFFD1A623FD614B57FC953465" box="[706,852,1051,1077]" pageId="6" pageNumber="415" value="1900-07-10">10-VII-1900</collectingDate>
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” / “
<collectingCountry id="0AEA7626FFD1A623FC274B57FC0D3465" box="[900,972,1051,1077]" name="China" pageId="6" pageNumber="415">China</collectingCountry>
, E
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, | Bayan Har Range, |
<locationDeviation id="63F02F94FFD1A623FA2E4B57FE523409" location="Chzhabu-vrun Pass" pageId="6" pageNumber="415">S of Chzhabu-vrun Pass</locationDeviation>
, | upper reaches of
<collectingMunicipality id="9226ACCCFFD1A623FDD14B73FCCC3409" box="[626,781,1087,1113]" pageId="6" pageNumber="415">By-chu River</collectingMunicipality>
, |
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<collectingDate id="1607E99EFFD1A623FC844B73FC693409" box="[807,936,1087,1113]" pageId="6" pageNumber="415" value="1900-07-10">10.vii.1900</collectingDate>
</date>
, |
<collectorName id="DF085360FFD1A623FC674B73FB8D3409" box="[964,1100,1087,1113]" pageId="6" pageNumber="415">P.K. Kozlov</collectorName>
leg.” / ‘Slide | AV7370³ |
<collectorName id="DF085360FFD1A623FA224B73FEC1342D" pageId="6" pageNumber="415">A. Volynkin</collectorName>
” (
<collectionCode id="14ECAE73FFD1A623FEBA4B2FFE99342D" box="[281,344,1123,1149]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="415">ZISP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="724236B6FFD1A623FF644BCBFB0D355D" blockId="6.[151,1437,1050,2013]" pageId="6" pageNumber="415">
<materialsCitation id="C2953CEBFFD1A623FF644BCBFE3934B9" collectionCode="ZISP" collectorName="E Kunlun Mts" country="China" location="Kunlun Mts" pageId="6" pageNumber="415" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai" typeStatus="paratype">
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<typeStatus id="AD468814FFD1A623FF644BCBFEFE34F1" box="[199,319,1159,1185]" pageId="6" pageNumber="415" type="paratype">Paratypes</typeStatus>
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.
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<collectingCountry id="0AEA7626FFD1A623FEEE4BCAFE6B34F0" box="[333,426,1158,1184]" name="China" pageId="6" pageNumber="415">CHINA</collectingCountry>
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:
<specimenCount id="64FBFD3FFFD1A623FE1B4BCBFDDD34F1" box="[440,540,1159,1185]" count="1" pageId="6" pageNumber="415" type="female">1 female</specimenCount>
, [
<collectingRegion id="B039F854FFD1A623FD8E4BCBFD3634F1" box="[557,759,1159,1185]" country="China" name="Qinghai" pageId="6" pageNumber="415">Qinghai Province</collectingRegion>
,
<collectorName id="DF085360FFD1A623FCA14BCBFC6334F1" box="[770,930,1159,1185]" pageId="6" pageNumber="415">
E
<location id="7722606DFFD1A623FCB94BCBFC6334F1" LSID="urn:lsid:plazi:treatment:FA5487A0FFD1A62DFF344CE2FAAF330D:7722606DFFD1A623FCB94BCBFC6334F1" box="[794,930,1159,1185]" country="China" name="Kunlun Mts" pageId="6" pageNumber="415" stateProvince="Qinghai">Kunlun Mts</location>
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, Bayan Har Range,
<locationDeviation id="63F02F94FFD1A623FB224BCBFA5734F1" box="[1153,1430,1159,1185]" location="Chzhabu-vrun Pass" pageId="6" pageNumber="415">N of Chzhabu-vrun Pass</locationDeviation>
] vall. of Dzhagyn-gol [River], E
<collectingRegion id="B039F854FFD1A623FE5C4BE7FDFD3495" box="[511,572,1195,1221]" country="China" name="Xizang" pageId="6" pageNumber="415">Tibet</collectingRegion>
, 14300
<emphasis id="4089EAA4FFD1A623FD2D4BE7FD563495" box="[654,663,1195,1221]" italics="true" pageId="6" pageNumber="415">’</emphasis>
, Kozlov [leg.]. 1–6 VII [19]00 (in Russian), gen. prep. No.: AV7372 (prepared by Volynkin) (
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)
</materialsCitation>
;
<materialsCitation id="C2953CEBFFD1A623FDA54B83FB09355D" collectingDate="1999-08-10" collectingDateMax="1999-08-11" collectingDateMin="1999-08-10" collectionCode="PGM" collectorName="P. Gyulai &amp; A. Garai &amp; No." country="China" elevation="3800" location="Anemaqing Mts" pageId="6" pageNumber="415" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai" typeStatus="paratype">
<specimenCount id="64FBFD3FFFD1A623FDA54B83FDAD34B9" box="[518,620,1231,1257]" count="1" pageId="6" pageNumber="415" type="female">1 female</specimenCount>
, Prov.
<collectingRegion id="B039F854FFD1A623FD1E4B83FCDB34B9" box="[701,794,1231,1257]" country="China" name="Qinghai" pageId="6" pageNumber="415">Qinghai</collectingRegion>
,
<quantity id="B5059B53FFD1A623FC844B83FCB934B9" box="[807,888,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="415" unit="m" value="3800.0">
<elevation id="F9D0D185FFD1A623FC844B83FCB934B9" box="[807,888,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="415" unit="m" value="3800.0">3800m</elevation>
</quantity>
,
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,
<locationDeviation id="63F02F94FFD1A623FBE84B83FA9B34B9" box="[1099,1370,1231,1257]" location="Xue Shan" pageId="6" pageNumber="415">
<quantity id="B5059B53FFD1A623FBE84B83FB5534B9" box="[1099,1172,1231,1257]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="415" unit="km" value="10.0">10 km</quantity>
SE of Xue Shan
</locationDeviation>
,
<locationDeviation id="63F02F94FFD1A623FAC44B83FE8F355D" location="Dong Qing Gou" pageId="6" pageNumber="415">N of Dong Qing Gou</locationDeviation>
,
<date id="06431076FFD1A623FEF94BBFFDD9355D" box="[346,536,1267,1293]" pageId="6" pageNumber="415" value="1999-08-10" valueMax="1999-08-11" valueMin="1999-08-10">
<collectingDate id="1607E99EFFD1A623FEF94BBFFDD9355D" box="[346,536,1267,1293]" pageId="6" pageNumber="415" value="1999-08-10" valueMax="1999-08-11" valueMin="1999-08-10">10–11.VIII.1999</collectingDate>
</date>
,
<collectorName id="DF085360FFD1A623FD804BBFFD48355D" box="[547,649,1267,1293]" pageId="6" pageNumber="415">P. Gyulai</collectorName>
&amp;
<collectorName id="DF085360FFD1A623FD0F4BBFFCCD355D" box="[684,780,1267,1293]" pageId="6" pageNumber="415">A. Garai</collectorName>
leg., gen. prep.
<collectorName id="DF085360FFD1A623FC634BBFFC2F355D" box="[960,1006,1267,1293]" pageId="6" pageNumber="415">No.</collectorName>
: GYP 1866 (
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)
</materialsCitation>
.
</paragraph>
<paragraph id="724236B6FFD1A623FF644A5BFD6E371D" blockId="6.[151,1437,1050,2013]" pageId="6" pageNumber="415">
<emphasis id="4089EAA4FFD1A623FF644A5BFE243560" bold="true" box="[199,485,1302,1329]" pageId="6" pageNumber="415">Note on the type locality</emphasis>
. The male specimen of the new species deposited in
<collectionCode id="14ECAE73FFD1A623FB934A5BFBA83561" box="[1072,1129,1303,1329]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="415">ZISP</collectionCode>
and designated as the
<typeStatus id="AD468814FFD1A623FAC24A5BFF063505" pageId="6" pageNumber="415" type="holotype">holotype</typeStatus>
originally had a handwritten label containing only the date “
<date id="06431076FFD1A623FCC54A77FC3B3505" box="[870,1018,1339,1365]" pageId="6" pageNumber="415" value="1900-07-10">10-VII-1900</date>
”, which is four days later the range of the dates on the label of the
<typeStatus id="AD468814FFD1A623FE774A13FDF53529" box="[468,564,1375,1401]" pageId="6" pageNumber="415" type="paratype">paratype</typeStatus>
female (1–6 VII [19]00”) collected by P.K. Kozlov in the valley of Dzhagyn-Gol River, Tibet. Unfortunately, in his book devoted to his three year journey to
<collectingCountry id="0AEA7626FFD1A623FBBB4ACFFB4435CD" box="[1048,1157,1411,1437]" name="Mongolia" pageId="6" pageNumber="415">Mongolia</collectingCountry>
and Tibet in 1899–1901, P.K.
<bibRefCitation id="166C4B47FFD1A623FF6D4AEBFEB53591" author="Kozlov, P. K." box="[206,372,1447,1473]" pageId="6" pageNumber="415" refId="ref8180" refString="Kozlov, P. K. (1913) Three-year long journey (1899 - 1901) through Mongolia and Tibet. Imperial Russian Geographical Society, St. Petersburg, 233 pp. [in Russian]" type="book" year="1913">Kozlov (1913)</bibRefCitation>
has not mentioned the exact dates describing the part of the journey between July 1–6, when they were traveling up the river Dzhagyn-Gol, and July 16, when the expedition set up camp in Khoshun. The only locality mentioned was Chzhabu-vrun Pass, after crossing which the expedition began its descent into the basin of the river Yangtze (called ‘Blue River’ by P.K. Kozlov). In P.K. Kozlov’s diaries housed in the Scientific Archive of the Russian Geographical Society (
<collectingRegion id="B039F854FFD1A623FD85497BFD063601" box="[550,711,1591,1617]" country="Russia" name="Saint Petersburg" pageId="6" pageNumber="415">St. Petersburg</collectingRegion>
,
<collectingCountry id="0AEA7626FFD1A623FD70497BFCE33601" box="[723,802,1591,1617]" name="Russia" pageId="6" pageNumber="415">Russia</collectingCountry>
), the range of the dates ‘July 10–12’ was also described generally and without exact localities being mentioned, the author only wrote that the expedition was descending into the Blue River basin. However, in the
<collectionCode id="14ECAE73FFD1A623FD214933FD7A36C9" box="[642,699,1663,1689]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="6" pageNumber="415">ZISP</collectionCode>
collection there are other specimens of
<taxonomicName id="B5FD4D35FFD1A623FB204933FACB36C9" authorityName="Linnaeus" authorityYear="1758" box="[1155,1290,1663,1689]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
collected by P.K. Kozlov on July 11, for example, the
<typeStatus id="AD468814FFD1A623FDD649EFFD1636ED" box="[629,727,1699,1725]" pageId="6" pageNumber="415" type="holotype">holotype</typeStatus>
of the tiger moth
<taxonomicName id="B5FD4D35FFD1A623FC0949EFFA9C36ED" authority="Dubatolov, 2008" authorityName="Dubatolov" authorityYear="2008" box="[938,1373,1699,1725]" class="Insecta" family="Erebidae" genus="Palerontobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="4089EAA4FFD1A623FC0949EFFB5436ED" box="[938,1173,1699,1725]" italics="true" pageId="6" pageNumber="415">Palerontobia kozlovi</emphasis>
<bibRefCitation id="166C4B47FFD1A623FB3D49EFFA9C36ED" author="Dubatolov, V. V." box="[1182,1373,1699,1725]" pageId="6" pageNumber="415" pagination="351 - 355" refId="ref7604" refString="Dubatolov, V. V. (2008) Palerontobia kozlovi - a new species and genus of high mountain tiger-moth from Qinghai (Lepidoptera, Arctiidae). Atalanta, 39 (1 / 4), 351 - 355." type="journal article" year="2008">Dubatolov, 2008</bibRefCitation>
</taxonomicName>
, with the locality data “riv. By chu (Namtso), basin of Blue [River], 14000
<emphasis id="4089EAA4FFD1A623FC05498BFC6D36B1" box="[934,940,1735,1761]" italics="true" pageId="6" pageNumber="415">’</emphasis>
” (
<bibRefCitation id="166C4B47FFD1A623FC73498BFB4C36B1" author="Dubatolov, V. V." box="[976,1165,1735,1761]" pageId="6" pageNumber="415" pagination="351 - 355" refId="ref7604" refString="Dubatolov, V. V. (2008) Palerontobia kozlovi - a new species and genus of high mountain tiger-moth from Qinghai (Lepidoptera, Arctiidae). Atalanta, 39 (1 / 4), 351 - 355." type="journal article" year="2008">Dubatolov 2008</bibRefCitation>
). Thus, summarising all of the above, it is possible to conclude that the
<typeStatus id="AD468814FFD1A623FD0649A7FCC63755" box="[677,775,1771,1797]" pageId="6" pageNumber="415" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="B5FD4D35FFD1A623FCC949A7FC113755" authority="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov, 2023" authorityName="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov" authorityYear="2023" box="[874,976,1771,1797]" class="Insecta" family="Erebidae" genus="Palerontobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. n.">
<emphasis id="4089EAA4FFD1A623FCC949A7FC113755" box="[874,976,1771,1797]" italics="true" pageId="6" pageNumber="415">I. kozlovi</emphasis>
</taxonomicName>
<emphasis id="4089EAA4FFD1A623FC7449A7FBD63755" bold="true" box="[983,1047,1771,1797]" pageId="6" pageNumber="415">
<taxonomicNameLabel id="5BBA57DFFFD1A623FC7449A7FBD63755" box="[983,1047,1771,1797]" pageId="6" pageNumber="415" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
was collected in the upper reaches of By-chu River south of the Chzhabu-vrun Pass (
<figureCitation id="EAC62A33FFD1A623FD644843FCDA3779" box="[711,795,1807,1833]" captionStart="FIGURE 34" captionStartId="12.[151,250,1681,1705]" captionTargetBox="[151,1436,1015,1658]" captionTargetId="figure-42@12.[151,1436,1015,1658]" captionTargetPageId="12" captionText="FIGURE 34. Distribution map of Isochlora spp. in Qinghai, China." figureDoi="http://doi.org/10.5281/zenodo.10152659" httpUri="https://zenodo.org/record/10152659/files/figure.png" pageId="6" pageNumber="415">Fig. 34</figureCitation>
). A printed label with the relevant data was pinned by the authors of the present paper under the
<typeStatus id="AD468814FFD1A623FDE6487FFD6A371D" box="[581,683,1843,1869]" pageId="6" pageNumber="415" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="3AE7653DFFD1A622FF64481AFB0C3111" lastPageId="7" lastPageNumber="416" pageId="6" pageNumber="415" type="diagnosis">
<paragraph id="724236B6FFD1A622FF64481AFB0C3111" blockId="6.[151,1437,1050,2013]" lastBlockId="7.[151,1437,151,321]" lastPageId="7" lastPageNumber="416" pageId="6" pageNumber="415">
<emphasis id="4089EAA4FFD1A623FF64481AFEFB3720" bold="true" box="[199,314,1878,1904]" pageId="6" pageNumber="415">Diagnosis</emphasis>
.
<taxonomicName id="B5FD4D35FFD1A623FEEB481BFDCB3721" authority="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov, 2023" authorityName="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov" authorityYear="2023" box="[328,522,1879,1905]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. nov.">
<emphasis id="4089EAA4FFD1A623FEEB481BFDCB3721" box="[328,522,1879,1905]" italics="true" pageId="6" pageNumber="415">Isochlora kozlovi</emphasis>
</taxonomicName>
<emphasis id="4089EAA4FFD1A623FDB2481BFD913721" bold="true" box="[529,592,1879,1905]" pageId="6" pageNumber="415">
<taxonomicNameLabel id="5BBA57DFFFD1A623FDB2481BFD913721" box="[529,592,1879,1905]" pageId="6" pageNumber="415" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="EAC62A33FFD1A623FDFC481BFD1F3721" box="[607,734,1879,1905]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Figs 13, 14</figureCitation>
) can be distinguished from the similar
<taxonomicName id="B5FD4D35FFD1A623FB36481BFB233721" authorityName="Alpheraky" authorityYear="1892" box="[1173,1250,1879,1905]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD1A623FB36481BFB233721" box="[1173,1250,1879,1905]" italics="true" pageId="6" pageNumber="415">I. flora</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD1A623FB52481BFAB13721" box="[1265,1392,1878,1905]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Figs 15–17</figureCitation>
) by its smaller size and the forewing upperside having a well-developed irregular reddish discal spot. The underside of both wings of the new species (
<figureCitation id="EAC62A33FFD1A623FE5448D3FD8937E9" box="[503,584,1951,1977]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Fig. 18</figureCitation>
) has more intense pink suffusion than in
<taxonomicName id="B5FD4D35FFD1A623FBB648D3FBA237E9" authorityName="Alpheraky" authorityYear="1892" box="[1045,1123,1951,1977]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD1A623FBB648D3FBA237E9" box="[1045,1123,1951,1977]" italics="true" pageId="6" pageNumber="415">I. flora</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD1A623FBD148D3FB0437E9" box="[1138,1221,1951,1977]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="6" pageNumber="415">Fig. 19</figureCitation>
), and the hindwing lacks the broad whitish area stretching along the costal margin and occupying nearly a half of the hindwing in the congener. The male genital capsules of the two species display no remarkable differences. Compared to
<taxonomicName id="B5FD4D35FFD0A622FAB74FDBFAA130E1" authorityName="Alpheraky" authorityYear="1892" box="[1300,1376,151,177]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD0A622FAB74FDBFAA130E1" box="[1300,1376,151,177]" italics="true" pageId="7" pageNumber="416">I. flora</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD0A622FACC4FDBFF773085" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 24–27. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="7" pageNumber="416">Fig. 27</figureCitation>
), the phallus of
<taxonomicName id="B5FD4D35FFD0A622FECD4FF7FE173085" authority="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov, 2023" authorityName="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov" authorityYear="2023" box="[366,470,187,213]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. n.">
<emphasis id="4089EAA4FFD0A622FECD4FF7FE173085" box="[366,470,187,213]" italics="true" pageId="7" pageNumber="416">I. kozlovi</emphasis>
</taxonomicName>
<emphasis id="4089EAA4FFD0A622FE7D4FF7FDDE3085" bold="true" box="[478,543,187,213]" pageId="7" pageNumber="416">
<taxonomicNameLabel id="5BBA57DFFFD0A622FE7D4FF7FDDE3085" box="[478,543,187,213]" pageId="7" pageNumber="416" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="EAC62A33FFD0A622FD8C4FF7FD403085" box="[559,641,187,213]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 24–27. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="7" pageNumber="416">Fig. 26</figureCitation>
) is shorter and has a shorter coecum. The vesica of
<taxonomicName id="B5FD4D35FFD0A622FB724FF7FAF83085" authority="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov, 2023" authorityName="Volynkin &amp; Titov &amp; Matov &amp; Tóth &amp; Saldaitis &amp; Rakhimov &amp; Egorov" authorityYear="2023" box="[1233,1337,187,213]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. n.">
<emphasis id="4089EAA4FFD0A622FB724FF7FAF83085" box="[1233,1337,187,213]" italics="true" pageId="7" pageNumber="416">I. kozlovi</emphasis>
</taxonomicName>
<emphasis id="4089EAA4FFD0A622FAE24FF7FA433085" bold="true" box="[1345,1410,187,213]" pageId="7" pageNumber="416">
<taxonomicNameLabel id="5BBA57DFFFD0A622FAE24FF7FA433085" box="[1345,1410,187,213]" pageId="7" pageNumber="416" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is considerably shorter and narrower than in
<taxonomicName id="B5FD4D35FFD0A622FDD94F93FD0830A9" authorityName="Alpheraky" authorityYear="1892" box="[634,713,223,249]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD0A622FDD94F93FD0830A9" box="[634,713,223,249]" italics="true" pageId="7" pageNumber="416">I. flora</emphasis>
</taxonomicName>
, and has a shorter medial diverticulum. In the female genitalia, the new species (
<figureCitation id="EAC62A33FFD0A622FEF44E4FFE68314D" box="[343,425,259,285]" captionStart="FIGURES 28–32" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 28–32. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="7" pageNumber="416">Fig. 31</figureCitation>
) differs from
<taxonomicName id="B5FD4D35FFD0A622FDE44E4FFD54314D" authorityName="Alpheraky" authorityYear="1892" box="[583,661,259,285]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD0A622FDE44E4FFD54314D" box="[583,661,259,285]" italics="true" pageId="7" pageNumber="416">I. flora</emphasis>
</taxonomicName>
(
<figureCitation id="EAC62A33FFD0A622FD064E4FFD36314D" box="[677,759,259,285]" captionStart="FIGURES 28–32" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 28–32. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="7" pageNumber="416">Fig. 32</figureCitation>
) in the longer sclerotised area of the posterior section of the corpus bursae, and the anterior section bearing three larger signa whereas
<taxonomicName id="B5FD4D35FFD0A622FC704E6BFBE03111" authorityName="Alpheraky" authorityYear="1892" box="[979,1057,295,321]" class="Insecta" family="Noctuidae" genus="Grumia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="flora">
<emphasis id="4089EAA4FFD0A622FC704E6BFBE03111" box="[979,1057,295,321]" italics="true" pageId="7" pageNumber="416">I. flora</emphasis>
</taxonomicName>
has four signa.
</paragraph>
</subSubSection>
<caption id="2682663EFFD0A622FF3449B6FC7F3743" ID-DOI="http://doi.org/10.5281/zenodo.10152647" ID-Zenodo-Dep="10152647" box="[151,958,1786,1811]" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="7" pageNumber="416" startId="7.[151,264,1786,1810]" targetBox="[189,1398,348,1750]" targetPageId="7" targetType="figure">
<paragraph id="724236B6FFD0A622FF3449B6FC7F3743" blockId="7.[151,958,1786,1811]" box="[151,958,1786,1811]" pageId="7" pageNumber="416">
<emphasis id="4089EAA4FFD0A622FF3449B6FE8F3743" bold="true" box="[151,334,1786,1811]" pageId="7" pageNumber="416">FIGURES 13–19</emphasis>
.
<taxonomicName id="B5FD4D35FFD0A622FEF949B7FE2E3742" box="[346,495,1787,1811]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4089EAA4FFD0A622FEF949B7FE7A3743" box="[346,443,1787,1811]" italics="true" pageId="7" pageNumber="416">Isochlora</emphasis>
spp.
</taxonomicName>
: adults. The specimens are deposited in ZISP.
</paragraph>
</caption>
<subSubSection id="3AE7653DFFD0A62DFF64480DFE0332A1" lastPageId="8" lastPageNumber="417" pageId="7" pageNumber="416" type="description">
<paragraph id="724236B6FFD0A62DFF64480DFE0332A1" blockId="7.[151,1437,1857,2027]" lastBlockId="8.[151,1437,151,861]" lastPageId="8" lastPageNumber="417" pageId="7" pageNumber="416">
<emphasis id="4089EAA4FFD0A622FF64480DFE90370B" bold="true" box="[199,337,1857,1883]" pageId="7" pageNumber="416">Description</emphasis>
.
<emphasis id="4089EAA4FFD0A622FEC3480DFE6E370B" bold="true" box="[352,431,1857,1883]" pageId="7" pageNumber="416">Adults</emphasis>
(
<figureCitation id="EAC62A33FFD0A622FE1C480DFDAD370B" box="[447,620,1857,1883]" captionStart="FIGURES 13–19" captionStartId="7.[151,264,1786,1810]" captionTargetBox="[189,1398,348,1750]" captionTargetId="figure-118@7.[176,1411,345,1763]" captionTargetPageId="7" captionText="FIGURES 13–19. Isochlora spp.: adults. The specimens are deposited in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152647" httpUri="https://zenodo.org/record/10152647/files/figure.png" pageId="7" pageNumber="416">Figs 13, 14, 18</figureCitation>
). Forewing length 9.0 mm in male
<typeStatus id="AD468814FFD0A622FC58480DFB9C370B" box="[1019,1117,1857,1883]" pageId="7" pageNumber="416" type="holotype">holotype</typeStatus>
and 9.5 mm in female
<typeStatus id="AD468814FFD0A622FAC0480DFF0A372F" pageId="7" pageNumber="416" type="paratype">paratype</typeStatus>
. Antenna brown, shortly bipectinate in male and filiform in female. Head and thorax grass green. Ground colour of forewing upperside, grass green with pink suffusion on costal margin. Cell with irregular reddish spot distally. Cilia green with admixture of pink. Legs pale ochreous with admixture of pink. Hindwing underside pale brown with intense pink suffusion along costal and outer margins, and elliptical cluster of whitish hair-like scales along anal margin proximally. Hindwing upperside greyish-brown with pink suffusion terminally, cilia pale brown with admixture of pink. Hindwing underside pale brown with pink suffusion becoming more intense distally, and diffuse with pale ochreous spot at costal margin postmedially. Abdomen grey with admixture of pale green, and with pink scales on male genitalia. In female ovipositor strongly elongate, telescopic.
<emphasis id="4089EAA4FFDFA62DFBD94E4FFAE6314C" bold="true" box="[1146,1319,258,285]" pageId="8" pageNumber="417">Male genitalia</emphasis>
(
<figureCitation id="EAC62A33FFDFA62DFA9A4E4FFA51314D" box="[1337,1424,259,285]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1929,1953]" captionTargetBox="[172,1414,194,1892]" captionTargetId="figure-23@10.[160,1427,181,1905]" captionTargetPageId="10" captionText="FIGURES 24–27. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 24, 26 and 27 in ZISP; 25 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152651" httpUri="https://zenodo.org/record/10152651/files/figure.png" pageId="8" pageNumber="417">Fig. 26</figureCitation>
). Uncus bar-like, weakly setose, almost straight, slightly dilated postmedially, distally tapered with triangular and downcurved tip. Tegumen with broadly triangular and apically rounded penicular lobe. Vinculum somewhat longer than tegumen, with V-shaped and apically rounded tip. Valva elongate, gradually tapering distally, with somewhat upcurved distal half bearing dense cluster of setae apically. Sacculus short (slightly shorter than 1/3 of valva length), broad proximally (ca. 2/3 of valva width) and distally tapered. Clasper thin, without harpe. Juxta bowl-shaped with deep medial depression. Anellus setose. Phallus cylindrical, slightly downcurved medially, with rounded coecum and granulose plate distally. Vesica recurved ventrally, medially dilated and distally tapered, weakly granulose medially and distally, and with short and conical medial diverticulum bearing thin spike-like cornutus apically.
<emphasis id="4089EAA4FFDFA62DFAE64D6FFF3F3230" bold="true" pageId="8" pageNumber="417">Female genitalia</emphasis>
(
<figureCitation id="EAC62A33FFDFA62DFEAE4D0BFEA13231" box="[269,352,583,609]" captionStart="FIGURES 28–32" captionStartId="11.[151,264,1942,1966]" captionTargetBox="[160,1434,186,1913]" captionTargetId="figure-18@11.[151,1435,181,1918]" captionTargetPageId="11" captionText="FIGURES 28–32. Isochlora spp.: female genitalia. Depositories of the specimens dissected: 28, 29, 31 and 32 in ZISP; 30 in MfN." figureDoi="http://doi.org/10.5281/zenodo.10152653" httpUri="https://zenodo.org/record/10152653/files/figure.png" pageId="8" pageNumber="417">Fig. 31</figureCitation>
). Ovipositor elongate and conical. Papilla analis elongate rectangular with rounded corners, with reduced setae. Apophyses strongly elongate, rod-like and distally flattened, apophysis posterioris somewhat longer than apophysis anterioris. Ductus bursae tubular and membranous. Corpus bursae medially constricted with narrower, rugose and weakly sclerotised posterior section and teardrop-shaped anterior section bearing three weakly sclerotised elliptical signa.
</paragraph>
</subSubSection>
<subSubSection id="3AE7653DFFDFA62DFF644DB6FC9B3345" box="[199,858,762,789]" pageId="8" pageNumber="417" type="distribution">
<paragraph id="724236B6FFDFA62DFF644DB6FC9B3345" blockId="8.[151,1437,151,861]" box="[199,858,762,789]" pageId="8" pageNumber="417">
<emphasis id="4089EAA4FFDFA62DFF644DB6FE983344" bold="true" box="[199,345,762,788]" pageId="8" pageNumber="417">Distribution</emphasis>
. Western
<collectingCountry id="0AEA7626FFDFA62DFE6A4DB7FDCD3345" box="[457,524,763,789]" name="China" pageId="8" pageNumber="417">China</collectingCountry>
(
<collectingRegion id="B039F854FFDFA62DFDB94DB7FD233345" box="[538,738,763,789]" country="China" name="Qinghai" pageId="8" pageNumber="417">Qinghai Province</collectingRegion>
) (
<figureCitation id="EAC62A33FFDFA62DFD594DB7FC8F3345" box="[762,846,763,789]" captionStart="FIGURE 34" captionStartId="12.[151,250,1681,1705]" captionTargetBox="[151,1436,1015,1658]" captionTargetId="figure-42@12.[151,1436,1015,1658]" captionTargetPageId="12" captionText="FIGURE 34. Distribution map of Isochlora spp. in Qinghai, China." figureDoi="http://doi.org/10.5281/zenodo.10152659" httpUri="https://zenodo.org/record/10152659/files/figure.png" pageId="8" pageNumber="417">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3AE7653DFFDFA62DFF644C53FAAF330D" pageId="8" pageNumber="417" type="etymology">
<paragraph id="724236B6FFDFA62DFF644C53FAAF330D" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="417">
<emphasis id="4089EAA4FFDFA62DFF644C53FE893369" bold="true" box="[199,328,799,825]" pageId="8" pageNumber="417">Etymology</emphasis>
. The new species is named after Pyotr Kuzmich Kozlov (1863–1935), a famous traveller and explorer of
<collectingCountry id="0AEA7626FFDFA62DFF5C4C0FFEAD330D" box="[255,364,835,861]" name="Mongolia" pageId="8" pageNumber="417">Mongolia</collectingCountry>
and Tibet, follower of Nikolai Przhevalsky, and collector of the
<typeStatus id="AD468814FFDFA62DFB9D4C0FFB61330D" box="[1086,1184,835,861]" pageId="8" pageNumber="417" type="holotype">holotype</typeStatus>
and
<specimenCount id="64FBFD3FFFDFA62DFB744C0FFAAB330D" box="[1239,1386,835,861]" count="1" pageId="8" pageNumber="417" type="generic" typeStatus="paratype">one paratype</specimenCount>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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