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<document id="B8F76F27EBD704E3BD8CD990715E22C9" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA249BBDC1FF37D8C91E13A63B" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma santanderensis Galeano &amp; Bernal 2002" docType="treatment" docVersion="7" lastPageNumber="135" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="134" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:title id="1B84D15E560D42E9157CBD983EAC70F2">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:name id="9B5036704ECECBB104B5FD4430D3E171" type="personal">
<mods:role id="441510F4E5A79C1A678DB3BBECA61402">
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<mods:namePart id="05FB56DE7DE9DC2F0EAC405F2EA94620">Henderson, Andrew</mods:namePart>
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<mods:title id="192BDF1B7E0C798E1EAF7A1FAC61D24A">Phytotaxa</mods:title>
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<mods:part id="84C2D89ED69BFB32A11454FC867C3AC3">
<mods:date id="6795444D7C414706D58D7777C18E5046">2011</mods:date>
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<mods:number id="A36E3CF80A494118B73EECA419BDCE51">2011-02-18</mods:number>
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<mods:identifier id="27BCE6DE9D78AFDDD5F3E9C83871525A" type="DOI">10.11646/phytotaxa.17.1</mods:identifier>
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<treatment id="FA4887FA249BBDC1FF37D8C91E13A63B" ID-DOI="http://doi.org/10.5281/zenodo.5609150" ID-GBIF-Taxon="160342372" ID-Zenodo-Dep="5609150" LSID="urn:lsid:plazi:treatment:FA4887FA249BBDC1FF37D8C91E13A63B" httpUri="http://treatment.plazi.org/id/FA4887FA249BBDC1FF37D8C91E13A63B" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134">
<subSubSection id="3AFB6567249BBDC2FF37D8C91F67A5F3" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="725E36EC249BBDC2FF37D8C91F67A5F3" blockId="133.[151,1436,1673,1778]" pageId="133" pageNumber="134">
<emphasis id="4095EAFE249BBDC2FF37D8C91B82A5A4" bold="true" box="[151,189,1675,1701]" pageId="133" pageNumber="134">54.</emphasis>
<taxonomicName id="B5E14D6F249BBDC2FF65D8C818BFA5A4" ID-CoL="3FQQ4" authority="Galeano &amp; Bernal (2002: 282)" authorityName="Galeano &amp; Bernal" authorityPageNumber="282" authorityYear="2002" box="[197,896,1673,1701]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="133" pageNumber="134" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="4095EAFE249BBDC2FF65D8C8193FA5A5" bold="true" box="[197,512,1673,1700]" italics="true" pageId="133" pageNumber="134">Geonoma santanderensis</emphasis>
<bibRefCitation id="16704B1D249BBDC2FDA8D8C918BFA5A4" author="Galeano, G. &amp; Bernal, R." box="[520,896,1675,1701]" journalOrPublisher="Caldasia" pageId="133" pageNumber="134" pagination="277 - 292" part="24" refId="ref122238" refString="Galeano, G. &amp; R. Bernal, R. (2002) New species and new records of Colombian palms. Caldasia 24: 277 - 292." title="New species and new records of Colombian palms" type="journal article" year="2002">Galeano &amp; Bernal (2002: 282)</bibRefCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C249BBDC2FC79D8C91F42A5A4" box="[985,1149,1675,1701]" name="Colombia" pageId="133" pageNumber="134">COLOMBIA</collectingCountry>
.
<collectingRegion id="B025F80E249BBDC2FB29D8C91E38A5A4" box="[1161,1287,1675,1701]" country="Colombia" name="Santander" pageId="133" pageNumber="134">Santander</collectingRegion>
: Suaita, San José de Suaita, ca.
<geoCoordinate id="17D5502B249BBDC2FEDBD8F31AE9A5CA" box="[379,470,1713,1739]" degrees="6" direction="north" minutes="10" orientation="latitude" pageId="133" pageNumber="134" precision="925" value="6.1666665">6°10N</geoCoordinate>
,
<geoCoordinate id="17D5502B249BBDC2FE43D8F3196EA5CA" box="[483,593,1713,1739]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="133" pageNumber="134" precision="925" value="-73.45">73°27W</geoCoordinate>
,
<quantity id="B5199B09249BBDC2FDFED8F31839A5CA" box="[606,774,1713,1739]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="133" pageNumber="134" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">17001900 m</quantity>
,
<date id="065F102C249BBDC2FCB3D8F3188EA5CA" box="[787,945,1713,1739]" pageId="133" pageNumber="134" value="2001-07-30">30 July 2001</date>
,
<emphasis id="4095EAFE249BBDC2FC1ED8F31AF7A5F3" italics="true" pageId="133" pageNumber="134">G. Galeano, J. Betancur, N. Castaño, L. Clavijo &amp; N. Garcia 6884</emphasis>
(
<typeStatus id="AD5A884E249BBDC2FE78D89A197FA5F3" box="[472,576,1752,1778]" pageId="133" pageNumber="134" type="holotype">holotype</typeStatus>
COL!, isotypes HUA
<emphasis id="4095EAFE249BBDC2FCE9D89B184EA5F3" box="[841,881,1753,1778]" italics="true" pageId="133" pageNumber="134">n.v.</emphasis>
, NY
<emphasis id="4095EAFE249BBDC2FC11D89B18E6A5F3" box="[945,985,1753,1778]" italics="true" pageId="133" pageNumber="134">n.v.</emphasis>
, UIS
<emphasis id="4095EAFE249BBDC2FBBED89B1F79A5F3" box="[1054,1094,1753,1778]" italics="true" pageId="133" pageNumber="134">n.v.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="3AFB6567249BBDC1FF37D96718C8A091" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134" type="description">
<paragraph id="725E36EC249BBDC1FF37D96718C8A091" blockId="133.[151,1437,1829,2010]" lastBlockId="134.[151,1438,152,1338]" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134">
<emphasis id="4095EAFE249BBDC2FF37D9671BDDA43E" box="[151,226,1829,1855]" italics="true" pageId="133" pageNumber="134">Plants</emphasis>
height no data; stems 1.3(1.01.5) m tall, 0.8(0.60.9) cm in diameter, solitary or clustered, cane-like; internodes 1.5(1.02.2) cm long, covered with reddish or brownish scales, especially in their distal part.
<emphasis id="4095EAFE249BBDC2FF37D9361BD6A48C" box="[151,233,1908,1933]" italics="true" pageId="133" pageNumber="134">Leaves</emphasis>
9(712) per stem, irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 8.0(6.011.0) cm long; petioles 15.9(8.029.0) cm long, drying green or yellowish; rachis 20.5(15.5 25.0) cm long, 1.6(1.42.2) mm in diameter; veins not raised or slightly raised and triangular in cross-section adaxially; pinnae 5(36) per side of rachis; basal pinna 10.7(9.711.5) cm long, 1.1(0.81.8) cm wide, forming an angle of 60(4572)° with the rachis; apical pinna 8.2(7.010.0) cm, 7.5(6.311.0) cm wide, forming an angle of 42(3848)° with the rachis.
<emphasis id="4095EAFE2498BDC1FD76DEA71840A3FE" box="[726,895,229,255]" italics="true" pageId="134" pageNumber="135">Inflorescences</emphasis>
unbranched; prophylls and peduncular bracts not ribbed with elongate, unbranched fibers, flattened, deciduous or persistent; prophylls 8.5(6.711.2) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 7.3(5.910.5) cm long, well-developed, inserted 1.1(0.52.3) cm above the prophyll; peduncles 8.7(6.014.5) cm long, 2.1(1.52.8) mm in diameter; rachillae 1, 8.4(5.710.0) cm long, 3.4(2.6 4.1) mm in diameter, the surfaces with spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits usually spirally arranged, glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted onto bifid and well-developed, non-jointed connectives; anthers short and curled over at anthesis; non-fertilized pistillate flowers deciduous after anthesis; staminodial tubes crenulate or shallowly lobed at the apex, those of non-fertilized pistillate flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE2498BDC1FD0ADD4019D5A01D" box="[682,746,770,796]" italics="true" pageId="134" pageNumber="135">fruits</emphasis>
8.5(7.89.9) mm long, 6.0(4.67.2) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis without operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672498BDC1FF66DDDF1E13A63B" pageId="134" pageNumber="135" type="distribution">
<paragraph id="725E36EC2498BDC1FF66DDDF184FA0DF" blockId="134.[151,1438,152,1338]" pageId="134" pageNumber="135">
<emphasis id="4095EAFE2498BDC1FF66DDDF1912A0B6" bold="true" box="[198,557,925,951]" pageId="134" pageNumber="135">Distribution and habitat:—</emphasis>
From 6°10
<geoCoordinate id="17D5502B2498BDC1FD69DDDF181EA0B6" box="[713,801,925,951]" degrees="7" direction="north" minutes="08" orientation="latitude" pageId="134" pageNumber="135" precision="925" value="7.133333">7°08N</geoCoordinate>
and 72°54
<geoCoordinate id="17D5502B2498BDC1FC68DDDF1F0CA0B6" box="[968,1075,925,951]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="134" pageNumber="135" precision="925" value="-73.45">73°27W</geoCoordinate>
in the Eastern Cordillera in
<collectingCountry id="0AF6767C2498BDC1FF37DD861A35A0DF" box="[151,266,964,990]" name="Colombia" pageId="134" pageNumber="135">Colombia</collectingCountry>
at
<quantity id="B5199B092498BDC1FE8EDD861AB8A0DF" box="[302,391,964,990]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="134" pageNumber="135" unit="m" value="1800.0">1800 m</quantity>
elevation in montane rainforest (
<figureCitation id="EADA2A692498BDC1FCABDD86185CA0DF" box="[779,867,964,990]" captionStart="FIGURE 36" captionStartId="132.[151,257,1960,1983]" captionTargetBox="[167,1417,925,1937]" captionTargetId="figure@132.[151,1436,905,1959]" captionTargetPageId="132" captionText="FIGURE 36. Distribution maps of Geonoma pohliana subsp. wittigiana, G. poiteauana, G. sanmartinensis, and G. santanderensis." figureDoi="http://doi.org/10.5281/zenodo.3538435" httpUri="https://zenodo.org/record/3538435/files/figure.png" pageId="134" pageNumber="135">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="725E36EC2498BDC1FF66DDA81FCAA7EC" blockId="134.[151,1438,152,1338]" pageId="134" pageNumber="135">
<emphasis id="4095EAFE2498BDC1FF66DDA81AFEA705" bold="true" box="[198,449,1002,1028]" pageId="134" pageNumber="135">Taxonomic notes:—</emphasis>
<taxonomicName id="B5E14D6F2498BDC1FE62DDA819D2A705" authorityName="Galeano &amp; Bernal" authorityYear="2002" box="[450,749,1002,1028]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="4095EAFE2498BDC1FE62DDA819D2A705" box="[450,749,1002,1028]" italics="true" pageId="134" pageNumber="135">Geonoma santanderensis</emphasis>
</taxonomicName>
was compared to
<taxonomicName id="B5E14D6F2498BDC1FC6BDDA81F42A705" authorityName="de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[971,1149,1002,1028]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="monospatha">
<emphasis id="4095EAFE2498BDC1FC6BDDA81F42A705" box="[971,1149,1002,1028]" italics="true" pageId="134" pageNumber="135">G. monospatha</emphasis>
</taxonomicName>
by
<bibRefCitation id="16704B1D2498BDC1FB0EDDA81BD4A72A" author="Galeano, G. &amp; Bernal, R." journalOrPublisher="Caldasia" pageId="134" pageNumber="135" pagination="277 - 292" part="24" refId="ref122238" refString="Galeano, G. &amp; R. Bernal, R. (2002) New species and new records of Colombian palms. Caldasia 24: 277 - 292." title="New species and new records of Colombian palms" type="journal article" year="2002">Galeano and Bernal (2002)</bibRefCitation>
. However, it appears closely related to a group of species within the
<taxonomicName id="B5E14D6F2498BDC1FB93DA531FA0A72A" authorityName="Kunth" authorityYear="1841" baseAuthorityName="Poiteau" box="[1075,1183,1041,1067]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="4095EAFE2498BDC1FB93DA531FA0A72A" box="[1075,1183,1041,1067]" italics="true" pageId="134" pageNumber="135">G. stricta</emphasis>
</taxonomicName>
clade comprising
<taxonomicName id="B5E14D6F2498BDC1FA23DA531A1FA753" authorityName="Spruce" authorityYear="1871" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="aspidiifolia">
<emphasis id="4095EAFE2498BDC1FA23DA531A1FA753" italics="true" pageId="134" pageNumber="135">G. aspidiifolia</emphasis>
</taxonomicName>
and
<taxonomicName id="B5E14D6F2498BDC1FEFCDA7A1AC1A753" authorityName="Trail" authorityYear="1876" box="[348,510,1080,1106]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="oligoclona">
<emphasis id="4095EAFE2498BDC1FEFCDA7A1AC1A753" box="[348,510,1080,1106]" italics="true" pageId="134" pageNumber="135">G. oligoclona</emphasis>
</taxonomicName>
. All three species share internodes covered with reddish or brownish scales, especially in their distal part, rachillae surfaces with spiky, fibrous projections or ridges, and staminodial tubes lobed at the apex with the lobes not spreading at anthesis and not acuminate.
<taxonomicName id="B5E14D6F2498BDC1FB80DAC71E76A79E" authorityName="Galeano &amp; Bernal" authorityYear="2002" box="[1056,1353,1157,1183]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="4095EAFE2498BDC1FB80DAC71E76A79E" box="[1056,1353,1157,1183]" italics="true" pageId="134" pageNumber="135">Geonoma santanderensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="B5E14D6F2498BDC1FF79DAEE1ABDA7C7" authorityName="Spruce" authorityYear="1871" box="[217,386,1196,1222]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="aspidiifolia">
<emphasis id="4095EAFE2498BDC1FF79DAEE1ABDA7C7" box="[217,386,1196,1222]" italics="true" pageId="134" pageNumber="135">G. aspidiifolia</emphasis>
</taxonomicName>
in its fruits which are bumpy from the numerous, subepidermal, tangential, short fibers present; and from
<taxonomicName id="B5E14D6F2498BDC1FECDDA911935A7EC" authorityName="Trail" authorityYear="1876" box="[365,522,1235,1261]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="134" pageNumber="135" phylum="Tracheophyta" rank="species" species="oligoclona">
<emphasis id="4095EAFE2498BDC1FECDDA911935A7EC" box="[365,522,1235,1261]" italics="true" pageId="134" pageNumber="135">G. oligoclona</emphasis>
</taxonomicName>
in its prophyll which is not short and asymmetrically apiculate.
</paragraph>
<paragraph id="725E36EC2498BDC1FF66DABB1E13A63B" blockId="134.[151,1438,152,1338]" pageId="134" pageNumber="135">
<emphasis id="4095EAFE2498BDC1FF66DABB1AC9A612" bold="true" box="[198,502,1273,1299]" pageId="134" pageNumber="135">Subspecific variation:—</emphasis>
No traits, except for stem branching, vary within this species. The specimens come from two separate areas, but there are too few specimens to test for differences between these.
</paragraph>
</subSubSection>
</treatment>
</document>