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<document id="6E47827F035698DFE5BA15C1C386FA97" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA245BBD01FF37DD121ABCA7EC" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma epetiolata Moore 1980" docType="treatment" docVersion="7" lastPageNumber="71" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="70" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:title id="EDB5E4DCB677D36661BBE8FCF8061227">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:role id="13807F853B606C98A33B3AEC5C766754">
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<mods:namePart id="D4932D75D750D27FDEBB41E80A5C552B">Henderson, Andrew</mods:namePart>
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<mods:title id="BB5CA5CD8F203FA5104B4E1DAD00A141">Phytotaxa</mods:title>
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<mods:part id="215C20E37CCABD2F4AE1AE95975BF380">
<mods:date id="E4B16AFCAE831647B35793A14B03F726">2011</mods:date>
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<mods:number id="FDF357B673AA905BE607F20CF85BC433">17</mods:number>
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<treatment id="FA4887FA245BBD01FF37DD121ABCA7EC" ID-DOI="http://doi.org/10.5281/zenodo.5608984" ID-GBIF-Taxon="160342452" ID-Zenodo-Dep="5608984" LSID="urn:lsid:plazi:treatment:FA4887FA245BBD01FF37DD121ABCA7EC" httpUri="http://treatment.plazi.org/id/FA4887FA245BBD01FF37DD121ABCA7EC" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<subSubSection id="3AFB6567245BBD02FF37DD121AE9A0B6" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="725E36EC245BBD02FF37DD121AE9A0B6" blockId="69.[151,1437,846,951]" pageId="69" pageNumber="70">
<emphasis id="4095EAFE245BBD02FF37DD121B80A06B" bold="true" box="[151,191,848,874]" pageId="69" pageNumber="70">20.</emphasis>
<taxonomicName id="B5E14D6F245BBD02FF68DD0D19FEA06B" ID-CoL="3FQJQ" authority="Moore (1980: 28)" authorityName="Moore" authorityPageNumber="28" authorityYear="1980" box="[200,705,846,874]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE245BBD02FF68DD0D1AF0A068" bold="true" box="[200,463,846,873]" italics="true" pageId="69" pageNumber="70">Geonoma epetiolata</emphasis>
<bibRefCitation id="16704B1D245BBD02FE78DD1219FEA06B" author="Moore, H." box="[472,705,848,874]" journalOrPublisher="Gentes Herbarum" pageId="69" pageNumber="70" pagination="25 - 29" part="12" refId="ref123610" refString="Moore, H. (1980) Two new species of Geonoma (Palmae). Gentes Herbarum 12: 25 - 29." title="Two new species of Geonoma (Palmae)" type="journal article" year="1980">Moore (1980: 28)</bibRefCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C245BBD02FC80DD121897A06B" box="[800,936,848,874]" name="Panama" pageId="69" pageNumber="70">PANAMA</collectingCountry>
.
<collectingRegion id="B025F80E245BBD02FC15DD121F10A06B" box="[949,1071,848,874]" country="Panama" name="Veraguas" pageId="69" pageNumber="70">Veraguas</collectingRegion>
: Guabal (Dos Bocas del Río Calovébora), about
<quantity id="B5199B09245BBD02FE20DD341AF4A091" box="[384,459,886,912]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="69" pageNumber="70" unit="km" value="16.0">16 km</quantity>
NW of Santa Fé,
<quantity id="B5199B09245BBD02FD02DD3419D0A091" box="[674,751,886,912]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="m" value="500.0">500 m</quantity>
,
<date id="065F102C245BBD02FD5CDD341F32A091" box="[764,1037,886,912]" pageId="69" pageNumber="70" value="1974-11-15" valueMax="1974-11-16" valueMin="1974-11-15">1516 November 1974</date>
,
<emphasis id="4095EAFE245BBD02FBBADD3A1FDDA091" box="[1050,1250,886,913]" italics="true" pageId="69" pageNumber="70">R. Dressler 4777</emphasis>
(
<typeStatus id="AD5A884E245BBD02FB52DD341E65A091" box="[1266,1370,886,912]" pageId="69" pageNumber="70" type="holotype">holotype</typeStatus>
BH!, isotypes MO!, PMA!
<collectingCountry id="0AF6767C245BBD02FE36DDDF1AFBA0B6" box="[406,452,925,951]" name="United States of America" pageId="69" pageNumber="70">US</collectingCountry>
!).
</paragraph>
</subSubSection>
<subSubSection id="3AFB6567245BBD02FF37DDA81882A4A4" pageId="69" pageNumber="70" type="description">
<paragraph id="725E36EC245BBD02FF37DDA81882A4A4" blockId="69.[151,1437,1002,2034]" pageId="69" pageNumber="70">
<emphasis id="4095EAFE245BBD02FF37DDA81BDDA705" box="[151,226,1002,1028]" italics="true" pageId="69" pageNumber="70">Plants</emphasis>
1.4(0.63.0) m tall; stems 1.6(0.53.0) m tall, 0.7(0.51.1) cm in diameter, solitary or clustered, canelike; internodes 1.2(0.62.6) cm long, yellowish and smooth.
<emphasis id="4095EAFE245BBD02FCC7DA501886A72A" box="[871,953,1042,1067]" italics="true" pageId="69" pageNumber="70">Leaves</emphasis>
9(810) per stem, undivided, not plicate, bases of blades recurved against the rachis; sheaths 5.8(3.08.2) cm long; petioles absent; rachis 34.1(18.8 50.5) cm long, 4.5(2.27.5) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae 1 per side of rachis; basal pinna forming an angle of 14(424)° with the rachis; apical pinna 9.0(5.214.0) cm long, forming an angle of 30(1645)° with the rachis.
<emphasis id="4095EAFE245BBD02FC80DAEE18F5A7C7" box="[800,970,1196,1222]" italics="true" pageId="69" pageNumber="70">Inflorescences</emphasis>
unbranched; prophylls and peduncular bracts ribbed with elongate, unbranched fibers, both bracts tubular, narrow, elongate, closely sheathing the peduncle, more or less persistent; prophylls 8.9(7.012.7) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 7.6(7.08.7) cm long, welldeveloped, inserted 2.5(1.05.6) cm above the prophyll; peduncles 10.2(5.015.0) cm long, 2.3(1.33.2) mm in diameter; rachillae 1, 20.3(9.038.0) cm long, 3.4(1.74.9) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits usually spirally arranged, sometimes tricussately, then the groups not closely spaced nor consistently arranged throughout the rachillae, glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers persistent after anthesis; staminodial tubes lobed, the lobes not spreading at anthesis, not acuminate, those of non-fertilized flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE245BBD02FEE2D9541ABFA431" box="[322,384,1814,1840]" italics="true" pageId="69" pageNumber="70">fruits</emphasis>
8.6(7.39.9) mm long, 5.1(4.75.3) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, not bumpy, not apiculate, ridged from the numerous, subepidermal, meridional, elongate fibers present, these coming to a point at fruit apices; locular epidermis without operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB6567245BBD01FF66D9F31ABCA7EC" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="distribution">
<paragraph id="725E36EC245BBD02FF66D9F31F08A4F3" blockId="69.[151,1437,1002,2034]" pageId="69" pageNumber="70">
<emphasis id="4095EAFE245BBD02FF66D9F3191AA4CA" bold="true" box="[198,549,1969,1995]" pageId="69" pageNumber="70">Distribution and habitat:—</emphasis>
From 8°31
<geoCoordinate id="17D5502B245BBD02FD1DD9F3181AA4CA" box="[701,805,1969,1995]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="69" pageNumber="70" precision="925" value="10.383333">10°23N</geoCoordinate>
and 79°16
<geoCoordinate id="17D5502B245BBD02FC61D9F31F10A4CA" box="[961,1071,1969,1995]" degrees="84" direction="west" minutes="06" orientation="longitude" pageId="69" pageNumber="70" precision="925" value="-84.1">84°06W</geoCoordinate>
in
<collectingCountry id="0AF6767C245BBD02FBFCD9F31FE0A4CA" box="[1116,1247,1969,1995]" name="Costa Rica" pageId="69" pageNumber="70">Costa Rica</collectingCountry>
and
<collectingCountry id="0AF6767C245BBD02FABFD9F31E43A4CA" box="[1311,1404,1969,1995]" name="Panama" pageId="69" pageNumber="70">Panama</collectingCountry>
at 557(3001200) m elevation in lowland to montane tropical rainforest (
<figureCitation id="EADA2A69245BBD02FC72D99A1F16A4F3" box="[978,1065,2008,2034]" captionStart="FIGURE 17" captionStartId="66.[151,255,1980,2003]" captionTargetBox="[164,1422,934,1950]" captionTargetId="figure@66.[161,1425,931,1954]" captionTargetPageId="66" captionText="FIGURE 17. Distribution maps of Geonoma dindoensis, G. divisa, G. elegans, and G. epetiolata." figureDoi="http://doi.org/10.5281/zenodo.3538396" httpUri="https://zenodo.org/record/3538396/files/figure.png" pageId="69" pageNumber="70">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="725E36EC2458BD01FF66DEDA1902A29B" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
<emphasis id="4095EAFE2458BD01FF66DEDA1AFFA3B3" bold="true" box="[198,448,152,178]" pageId="70" pageNumber="71">Taxonomic notes:—</emphasis>
<taxonomicName id="B5E14D6F2458BD01FE60DEDA198FA3B3" authorityName="Moore" authorityYear="1980" box="[448,688,152,178]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE2458BD01FE60DEDA198FA3B3" box="[448,688,152,178]" italics="true" pageId="70" pageNumber="71">Geonoma epetiolata</emphasis>
</taxonomicName>
is a member of a group of four Central American species, part of the
<taxonomicName id="B5E14D6F2458BD01FF44DEFC1A5CA3D8" authorityName="Wendland ex Spruce" authorityYear="1871" box="[228,355,190,217]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="cuneata">
<emphasis id="4095EAFE2458BD01FF44DEFC1A5CA3D8" box="[228,355,190,217]" italics="true" pageId="70" pageNumber="71">G. cuneata</emphasis>
</taxonomicName>
clade, including
<emphasis id="4095EAFE2458BD01FD92DEFC1844A3D9" box="[562,891,190,216]" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="B5E14D6F2458BD01FD92DEFC198AA3D9" authorityName="Grayum" authorityYear="1998" box="[562,693,190,216]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="brenesii">G. brenesii</taxonomicName>
,
<taxonomicName id="B5E14D6F2458BD01FD62DEFC1848A3D9" authorityName="de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[706,887,190,216]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="monospatha">G. monospatha</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B5E14D6F2458BD01FC17DEFC1F09A3D9" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[951,1078,190,216]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="hugonis">
<emphasis id="4095EAFE2458BD01FC17DEFC1F09A3D9" box="[951,1078,190,216]" italics="true" pageId="70" pageNumber="71">G. hugonis</emphasis>
</taxonomicName>
. They all have unbranched or few-branched inflorescences and share the character state of the staminodial tubes being lobed at the apex, but the lobes are not spreading at anthesis and are not acuminate. Although petiole length is treated as a quantitative variable here, the short or virtually absent petiole of
<taxonomicName id="B5E14D6F2458BD01FC2FDF701F1BA24D" authorityName="Moore" authorityYear="1980" box="[911,1060,306,332]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE2458BD01FC2FDF701F1BA24D" box="[911,1060,306,332]" italics="true" pageId="70" pageNumber="71">G. epetiolata</emphasis>
</taxonomicName>
is characteristic. This species is also characterized by its mottled leaves (
<bibRefCitation id="16704B1D2458BD01FD26DF1B1F11A272" author="Blanco, M. &amp; Marten-Rodriguez, S." box="[646,1070,345,371]" journalOrPublisher="Palms" pageId="70" pageNumber="71" pagination="139 - 146" part="51" refId="ref120866" refString="Blanco, M. &amp; Marten-Rodriguez, S. (2007) The stained - glass palm, Geonoma epetiolata. Palms 51: 139 - 146." title="The stained - glass palm, Geonoma epetiolata" type="journal article" year="2007">Blanco &amp; Martén-Rodríguez 2007</bibRefCitation>
), although this mottling is not obvious from herbarium specimens.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DFE41F6FA2E6" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
<emphasis id="4095EAFE2458BD01FF66DFE41AC9A2C1" bold="true" box="[198,502,422,448]" pageId="70" pageNumber="71">Subspecific variation:—</emphasis>
There is no variation in traits, except for stem branching. There is geographic discontinuity, and specimens come from
<collectingCountry id="0AF6767C2458BD01FDD7DF8F19C8A2E6" box="[631,759,461,487]" name="Costa Rica" pageId="70" pageNumber="71">Costa Rica</collectingCountry>
and several areas in
<collectingCountry id="0AF6767C2458BD01FC4BDF8F1F73A2E6" box="[1003,1100,461,487]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DFB618F8A1A9" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Specimens from
<collectingCountry id="0AF6767C2458BD01FE2EDFB61930A10F" box="[398,527,500,526]" name="Costa Rica" pageId="70" pageNumber="71">Costa Rica</collectingCountry>
and Santa Fé,
<collectingCountry id="0AF6767C2458BD01FD1EDFB61824A10F" box="[702,795,500,526]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
differ significantly from other Panamanian specimens in nine quantitative variables (stem diameter, rachis length, rachis width, basal pinna angle, apical pinna length, apical pinna angle, interbract distance, peduncle width, rachilla length)(
<emphasis id="4095EAFE2458BD01FBFFDC001F58A15A" box="[1119,1127,578,603]" italics="true" pageId="70" pageNumber="71">t</emphasis>
-test,
<emphasis id="4095EAFE2458BD01FB0CDC001F81A15A" box="[1196,1214,578,603]" italics="true" pageId="70" pageNumber="71">P</emphasis>
&lt;0.05). However, there is variation in both depth of staminodial tube lobing (
<bibRefCitation id="16704B1D2458BD01FC23DC2A1FE5A183" author="de Nevers, G. &amp; Grayum, M." box="[899,1242,616,642]" journalOrPublisher="Principes" pageId="70" pageNumber="71" pagination="94 - 103" part="42" refId="ref121952" refString="de Nevers, G. &amp; Grayum, M. (1998) Notes on Geonoma in Mesoamerica. Principes 42: 94 - 103." title="Notes on Geonoma in Mesoamerica" type="journal article" year="1998">de Nevers &amp; Grayum 1998</bibRefCitation>
) and flower pit arrangement, and this does not correspond to the geographic division.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DCF718E8A1F7" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Most specimens from
<collectingCountry id="0AF6767C2458BD01FE79DCF71961A1CE" box="[473,606,693,719]" name="Costa Rica" pageId="70" pageNumber="71">Costa Rica</collectingCountry>
have spirally arranged flower pits but some tend to be tricussately arranged. Depth of lobes of staminodial tubes varies from
<quantity id="B5199B092458BD01FCE2DC9E18EDA1F7" box="[834,978,732,758]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="70" pageNumber="71" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DD401A67A042" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Specimens from Santa Fé,
<collectingCountry id="0AF6767C2458BD01FDAADD401958A01D" box="[522,615,770,796]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
have spirally arranged flower pits and depth of lobes of staminodial tubes is
<quantity id="B5199B092458BD01FF56DD6B1A6BA042" box="[246,340,809,835]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="70" pageNumber="71" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DD1218B2A091" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Specimens from Cerro Tife and El Copé,
<collectingCountry id="0AF6767C2458BD01FD0DDD121835A06B" box="[685,778,848,874]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
have spirally to almost tricussately arranged flower pits and depth of lobes of staminodial tubes varies from
<quantity id="B5199B092458BD01FD58DD3418B6A091" box="[760,905,886,912]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="70" pageNumber="71" unit="mm" value="0.55" valueMax="0.9" valueMin="0.2">0.20.9 mm</quantity>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DDDF182DA0DF" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Specimens from Llano Grande,
<collectingCountry id="0AF6767C2458BD01FD9BDDDF19A7A0B6" box="[571,664,925,951]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
have spirally to almost tricussately arranged flower pits and depth of lobes of staminodial tubes varies from
<quantity id="B5199B092458BD01FDDEDD861831A0DF" box="[638,782,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="70" pageNumber="71" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DDA81881A72A" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
East of the Canal Zone, specimens from Cerro Brewster and Cerro Bruja have spirally arranged flower pits and depth of lobes of staminodial tubes varies from
<quantity id="B5199B092458BD01FC89DA531885A72A" box="[809,954,1041,1067]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="70" pageNumber="71" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.40.7 mm</quantity>
.
</paragraph>
<paragraph id="725E36EC2458BD01FF66DA7A1ABCA7EC" blockId="70.[151,1437,152,1261]" pageId="70" pageNumber="71">
Independently of staminodial tube lobing, there are at least four distinct populations of
<taxonomicName id="B5E14D6F2458BD01FB47DA7A1E41A753" authorityName="Moore" authorityYear="1980" box="[1255,1406,1080,1106]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE2458BD01FB47DA7A1E41A753" box="[1255,1406,1080,1106]" italics="true" pageId="70" pageNumber="71">G. epetiolata</emphasis>
</taxonomicName>
<emphasis id="4095EAFE2458BD01FADEDA7A1EA4A753" bold="true" box="[1406,1435,1080,1106]" pageId="70" pageNumber="71"></emphasis>
<collectingCountry id="0AF6767C2458BD01FF37DA1C1A24A779" box="[151,283,1118,1144]" name="Costa Rica" pageId="70" pageNumber="71">Costa Rica</collectingCountry>
; Santa Fé; El Copé/Llano Grande/Cerro Tife; and Cerro Brewster/Cerro Bruja. However, because of the complexity in staminodial tube lobing and triad arrangement, and because there are few specimens from most sites, and the gaps between the sites in
<collectingCountry id="0AF6767C2458BD01FD05DAEE183DA7C7" box="[677,770,1196,1222]" name="Panama" pageId="70" pageNumber="71">Panama</collectingCountry>
may be a result of insufficient collecting, no subspecific taxa are recognized.
</paragraph>
</subSubSection>
</treatment>
</document>