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<document id="F09B0A4CAE6B5593AE693920D55FBEB8" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA2450BD08FF37D86C18F6A47E" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma hugonis Grayum &amp; de Nevers in de Nevers &amp; Grayum 1998" docType="treatment" docVersion="7" lastPageNumber="80" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="79" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:title id="19E8F356138140B951BC08D495F0FE76">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:namePart id="5E47FE20E1A0A3A7C16BE075B0BEE41E">Henderson, Andrew</mods:namePart>
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<mods:title id="733C19EAE76E4B559EE47A300B959A8B">Phytotaxa</mods:title>
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<mods:part id="2774C5EC7BBF7BC7272C7EF762C1CD52">
<mods:date id="59AA680A6B2ADADDDBAE9956F0D723DE">2011</mods:date>
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<mods:number id="CE079725D22E62911E54A4D35B0E92BE">2011-02-18</mods:number>
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<treatment id="FA4887FA2450BD08FF37D86C18F6A47E" ID-DOI="http://doi.org/10.5281/zenodo.5609007" ID-GBIF-Taxon="160342467" ID-Zenodo-Dep="5609007" LSID="urn:lsid:plazi:treatment:FA4887FA2450BD08FF37D86C18F6A47E" httpUri="http://treatment.plazi.org/id/FA4887FA2450BD08FF37D86C18F6A47E" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<subSubSection id="3AFB65672450BD09FF37D86C1F6BA549" box="[151,1108,1581,1608]" pageId="78" pageNumber="79" type="nomenclature">
<paragraph id="725E36EC2450BD09FF37D86C1F6BA549" blockId="78.[151,1436,1581,1725]" box="[151,1108,1581,1608]" pageId="78" pageNumber="79">
<emphasis id="4095EAFE2450BD09FF37D86C1B83A549" bold="true" box="[151,188,1582,1608]" pageId="78" pageNumber="79">28.</emphasis>
<taxonomicName id="B5E14D6F2450BD09FF63D86C1F6FA549" ID-CoL="3FQKZ" authority="Grayum &amp; de Nevers in de Nevers &amp; Grayum (1998: 94)" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityPageNumber="94" authorityYear="1998" box="[195,1104,1581,1608]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="species" species="hugonis">
<emphasis id="4095EAFE2450BD09FF63D86C1A9DA549" bold="true" box="[195,418,1581,1608]" italics="true" pageId="78" pageNumber="79">Geonoma hugonis</emphasis>
Grayum &amp; de Nevers in
<bibRefCitation id="16704B1D2450BD09FD6DD86C1F6FA549" author="de Nevers, G. &amp; Grayum, M." box="[717,1104,1582,1608]" journalOrPublisher="Principes" pageId="78" pageNumber="79" pagination="94 - 103" part="42" refId="ref121952" refString="de Nevers, G. &amp; Grayum, M. (1998) Notes on Geonoma in Mesoamerica. Principes 42: 94 - 103." title="Notes on Geonoma in Mesoamerica" type="journal article" year="1998">de Nevers &amp; Grayum (1998: 94)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672450BD08FBFCD86C185BA06B" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="725E36EC2450BD09FBFCD86C1A4CA5BC" blockId="78.[151,1436,1581,1725]" pageId="78" pageNumber="79">
Type:
<collectingCountry id="0AF6767C2450BD09FB06D86C1E18A549" box="[1190,1319,1582,1608]" name="Panama" pageId="78" pageNumber="79">PANAMA</collectingCountry>
.
<collectingRegion id="B025F80E2450BD09FA92D86C1EA7A549" box="[1330,1432,1582,1608]" country="Panama" name="Chiriqui" pageId="78" pageNumber="79">Chiriquí</collectingRegion>
: Fortuna Dam Area, between Quebrada Los Chorros and Quebrada Hondo, to
<collectionCode id="14F0AE292450BD09FBE0D8171F6AA56E" box="[1088,1109,1621,1647]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="78" pageNumber="79" type="Herbarium">N</collectionCode>
of reservoir, in forest
<collectionCode id="14F0AE292450BD09FAC5D8171E45A56E" box="[1381,1402,1621,1647]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="78" pageNumber="79" type="Herbarium">N</collectionCode>
of road,
<geoCoordinate id="17D5502B2450BD09FF7ED83E1A05A597" box="[222,314,1660,1686]" degrees="8" direction="north" minutes="45" orientation="latitude" pageId="78" pageNumber="79" precision="925" value="8.75">8°45N</geoCoordinate>
,
<geoCoordinate id="17D5502B2450BD09FEE8D83E1A87A597" box="[328,440,1660,1686]" degrees="82" direction="west" minutes="14" orientation="longitude" pageId="78" pageNumber="79" precision="925" value="-82.23333">82°14W</geoCoordinate>
,
<quantity id="B5199B092450BD09FE66D83E191AA597" box="[454,549,1660,1686]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="78" pageNumber="79" unit="m" value="1100.0">1100 m</quantity>
,
<date id="065F102C2450BD09FD93D83E181AA597" box="[563,805,1660,1686]" pageId="78" pageNumber="79" value="1984-09-20">20 September 1984</date>
,
<emphasis id="4095EAFE2450BD09FC92D83F1FECA597" box="[818,1235,1660,1686]" italics="true" pageId="78" pageNumber="79">
<collectionCode id="14F0AE292450BD09FC92D83F1873A597" box="[818,844,1661,1686]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="78" pageNumber="79" type="Herbarium">H</collectionCode>
. Churchill &amp;
<collectionCode id="14F0AE292450BD09FC56D83E1F32A597" box="[1014,1037,1660,1686]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="78" pageNumber="79" type="Herbarium">A</collectionCode>
. Churchill 6185
</emphasis>
(
<typeStatus id="AD5A884E2450BD09FB44D83E1E6EA597" box="[1252,1361,1660,1686]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
<collectionCode id="14F0AE292450BD09FAFAD83E1EABA597" box="[1370,1428,1660,1686]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="78" pageNumber="79" type="Herbarium">MO</collectionCode>
!, isotype
<collectionCode id="14F0AE292450BD09FF53D8E11A10A5BC" box="[243,303,1699,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="78" pageNumber="79" type="Museum">CAS</collectionCode>
,
<emphasis id="4095EAFE2450BD09FE9AD8E61A5DA5BC" box="[314,354,1700,1725]" italics="true" pageId="78" pageNumber="79">n.v.</emphasis>
).
</paragraph>
<paragraph id="725E36EC2450BD08FF37D8B2185BA06B" blockId="78.[151,1437,1776,2034]" lastBlockId="79.[151,1437,152,951]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<emphasis id="4095EAFE2450BD09FF37D8B21BDDA40B" box="[151,226,1776,1802]" italics="true" pageId="78" pageNumber="79">Plants</emphasis>
0.8(0.31.5) m tall; stems 0.7(0.11.2) m tall, 0.6(0.40.8) cm in diameter, solitary or clustered, canelike; internodes 1.9(1.03.5) cm long, covered with reddish or brownish scales, especially in their distal part.
<emphasis id="4095EAFE2450BD09FF37D97D1BD5A459" box="[151,234,1855,1880]" italics="true" pageId="78" pageNumber="79">Leaves</emphasis>
8(412) per stem, undivided, not plicate, bases of blades running diagonally into the rachis; sheaths 7.2(3.514.0) cm long; petioles 7.0(1.012.0) cm long, drying green or yellowish; rachis 12.7(7.020.0) cm long, 1.8(1.32.7) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae 1 per side of rachis; basal pinna length and width not applicable, forming an angle of 29(2240)° with the rachis; apical pinna 10.4(5.817.2) cm long, width not applicable, forming an angle of 32(2242)° with the rachis.
<emphasis id="4095EAFE2451BD08FF37DEDA1A7BA3B3" box="[151,324,152,178]" italics="true" pageId="79" pageNumber="80">Inflorescences</emphasis>
unbranched; prophylls and peduncular bracts ribbed with elongate, unbranched fibers, both bracts tubular, narrow, elongate, closely sheathing the peduncle, more or less persistent; prophylls 12.8(11.0 17.0) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 11.4(1.518.4) cm long, well-developed, inserted 20.4(17.823.0) cm above the prophyll; peduncles 29.9(13.543.5) cm long, 1.3(0.92.0) mm in diameter; rachillae1, 7.1(4.612.5) cm long, 2.4(1.7 3.5) mm in diameter the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips with a central notch before anthesis, often the two sides of the notch overlapping, not recurved after anthesis, not hood-shaped at anthesis; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers deciduous after anthesis; staminodial tubes lobed, the lobes not spreading at anthesis, not acuminate; staminodial tubes of non-fertilized pistillate flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE2451BD08FE6EDC9E192FA1F7" box="[462,528,732,758]" italics="true" pageId="79" pageNumber="80">fruits</emphasis>
6.2(5.46.9) mm long, 5.4(4.76.3) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis without operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672451BD08FF66DD3418F6A47E" pageId="79" pageNumber="80" type="distribution">
<paragraph id="725E36EC2451BD08FF66DD34184FA0B6" blockId="79.[151,1437,152,951]" pageId="79" pageNumber="80">
<emphasis id="4095EAFE2451BD08FF66DD341902A091" bold="true" box="[198,573,886,912]" pageId="79" pageNumber="80">Distribution and habitat:—</emphasis>
From 8°30
<geoCoordinate id="17D5502B2451BD08FD43DD341805A091" box="[739,826,886,912]" degrees="8" direction="north" minutes="46" orientation="latitude" pageId="79" pageNumber="80" precision="925" value="8.766666">8°46N</geoCoordinate>
and 81°45
<geoCoordinate id="17D5502B2451BD08FC50DD341F63A091" box="[1008,1116,886,912]" degrees="82" direction="west" minutes="17" orientation="longitude" pageId="79" pageNumber="80" precision="925" value="-82.28333">82°17W</geoCoordinate>
in western
<collectingCountry id="0AF6767C2451BD08FAB3DD341E4FA091" box="[1299,1392,886,912]" name="Panama" pageId="79" pageNumber="80">Panama</collectingCountry>
at 1243(11001450) m elevation in montane rainforest (
<figureCitation id="EADA2A692451BD08FCAADDDF185DA0B6" box="[778,866,925,951]" captionStart="FIGURE 20" captionStartId="79.[151,255,1553,1576]" captionTargetBox="[169,1393,1029,1527]" captionTargetId="figure@79.[151,1436,1009,1527]" captionTargetPageId="79" captionText="FIGURE 20. Distribution maps of Geonoma hollinensis and G. hugonis." figureDoi="http://doi.org/10.5281/zenodo.3538402" httpUri="https://zenodo.org/record/3538402/files/figure.png" pageId="79" pageNumber="80">Fig. 20</figureCitation>
).
</paragraph>
<caption id="269E66642451BD08FF37D85318A0A529" ID-DOI="http://doi.org/10.5281/zenodo.3538402" ID-Zenodo-Dep="3538402" box="[151,927,1553,1576]" httpUri="https://zenodo.org/record/3538402/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,255,1553,1576]" targetBox="[169,1393,1029,1527]" targetPageId="79">
<paragraph id="725E36EC2451BD08FF37D85318A0A529" blockId="79.[151,927,1553,1576]" box="[151,927,1553,1576]" pageId="79" pageNumber="80">
<emphasis id="4095EAFE2451BD08FF37D8531A1FA529" bold="true" box="[151,288,1553,1576]" pageId="79" pageNumber="80">FIGURE 20</emphasis>
. Distribution maps of
<taxonomicName id="B5E14D6F2451BD08FDB0D85319CDA529" authorityName="Henderson, Borchsenius &amp; Balslev" authorityYear="2008" box="[528,754,1553,1576]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="hollinensis">
<emphasis id="4095EAFE2451BD08FDB0D85319CDA529" box="[528,754,1553,1576]" italics="true" pageId="79" pageNumber="80">Geonoma hollinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B5E14D6F2451BD08FC86D85318A8A529" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[806,919,1553,1576]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="hugonis">
<emphasis id="4095EAFE2451BD08FC86D8531809A529" box="[806,822,1553,1576]" italics="true" pageId="79" pageNumber="80">G</emphasis>
.
<emphasis id="4095EAFE2451BD08FCE3D85318A8A529" box="[835,919,1553,1576]" italics="true" pageId="79" pageNumber="80">hugonis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="725E36EC2451BD08FF66D83F1807A430" blockId="79.[151,1437,1661,1919]" pageId="79" pageNumber="80">
<emphasis id="4095EAFE2451BD08FF66D83F1A80A596" bold="true" box="[198,447,1661,1687]" pageId="79" pageNumber="80">Taxonomic notes:—</emphasis>
<taxonomicName id="B5E14D6F2451BD08FE1FD83F19A9A596" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[447,662,1661,1687]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="hugonis">
<emphasis id="4095EAFE2451BD08FE1FD83F19A9A596" box="[447,662,1661,1687]" italics="true" pageId="79" pageNumber="80">Geonoma hugonis</emphasis>
</taxonomicName>
is a member of a group of four Central American species, part of the
<taxonomicName id="B5E14D6F2451BD08FF63D8E61A7FA5BF" authorityName="Wendland ex Spruce" authorityYear="1871" box="[195,320,1700,1726]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="cuneata">
<emphasis id="4095EAFE2451BD08FF63D8E61A7FA5BF" box="[195,320,1700,1726]" italics="true" pageId="79" pageNumber="80">G. cuneata</emphasis>
</taxonomicName>
clade, also including
<taxonomicName id="B5E14D6F2451BD08FDE4D8E619FDA5BF" authorityName="Grayum" authorityYear="1998" box="[580,706,1700,1726]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="brenesii">
<emphasis id="4095EAFE2451BD08FDE4D8E619FDA5BF" box="[580,706,1700,1726]" italics="true" pageId="79" pageNumber="80">G. brenesii</emphasis>
</taxonomicName>
,
<emphasis id="4095EAFE2451BD08FD72D8E618B6A5BF" box="[722,905,1700,1726]" italics="true" pageId="79" pageNumber="80">
<taxonomicName id="B5E14D6F2451BD08FD72D8E618BAA5BF" authorityName="de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[722,901,1700,1726]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="monospatha">G. monospatha</taxonomicName>
,
</emphasis>
and
<taxonomicName id="B5E14D6F2451BD08FC64D8E61F65A5BF" authorityName="Moore" authorityYear="1980" box="[964,1114,1700,1726]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE2451BD08FC64D8E61F65A5BF" box="[964,1114,1700,1726]" italics="true" pageId="79" pageNumber="80">G. epetiolata</emphasis>
</taxonomicName>
. They all have unbranched or few-branched inflorescences and share the character state of the staminodial tubes being lobed at the apex, but the lobes are not spreading at anthesis and are not acuminate.
<taxonomicName id="B5E14D6F2451BD08FC34D8B31F55A40A" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[916,1130,1777,1803]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="hugonis">
<emphasis id="4095EAFE2451BD08FC34D8B31F55A40A" box="[916,1130,1777,1803]" italics="true" pageId="79" pageNumber="80">Geonoma hugonis</emphasis>
</taxonomicName>
differs from these species in its internodes covered with reddish or brownish scales.
</paragraph>
<paragraph id="725E36EC2451BD08FF66D97C18F6A47E" blockId="79.[151,1437,1661,1919]" pageId="79" pageNumber="80">
<emphasis id="4095EAFE2451BD08FF66D97C1AC5A459" bold="true" box="[198,506,1854,1880]" pageId="79" pageNumber="80">Subspecific variation:—</emphasis>
Only one trait (stem branching) varies in this species. There is geographic disjunction but this is likely to be an artifact of insufficient collecting.
</paragraph>
</subSubSection>
</treatment>
</document>