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<document id="FCD55C2D6C772E866FF734D68909EF65" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA2436BD6EFF37DB7F1842A0B6" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma brenesii Grayum 1998" docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="41" updateTime="1698765326825" updateUser="ExternalLinkService">
<mods:mods id="CD67C1F0331F6A791FE57DA0A1192E7E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="83AB06D47DCF6B48ECFAA8C297562DEE">
<mods:title id="6FD17307A1D30A26F4172EE901738145">A revision of Geonoma (Arecaceae)</mods:title>
</mods:titleInfo>
<mods:name id="5A6B75B08FA703CF537249BAB3815088" type="personal">
<mods:role id="87597810A7306E497249174C94052A93">
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<mods:namePart id="215B7586CCF0C0C3DBA2BCD1A930C126">Henderson, Andrew</mods:namePart>
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<mods:title id="AA8FA1A1B0CA9A3F3440FC1976F0CDAC">Phytotaxa</mods:title>
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<mods:part id="677A585FD716A84A390CF483FB487C49">
<mods:date id="FB9DC2946A82C61F9B11324E9C2A818F">2011</mods:date>
<mods:detail id="C7D8CC22F414927A7D23C18F00EA1182" type="pubDate">
<mods:number id="88D6419BDF7A72105418D304D4763457">2011-02-18</mods:number>
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<mods:number id="C2A5A1A3367EF30AD7942418CB9D7077">17</mods:number>
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<mods:identifier id="A6D89057B7710C802DDC5208B320644A" type="DOI">10.11646/phytotaxa.17.1</mods:identifier>
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<treatment id="FA4887FA2436BD6EFF37DB7F1842A0B6" ID-DOI="http://doi.org/10.5281/zenodo.5608895" ID-GBIF-Taxon="160342508" ID-Zenodo-Dep="5608895" LSID="urn:lsid:plazi:treatment:FA4887FA2436BD6EFF37DB7F1842A0B6" httpUri="http://treatment.plazi.org/id/FA4887FA2436BD6EFF37DB7F1842A0B6" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<subSubSection id="3AFB65672436BD6FFF37DB7F1897A6A4" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="725E36EC2436BD6FFF37DB7F1897A6A4" blockId="40.[151,1436,1340,1445]" pageId="40" pageNumber="41">
<emphasis id="4095EAFE2436BD6FFF37DB7F1B92A656" bold="true" box="[151,173,1341,1367]" pageId="40" pageNumber="41">5.</emphasis>
<taxonomicName id="B5E14D6F2436BD6FFF14DB7F19B3A656" ID-CoL="3FQH2" authority="Grayum (1998: 322)" authorityName="Grayum" authorityPageNumber="322" authorityYear="1998" box="[180,652,1340,1367]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="brenesii">
<emphasis id="4095EAFE2436BD6FFF14DB7F1AAFA656" bold="true" box="[180,400,1340,1367]" italics="true" pageId="40" pageNumber="41">Geonoma brenesii</emphasis>
<bibRefCitation id="16704B1D2436BD6FFE37DB7F19B3A656" author="Grayum, M." box="[407,652,1341,1367]" journalOrPublisher="Phytologia" pageId="40" pageNumber="41" pagination="307 - 327" part="84" refId="ref122440" refString="Grayum, M. (1998) Nomenclatural and taxonomic notes on Costa Rican palms (Arecaceae), with five new species. Phytologia 84: 307 - 327." title="Nomenclatural and taxonomic notes on Costa Rican palms (Arecaceae), with five new species" type="journal article" year="1998">Grayum (1998: 322)</bibRefCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C2436BD6FFD40DB7F18B0A656" box="[736,911,1341,1367]" name="Costa Rica" pageId="40" pageNumber="41">COSTA RICA</collectingCountry>
.
<collectingRegion id="B025F80E2436BD6FFC39DB7F1F3EA656" box="[921,1025,1341,1367]" country="Costa Rica" name="Alajuela" pageId="40" pageNumber="41">Alajuela</collectingRegion>
: Reserva Biológica de San Ramón, road from Las Lagunas to Colonia Palmareña,
<geoCoordinate id="17D5502B2436BD6FFD61DB261816A67F" box="[705,809,1380,1406]" degrees="10" direction="north" minutes="04" orientation="latitude" pageId="40" pageNumber="41" precision="925" value="10.066667">10°04N</geoCoordinate>
,
<geoCoordinate id="17D5502B2436BD6FFC96DB26189EA67F" box="[822,929,1380,1406]" degrees="84" direction="west" minutes="32" orientation="longitude" pageId="40" pageNumber="41" precision="925" value="-84.53333">84°32W</geoCoordinate>
,
<quantity id="B5199B092436BD6FFC0EDB261F7AA67F" box="[942,1093,1380,1406]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="11.0" metricValueMin="8.5" pageId="40" pageNumber="41" unit="m" value="975.0" valueMax="1100.0" valueMin="850.0">8501100 m</quantity>
,
<date id="065F102C2436BD6FFBF1DB261FCDA67F" box="[1105,1266,1380,1406]" pageId="40" pageNumber="41" value="1986-05-30">30 May 1986</date>
,
<emphasis id="4095EAFE2436BD6FFB5EDB261ACAA6A4" italics="true" pageId="40" pageNumber="41">G. de Nevers, B. Hammel &amp; C. Gómez 7789</emphasis>
(
<typeStatus id="AD5A884E2436BD6FFDA5DBC91952A6A4" box="[517,621,1419,1445]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
MO!, isotypes CR!, NY!).
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672436BD6EFF37DB9A19AFA15A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph id="725E36EC2436BD6EFF37DB9A19AFA15A" blockId="40.[151,1437,1496,2025]" lastBlockId="41.[151,1437,152,951]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="4095EAFE2436BD6FFF37DB9A1BDDA6F3" box="[151,226,1496,1522]" italics="true" pageId="40" pageNumber="41">Plants</emphasis>
0.8(0.51.0) m tall; stems 0.3(0.20.4) m tall,
<quantity id="B5199B092436BD6FFCA4DB9A186CA6F3" box="[772,851,1496,1522]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="40" pageNumber="41" unit="cm" value="1.2">1.2 cm</quantity>
in diameter, solitary, not cane-like; internodes
<quantity id="B5199B092436BD6FFAD8DB9A1B82A518" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="40" pageNumber="41" unit="cm" value="0.5">0.5 cm</quantity>
long, not scaly.
<emphasis id="4095EAFE2436BD6FFE31D8421AD7A518" box="[401,488,1536,1561]" italics="true" pageId="40" pageNumber="41">Leaves</emphasis>
9(811) per stem, irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 8.2(6.510.0) cm long; petioles 31.8(21.038.5) cm long, drying green or yellowish; rachis 24.6(18.829.8) cm long, 2.8(1.94.3) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae 3(34) per side of rachis; basal pinna 28.3(21.536.5) cm long, 4.1(2.010.2) cm wide, forming an angle of 43(3053)° with the rachis; apical pinna 16.7(13.521.0) cm long, 9.5(6.5 12.0) cm wide, forming an angle of 29(2532)° with the rachis.
<emphasis id="4095EAFE2436BD6FFC04D8821F6EA5DB" box="[932,1105,1728,1754]" italics="true" pageId="40" pageNumber="41">Inflorescences</emphasis>
unbranched; prophylls and peduncular bracts ribbed with elongate, unbranched fibers, both bracts tubular, narrow, elongate, closely sheathing the peduncle, more or less persistent; prophylls 13.5(10.017.5) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 3.9(1.5 7.1) cm long, vestigial, inserted 4.9(3.47.0) cm above the prophyll; peduncles 24.3(15.037.5) cm long, 2.1(1.42.8) mm in diameter; rachillae 1, 12.5(8.516.0) cm long, 3.6(2.85.0) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips with a central notch before anthesis, often the two sides of the notch overlapping, the lip, more or less heartshaped, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers persistent after anthesis; staminodial tubes lobed, the lobes not spreading at anthesis, not acuminate, those of non-fertilized flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE2437BD6EFEA6DF8F1A7AA2E6" box="[262,325,461,487]" italics="true" pageId="41" pageNumber="42">fruits</emphasis>
7.3(6.97.6) mm long, 6.1(6.06.2) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis without operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672437BD6EFF66DC2A1842A0B6" pageId="41" pageNumber="42" type="distribution">
<paragraph id="725E36EC2437BD6EFF66DC2A19DCA1A9" blockId="41.[151,1437,152,951]" pageId="41" pageNumber="42">
<emphasis id="4095EAFE2437BD6EFF66DC2A1919A183" bold="true" box="[198,550,616,642]" pageId="41" pageNumber="42">Distribution and habitat:—</emphasis>
From 10°02
<geoCoordinate id="17D5502B2437BD6EFD70DC2A180BA183" box="[720,820,616,642]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="10.383333">10°23N</geoCoordinate>
and 83°59
<geoCoordinate id="17D5502B2437BD6EFC74DC2A1F7FA183" box="[980,1088,616,642]" degrees="84" direction="west" minutes="51" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-84.85">84°51W</geoCoordinate>
in
<collectingCountry id="0AF6767C2437BD6EFBCDDC2A1FCEA183" box="[1133,1265,616,642]" name="Costa Rica" pageId="41" pageNumber="42">Costa Rica</collectingCountry>
at 1293(967 1550) m elevation in montane rainforest (
<figureCitation id="EADA2A692437BD6EFDDEDCCC19E9A1A9" box="[638,726,654,680]" captionStart="FIGURE 10" captionStartId="40.[159,269,1195,1218]" captionTargetBox="[170,1417,161,1170]" captionTargetId="figure@40.[151,1436,141,1190]" captionTargetPageId="40" captionText="FIGURE 10. Distribution maps of Geonoma braunii, G. brenesii, G. brongniartii subsp. brongniartii, and G. brongniartii subsp. pascoensis." figureDoi="http://doi.org/10.5281/zenodo.3538382" httpUri="https://zenodo.org/record/3538382/files/figure.png" pageId="41" pageNumber="42">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="725E36EC2437BD6EFF66DCF71946A091" blockId="41.[151,1437,152,951]" pageId="41" pageNumber="42">
<emphasis id="4095EAFE2437BD6EFF66DCF71A80A1CE" bold="true" box="[198,447,693,719]" pageId="41" pageNumber="42">Taxonomic notes:—</emphasis>
<taxonomicName id="B5E14D6F2437BD6EFE1FDCF719A9A1CE" authorityName="Grayum" authorityYear="1998" box="[447,662,693,719]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="brenesii">
<emphasis id="4095EAFE2437BD6EFE1FDCF719A9A1CE" box="[447,662,693,719]" italics="true" pageId="41" pageNumber="42">Geonoma brenesii</emphasis>
</taxonomicName>
is a member of a group of four Central American species, part of the
<taxonomicName id="B5E14D6F2437BD6EFF64DC9E1A7BA1F7" authorityName="Wendland ex Spruce" authorityYear="1871" box="[196,324,732,758]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="cuneata">
<emphasis id="4095EAFE2437BD6EFF64DC9E1A7BA1F7" box="[196,324,732,758]" italics="true" pageId="41" pageNumber="42">G. cuneata</emphasis>
</taxonomicName>
clade, including
<emphasis id="4095EAFE2437BD6EFDB4DC9E1869A1F7" box="[532,854,732,758]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="B5E14D6F2437BD6EFDB4DC9E19F5A1F7" authorityName="de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[532,714,732,758]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="monospatha">G. monospatha</taxonomicName>
,
<taxonomicName id="B5E14D6F2437BD6EFD77DC9E1869A1F7" authorityName="Grayum &amp; de Nevers in de Nevers &amp; Grayum" authorityYear="1998" box="[727,854,732,758]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="hugonis">G. hugonis</taxonomicName>
</emphasis>
, and
<taxonomicName id="B5E14D6F2437BD6EFC3ADC9E1F0CA1F7" authorityName="Moore" authorityYear="1980" box="[922,1075,732,758]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="epetiolata">
<emphasis id="4095EAFE2437BD6EFC3ADC9E1F0CA1F7" box="[922,1075,732,758]" italics="true" pageId="41" pageNumber="42">G. epetiolata</emphasis>
</taxonomicName>
. They all have unbranched or few-branched inflorescences and share the character state of the staminodial tubes being lobed at the apex, but the lobes are not spreading at anthesis and are not acuminate.
<taxonomicName id="B5E14D6F2437BD6EFC0DDD6B1FACA042" authorityName="Grayum" authorityYear="1998" box="[941,1171,809,835]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="brenesii">
<emphasis id="4095EAFE2437BD6EFC0DDD6B1FACA042" box="[941,1171,809,835]" italics="true" pageId="41" pageNumber="42">Geonoma brenesii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="B5E14D6F2437BD6EFA22DD6B1A17A06B" authorityName="de Nevers in de Nevers &amp; Grayum" authorityYear="1998" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="monospatha">
<emphasis id="4095EAFE2437BD6EFA22DD6B1A17A06B" italics="true" pageId="41" pageNumber="42">G. monospatha</emphasis>
</taxonomicName>
, differing in its tubular, narrow, elongate, sheathing, persistent prophylls and peduncular bracts which are ribbed with unbranched fibers.
</paragraph>
<paragraph id="725E36EC2437BD6EFF66DDDF1842A0B6" blockId="41.[151,1437,152,951]" box="[198,893,925,951]" pageId="41" pageNumber="42">
<emphasis id="4095EAFE2437BD6EFF66DDDF1ACBA0B6" bold="true" box="[198,500,925,951]" pageId="41" pageNumber="42">Subspecific variation:—</emphasis>
No traits vary within this species.
</paragraph>
</subSubSection>
</treatment>
</document>