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<document id="1B264F258FD7DC05343CF066D33D29D7" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA2432BD6AFF37D8DE185FA456" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma camana Trail 1876" docType="treatment" docVersion="7" lastPageNumber="46" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="45" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:title id="DFD0695795015EF26275A09B3DC89D58">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:namePart id="4A5C86ECA307FC6230D27F3EC0F02BF0">Henderson, Andrew</mods:namePart>
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<mods:title id="72E8968F2802A32F13892DF5226A5DDA">Phytotaxa</mods:title>
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<mods:date id="E86083AD3CF118DC8C225ADBC2ABAE72">2011</mods:date>
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<treatment id="FA4887FA2432BD6AFF37D8DE185FA456" ID-DOI="http://doi.org/10.5281/zenodo.5608908" ID-GBIF-Taxon="160342500" ID-Zenodo-Dep="5608908" LSID="urn:lsid:plazi:treatment:FA4887FA2432BD6AFF37D8DE185FA456" httpUri="http://treatment.plazi.org/id/FA4887FA2432BD6AFF37D8DE185FA456" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="3AFB65672432BD6BFF37D8DE1F8DA5B7" box="[151,1202,1691,1718]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="725E36EC2432BD6BFF37D8DE1F8DA5B7" blockId="44.[151,1437,1691,1851]" box="[151,1202,1691,1718]" pageId="44" pageNumber="45">
<emphasis id="4095EAFE2432BD6BFF37D8DE1B92A5B7" bold="true" box="[151,173,1692,1718]" pageId="44" pageNumber="45">8.</emphasis>
<taxonomicName id="B5E14D6F2432BD6BFF14D8D9195BA5B7" ID-CoL="3FQH8" authority="Trail (1876: 324)" authorityName="Trail" authorityPageNumber="324" authorityYear="1876" box="[180,612,1691,1718]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2432BD6BFF14D8D91AAEA5B4" bold="true" box="[180,401,1691,1717]" italics="true" pageId="44" pageNumber="45">Geonoma camana</emphasis>
<bibRefCitation id="16704B1D2432BD6BFE38D8DE195BA5B7" author="Trail, W." box="[408,612,1692,1718]" journalOrPublisher="Journal of Botany" pageId="44" pageNumber="45" pagination="323 - 333" part="5" refId="ref124658" refString="Trail, W. (1876) Descriptions of new species and varieties of palm collected in the valley of the Amazon in north Brazil, in 1874. Journal of Botany 5: 323 - 333." title="Descriptions of new species and varieties of palm collected in the valley of the Amazon in north Brazil" type="journal article" year="1876">Trail (1876: 324)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="B5E14D6F2432BD6BFDCFD8DE1F8DA5B7" authority="(Trail) Burret (1930 a: 270)" authorityName="Burret" authorityYear="1930" baseAuthorityName="Trail" box="[623,1202,1692,1718]" class="Liliopsida" family="Arecaceae" genus="Taenianthera" higherTaxonomySource="CoL" kingdom="Plantae" order="Arecales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2432BD6BFDCFD8DE1850A5B7" box="[623,879,1692,1718]" italics="true" pageId="44" pageNumber="45">Taenianthera camana</emphasis>
(Trail)
<bibRefCitation id="16704B1D2432BD6BFC69D8DE1F8DA5B7" author="Burret, M." box="[969,1202,1692,1718]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="44" pageNumber="45" pagination="123 - 270" part="63" refId="ref121027" refString="Burret, M. (1930 a) Geonomeae americanae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 63: 123 - 270." title="Geonomeae americanae" type="journal article" year="1930">Burret (1930a: 270)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672432BD6BFF37D8801A4CA5F8" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="725E36EC2432BD6BFF37D8801A4CA5F8" blockId="44.[151,1437,1691,1851]" pageId="44" pageNumber="45">
Type:
<collectingCountry id="0AF6767C2432BD6BFF7AD8801A01A5D8" box="[218,318,1730,1753]" name="Brazil" pageId="44" pageNumber="45">BRAZIL</collectingCountry>
. Amazonas: San Antonia da Boa Vista, Rio Javari,
<date id="065F102C2432BD6BFCC3D8801F1CA5D8" box="[867,1059,1730,1753]" pageId="44" pageNumber="45" value="1874-12-04">4 December 1874</date>
,
<emphasis id="4095EAFE2432BD6BFB8ED8861E2AA5D8" box="[1070,1301,1730,1755]" italics="true" pageId="44" pageNumber="45">
<collectionCode id="14F0AE292432BD6BFB8ED8861F02A5DA" box="[1070,1085,1732,1755]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="44" pageNumber="45" type="Herbarium">J</collectionCode>
. Trail 977/CLXXXII
</emphasis>
(
<typeStatus id="AD5A884E2432BD6BFA83D8801EBDA5D8" box="[1315,1410,1730,1753]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
<collectionCode id="14F0AE292432BD6BFA29D8801EA3A5D8" box="[1417,1436,1730,1753]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="44" pageNumber="45" type="Herbarium">K</collectionCode>
<emphasis id="4095EAFE2432BD6BFF67D8A61BD4A5FA" box="[199,235,1764,1787]" italics="true" pageId="44" pageNumber="45">n.v.</emphasis>
, isotype
<collectionCode id="14F0AE292432BD6BFEECD8A01A5EA5F8" box="[332,353,1762,1785]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="44" pageNumber="45" type="Herbarium">P</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672432BD6AFF37D9401887A0DF" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="725E36EC2432BD6BFF37D9401FD0A438" blockId="44.[151,1437,1691,1851]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="52F111C22432BD6BFF37D9401FD0A438" infered="true" pageId="44" pageNumber="45">
<taxonomicName id="B5E14D6F2432BD6BFF37D9401955A418" ID-CoL="54GVD" authority="Dammer (1907: 121)" authorityName="Dammer" authorityPageNumber="121" authorityYear="1907" box="[151,618,1794,1817]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="lagesiana">
<emphasis id="4095EAFE2432BD6BFF37D9401A4DA418" box="[151,370,1794,1817]" italics="true" pageId="44" pageNumber="45">Geonoma lagesiana</emphasis>
<treatmentCitation id="F34010FD2432BD6BFEDCD9401955A418" author="Dammer, U." box="[380,618,1794,1817]" page="121" pageId="44" pageNumber="45" year="1907">
<bibRefCitation id="16704B1D2432BD6BFEDCD9401955A418" author="Dammer, U." box="[380,618,1794,1817]" journalOrPublisher="Verhandlungen des Botanischen Vereins der Provinz Brandenburg" pageId="44" pageNumber="45" pagination="118 - 129" part="48" refId="ref121798" refString="Dammer, U. (1907) Palmae. Verhandlungen des Botanischen Vereins der Provinz Brandenburg 48: 118 - 129." title="Palmae" type="journal article" year="1907">Dammer (1907: 121)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="B5E14D6F2432BD6BFDD9D9401FE0A418" authority="(Dammer) Burret (1930 a: 270)" authorityName="Burret" authorityYear="1930" baseAuthorityName="Dammer" box="[633,1247,1794,1817]" class="Liliopsida" family="Arecaceae" genus="Taenianthera" higherTaxonomySource="CoL" kingdom="Plantae" order="Arecales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="lagesiana">
<emphasis id="4095EAFE2432BD6BFDD9D9401846A418" box="[633,889,1794,1817]" italics="true" pageId="44" pageNumber="45">Taenianthera lagesiana</emphasis>
(Dammer)
<treatmentCitation id="F34010FD2432BD6BFC5FD9401FE0A418" author="Burret, M." box="[1023,1247,1794,1817]" page="270" pageId="44" pageNumber="45" year="1930">
<bibRefCitation id="16704B1D2432BD6BFC5FD9401FE0A418" author="Burret, M." box="[1023,1247,1794,1817]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="44" pageNumber="45" pagination="123 - 270" part="63" refId="ref121027" refString="Burret, M. (1930 a) Geonomeae americanae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 63: 123 - 270." title="Geonomeae americanae" type="journal article" year="1930">Burret (1930a: 270)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C2432BD6BFA95D9401EA6A418" box="[1333,1433,1794,1817]" name="Brazil" pageId="44" pageNumber="45">BRAZIL</collectingCountry>
. Amazonas: Rio Juruá, Juruámirim,
<date id="065F102C2432BD6BFDEFD96019E5A438" box="[591,730,1826,1849]" pageId="44" pageNumber="45" value="1901-08">August 1901</date>
,
<emphasis id="4095EAFE2432BD6BFD45D9661859A438" box="[741,870,1826,1851]" italics="true" pageId="44" pageNumber="45">E. Ule 5745</emphasis>
(
<typeStatus id="AD5A884E2432BD6BFCD3D96018EDA438" box="[883,978,1826,1849]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
not known, isotype MG!).
</treatmentCitationGroup>
</paragraph>
<paragraph id="725E36EC2432BD6AFF37D9211887A0DF" blockId="44.[151,1437,1891,2032]" lastBlockId="45.[151,1438,152,1879]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="4095EAFE2432BD6BFF37D9211BDAA47C" box="[151,229,1891,1917]" italics="true" pageId="44" pageNumber="45">Plants</emphasis>
2.0(0.84.0) m tall; stems 0.5(0.11.3) m tall, 2.3(1.72.8) cm in diameter, solitary, not cane-like; internodes 0.6(0.50.6) cm long, not scaly.
<emphasis id="4095EAFE2432BD6BFD3BD9C819D2A4A2" box="[667,749,1930,1955]" italics="true" pageId="44" pageNumber="45">Leaves</emphasis>
9(514) per stem, irregularly pinnate or regularly pinnate and the pinnae with 1 main vein only, not plicate, the bases of blades running diagonally into the rachis; sheaths 19.2(5.050.0) cm long; petioles 71.0(24.0140.0) cm long, drying green or yellowish; rachis 83.1(46.0137.0) cm long, 5.7(2.89.4) mm in diameter; veins not raised or slightly raised and triangular in cross-section adaxially; pinnae 17(238) per side of rachis; basal pinna 30.5(13.457.0) cm long, 4.3(0.3 24.0) cm wide, forming an angle of 53(2095)° with the rachis; apical pinna 19.9(6.836.0) cm long, 12.6(1.731.0) cm wide, forming an angle of 44(3160)° with the rachis.
<emphasis id="4095EAFE2433BD6AFBE3DF4E1FC6A227" box="[1091,1273,268,294]" italics="true" pageId="45" pageNumber="46">Inflorescences</emphasis>
unbranched; prophylls and peduncular bracts ribbed with elongate, unbranched fibers, both bracts tubular, narrow, elongate, closely sheathing the peduncle, more or less persistent; prophylls 11.6(4.525.5) cm long, not short and asymmetrically apiculate, the surfaces not ridged, without unequally wide ridges; peduncular bracts 20.4(12.028.5) cm long, well-developed, inserted 2.7(0.64.4) cm above the prophyll; peduncles 44.9(21.5 100.0) cm long, 3.9(2.15.8) mm in diameter; rachillae 1, 23.8(10.536.0) cm long, 5.8(2.88.3) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown or yellow-brown, without short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips pits with a central notch before anthesis, often the two sides of the notch overlapping, not recurved after anthesis, not hood-shaped; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips well-developed; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted directly onto the apiculate filament apices; anthers not short and curled at anthesis, usually elongate, spiraled and twisted or sometimes remaining straight; non-fertilized pistillate flowers deciduous after anthesis; staminodial tubes lobed at the apex, the lobes spreading at anthesis, acuminate, those of non-fertilized flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE2433BD6AFEE8DD341AB7A091" box="[328,392,886,912]" italics="true" pageId="45" pageNumber="46">fruits</emphasis>
10.7(8.713.5) mm long, 6.9(6.08.9) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, not bumpy, not apiculate; locular epidermis with operculum, smooth, without pores.
</paragraph>
</subSubSection>
<subSubSection id="3AFB65672433BD6AFF66DDA8185FA456" pageId="45" pageNumber="46" type="distribution">
<paragraph id="725E36EC2433BD6AFF66DDA81E3EA72A" blockId="45.[151,1438,152,1879]" pageId="45" pageNumber="46">
<emphasis id="4095EAFE2433BD6AFF66DDA81922A705" bold="true" box="[198,541,1002,1028]" pageId="45" pageNumber="46">Distribution and habitat:—</emphasis>
From 0°08
<geoCoordinate id="17D5502B2433BD6AFD17DDA8182CA705" box="[695,787,1002,1028]" degrees="13" direction="south" minutes="21" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="-13.35">13°21S</geoCoordinate>
and 68°46
<geoCoordinate id="17D5502B2433BD6AFC0BDDA81F28A705" box="[939,1047,1002,1028]" degrees="78" direction="west" minutes="19" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-78.316666">78°19W</geoCoordinate>
in the western Amazon region of
<collectingCountry id="0AF6767C2433BD6AFF37DA531A31A72A" box="[151,270,1041,1067]" name="Colombia" pageId="45" pageNumber="46">Colombia</collectingCountry>
,
<collectingCountry id="0AF6767C2433BD6AFEB9DA531A43A72A" box="[281,380,1041,1067]" name="Ecuador" pageId="45" pageNumber="46">Ecuador</collectingCountry>
,
<collectingCountry id="0AF6767C2433BD6AFE28DA531AFEA72A" box="[392,449,1041,1067]" name="Peru" pageId="45" pageNumber="46">Peru</collectingCountry>
, and
<collectingCountry id="0AF6767C2433BD6AFE5EDA531979A72A" box="[510,582,1041,1067]" name="Brazil" pageId="45" pageNumber="46">Brazil</collectingCountry>
at 210(87850) m elevation in lowland rainforest (
<figureCitation id="EADA2A692433BD6AFB3DDA531FCDA72A" box="[1181,1266,1041,1067]" captionStart="FIGURE 11" captionStartId="46.[151,256,1189,1212]" captionTargetBox="[167,1417,158,1169]" captionTargetId="figure@46.[151,1436,143,1184]" captionTargetPageId="46" captionText="FIGURE 11. Distribution maps of Geonoma calyptrogynoidea, G. camana, G. chlamydostachys, G. chococola subsp. chococola, and G. chococola subsp. awaensis." figureDoi="http://doi.org/10.5281/zenodo.3538384" httpUri="https://zenodo.org/record/3538384/files/figure.png" pageId="45" pageNumber="46">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="725E36EC2433BD6AFF66DA7A1ACAA612" blockId="45.[151,1438,152,1879]" pageId="45" pageNumber="46">
<emphasis id="4095EAFE2433BD6AFF66DA7A1AFFA753" bold="true" box="[198,448,1080,1106]" pageId="45" pageNumber="46">Taxonomic notes:—</emphasis>
In previous treatments (e.g.,
<bibRefCitation id="16704B1D2433BD6AFCB3DA7A1F1CA753" author="Henderson, A. &amp; Galeano, G. &amp; Bernal, R." box="[787,1059,1080,1106]" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="45" pageNumber="46" pagination="1 - 352" refId="ref122861" refString="Henderson, A., Galeano, G. &amp; Bernal, R. (1995) A field guide to the palms of the Americas. Princeton University Press, Princeton, New Jersey, pp. 1 - 352." title="A field guide to the palms of the Americas" type="book chapter" year="1995">
Henderson
<emphasis id="4095EAFE2433BD6AFC3ADA7B18E2A753" box="[922,989,1080,1106]" italics="true" pageId="45" pageNumber="46">et al.,</emphasis>
1995
</bibRefCitation>
),
<taxonomicName id="B5E14D6F2433BD6AFB99DA7A1E2FA753" authorityName="Trail" authorityYear="1876" box="[1081,1296,1080,1106]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2433BD6AFB99DA7A1E2FA753" box="[1081,1296,1080,1106]" italics="true" pageId="45" pageNumber="46">Geonoma camana</emphasis>
</taxonomicName>
was said to be characterized by its thick, smooth pinnae with conspicuous, submarginal veins. In the present work, these potential characters have been found to be inconsistent and difficult to score, and are therefore not used.
<taxonomicName id="B5E14D6F2433BD6AFF37DAEE1A43A7C7" authorityName="Trail" authorityYear="1876" box="[151,380,1196,1222]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2433BD6AFF37DAEE1A43A7C7" box="[151,380,1196,1222]" italics="true" pageId="45" pageNumber="46">Geonoma camana</emphasis>
</taxonomicName>
differs from similar species in the
<taxonomicName id="B5E14D6F2433BD6AFCE3DAEE1F2BA7C7" authorityName="Martius" authorityYear="1823" box="[835,1044,1196,1222]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="macrostachys">
<emphasis id="4095EAFE2433BD6AFCE3DAEE1F2BA7C7" box="[835,1044,1196,1222]" italics="true" pageId="45" pageNumber="46">G. macrostachys</emphasis>
</taxonomicName>
clade (
<taxonomicName id="B5E14D6F2433BD6AFBD4DAEE1E4FA7C7" authorityName="Galeano" authorityYear="1986" box="[1140,1392,1196,1222]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="chlamydostachys">
<emphasis id="4095EAFE2433BD6AFBD4DAEE1E4FA7C7" box="[1140,1392,1196,1222]" italics="true" pageId="45" pageNumber="46">G. chlamydostachys</emphasis>
</taxonomicName>
,
<emphasis id="4095EAFE2433BD6AFA22DAEE1AF3A7EC" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="B5E14D6F2433BD6AFA22DAEE1A25A7EC" authorityName="Wessels Boer" authorityYear="1968" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="chococola">G. chococola</taxonomicName>
,
<taxonomicName id="B5E14D6F2433BD6AFE88DA911AF3A7EC" authorityName="Henderson" authorityYear="2011" box="[296,460,1235,1261]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="deneversii">G. deneversii</taxonomicName>
</emphasis>
, and
<taxonomicName id="B5E14D6F2433BD6AFDB2DA9119ACA7EC" box="[530,659,1235,1261]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="maxima">
<emphasis id="4095EAFE2433BD6AFDB2DA9119ACA7EC" box="[530,659,1235,1261]" italics="true" pageId="45" pageNumber="46">G. maxima</emphasis>
</taxonomicName>
) in its fruit surfaces not splitting, and locular epidermis with operculum and without pores.
</paragraph>
<paragraph id="725E36EC2433BD6AFF66DB621AA1A6AF" blockId="45.[151,1438,152,1879]" pageId="45" pageNumber="46">
<emphasis id="4095EAFE2433BD6AFF66DB621AC5A63B" bold="true" box="[198,506,1312,1338]" pageId="45" pageNumber="46">Subspecific variation:—</emphasis>
No traits vary within this species. There is a gap in the distribution of
<taxonomicName id="B5E14D6F2433BD6AFA23DB621BCBA660" authorityName="Trail" authorityYear="1876" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2433BD6AFA23DB621BCBA660" italics="true" pageId="45" pageNumber="46">G. camana</emphasis>
</taxonomicName>
, between
<collectingCountry id="0AF6767C2433BD6AFECBDB041AF2A661" box="[363,461,1350,1376]" name="Ecuador" pageId="45" pageNumber="46">Ecuador</collectingCountry>
and adjacent
<collectingCountry id="0AF6767C2433BD6AFDCDDB0419DEA661" box="[621,737,1350,1376]" name="Colombia" pageId="45" pageNumber="46">Colombia</collectingCountry>
and
<collectingCountry id="0AF6767C2433BD6AFCB9DB041870A661" box="[793,847,1350,1376]" name="Peru" pageId="45" pageNumber="46">Peru</collectingCountry>
in the west, and
<collectingCountry id="0AF6767C2433BD6AFBB4DB041FB4A661" box="[1044,1163,1350,1376]" name="Colombia" pageId="45" pageNumber="46">Colombia</collectingCountry>
,
<collectingCountry id="0AF6767C2433BD6AFB36DB041FF0A661" box="[1174,1231,1350,1376]" name="Peru" pageId="45" pageNumber="46">Peru</collectingCountry>
, and
<collectingCountry id="0AF6767C2433BD6AFAACDB041E6BA661" box="[1292,1364,1350,1376]" name="Brazil" pageId="45" pageNumber="46">Brazil</collectingCountry>
in the east. However, the intervening area is poorly collected for palms, and this gap may be an artifact of insufficient collecting.
</paragraph>
<paragraph id="725E36EC2433BD6AFF66DBF91EA2A5BC" blockId="45.[151,1438,152,1879]" pageId="45" pageNumber="46">
There is geographical variation in this species. Regression shows there are significant associations between longitude and two plant, seven leaf, and three inflorescence variables. Squared multiple
<emphasis id="4095EAFE2433BD6AFA91DBA01E7CA6FA" box="[1329,1347,1506,1531]" italics="true" pageId="45" pageNumber="46">R</emphasis>
for the regression of plant height on longitude is 0.36, stem height 0.24, petiole length 0.40, rachis width 0.54, basal pinna length 0.31, basal pinna angle 0.12, apical pinna length 0.50, apical pinna width 0.10, apical pinna angle 0.19, peduncle width 0.25, rachilla length 0.19, and rachilla width 0.18. From east to west, specimens become smaller. In particular, there is a change in leaf size and shape, from specimens in the east having longer basal and apical pinnae with narrower angles, to those in the west shorter basal and apical pinnae with wider angles.
</paragraph>
<paragraph id="725E36EC2433BD6AFF66D88B185FA456" blockId="45.[151,1438,152,1879]" pageId="45" pageNumber="46">
Four specimens from
<collectingRegion id="B025F80E2433BD6AFE6DD88B1936A5E2" box="[461,521,1737,1763]" country="Brazil" name="Acre" pageId="45" pageNumber="46">Acre</collectingRegion>
,
<collectingCountry id="0AF6767C2433BD6AFDB6D88B1962A5E2" box="[534,605,1737,1763]" name="Brazil" pageId="45" pageNumber="46">Brazil</collectingCountry>
(
<emphasis id="4095EAFE2433BD6AFDD1D88818F5A5E2" box="[625,970,1737,1763]" italics="true" pageId="45" pageNumber="46">Ferreira 125, 136, 163, 165)</emphasis>
are much smaller than others and may represent hybrids between
<taxonomicName id="B5E14D6F2433BD6AFE72D8B21974A40B" authorityName="Trail" authorityYear="1876" box="[466,587,1776,1802]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="camana">
<emphasis id="4095EAFE2433BD6AFE72D8B21974A40B" box="[466,587,1776,1802]" italics="true" pageId="45" pageNumber="46">G. camana</emphasis>
</taxonomicName>
and
<taxonomicName id="B5E14D6F2433BD6AFD28D8B21876A40B" authorityName="Martius" authorityYear="1823" box="[648,841,1776,1802]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="macrostachys">
<emphasis id="4095EAFE2433BD6AFD28D8B21876A40B" box="[648,841,1776,1802]" italics="true" pageId="45" pageNumber="46">G. macrostachys</emphasis>
</taxonomicName>
. Other specimens from
<collectingRegion id="B025F80E2433BD6AFBC4D8B21F8AA40B" box="[1124,1205,1776,1802]" country="Peru" name="Loreto" pageId="45" pageNumber="46">Loreto</collectingRegion>
,
<collectingCountry id="0AF6767C2433BD6AFB60D8B21FC9A40B" box="[1216,1270,1776,1802]" name="Peru" pageId="45" pageNumber="46">Peru</collectingCountry>
(
<emphasis id="4095EAFE2433BD6AFAA7D8B21EADA40B" box="[1287,1426,1776,1802]" italics="true" pageId="45" pageNumber="46">Balick 1131</emphasis>
) and from Amazonas,
<collectingCountry id="0AF6767C2433BD6AFE34D9541AE3A431" box="[404,476,1814,1840]" name="Brazil" pageId="45" pageNumber="46">Brazil</collectingCountry>
(
<emphasis id="4095EAFE2433BD6AFE4DD9541955A431" box="[493,618,1814,1840]" italics="true" pageId="45" pageNumber="46">Pardini 45</emphasis>
) also appear intermediate between these two species. These potential hybrids are not included in the above description or analysis.
</paragraph>
</subSubSection>
</treatment>
</document>