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<document ID-DOI="10.5733/afin.051.0212" ID-ISSN="2305-2562" ID-Zenodo-Dep="7913777" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683579024145" checkinUser="felipe" docAuthor="Londt, Jason G. H." docDate="2010" docId="FA455017FFFEAD60FE5477A14DAAFC79" docLanguage="en" docName="AfrInvertebr.51.2.431-482.pdf" docOrigin="African Invertebrates 51 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.051.0212" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Acnephalomyia probolos Londt 2010, sp. n." docType="treatment" docVersion="3" lastPageNumber="458" masterDocId="067C286FFFE4AD7BFFED75294D5FFFC2" masterDocTitle="A review of Afrotropical Acnephalum Macquart, 1838, including the reinstatement of Sporadothrix Hermann, 1907 and descriptions of two new genera (Diptera: Asilidae: Stenopogoninae)" masterLastPageNumber="431" masterPageNumber="431" pageNumber="457" updateTime="1683633860393" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Afrotropical Acnephalum Macquart, 1838, including the reinstatement of Sporadothrix Hermann, 1907 and descriptions of two new genera (Diptera: Asilidae: Stenopogoninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Londt, Jason G. H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>431</mods:start>
<mods:end>431</mods:end>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.051.0212</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.051.0212</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7913777</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7913865" ID-Zenodo-Dep="7913865" LSID="urn:lsid:plazi:treatment:FA455017FFFEAD60FE5477A14DAAFC79" httpUri="http://treatment.plazi.org/id/FA455017FFFEAD60FE5477A14DAAFC79" lastPageId="27" lastPageNumber="458" pageId="26" pageNumber="457">
<subSubSection box="[441,806,648,675]" pageId="26" pageNumber="457" type="nomenclature">
<paragraph blockId="26.[144,1104,648,1674]" box="[441,806,648,675]" pageId="26" pageNumber="457">
<heading box="[441,806,648,675]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="457" reason="2">
<taxonomicName ID-CoL="89TV8" authority="Londt, 2010" authorityName="Londt" authorityYear="2010" box="[441,739,648,674]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="457" phylum="Arthropoda" rank="species" species="probolos" status="sp. n.">
<emphasis bold="true" box="[441,739,648,674]" pageId="26" pageNumber="457">Acnephalomyia probolos</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[746,806,649,675]" pageId="26" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="458" pageId="26" pageNumber="457" type="description">
<paragraph blockId="26.[144,1104,648,1674]" box="[540,707,688,715]" pageId="26" pageNumber="457">
<figureCitation box="[540,667,688,715]" captionStart="Figs 4451" captionStartId="46.[144,183,1495,1517]" captionTargetBox="[162,1087,331,1473]" captionTargetId="figure-9@46.[144,1104,313,1480]" captionTargetPageId="46" captionText="Figs 4451.Acnephalomyia male terminalia:(4446) A. leukoros sp. n.paratype, lateral,dorsal and ventral; (4749) A.platygaster (Loew, 1858) topotypic ♂ lateral, dorsal and ventral; (50, 51) A. probolos sp. n. paratype, lateral and ventral. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7913795" httpUri="https://zenodo.org/record/7913795/files/figure.png" pageId="26" pageNumber="457">Figs 50, 51</figureCitation>
,
<figureCitation box="[679,707,688,714]" captionStart="Fig" captionStartId="50.[144,177,852,874]" captionTargetBox="[287,981,263,828]" captionTargetId="figure-9@50.[286,982,256,834]" captionTargetPageId="50" captionText="Fig. 68. Distribution of species: Acnephalomyia iota sp. n. diamonds; A. probolos sp. n. star; Sporadothrix gracilis Hermann, 1907 circles; Ammodaimon platythrix sp. n. triangle; Astiptomyia bikos sp. n. square." figureDoi="http://doi.org/10.5281/zenodo.7913813" httpUri="https://zenodo.org/record/7913813/files/figure.png" pageId="26" pageNumber="457">68</figureCitation>
</paragraph>
<paragraph blockId="26.[144,1104,648,1674]" pageId="26" pageNumber="457">
Etymology: From Greek
<taxonomicName authorityName="Londt" authorityYear="2010" box="[433,532,728,754]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="457" phylum="Arthropoda" rank="species" species="probolos">
<emphasis box="[433,532,728,754]" italics="true" pageId="26" pageNumber="457">probolos</emphasis>
</taxonomicName>
(any protruding or jutting object or prominence); noun in apposition. The name refers to the long, projecting hypandrium.
</paragraph>
<paragraph blockId="26.[144,1104,648,1674]" pageId="26" pageNumber="457">
Description (Based on
<typeStatus box="[402,504,800,826]" pageId="26" pageNumber="457">holotype</typeStatus>
. Condition: Good; double mounted, right prothoracic leg missing terminal four tarsomeres, left mesothoracic leg broken off and glued to mounting strip.):
</paragraph>
<paragraph blockId="26.[144,1104,648,1674]" pageId="26" pageNumber="457">
<emphasis box="[144,205,904,930]" italics="true" pageId="26" pageNumber="457">Head</emphasis>
: Dark red-brown to black, white setose, weakly silver pruinose.Antenna:Dark red-brown except for tip of style which is pale yellow. Segmental ratios 1.0:0.9:3.0:0.2:1.2, scape and pedicel subequal in length, pale whitish setose, macroseta on ventral aspect of pedicel longer than postpedicel. Face dark red-brown to black, apruinose except for narrow strips adjacent to eye margins, mystax white, covering entire face. Frons, vertex and postocular region dark red-brown to black, apruinose except for central part of postocular region, mainly white setose (some dorsal setae yellowish); angle subtended by eye margins at level of frons/vertex
<emphasis box="[713,725,1129,1154]" italics="true" pageId="26" pageNumber="457">c</emphasis>
. 17°. Proboscis red-brown, white setose. Palpus dark red-brown, 2-segmented, pale yellowish setose.
</paragraph>
<paragraph blockId="26.[144,1104,648,1674]" pageId="26" pageNumber="457">
<emphasis box="[144,225,1200,1226]" italics="true" pageId="26" pageNumber="457">Thorax</emphasis>
: Dark red-brown to black, apruinose, mostly white setose. Pronotum mostly white setose, but with group of pale yellowish macrosetae medially. Mesonotum mostly white setose interspersed with some yellow and dark red-brown ones. Lateral macrosetae moderately well developed, yellow (3
<emphasis box="[658,694,1296,1322]" italics="true" pageId="26" pageNumber="457">npl</emphasis>
, 3
<emphasis box="[735,782,1296,1322]" italics="true" pageId="26" pageNumber="457">spal</emphasis>
, 5 or 6
<emphasis box="[882,918,1296,1322]" italics="true" pageId="26" pageNumber="457">pal</emphasis>
). Pleura largely asetose except for numerous, long, somewhat wavy, white katatergals, dorsally situated anepisternals and katepisternals. Scutellum shiny dark red-brown apruinose with poorly-developed transverse, subapical groove. About 10 moderately developed yellowish apical macrosetae accompanied by slightly shorter, fine, white setae; disc sparsely white setose. Legs: Mostly dark red-brown, but
<emphasis box="[786,826,1456,1482]" italics="true" pageId="26" pageNumber="457">fem</emphasis>
orange-brown ventrally. Major setae erect, whitish, minor setae white. Ventral parts of
<emphasis box="[877,910,1489,1514]" italics="true" pageId="26" pageNumber="457">tar</emphasis>
and distoventral tip of
<emphasis box="[214,244,1520,1546]" italics="true" pageId="26" pageNumber="457">tib</emphasis>
black setose. Claws black, longish (but shorter than tarsomere 5); empodia slender, yellow, about two-thirds length of claws; pulvilli tiny (about one-third length of empodium). Haltere yellow, base yellow-brown. Wing: 3.7×
<quantity box="[855,944,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="457" unit="mm" value="1.5">1.5 mm</quantity>
. Veins brown, membrane lacking microtrichiae; entirely transparent, unstained. Vein R
<subScript attach="left" box="[959,967,1634,1648]" fontSize="6" pageId="26" pageNumber="457">4</subScript>
with only a trace of a basal stump-vein.
</paragraph>
<paragraph blockId="27.[181,1142,204,745]" pageId="27" pageNumber="458">
<emphasis box="[181,287,204,230]" italics="true" pageId="27" pageNumber="458">Abdomen</emphasis>
: Terga broader than long, dark red-brown to black. Terga apruinose, but entirely pitted by setal sockets. Terga with longish white setae laterally. White setae extend for a short distance along distal margins of terga, becoming shorter medially. Large areas of terga appear asetose, but are covered with tiny reddish brown setae. Sterna dark red-brown, apruinose, longish white setose (wavy anteriorly). Terminalia largely withdrawn between T6 and S6, distal parts somewhat obscured by setae. Genitalia (
<figureCitation box="[248,386,395,422]" captionStart="Figs 4451" captionStartId="46.[144,183,1495,1517]" captionTargetBox="[162,1087,331,1473]" captionTargetId="figure-9@46.[144,1104,313,1480]" captionTargetPageId="46" captionText="Figs 4451.Acnephalomyia male terminalia:(4446) A. leukoros sp. n.paratype, lateral,dorsal and ventral; (4749) A.platygaster (Loew, 1858) topotypic ♂ lateral, dorsal and ventral; (50, 51) A. probolos sp. n. paratype, lateral and ventral. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7913795" httpUri="https://zenodo.org/record/7913795/files/figure.png" pageId="27" pageNumber="458">Figs 50, 51</figureCitation>
):
<emphasis box="[410,483,396,422]" italics="true" pageId="27" pageNumber="458">Epand</emphasis>
highly reduced.
<emphasis box="[683,736,396,421]" italics="true" pageId="27" pageNumber="458">Proc</emphasis>
in lateral view longer than
<emphasis box="[1066,1134,396,422]" italics="true" pageId="27" pageNumber="458">epand</emphasis>
. Exterior lobe of
<emphasis box="[370,437,428,453]" italics="true" pageId="27" pageNumber="458">goncx</emphasis>
tapering to a fairly acute tip; interior lobe longish, finger­like and jutting out to about same level as achieved by aedeagal tip.
<emphasis box="[902,969,460,486]" italics="true" pageId="27" pageNumber="458">Gonst</emphasis>
small.
<emphasis box="[1052,1113,460,486]" italics="true" pageId="27" pageNumber="458">Hypd</emphasis>
in ventral view somewhat truncate basally, tapering rapidly to long finger­like, distally projecting medial lobe.
<emphasis box="[445,506,524,550]" italics="true" pageId="27" pageNumber="458">Hypd</emphasis>
basodorsally with a disc-like projection that lies between gonocoxites.
</paragraph>
<paragraph blockId="27.[181,1142,204,745]" pageId="27" pageNumber="458">
Variation: The female
<typeStatus box="[440,536,598,624]" pageId="27" pageNumber="458">paratype</typeStatus>
agrees well with the
<typeStatus box="[788,889,597,623]" pageId="27" pageNumber="458">holotype</typeStatus>
, but is slightly bigger (wing length
<quantity box="[331,418,629,655]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="27" pageNumber="458" unit="mm" value="4.7">4.7 mm</quantity>
) and stump-veins are moderately well developed.
</paragraph>
<paragraph blockId="27.[181,1142,204,745]" pageId="27" pageNumber="458">
<materialsCitation collectingDate="1983-09-02" collectionCode="NMSA" collectorName="Londt &amp; Stuckenberg" location="Ochta Mine" pageId="27" pageNumber="458" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus box="[181,268,667,689]" pageId="27" pageNumber="458">Holotype</typeStatus>
:
<specimenCount box="[275,292,668,689]" pageId="27" pageNumber="458" type="male"></specimenCount>
SOUTH ARICA: Sth Africa
<collectingRegion box="[559,605,667,688]" country="South Africa" name="Western Cape" pageId="27" pageNumber="458">Cape</collectingRegion>
Prov / Richtersveld 2816BD /
<quantity box="[874,925,667,689]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="458" unit="km" value="40.0">40km</quantity>
S of
<location LSID="urn:lsid:plazi:treatment:FA455017FFFEAD60FE5477A14DAAFC79:7733B7DAFFFFAD60FC2577B2496EFD73" box="[968,1073,667,689]" name="Ochta Mine" pageId="27" pageNumber="458" stateProvince="Western Cape">Ochta Mine</location>
/
<collectorName box="[1087,1141,667,689]" pageId="27" pageNumber="458">Londt</collectorName>
&amp;
<collectorName box="[205,318,691,713]" pageId="27" pageNumber="458">Stuckenberg</collectorName>
/
<date box="[335,424,691,713]" pageId="27" pageNumber="458" value="1983-09-02">
<collectingDate box="[335,424,691,713]" pageId="27" pageNumber="458" value="1983-09-02">2.ix.1983</collectingDate>
</date>
. Mixed Karoo / bush with few flowers (
<collectionCode box="[790,861,691,712]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="27" pageNumber="458" type="Museum">NMSA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="27.[181,1142,204,745]" box="[181,599,723,745]" pageId="27" pageNumber="458">
<materialsCitation box="[181,595,723,745]" collectingDate="1983-09-02" collectionCode="NMSA" collectorName="Londt &amp; Stuckenberg" location="Ochta Mine" pageId="27" pageNumber="458" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus box="[181,264,724,745]" pageId="27" pageNumber="458">Paratype</typeStatus>
:
<specimenCount box="[273,301,723,744]" pageId="27" pageNumber="458" type="female">1♀</specimenCount>
same data as holotype (
<collectionCode box="[518,589,723,744]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="27" pageNumber="458" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="27.[181,1141,761,955]" pageId="27" pageNumber="458">
Distribution and phenology (Table 3): Known only from the
<typeStatus box="[853,901,761,787]" pageId="27" pageNumber="458">type</typeStatus>
locality (
<figureCitation box="[1006,1087,761,787]" captionStart="Fig" captionStartId="50.[144,177,852,874]" captionTargetBox="[287,981,263,828]" captionTargetId="figure-9@50.[286,982,256,834]" captionTargetPageId="50" captionText="Fig. 68. Distribution of species: Acnephalomyia iota sp. n. diamonds; A. probolos sp. n. star; Sporadothrix gracilis Hermann, 1907 circles; Ammodaimon platythrix sp. n. triangle; Astiptomyia bikos sp. n. square." figureDoi="http://doi.org/10.5281/zenodo.7913813" httpUri="https://zenodo.org/record/7913813/files/figure.png" pageId="27" pageNumber="458">Fig. 68</figureCitation>
) and collected only in September.
</paragraph>
<paragraph blockId="27.[181,1141,761,955]" pageId="27" pageNumber="458">
Similar species: Male genitalia are similar to those of
<taxonomicName authorityName="Londt" authorityYear="2010" baseAuthorityName="Macquart" baseAuthorityYear="1838" box="[794,884,833,859]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="458" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis box="[794,884,833,859]" italics="true" pageId="27" pageNumber="458">dorsalis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Londt" authorityYear="2010" box="[899,976,834,859]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="458" phylum="Arthropoda" rank="species" species="eremia">
<emphasis box="[899,976,834,859]" italics="true" pageId="27" pageNumber="458">eremia</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Londt" authorityYear="2010" box="[991,1086,833,859]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="458" phylum="Arthropoda" rank="species" species="leukoros">
<emphasis box="[991,1086,833,859]" italics="true" pageId="27" pageNumber="458">leukoros</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName=": Loew" authorityYear="1858" box="[181,307,865,891]" class="Insecta" family="Asilidae" genus="Acnephalum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="458" phylum="Arthropoda" rank="species" species="platygaster">
<emphasis box="[181,307,865,891]" italics="true" pageId="27" pageNumber="458">platygaster</emphasis>
</taxonomicName>
, but the form of the epandrium and hypandrium is distinctive. Identification of females of these species is sometimes problematic as they so closely resemble each other.
</paragraph>
</subSubSection>
</treatment>
</document>