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<document id="CA9B3FF73F771FC1EDDE70059F5BFD03" ID-DOI="10.5281/zenodo.186449" ID-GBIF-Dataset="eaadaeff-41cd-4d7e-bb99-45c9d56ee9f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="186449" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051969226" checkinUser="plazi" docAuthor="Lapolla, John S." docDate="2009" docId="FA3BA029FFA4FF959BA9FF584AD0FE31" docLanguage="en" docName="zt02046p025.pdf" docOrigin="Zootaxa 2046" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Euprenolepis procera Emery 1900" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="0602D851FFA8FF859B3EFFCF4860FFAB" masterDocTitle="Taxonomic Revision of the Southeast Asian Ant Genus Euprenolepis" masterLastPageNumber="25" masterPageNumber="1" pageNumber="13" updateTime="1698579287766" updateUser="plazi">
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<mods:title id="11A3C994F62CE966C5DAE6B115BE782F">Taxonomic Revision of the Southeast Asian Ant Genus Euprenolepis</mods:title>
</mods:titleInfo>
<mods:name id="B38415A9BFBFCA2CC0CD4ABA78D05007" type="personal">
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<mods:namePart id="C414828743C77DA1E2A64076975A4DF4">Lapolla, John S.</mods:namePart>
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<mods:date id="0A16917656A51D872FBA19B8F84B3A40">2009</mods:date>
<mods:detail id="5D5D6BA6334E3FB2CFA02FA0183ED50C" type="volume">
<mods:number id="569B88F970127AC7CC8A09753761C9B4">2046</mods:number>
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<mods:classification id="A94BD8722663F3B660434427D44392EA">journal article</mods:classification>
<mods:identifier id="121A25309C5BDA55EF9C4E2719A724BE" type="DOI">10.5281/zenodo.186449</mods:identifier>
<mods:identifier id="7B6151F0A73F9C10DCE52E0655BEDC34" type="GBIF-Dataset">eaadaeff-41cd-4d7e-bb99-45c9d56ee9f2</mods:identifier>
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<mods:identifier id="F0E4F9DD884A4DE392B05FC785F315F7" type="Zenodo-Dep">186449</mods:identifier>
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<treatment id="FA3BA029FFA4FF959BA9FF584AD0FE31" ID-DOI="http://doi.org/10.5281/zenodo.6214322" ID-GBIF-Taxon="119612246" ID-Zenodo-Dep="6214322" LSID="urn:lsid:plazi:treatment:FA3BA029FFA4FF959BA9FF584AD0FE31" httpUri="http://treatment.plazi.org/id/FA3BA029FFA4FF959BA9FF584AD0FE31" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
<subSubSection id="3A8842B4FFA4FF899BA9FF584BE5FF7F" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="722D113FFFA4FF899BA9FF584A33FF1A" blockId="12.[151,901,151,212]" box="[151,595,151,177]" pageId="12" pageNumber="13">
<heading id="2965A653FFA4FF899BA9FF584A33FF1A" bold="true" box="[151,595,151,177]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="B5926ABCFFA4FF899BA9FF584A33FF1A" ID-CoL="3CX6D" authority="Emery, 1900" authorityName="Emery" authorityYear="1900" box="[151,595,151,177]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFA4FF899BA9FF584A33FF1A" bold="true" box="[151,595,151,177]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BA9FF5849FCFF1A" bold="true" box="[151,412,151,177]" italics="true" pageId="12" pageNumber="13">Euprenolepis procera</emphasis>
(
<bibRefCitation id="16036CCEFFA4FF899A95FF584A2AFF1A" author="Emery" box="[427,586,151,177]" pageId="12" pageNumber="25" refString="Emery, C. (1900) Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. [part]. Annali del Museo Civico di Storia Naturale di Genova, 40 [= (2 (20)], 689 - 720." type="journal article" year="1900">Emery, 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="722D113FFFA4FF899BA9FF754BE5FF7F" blockId="12.[151,901,151,212]" box="[151,901,186,212]" pageId="12" pageNumber="13">
(
<figureCitation id="EAA90DBAFFA4FF899BA1FF75496DFF7F" box="[159,269,186,212]" captionStart="FIGURE 7" captionStartId="13.[151,257,1964,1988]" captionTargetBox="[172,1415,184,1940]" captionTargetId="figure@13.[172,1416,184,1940]" captionTargetPageId="13" captionText="FIGURE 7 A-D. E. procera major worker. A) lateral view; B) gaster in dorsal view; C) head in full-frontal view; D) scape. Malaysia, Sarawak (BMNH)" httpUri="https://zenodo.org/record/186456/files/figure.png" pageId="12" pageNumber="13">Figures 7</figureCitation>
A–D [major]; 8A–B [minor]; 9A–I [queen and male])
</paragraph>
</subSubSection>
<subSubSection id="3A8842B4FFA4FF899BA9FEC74C74FE74" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="722D113FFFA4FF899BA9FEC74BEDFED4" blockId="12.[151,1436,264,479]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="52823611FFA4FF899BA9FEC74BEDFED4" pageId="12" pageNumber="13">
<taxonomicName id="B5926ABCFFA4FF899BA9FEC74A48FEB4" authority="Emery, 1900: 699" authorityName="Emery" authorityPageNumber="699" authorityYear="1900" box="[151,552,264,287]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFA4FF899BA9FEC74903FEB4" box="[151,355,264,287]" italics="true" pageId="12" pageNumber="13">Prenolepis procera</emphasis>
<treatmentCitation id="F333372EFFA4FF899A54FEC74A48FEB4" author="Emery" box="[362,552,264,287]" page="699" pageId="12" pageNumber="13" year="1900">
<bibRefCitation id="16036CCEFFA4FF899A54FEC74995FEB4" author="Emery" box="[362,501,264,287]" pageId="12" pageNumber="25" refString="Emery, C. (1900) Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. [part]. Annali del Museo Civico di Storia Naturale di Genova, 40 [= (2 (20)], 689 - 720." type="journal article" year="1900">Emery, 1900</bibRefCitation>
: 699
</treatmentCitation>
</taxonomicName>
(worker – major and minor).
<typeStatus id="AD29AF9DFFA4FF899858FEC74BDDFEB4" box="[870,957,264,287]" pageId="12" pageNumber="13" type="syntype">Syntype</typeStatus>
workers,
<collectingCountry id="0A8551AFFFA4FF899F19FEC74CDDFEB4" box="[1063,1213,264,287]" name="Indonesia" pageId="12" pageNumber="13">INDONESIA</collectingCountry>
: Sumatra, Si Rembé, Panherang Pisang. (
<typeStatus id="AD29AF9DFFA4FF899A9EFEE749AFFE94" box="[416,463,296,319]" pageId="12" pageNumber="13">type</typeStatus>
depository unknown, not in MCSN) [not examined].
<treatmentCitation id="F333372EFFA4FF899F1AFEE74C8AFE94" author="Emery" box="[1060,1258,296,319]" page="134" pageId="12" pageNumber="13" year="1906">
<bibRefCitation id="16036CCEFFA4FF899F1AFEE74CD3FE94" author="Emery" box="[1060,1203,296,319]" pageId="12" pageNumber="25" refString="Emery, C. (1906) Note sur Prenolepis vividula Nyl. et sur la classification des especes du genre Prenolepis. Annales de la Societe Entomologique de Belgique, 50, 130 - 134." type="journal article" year="1906">Emery, 1906</bibRefCitation>
: 134
</treatmentCitation>
, combination in
<taxonomicName id="B5926ABCFFA4FF899BF9FE874986FEF4" ID-CoL="6VK5" authority="(Euprenolepis)" baseAuthorityName="Euprenolepis" box="[199,486,328,351]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFA4FF899BF9FE87495AFEF4" box="[199,314,328,351]" italics="true" pageId="12" pageNumber="13">Prenolepis</emphasis>
(
<emphasis id="40E6CD2DFFA4FF899A73FE8749BAFEF4" box="[333,474,328,351]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
)
</taxonomicName>
;
<treatmentCitation id="F333372EFFA4FF899AC8FE874AA1FEF4" author="Emery" box="[502,705,328,351]" page="224" pageId="12" pageNumber="13" year="1925">
<bibRefCitation id="16036CCEFFA4FF899AC8FE874AE7FEF4" author="Emery" box="[502,647,328,351]" pageId="12" pageNumber="25" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery, 1925</bibRefCitation>
: 224
</treatmentCitation>
, combination in
<taxonomicName id="B5926ABCFFA4FF8998BEFE874CDAFEF4" authority="(Euprenolepis)" baseAuthorityName="Euprenolepis" box="[896,1210,328,351]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Paratrechina">
<emphasis id="40E6CD2DFFA4FF8998BEFE874C6DFEF4" box="[896,1037,328,351]" italics="true" pageId="12" pageNumber="13">Paratrechina</emphasis>
(
<emphasis id="40E6CD2DFFA4FF899F1CFE874CCFFEF4" box="[1058,1199,328,351]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
)
</taxonomicName>
;
<treatmentCitation id="F333372EFFA4FF899FF5FE874DECFEF4" author="Bolton" box="[1227,1420,328,351]" page="189" pageId="12" pageNumber="13" year="1995">
<bibRefCitation id="16036CCEFFA4FF899FF5FE874D3EFEF4" author="Bolton" box="[1227,1374,328,351]" pageId="12" pageNumber="24" refString="Bolton, B. (1995) A new general catalogue of the ants of the world. Harvard University Press, Cambridge, Massachusetts. 504 pp." type="book" year="1995">Bolton, 1995</bibRefCitation>
:189
</treatmentCitation>
, combination in
<taxonomicName id="B5926ABCFFA4FF899A50FEA7499BFED4" ID-CoL="4GFF" box="[366,507,360,383]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFA4FF899A50FEA7499BFED4" box="[366,507,360,383]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
</taxonomicName>
;
<bibRefCitation id="16036CCEFFA4FF899937FEA74AB3FED4" author="Bolton" box="[521,723,360,383]" pageId="12" pageNumber="24" refString="Bolton, B., Alpert, G., Ward, P. S. &amp; Naskrecki, P. (2006) Bolton's catalogue of ants of the world: 1758 - 2005 [CD-ROM]. Havard University Press, Cambridge, Massachusetts." type="book chapter" year="2006">
Bolton
<emphasis id="40E6CD2DFFA4FF899967FEA74AE7FED4" box="[601,647,360,383]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2006
</bibRefCitation>
, in
<taxonomicName id="B5926ABCFFA4FF8999C7FEA74BE6FED4" ID-CoL="4GFF" box="[761,902,360,383]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFA4FF8999C7FEA74BE6FED4" box="[761,902,360,383]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="722D113FFFA4FF899BA9FE474C74FE74" blockId="12.[151,1436,264,479]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="52823611FFA4FF899BA9FE474C74FE74" pageId="12" pageNumber="13">
<taxonomicName id="B5926ABCFFA4FF899BA9FE474B5FFE34" authority="Forel, 1913: 130" authorityName="Forel" authorityPageNumber="130" authorityYear="1913" box="[151,831,392,415]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="antespectans" subGenus="Myrmocsphincta">
<emphasis id="40E6CD2DFFA4FF899BA9FE47497EFE34" box="[151,286,392,415]" italics="true" pageId="12" pageNumber="13">Camponotus</emphasis>
(
<emphasis id="40E6CD2DFFA4FF899A0EFE474983FE34" box="[304,483,392,415]" italics="true" pageId="12" pageNumber="13">Myrmocsphincta</emphasis>
)
<emphasis id="40E6CD2DFFA4FF899ACBFE474A1FFE34" box="[501,639,392,415]" italics="true" pageId="12" pageNumber="13">antespectans</emphasis>
<treatmentCitation id="F333372EFFA4FF8999B6FE474B5FFE34" author="Forel" box="[648,831,392,415]" page="130" pageId="12" pageNumber="13" year="1913">
<bibRefCitation id="16036CCEFFA4FF8999B6FE474B6AFE34" author="Forel" box="[648,778,392,415]" pageId="12" pageNumber="25" refString="Forel, A. (1913) Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 36, 1 - 148." type="journal article" year="1913">Forel, 1913</bibRefCitation>
: 130
</treatmentCitation>
</taxonomicName>
(worker).
<typeStatus id="AD29AF9DFFA4FF89988BFE474C78FE34" box="[949,1048,392,415]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="0A8551AFFFA4FF899F43FE474D73FE34" box="[1149,1299,392,415]" name="Indonesia" pageId="12" pageNumber="13">INDONESIA</collectingCountry>
: Sumatra (v. Buttel) (MNHB) [examined].
<treatmentCitation id="F333372EFFA4FF899931FE674AB6FE14" author="Emery" box="[527,726,424,447]" page="223" pageId="12" pageNumber="13" year="1925">
<bibRefCitation id="16036CCEFFA4FF899931FE674AFEFE14" author="Emery" box="[527,670,424,447]" pageId="12" pageNumber="25" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery, 1925</bibRefCitation>
: 223
</treatmentCitation>
, combination in
<taxonomicName id="B5926ABCFFA4FF8998AFFE674CA9FE14" authority="(Euprenolepis)" baseAuthorityName="Euprenolepis" box="[913,1225,424,447]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subTribe" subTribe="Paratrechina">
<emphasis id="40E6CD2DFFA4FF8998AFFE674C7EFE14" box="[913,1054,424,447]" italics="true" pageId="12" pageNumber="13">Paratrechina</emphasis>
(
<emphasis id="40E6CD2DFFA4FF899F0FFE674CDEFE14" box="[1073,1214,424,447]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
)
</taxonomicName>
;
<treatmentCitation id="F333372EFFA4FF899FE6FE674DF7FE14" author="Bolton" box="[1240,1431,424,447]" page="189" pageId="12" pageNumber="13" year="1995">
<bibRefCitation id="16036CCEFFA4FF899FE6FE674D0AFE14" author="Bolton" box="[1240,1386,424,447]" pageId="12" pageNumber="24" refString="Bolton, B. (1995) A new general catalogue of the ants of the world. Harvard University Press, Cambridge, Massachusetts. 504 pp." type="book" year="1995">Bolton, 1995</bibRefCitation>
:189
</treatmentCitation>
, combination in
<taxonomicName id="B5926ABCFFA4FF899A50FE07499BFE74" ID-CoL="4GFF" box="[366,507,456,479]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFA4FF899A50FE07499BFE74" box="[366,507,456,479]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
</taxonomicName>
;
<bibRefCitation id="16036CCEFFA4FF899937FE074AB3FE74" author="Bolton" box="[521,723,456,479]" pageId="12" pageNumber="24" refString="Bolton, B., Alpert, G., Ward, P. S. &amp; Naskrecki, P. (2006) Bolton's catalogue of ants of the world: 1758 - 2005 [CD-ROM]. Havard University Press, Cambridge, Massachusetts." type="book chapter" year="2006">
Bolton
<emphasis id="40E6CD2DFFA4FF899967FE074AE7FE74" box="[601,647,456,479]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
., 2006
</bibRefCitation>
, in
<taxonomicName id="B5926ABCFFA4FF8999C7FE074BE6FE74" ID-CoL="4GFF" box="[761,902,456,479]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFA4FF8999C7FE074BE6FE74" box="[761,902,456,479]" italics="true" pageId="12" pageNumber="13">Euprenolepis</emphasis>
</taxonomicName>
.
<emphasis id="40E6CD2DFFA4FF8998AAFE084C74FE74" bold="true" box="[916,1044,455,479]" pageId="12" pageNumber="13">SYN. NOV.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3A8842B4FFA4FF8B9BA9FDC14B08F95C" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="722D113FFFA4FF899BA9FDC14B37FDE4" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BA9FDC1489EFD83" bold="true" box="[151,254,526,552]" pageId="12" pageNumber="13">Worker.</emphasis>
Measurements:
<emphasis id="40E6CD2DFFA4FF899AFAFDC04A7AFD83" box="[452,538,527,552]" italics="true" pageId="12" pageNumber="13">Majors</emphasis>
(n=4): TL: 5.08–6.25; HW: 1.35–1.61; HL: 1.43–1.62; SL: 1.8–2.03; WL: 1.9–2.22; GL: 1.75–2.41.
<emphasis id="40E6CD2DFFA4FF899AF9FDFA4A7CFDE5" box="[455,540,565,590]" italics="true" pageId="12" pageNumber="13">Indices</emphasis>
: CI: 94–101; SI: 126–139.
</paragraph>
<paragraph id="722D113FFFA4FF899BFBFD934AEDFD37" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BFBFD93497FFDDE" box="[197,287,604,629]" italics="true" pageId="12" pageNumber="13">Minors</emphasis>
(n=7): TL: 3.53–4.39; HW: 0.82–1.05; HL: 0.94–1.14; SL: 1.4–1.67; WL: 1.49–1.67; GL: 1.1–1.57.
<emphasis id="40E6CD2DFFA4FF899A32FD4D4901FD30" box="[268,353,642,667]" italics="true" pageId="12" pageNumber="13">Indices</emphasis>
: CI: 87–95; SI: 153–178.
</paragraph>
<paragraph id="722D113FFFA4FF899BFBFD664B78FBED" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BFBFD664976FD69" box="[197,278,681,706]" italics="true" pageId="12" pageNumber="13">Major.</emphasis>
Head dark reddish-brown, roughly heart-shaped; head distinctly broader than long. Antennal funiculi slightly lighter in color. Head with scattered erect setae and a sparse layer of pubescence; cuticular surface strongly rugoreticulate. Posterior margin strongly emarginate medially. Scapes surpass posterior margin by about length of first three funicular segments; scapes with erect setae and sparse pubescence. Anterior clypeal margin strongly emarginate. Mesosoma dark reddish-brown, legs becoming lighter distally; cuticular surface rugoreticulate. Mesosomal dorsum with scattered erect setae of varying lengths on the posterior pronotum and mesonotum; sparse pubescence scattered across mesosoma, with denser patches on the anterior pronotum, metanotal area and propodeum. Pronotum rises at about 45° toward mesonotum; mesonotum and mesonotal area flat; propodeum dorsum rounded, dome-like; declivity steep. Petiole triangular with posterior face longer than anterior face; gaster dark reddish-brown, covered in erect setae with a layer of pubescence; cuticular surface rugoreticulate.
</paragraph>
<paragraph id="722D113FFFA4FF899BFBFB9C488EFB11" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BFBFB9C4974FBC7" box="[197,276,1107,1132]" italics="true" pageId="12" pageNumber="13">Minor.</emphasis>
As in major, with modifications expected for caste. Scapes long, surpassing posterior margin by about length of the first four funicular segments. The legs of minor workers are slightly lighter than found in majors.
</paragraph>
<paragraph id="722D113FFFA4FF899BF8FB0948AAFAAC" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BF8FB09497FFB4B" bold="true" box="[198,287,1222,1248]" pageId="12" pageNumber="13">Queen.</emphasis>
Measurements (n=1): TL: 6.3; HW: 1.1; HL: 1.0; SL: 1.6; WL: 2.5; GL: 2.8.
<emphasis id="40E6CD2DFFA4FF899F91FB084D63FB4B" box="[1199,1283,1223,1248]" italics="true" pageId="12" pageNumber="13">Indices</emphasis>
: CI: 110; SI: 145.
</paragraph>
<paragraph id="722D113FFFA4FF899BF8FADB4BFCFA09" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">As in worker, with modifications expected for caste and the following differences: Overall color similar to worker castes, but mottled dark brown, with lighter brown patches especially on head and mesosoma. Entire body covered in a dense layer of pubescence; short erect setae cover head, mesosoma notum, legs and gaster. Erect setae most abundant on gaster, becoming denser posteriorly.</paragraph>
<paragraph id="722D113FFFA4FF899BF8FA614B37FA44" blockId="12.[151,1437,526,2022]" pageId="12" pageNumber="13">
<emphasis id="40E6CD2DFFA4FF899BF8FA61496DFA63" bold="true" box="[198,269,1454,1480]" pageId="12" pageNumber="13">Male.</emphasis>
Measurements (n=2): TL: 4.54–4.85; HW: 0.70–0.75; HL: 0.86–0.89; SL: 1.54–1.63; WL: 1.77–1.81; GL: 1.88–2.18.
<emphasis id="40E6CD2DFFA4FF899AE8FA1A4A4AFA45" box="[470,554,1493,1518]" italics="true" pageId="12" pageNumber="13">Indices</emphasis>
: CI: 81–84; SI: 218–219.
</paragraph>
<paragraph id="722D113FFFA4FF8B9BFBFA334B08F95C" blockId="12.[151,1437,526,2022]" lastBlockId="14.[151,1437,1719,2018]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">Overall color brown to brownish-yellow, lighter brown on legs and funiculi; head longer than broad. Cuticular surface shiny, covered in a dense layer of pubescence. Scapes long, surpassing posterior margin by about length of first four funicular segments; scapes with scattered erect setae and a dense layer of pubescence. Clypeus broad, with anterior margin emarginated. Mandible with a basal angle and a distinct apical tooth that curves in toward body midline. Mesosoma enlarged to accommodate flight muscles; covered in a layer of pubescence, with erect setae dorsally and on legs. Pronotum small and distinctly collar-like; mesonotum offset from pronotum at sulcus; mesonotum rises sharply above height of pronotum, giving pronotum a shelf-like appearance in lateral view. Mesonotum flat dorsally with scattered erect setae; propodeum indistinct from remainder of mesosoma, with declivity shallow, gently sloping toward petiole. Petiole triangular, upright, with posterior face only slightly longer than anterior face. Gaster with a dense layer of pubescence and scattered erect setae, becoming denser posteriorly toward terminalia; terminalia especially setose. Parameres roughly triangular, turning slightly mesad posteriorly; long setae extend off of parameres. Cuspi small and tubular, reaching digiti dorsally; digiti anvil-shaped, directed ventrally; digiti visible in lateral view. Volsellar lobes with large bulbous extensions; appressed setae cover surface. Penis valves with apodeme projecting dorsally; in lateral view the valves extend dorsally.</paragraph>
</subSubSection>
<caption id="26ED41B7FFA5FF889BA9F8634A72F84D" httpUri="https://zenodo.org/record/186456/files/figure.png" pageId="13" pageNumber="14" targetBox="[172,1415,184,1940]" targetPageId="13">
<paragraph id="722D113FFFA5FF889BA9F8634A72F84D" blockId="13.[151,1436,1964,2022]" pageId="13" pageNumber="14">
<emphasis id="40E6CD2DFFA5FF889BA9F863492DF86F" bold="true" box="[151,333,1964,1988]" pageId="13" pageNumber="14">FIGURE 7A-D.</emphasis>
<taxonomicName id="B5926ABCFFA5FF889A6AF86349AAF868" box="[340,458,1964,1987]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFA5FF889A6AF86349AAF868" box="[340,458,1964,1987]" italics="true" pageId="13" pageNumber="14">E. procera</emphasis>
</taxonomicName>
major worker. A) lateral view; B) gaster in dorsal view; C) head in full-frontal view; D) scape. Malaysia, Sarawak (BMNH)
</paragraph>
</caption>
<caption id="26ED41B7FFA6FF8B9BA9F9BE4D64F923" box="[151,1284,1649,1673]" httpUri="https://zenodo.org/record/186457/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1436,194,1625]" targetPageId="14">
<paragraph id="722D113FFFA6FF8B9BA9F9BE4D64F923" blockId="14.[151,1284,1649,1673]" box="[151,1284,1649,1673]" pageId="14" pageNumber="15">
<emphasis id="40E6CD2DFFA6FF8B9BA9F9BE492DF922" bold="true" box="[151,333,1649,1673]" pageId="14" pageNumber="15">FIGURE 8A–D.</emphasis>
<taxonomicName id="B5926ABCFFA6FF8B9A6AF9BD49A4F922" box="[340,452,1650,1673]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFA6FF8B9A6AF9BD49A4F922" box="[340,452,1650,1673]" italics="true" pageId="14" pageNumber="15">E. procera</emphasis>
</taxonomicName>
minor worker. A) lateral view; B) head in full-frontal view. Malaysia (SYPC)
</paragraph>
</caption>
<subSubSection id="3A8842B4FFA6FF8B9BFBF8CB4B82F849" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="722D113FFFA6FF8B9BFBF8CB4B82F849" blockId="14.[151,1437,1719,2018]" pageId="14" pageNumber="15">
<emphasis id="40E6CD2DFFA6FF8B9BFBF8CB4A27F8B5" bold="true" box="[197,583,1796,1822]" pageId="14" pageNumber="15">Additional material examined:</emphasis>
<collectingCountry id="0A8551AFFFA6FF8B9971F8CB4A94F8B5" box="[591,756,1796,1822]" name="Indonesia" pageId="14" pageNumber="15">INDONESIA</collectingCountry>
: South Sumatra, Lampung, Wai Lima
<date id="062C37FFFFA6FF8B9F87F8CB4D54F8B5" box="[1209,1332,1796,1822]" pageId="14" pageNumber="15" value="1921-11-12">12.xi.1921</date>
(Karny); Sumatra, Prov. Lampung, Sumberaja,
<geoCoordinate id="17A677F8FFA6FF8B9945F8E44B7EF8EE" box="[635,798,1835,1861]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="-4.991667">4° 59’ 30” S</geoCoordinate>
,
<geoCoordinate id="17A677F8FFA6FF8B9813F8E44B90F8EE" box="[813,1008,1835,1861]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="104.525">104° 31’ 30” E</geoCoordinate>
,
<quantity id="B56ABCDAFFA6FF8B98C1F8E44C31F8EE" box="[1023,1105,1835,1861]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="14" pageNumber="15" unit="m" value="735.0">735 m</quantity>
,
<date id="062C37FFFFA6FF8B9F5EF8E44D66F8EE" box="[1120,1286,1835,1861]" pageId="14" pageNumber="15" value="2004-01-15" valueMax="2004-01-30" valueMin="2004-01-15">15–30.i.2004</date>
(F. Susilo);
<collectingCountry id="0A8551AFFFA6FF8B9BA9F89B4954F8C5" box="[151,308,1876,1902]" name="Malaysia" pageId="14" pageNumber="15">MALAYSIA</collectingCountry>
: Kinabalu Nat. Park, Poring Hot Spring, East Ridge, 11.07.–
<date id="062C37FFFFA6FF8B98C8F89B4C1EF8C5" box="[1014,1150,1876,1902]" pageId="14" pageNumber="15" value="1995-07-20">20.07.1995</date>
(C. Brühl);
<collectingCountry id="0A8551AFFFA6FF8B9E33F89B4D15F8C5" box="[1293,1397,1876,1902]" name="Malaysia" pageId="14" pageNumber="15">Sarawak</collectingCountry>
, 4th Div. G. Mulu Nat. Park,
<date id="062C37FFFFA6FF8B9A87F8B44A4EF83E" box="[441,558,1915,1941]" pageId="14" pageNumber="15" value="1977-10-14">14.x.1977</date>
(B. Bolton); Neg. Sembilian, Pasoh Forest Reserve,
<date id="062C37FFFFA6FF8B9F9CF8B44C9BF83E" box="[1186,1275,1915,1941]" pageId="14" pageNumber="15" value="1994-11">xi.1994</date>
(M. Brendell et al.); Pinang-Pinang, Ulu Gadut, Padang,
<date id="062C37FFFFA6FF8B99A4F86D4B70F817" box="[666,784,1954,1980]" pageId="14" pageNumber="15" value="2004-02-11">11.ii.2004</date>
(S. Yamane); Perak, Trolac Forest Res.,
<date id="062C37FFFFA6FF8B9FC9F86D4D0FF817" box="[1271,1391,1954,1980]" pageId="14" pageNumber="15" value="1967-07-06">6.vii.1967</date>
(R. Crozier); Selangor, Ulu Gombak For. Reserve,
<date id="062C37FFFFA6FF8B9981F8074B24F849" box="[703,836,1992,2018]" pageId="14" pageNumber="15" value="1967-07-24">24.vii.1967</date>
(R. Crozier).
</paragraph>
</subSubSection>
<caption id="26ED41B7FFA7FF8A9BA9F85A4C7DF859" httpUri="https://zenodo.org/record/186458/files/figure.png" pageId="15" pageNumber="16" targetBox="[172,1415,182,1917]" targetPageId="15">
<paragraph id="722D113FFFA7FF8A9BA9F85A4C7DF859" blockId="15.[151,1436,1941,2034]" pageId="15" pageNumber="16">
<emphasis id="40E6CD2DFFA7FF8A9BA9F85A4928F806" bold="true" box="[151,328,1941,1965]" pageId="15" pageNumber="16">FIGURE 9A–I.</emphasis>
<taxonomicName id="B5926ABCFFA7FF8A9A71F85A49A3F807" box="[335,451,1941,1964]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFA7FF8A9A71F85A49A3F807" box="[335,451,1941,1964]" italics="true" pageId="15" pageNumber="16">E. procera</emphasis>
</taxonomicName>
queen and male. A) queen in lateral view; B) queen head in full-frontal view; C) male in lateral view; D) male head in full-frontal view; E–I male genitalia: E) dorsal view; F) ventral view; G) lateral view; H) volsellar lobe; I) penis valve. Queen and male are both from South Thailand (SYPC)
</paragraph>
</caption>
<subSubSection id="3A8842B4FFB8FF959BFBFF584AD0FE31" pageId="16" pageNumber="17" type="discussion">
<paragraph id="722D113FFFB8FF959BFBFF584AD0FE31" blockId="16.[151,1437,151,410]" pageId="16" pageNumber="17">
<emphasis id="40E6CD2DFFB8FF959BFBFF584933FF1A" bold="true" box="[197,339,151,177]" pageId="16" pageNumber="17">Discussion:</emphasis>
This is the only known polymorphic
<taxonomicName id="B5926ABCFFB8FF959836FF574BC3FF1A" box="[776,931,152,177]" class="Insecta" family="Formicidae" genus="Euprenolepis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFB8FF959836FF574BC3FF1A" box="[776,931,152,177]" italics="true" pageId="16" pageNumber="17">Euprenolepis</emphasis>
</taxonomicName>
species. Majors of
<taxonomicName id="B5926ABCFFB8FF959FB8FF574D62FF1A" box="[1158,1282,152,177]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFB8FF959FB8FF574D62FF1A" box="[1158,1282,152,177]" italics="true" pageId="16" pageNumber="17">E. procera</emphasis>
</taxonomicName>
superficially resemble
<taxonomicName id="B5926ABCFFB8FF959A2FFF7049D7FF73" box="[273,439,191,216]" class="Insecta" family="Formicidae" genus="Pseudolasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="40E6CD2DFFB8FF959A2FFF7049D7FF73" box="[273,439,191,216]" italics="true" pageId="16" pageNumber="17">Pseudolasius</emphasis>
</taxonomicName>
, but can be separated from that genus by the characters listed under the genus diagnosis section. This species possesses several unique characters other than worker polymorphism.
<taxonomicName id="B5926ABCFFB8FF959EBCFF2A4893FE8E" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFB8FF959EBCFF2A4893FE8E" italics="true" pageId="16" pageNumber="17">E. procera</emphasis>
</taxonomicName>
has especially long scapes, although not as long as in
<taxonomicName id="B5926ABCFFB8FF959855FEC34C75FE8E" box="[875,1045,268,293]" class="Insecta" genus="Chapmanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="negrosensis">
<emphasis id="40E6CD2DFFB8FF959855FEC34C75FE8E" box="[875,1045,268,293]" italics="true" pageId="16" pageNumber="17">E. negrosensis</emphasis>
</taxonomicName>
. The eyes of
<taxonomicName id="B5926ABCFFB8FF959F8CFEC34D4EFE8E" box="[1202,1326,268,293]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFB8FF959F8CFEC34D4EFE8E" box="[1202,1326,268,293]" italics="true" pageId="16" pageNumber="17">E. procera</emphasis>
</taxonomicName>
are more forward toward the midline and lower in the head than observed in other species. The medially emarginated clypeus is most pronounced in
<taxonomicName id="B5926ABCFFB8FF959936FE964AE6FED9" box="[520,646,345,370]" class="Insecta" family="Formicidae" genus="Prenolepis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procera">
<emphasis id="40E6CD2DFFB8FF959936FE964AE6FED9" box="[520,646,345,370]" italics="true" pageId="16" pageNumber="17">E. procera</emphasis>
</taxonomicName>
. It is by far the darkest colored species and possesses the thickest, most distinct rugoreticulae covering the body.
</paragraph>
</subSubSection>
</treatment>
</document>
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