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<document ID-CLB-Dataset="55738" ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-GBIF-Dataset="8b26ea49-e662-41f7-a213-47f4b598c91c" ID-PMC="PMC9848896" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-PubMed="36761078" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="FA389A214FB55C6D9B677EFD2420F1A8" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus sambiranoensis Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="5A84A1E2-D64E-4665-BACA-83CEAC5DB8BF" docUuidSource="ZooBank" docVersion="5" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1732660529870" updateUser="ExternalLinkService">
<mods:mods id="8CD9ECCC92BD0A8BAC4A31425001E866" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5625264C4F4C0EC42CAF0D9B824DF6FE">
<mods:title id="0BBFFFC36306E405AEB6CE4BEF2DC7BA">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
</mods:titleInfo>
<mods:name id="DB6E401D3A067F88888DC48A9C22AB86" type="personal">
<mods:role id="FDF3F61C215F4117AD3BEEE6CD981954">
<mods:roleTerm id="53FF3847E70C89E752DDC06E9DC9A721">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4837B45F0889163445EDE65BA4FC708A">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="12B2CA62E239A3C913C8554FFBBE9753">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier id="CFBABB3BC8D3B60C9FC137738B01F484" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="98B68798155C1640075C6D15715282F7" type="personal">
<mods:role id="F38E779AF38F050DF38F31893F848681">
<mods:roleTerm id="79F0E84E31A0F084961D1E61F288F511">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F37E82467B8E8209B6A3761DB503170">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="58F25FD48CAEF6994A62E19F18ECB97B" type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation id="F340015B74C8FF71E547A8C3E44F03FC">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="077CFB19C686BC921F8A7D65B05C6C85">text</mods:typeOfResource>
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<mods:titleInfo id="18A25364CCFE5F7C480A031548C65E68">
<mods:title id="D623E2F57EA831E7C9A233551E76BA0B">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="7170DE0D12078CEEC74274C3F1D88BAA">
<mods:date id="B5F84554617E05405E1D4E4EE3109D25">2022</mods:date>
<mods:detail id="A537E23D0999CA367C35ED5785A6AB5D" type="pubDate">
<mods:number id="1BEA9216BA6E97E2D5332EF6FA4BD3D1">2022-05-03</mods:number>
</mods:detail>
<mods:detail id="2A629DB444582C347A0964002744944C" type="volume">
<mods:number id="25BCA9F6FD65B3BE7ACEAD02F750574D">1098</mods:number>
</mods:detail>
<mods:extent id="B4DF618DBBECF0D2CB6063E2DA5279FE" unit="page">
<mods:start id="C195D4BFEA34B5FB5CA468DE3E04B2E4">1</mods:start>
<mods:end id="4322B00D260FAB22FD8A510B587FC2AC">180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4D28D6FE76D93AC5202664A4696862AE">
<mods:url id="AAB9A470634BF321107E13AD89FC7713">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:url>
</mods:location>
<mods:classification id="92F20E2F17F59D3E83D607D53F64A4B4">journal article</mods:classification>
<mods:identifier id="B1732DBC5E6FCA1DA8B5429545E88D26" type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
<mods:identifier id="D28E3C765C7C9C9B08DAC6DC770F1D79" type="Pensoft-Pub">1313-2970-1098-1</mods:identifier>
<mods:identifier id="0018E59DF1BA24BC9FD125F23FAB8EAF" type="ZooBank">B4F4033F296E43CCBE54B9413BC19268</mods:identifier>
<mods:identifier id="C116ECD91198B76081F9E749D663F0E9" type="Pensoft-UUID">B22184E75B1A59C9B985C0DD9F092196</mods:identifier>
</mods:mods>
<treatment id="FA389A214FB55C6D9B677EFD2420F1A8" ID-GBIF-Taxon="195190277" LSID="urn:lsid:zoobank.org:act:5A84A1E2-D64E-4665-BACA-83CEAC5DB8BF" httpUri="http://treatment.plazi.org/id/FA389A214FB55C6D9B677EFD2420F1A8" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="9FFDC0A5E1E0DE06638C3D854A53D6F2" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="55E4D46C6B738C803D2C52673124701B" pageId="0" pageNumber="1">
<taxonomicName id="C16A638A16918294C9E08B80FF689888" LSID="http://zoobank.org/5A84A1E2-D64E-4665-BACA-83CEAC5DB8BF" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus sambiranoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sambiranoensis" status="sp. nov.">Camponotus sambiranoensis</taxonomicName>
<taxonomicNameLabel id="361543D716C8ACE5DD2F7E107B490E3C" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ADFA569BD2D701A7C70E8AE53A315545" pageId="0" pageNumber="1" type="description">
<paragraph id="EBC9955ABBC4EF030D32422F6AE76EF4" pageId="0" pageNumber="1">
<figureCitation id="162F6C47B6A9BF05A044829577AF6117" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head in full-face view A C. sambiranoensis (CASENT 0231845) B C. radamae (CASENT 0066777) C C. immaculatus (CASENT 0179441)." figureDoi="10.3897/zookeys.1098.73223.figure6" httpUri="https://binary.pensoft.net/fig/678065" pageId="0" pageNumber="1">Figs 6A</figureCitation>
<figureCitation id="BEBDC639FCE56D498B1AD07BEB72A5EA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Head in full-face view A C. sambiranoensis (CASENT 0231845) B C. cervicalis (CASENT 0217303)." figureDoi="10.3897/zookeys.1098.73223.figure9" httpUri="https://binary.pensoft.net/fig/678068" pageId="0" pageNumber="1">, 9A</figureCitation>
<figureCitation id="7FF607E2E6BDC0525B313DD50C052455" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Camponotus sambiranoensis A lateral view B head in full-face view C dorsal view of minor worker CASENT 0231845 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure73" httpUri="https://binary.pensoft.net/fig/678132" pageId="0" pageNumber="1">, 73</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CC00664EA48A22ECCAB28A555A609F7E" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="4C9A16F247DEC8D99D7B7C11DC3A02FD" pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph id="16E03DE6006790175D96381190FC0719" pageId="0" pageNumber="1">
<emphasis id="3BFFB0110429C827AD9BCD1880919432" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis id="A31A39E14ACF4C4B66D5BA72FE60E08B" bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: RS Ankarana, 22.9 km 224° SW Anivorano Nord,
<geoCoordinate id="74D1CF37584768B1502EC3228B9C4EB4" degrees="12.90889" direction="south" orientation="latitude" precision="1" value="-12.90889">-12.90889</geoCoordinate>
,
<geoCoordinate id="B9C66143810C0ED5E0BD3B99BD770B8F" degrees="49.10983" direction="east" orientation="longitude" precision="1" value="49.10983">49.10983</geoCoordinate>
, 80 m, tropical dry forest, ex rotten log, 10-16 Feb 2001, (Fisher, Griswold et al.) collection code: BLF02880, collection code: CASENT0428077 (CAS).
</paragraph>
</subSubSection>
<subSubSection id="DE5578424133037594E2DBF2A43855C0" pageId="0" pageNumber="1" type="paratypes">
<paragraph id="F264B8FFE1D052F0F8948A5A2B1848AB" pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph id="ED7D41FD89B51C82D4FE6C1BD9B17884" pageId="0" pageNumber="1">3 minor workers of the same data as holotype but with the following specimen codes: CASENT0428078, CASENT0428080, CASENT0428079 (NHMUK, MHNG, CAS).</paragraph>
</subSubSection>
<subSubSection id="E7A6C36992D4AAEDEBEE2D9140E9404E" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="4D375C53F3FF6D14F6FC26AEE8DE3FAC" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="18512A094D301E6754DC7E092623B5CB" pageId="0" pageNumber="1">
<materialsCitation id="D1C2573FEAFB5F28EEF1EF82DBE19D84" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="B. L. Fisher, de Binara, Hell-Ville S., D. Lees, R. Ranaivosolo, RS Manongarivo, RS Ambre, RS Ankarana" country="Madagascar" county="Ampasindava" elevation="600" latitude="-13.2505" location="Binara Forest" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="B2679AA4CA3E8B63FE4BDF7A0B8F60A3" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="115ADF9B13392A3FF1F76B316159101E" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion id="FCE9A5A65C0824D93C67D7B16A899DB3" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
</emphasis>
:
<collectingCounty id="938584F6F385526E5E2E58EC6BA510CF">Ampasindava</collectingCounty>
,
<collectingMunicipality id="972DB97FA331821AF02FAD67EA4750F0">
<normalizedToken id="B4A66800CF8F2079303C6ADECF2E5F4C" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="4B75BAC460820746299109BD37B65603" originalValue="d’Ambilanivy">d'Ambilanivy</normalizedToken>
</collectingMunicipality>
,
<locationDeviation id="5587420BB30C4588AB51E79873B2B16F" location="Ambaliha">
<quantity id="79AE38BAE84245442F08028772E43B18" metricMagnitude="3" metricUnit="m" metricValue="3.9" unit="km" value="3.9">3.9 km</quantity>
181° S Ambaliha
</locationDeviation>
,
<geoCoordinate id="68ABCAE26277DFD4E02BD1E169334FF7" degrees="13.79861" direction="south" orientation="latitude" precision="1" value="-13.79861">-13.79861</geoCoordinate>
,
<geoCoordinate id="90098A4BBC7AF331391494C9FA7BDEFF" degrees="48.16167" direction="east" orientation="longitude" precision="1" value="48.16167">48.16167</geoCoordinate>
,
<quantity id="DF041F54013DD3E6571B4A7916A227BA" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation id="9D75156D68290348185BFA83A72B45FF" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
, rainforest (Fisher, Griswold et al.) (CAS);
<location id="435B86B59D0461F69BD6EDBE0D7C9D2E" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:435B86B59D0461F69BD6EDBE0D7C9D2E" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="Binara Forest" stateProvince="Antsiranana">Binara Forest</location>
,
<geoCoordinate id="F3956BA9955F4CC2BC21B3F8B0C2D128" degrees="13.26388" direction="south" orientation="latitude" precision="1" value="-13.26388">-13.26388</geoCoordinate>
,
<geoCoordinate id="F2CE173ABF7721C9E588B0D204FE8A68" degrees="49.60141" direction="east" orientation="longitude" precision="1" value="49.60141">49.60141</geoCoordinate>
,
<quantity id="9F19671B983819C020A3BA977183C556" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation id="AB8525CC2065C77F8C7BE9F03B464141" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
, rainforest (
<collectorName id="EFF94B957E7C87CE194D127667BDEE11">B.L. Fisher</collectorName>
et al.) (CAS);
<location id="76D7E8D96BDF8DA87CE6019284291CF0" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:76D7E8D96BDF8DA87CE6019284291CF0" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="District of Vohemar" stateProvince="Antsiranana">District of Vohemar</location>
, Analabe Sahaka
<locationDeviation id="1A0B7A37DA09EB11E431A2EC312CB710" location="Daraina">
<quantity id="282D230ACFDC470D2E5E1C2F9396584B" metricMagnitude="4" metricUnit="m" metricValue="4.3" unit="km" value="43.0">43 km</quantity>
E of Daraina
</locationDeviation>
,
<geoCoordinate id="78FEB1ECA72776AEC71BD5759275D963" degrees="13.07933" direction="south" orientation="latitude" precision="1" value="-13.07933">-13.07933</geoCoordinate>
,
<geoCoordinate id="8B4FFFA2AD97A4F7FD36E208818F0570" degrees="49.90233" direction="east" orientation="longitude" precision="1" value="49.90233">49.90233</geoCoordinate>
,
<quantity id="22CAEBFEB28DACE57AD5986AF4F99902" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">
<elevation id="8BF63B09D26A93215576060AE3AB9C0C" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</elevation>
</quantity>
, wooded grassland-bushland (Mike,
<normalizedToken id="CB0E77AA064F1C7B82D92DCAE12AE6B0" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS);
<location id="EB0FD5126F7A5E7DED25622BB55DE124" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:EB0FD5126F7A5E7DED25622BB55DE124" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="Foret de Binara" stateProvince="Antsiranana">
<normalizedToken id="360C45230C330B471A9506BC9C0859F4" originalValue="Forêt">Foret</normalizedToken>
<collectorName id="2B210D916DE2976EAFEADC556486A5FA">de Binara</collectorName>
</location>
,
<locationDeviation id="B0746DD2C1873709DF0F78EFB6A24690" location="Daraina">
<quantity id="7EC353FD021B78F034D3E8ED218F866A" metricMagnitude="3" metricUnit="m" metricValue="7.5" unit="km" value="7.5">7.5 km</quantity>
230° SW Daraina
</locationDeviation>
,
<geoCoordinate id="26DF6A98D38C07314637D8FFA6D1E345" degrees="13.255" direction="south" orientation="latitude" precision="55" value="-13.255">-13.255</geoCoordinate>
,
<geoCoordinate id="A62D8F9C9F3E3ECFED519322E367627A" degrees="49.61667" direction="east" orientation="longitude" precision="1" value="49.61667">49.61667</geoCoordinate>
,
<quantity id="33E39CDD5F65FDC47DDFE40D0F9C9B4C" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">
<elevation id="3F17E0B4D5066644D9CE6BA55DEFC72B" metricMagnitude="2" metricUnit="m" metricValue="3.75" unit="m" value="375.0">375 m</elevation>
</quantity>
, tropical dry forest (
<collectorName id="C5DD4269F162B4CCF1ABB0AEFE429183">B.L. Fisher</collectorName>
et al.) (CAS);
<location id="B616B48AF6BC9CFBCADAC26154EA38AA" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:B616B48AF6BC9CFBCADAC26154EA38AA" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="Makirovana forest" stateProvince="Antsiranana">Makirovana forest</location>
,
<geoCoordinate id="D2357303F7264642BACB4C61D7F36836" degrees="14.17066" direction="south" orientation="latitude" precision="1" value="-14.17066">-14.17066</geoCoordinate>
,
<geoCoordinate id="19114E942320C7CC851AAACFA57D35BA" degrees="49.95409" direction="east" orientation="longitude" precision="1" value="49.95409">49.95409</geoCoordinate>
,
<quantity id="833994CBA089DCB26D54BC279656052D" metricMagnitude="2" metricUnit="m" metricValue="4.15" unit="m" value="415.0">
<elevation id="FE97E48EB50B6A0D10DF04DCBDFE48AA" metricMagnitude="2" metricUnit="m" metricValue="4.15" unit="m" value="415.0">415 m</elevation>
</quantity>
, rainforest (
<collectorName id="B136A1009BF77C16CCA59585D267327D">B.L. Fisher</collectorName>
et al.) (CAS); Nosy-Be Pref., Antsirambazaha,
<collectorName id="73E37ADB0656B90875A41E68E1041505">Hell-Ville S.</collectorName>
-Pref.: Nosy Be, Site 3,
<geoCoordinate id="210CFCDB445FFFCECF5CD95A4BBB0B93" degrees="13.41345" direction="south" orientation="latitude" precision="1" value="-13.41345">-13.41345</geoCoordinate>
,
<geoCoordinate id="146162915BF7C7991B1B30042F0D34B3" degrees="48.3113" direction="east" orientation="longitude" precision="5" value="48.3113">48.3113</geoCoordinate>
,
<quantity id="6B499750B35C7BFDB3E0E9F834E9B60D" metricMagnitude="2" metricUnit="m" metricValue="1.43" unit="m" value="143.0">
<elevation id="7F487DF921E3E97F2AA0CF513A2494E5" metricMagnitude="2" metricUnit="m" metricValue="1.43" unit="m" value="143.0">143 m</elevation>
</quantity>
, in degraded primary rainforest on slope with Nastus (
<collectorName id="A7FD93F0BFEEFB31E387FF3016A3591F">D. Lees</collectorName>
&amp;
<collectorName id="7D96BC75416D164DD59255A8A866093B">R. Ranaivosolo</collectorName>
) (CAS);
<collectorName id="7AB5D483CEFCE4F26AD89F28C6FF7612">RS Manongarivo</collectorName>
,
<locationDeviation id="DE2E3DA4B46FC2129E7736F510F75A82" location="Antanambao">
<quantity id="4E8F15D49940D2C31CA121F63D906C3E" metricMagnitude="4" metricUnit="m" metricValue="1.08" unit="km" value="10.8">10.8 km</quantity>
229° SW Antanambao
</locationDeviation>
,
<geoCoordinate id="8F23DB10672143F7B46B8BB873EB26BB" degrees="13.96167" direction="south" orientation="latitude" precision="1" value="-13.96167">-13.96167</geoCoordinate>
,
<geoCoordinate id="0E0999022F631902C8C1E8142894EF1C" degrees="48.43333" direction="east" orientation="longitude" precision="1" value="48.43333">48.43333</geoCoordinate>
,
<quantity id="811CB6CEE098BA1CF21AE51283AA1615" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="95013865AFC5AE0971C391FF8C72BCD7" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
, rainforest (
<collectorName id="86672DD12F4E6A1E3A1B1F6D5871BA72">B.L. Fisher</collectorName>
) (CAS);
<collectorName id="89E2D3FDBC3ABC61AAA61650BBC07BE3">RS Manongarivo</collectorName>
,
<locationDeviation id="11F2C4173CB3916BA886CE505488B53A" location="Antanambao">
<quantity id="8F350F137F70A129FEB1355EDF01493D" metricMagnitude="4" metricUnit="m" metricValue="1.45" unit="km" value="14.5">14.5 km</quantity>
220° SW Antanambao
</locationDeviation>
, -14, 48.43167,
<quantity id="944634EF1A15B81D798BB77C3199DFE3" metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">
<elevation id="177192050DE8A158388A9504E270293F" metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">1220 m</elevation>
</quantity>
, montane rainforest (
<collectorName id="98021FFE9884FBA0081B82211500F740">B.L. Fisher</collectorName>
) (CAS);
<collectorName id="98811FD0C5F5F0BF6C3154C443686AAD">RS Ambre</collectorName>
,
<locationDeviation id="EAEB86BFAC5F4BB6A0B4A4CF134582F3" location="Sakaramy">
<quantity id="E684F1078FAD647DB1D511DC20C8300B" metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="km" value="3.5">3.5 km</quantity>
235° SW Sakaramy
</locationDeviation>
,
<geoCoordinate id="1A1720BF2C38F2F58A1805A8A942B706" degrees="12.46889" direction="south" orientation="latitude" precision="1" value="-12.46889">-12.46889</geoCoordinate>
,
<geoCoordinate id="325A7E0E3B07AA0BF946E78C463EC220" degrees="49.24217" direction="east" orientation="longitude" precision="1" value="49.24217">49.24217</geoCoordinate>
,
<quantity id="B11C1D2D1E231718B3E1AB215D70BBBE" metricMagnitude="2" metricUnit="m" metricValue="3.25" unit="m" value="325.0">
<elevation id="43C074290E2B842B91EE25DEC7512C86" metricMagnitude="2" metricUnit="m" metricValue="3.25" unit="m" value="325.0">325 m</elevation>
</quantity>
, tropical dry forest edge (Fisher, Griswold et al.) (CAS);
<collectorName id="E9BAC76F6E06DCA922D4F1D26E58F885">RS Ankarana</collectorName>
,
<locationDeviation id="A54ACD35D1879DB2933F9B17D16F216A" location="Anivorano Nord">
<quantity id="D3C7D5978DDE6A8274B1013A2A8BAAEE" metricMagnitude="4" metricUnit="m" metricValue="2.29" unit="km" value="22.9">22.9 km</quantity>
224° SW Anivorano Nord
</locationDeviation>
,
<geoCoordinate id="A2369B2D015F459FB394BB57F681EE99" degrees="12.90889" direction="south" orientation="latitude" precision="1" value="-12.90889">-12.90889</geoCoordinate>
,
<geoCoordinate id="07FE2B6C84D8A922C8F24ECECCCC670A" degrees="49.10983" direction="east" orientation="longitude" precision="1" value="49.10983">49.10983</geoCoordinate>
,
<quantity id="84167D339D1BC70849B9B308CC75DA2E" metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">
<elevation id="9C5026FD4305002F71F3B3051009CF81" metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">80 m</elevation>
</quantity>
, tropical dry forest (Fisher, Griswold et al.) (CAS). SAVA Region,
<location id="96064B6E23F3AEF5697F3B284B388BCF" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:96064B6E23F3AEF5697F3B284B388BCF" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="District of Vohemar" stateProvince="Antsiranana">District of Vohemar</location>
,
<location id="48E594AAEEA96A819E2E845C09703129" LSID="urn:lsid:plazi:treatment:FA389A214FB55C6D9B677EFD2420F1A8:48E594AAEEA96A819E2E845C09703129" country="Madagascar" county="Ampasindava" latitude="-13.2505" longLatPrecision="5" longitude="49.61667" municipality="Foret d'Ambilanivy" name="Antsahabelela Rain Forest" stateProvince="Antsiranana">Antsahabelela Rain Forest</location>
,
<locationDeviation id="7300D667DB71F8846A88E32959EC0C97" location="Daraina">
<quantity id="B9DF3A0051F0A766940BFE9CAE99CBA6" metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
SW of Daraina
</locationDeviation>
,
<geoCoordinate id="4CD2EFB6E311C391316B8ED5F868E925" degrees="13.2505" direction="south" orientation="latitude" precision="5" value="-13.2505">-13.2505</geoCoordinate>
,
<geoCoordinate id="33F8B7BE1BA2B082F1AAAB7A55E9BE4F" degrees="49.61667" direction="east" orientation="longitude" precision="1" value="49.61667">49.61667</geoCoordinate>
,
<quantity id="65957AA8F140FFBF8F35E668587151E2" metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" unit="m" value="182.0">
<elevation id="750B269A07C85924913D11E293E9DF79" metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" unit="m" value="182.0">182 m</elevation>
</quantity>
, humid
<normalizedToken id="E1EEC4D807EFA2333C5B414AAB46856C" originalValue="Forêt">Foret</normalizedToken>
(Mike,
<normalizedToken id="BC469B7BCC6D3DC9C1876881B314A0B1" originalValue="Rin’ha">Rin'ha</normalizedToken>
) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9F4364D4860DF80339DCD43304D6A211" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="64B5CF31E6DF18123831A3587915BD17" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="1802DEDCE1FF3717958EFA00FD168E8C" pageId="0" pageNumber="1">With head in full-face view, lateral cephalic margins converging posteriorly towards eye level; anteromedian margin of clypeus broadly convex; two apical teeth of mandible normally spaced; lateral cephalic margin anterior to eye level without erect hairs.</paragraph>
</subSubSection>
<subSubSection id="6B70DA3CFF815C81511236B9D8F201E6" pageId="0" pageNumber="1" type="description">
<paragraph id="A1495EEE6961FD5AB695634DF2052E9D" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="180D015DE47806B7D4945885D9CA4404" pageId="0" pageNumber="1">
<emphasis id="C3C17795FB8347A4766E6CB625835103" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, lateral margins of head converging progressively to level of anterior ocular margin, strongly converging to a short posterior margin behind eye level; eye protruding and large (EL/CS: 0.28
<normalizedToken id="F6F9F55FC852768462E8DF1D7B09BBDF" originalValue="±">+/-</normalizedToken>
0.01; 0.27-0.29), breaking lateral cephalic border, level of its posterior margin at least at posterior 1/3 of head (PoOc/CL; 0.32
<normalizedToken id="39B482677A39D90EC34624B9C019CCBC" originalValue="±">+/-</normalizedToken>
0.04; 0.30-0.42); frontal carinae wide (FR/CS: 0.22
<normalizedToken id="441FC86253AF8E161BD002F0B5FB09F4" originalValue="±">+/-</normalizedToken>
0.0; 0.21-0.23), posteriorly parallel, distance between them larger than smallest distance to eye; clypeus without well-defined anterolateral angle, its anteromedian margin broadly convex or triangular; mandible with two apical teeth closely spaced; antennal scape relatively long (SL/CS: 2.04
<normalizedToken id="2181D6E2BB297D31BCD1EEB962FB15F5" originalValue="±">+/-</normalizedToken>
0.07; 1.97-2.18). Mesosoma low and long (MPH/ML: 0.33
<normalizedToken id="ABD0DC1ADC89A490C648628DB4484D62" originalValue="±">+/-</normalizedToken>
0.03; 0.28-0.36), with weakly convex promesonotum and approximately flat mesopropodeum; metanotal groove barely visible; propodeal dorsum almost straight or anteriorly slightly convex and posteriorly flat; dorsal margin of propodeum and declivity joined at blunt angle; propodeal declivity 1/3 of length of dorsum. Petiole nodiform, dorsal margin inclined posteriorly, its junction to anterior face bluntly angulate; anterior face of petiolar node 1/2 height of posterior face; femur of hind leg rounded axially, not twisted basally.
</paragraph>
<paragraph id="E10B16801893E1FEC144756905AC4563" pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; lateral margin of head without erect hairs; two erect hairs present near posterior cephalic margin; antennal scape without erect hairs but covered with numerous appressed hairs; pronotum with a pair of erect hairs; posterodorsal angle of propodeum without erect hairs.</paragraph>
<paragraph id="4C1F5CFEC86317FF431ADCB44750DC7E" pageId="0" pageNumber="1">
<emphasis id="6AAF84938A6A0288DD194B1844B77B8B" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: larger head (CS: 4.45; CWb/CL: 0.98); apical 1/4 of antennal scape extending beyond posterior cephalic margin; very robust, short, and high mesosoma, with promesonotum and metanotum forming an even convexity; and propodeal dorsum sloping and joining declivity with blunt angle. Petiolar node much compressed anteroposteriorly, anterior face shorter than posterior.
</paragraph>
</subSubSection>
<subSubSection id="855D93935FAD7A5B72FEEEE9CE0C6FD1" pageId="0" pageNumber="1" type="distribution">
<paragraph id="C2643905C19BC4D7B322BAA1BDF06372" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="B080CEA8C97B62EC9BA00BADC11E51AE" pageId="0" pageNumber="1">
<taxonomicName id="46FE08247DBC1724482FC4E8189B7187" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus sambiranoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sambiranoensis">
<emphasis id="763239D41840B74C9A890B7BD388BDB7" italics="true" pageId="0" pageNumber="1">Camponotus sambiranoensis</emphasis>
</taxonomicName>
is only known to occur in the north of Madagascar. It colonizes the dry forests of Binara, RS Ambre, and RS Ankarana, the transitional forests of Ampasindava and Binara, the rainforests of Makirovana and RS Manongarivo, and the montane forest of RS Manongarivo (Fig.
<figureCitation id="E68D771A3EC007AF89FD1EDE964D2A61" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Camponotus sambiranoensis A lateral view B head in full-face view C dorsal view of minor worker CASENT 0231845 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure73" httpUri="https://binary.pensoft.net/fig/678132" pageId="0" pageNumber="1">73D</figureCitation>
). This species nests in rotten logs and in the ground, and forages on the ground, in leaf litter, and on low vegetation.
</paragraph>
<caption id="6274E3DE730B938E82952ACBB29337FA" doi="10.3897/zookeys.1098.73223.figure73" httpUri="https://binary.pensoft.net/fig/678132" pageId="0" pageNumber="1" start="Figure 73" startId="F73">
<paragraph id="765A63F39FF49703344FFDF9497337C8" pageId="0" pageNumber="1">
<emphasis id="49C24239719F5DB70326D290FC94D08F" bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
<taxonomicName id="5C67653E2AE712A0D05FDBA8711A93E9" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus sambiranoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sambiranoensis">
<emphasis id="3C9AF14A27F575850CF3564FD2261192" italics="true" pageId="0" pageNumber="1">Camponotus sambiranoensis</emphasis>
</taxonomicName>
<emphasis id="D58887274C37D3092E6A16CADCED36E9" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis id="DA1D710249C219BF9E724A1C533DD48F" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis id="A0C9BAC9DB3E8303D2AD3FD7EADAE1F7" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of minor worker CASENT0231845
<emphasis id="22FDE918833DD16918498555554AF6C5" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="37566D439435BD23FCFD124915DE6DF8" pageId="0" pageNumber="1" type="discussion">
<paragraph id="0A298B30879073CBCD03E96A0AA5CC5A" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="A84F594F6919E658A3189EEFCA076ABF" pageId="0" pageNumber="1">
<taxonomicName id="22543AF048015EA5F9FDF1D77B430FA8" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus sambiranoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sambiranoensis">
<emphasis id="A434DC1E603FF0F1583D2B6BAD4829D1" italics="true" pageId="0" pageNumber="1">Camponotus sambiranoensis</emphasis>
</taxonomicName>
shares a broadly convex anteromedian margin of clypeus with
<taxonomicName id="E79258364B1AFC931D8770A55D1CC89D" lsidName="C. niavo" pageId="0" pageNumber="1" rank="species" species="niavo">
<emphasis id="5DD6A66661A20DD96FF2CB8BBC2A7021" italics="true" pageId="0" pageNumber="1">C. niavo</emphasis>
</taxonomicName>
, and
<taxonomicName id="84654CBF2573CB61EFA46817ACCB4094" lsidName="C. cervicalis" pageId="0" pageNumber="1" rank="species" species="cervicalis">
<emphasis id="E43CF0AB631D40EC0664F2ECDADA104D" italics="true" pageId="0" pageNumber="1">C. cervicalis</emphasis>
</taxonomicName>
, but in the two latter species the lateral cephalic margin anterior to the eye level is covered with erect hairs.
</paragraph>
<paragraph id="5EE3372F7B2DFF41141571327F6C3CFB" pageId="0" pageNumber="1">
The qualitative, morphology-based delimitation of
<taxonomicName id="40452D2DF988D9469FF607CC1B7E7937" lsidName="C. sambiranoensis" pageId="0" pageNumber="1" rank="species" species="sambiranoensis">
<emphasis id="02D76FCE597F8D33365E4CDA69BE1ACA" italics="true" pageId="0" pageNumber="1">C. sambiranoensis</emphasis>
</taxonomicName>
is confirmed by the multivariate morphometric technique. The validity of the species is also confirmed by LDA at 100% classification success.
</paragraph>
</subSubSection>
<subSubSection id="9F2A7930873D7927B8DE9CF4AB17CF57" pageId="0" pageNumber="1" type="etymology">
<paragraph id="4F4EE193A4186E097D4580264EC0C4CC" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="EE1070539E00EA5122230936A03C74BE" pageId="0" pageNumber="1">
The species name
<emphasis id="E4F398AF52979E82CEEEF0F9A7D3E678" italics="true" pageId="0" pageNumber="1">sambiranoensis</emphasis>
is a Latin singular adjective in the nominative case of masculine gender. This species name refers to the region of Sambirano where the species was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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