307 lines
38 KiB
XML
307 lines
38 KiB
XML
<document id="82E7CF678A0BB848A6F6445F3FE8C288" ID-DOI="10.5281/zenodo.275690" ID-GBIF-Dataset="fd14b593-5c84-4314-806e-e288bf3415f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="275690" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460158641716" checkinUser="plazi" docAuthor="Hall, Eugene & Short, Andrew E. Z." docDate="2010" docId="FA3687EEDF408319A5D8FF58C86BC91D" docLanguage="en" docName="zt02358p056.pdf" docOrigin="Zootaxa 2358" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hydroscapha perijaensis Hall & Short, 2010, sp. n." docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="060FFF96DF41831CA54FFFCFCC4DCC0F" masterDocTitle="A new species of Hydroscapha LeConte from northwestern Venezuela (Coleoptera: Myxophaga: Hydroscaphidae)" masterLastPageNumber="56" masterPageNumber="49" pageNumber="50" updateTime="1698235668619" updateUser="plazi">
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<mods:mods id="E6492FA91BF78599F6319F2FE731182B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="63B7D5C4F7BFA500D1C6D2673C0D5588">
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<mods:title id="FBE8DB19D64C8E056E99A32B86854753">A new species of Hydroscapha LeConte from northwestern Venezuela (Coleoptera: Myxophaga: Hydroscaphidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="4EFE4691EC19DFA2032987189AC2BA22" type="personal">
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<mods:roleTerm id="58905813D62D1C3CD33D8507DECB9BAD">Author</mods:roleTerm>
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<mods:namePart id="AC31F367AFFAEB0346537D4A5F9FC85E">Hall, Eugene</mods:namePart>
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<mods:name id="ADC120EBF9E0D0A5A14911E5058EDFA0" type="personal">
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<mods:roleTerm id="0109A6B3451C3B81DD4FAA38FDE9BDD5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07C353D2F6BD10E736933D076E3955F7">Short, Andrew E. Z.</mods:namePart>
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<mods:typeOfResource id="760B085A0F7276AA0321A911BCB7B673">text</mods:typeOfResource>
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<mods:titleInfo id="CAE43D48492F35FCD4138CC6C413167F">
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<mods:title id="E717D3EF594BA516916DA6C5B2DC8C39">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BBD02246DEE9138A9C3F21B3323EFC0B">
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<mods:date id="EA627A1CCBCC3924858DBC1D71F6C995">2010</mods:date>
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<mods:detail id="DE28CD26567F8A2E905C1BFDAD4D0B0E" type="volume">
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<mods:number id="74730FACB045DA0D8FA5A0615E84DF68">2358</mods:number>
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<mods:start id="0E4609B4671907A45D4EB2F0425EA1CC">49</mods:start>
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<mods:classification id="4F4850A4022255FC0FB21D9EEB2B0AB4">journal article</mods:classification>
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<mods:identifier id="D38C81415BFA02239BA34EC889FABC5E" type="DOI">10.5281/zenodo.275690</mods:identifier>
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<mods:identifier id="95DC31D13B9F75A8F3DE4653B47DBFD5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="648B886FC41A126B59B22D9A415007FD" type="Zenodo-Dep">275690</mods:identifier>
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<treatment id="FA3687EEDF408319A5D8FF58C86BC91D" ID-DOI="http://doi.org/10.5281/zenodo.6211219" ID-GBIF-Taxon="127654833" ID-Zenodo-Dep="6211219" LSID="urn:lsid:plazi:treatment:FA3687EEDF408319A5D8FF58C86BC91D" httpUri="http://treatment.plazi.org/id/FA3687EEDF408319A5D8FF58C86BC91D" lastPageId="5" lastPageNumber="54" pageId="1" pageNumber="50">
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<subSubSection id="3A856573DF40831DA5D8FF58CD5CCCD8" pageId="1" pageNumber="50" type="nomenclature">
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<paragraph id="722036F8DF40831DA5D8FF58CE46CCBE" blockId="1.[151,523,151,215]" box="[151,523,151,177]" pageId="1" pageNumber="50">
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<heading id="29688194DF40831DA5D8FF58CE46CCBE" bold="true" box="[151,523,151,177]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
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<emphasis id="40EBEAEADF40831DA5D8FF58CE46CCBE" bold="true" box="[151,523,151,177]" pageId="1" pageNumber="50">
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<taxonomicName id="B59F4D7BDF40831DA5D8FF58CD8CCCBE" box="[151,449,151,177]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
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<emphasis id="40EBEAEADF40831DA5D8FF58CD8CCCBE" bold="true" box="[151,449,151,177]" italics="true" pageId="1" pageNumber="50">Hydroscapha perijaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5BD85791DF40831DA486FF58CE46CCBE" box="[457,523,151,177]" pageId="1" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="722036F8DF40831DA5D8FF72CD5CCCD8" blockId="1.[151,523,151,215]" box="[151,273,189,215]" pageId="1" pageNumber="50">
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(
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<figureCitation id="EAA42A7DDF40831DA5D0FF72CD4ACCD8" box="[159,263,189,215]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2. H" captionStart-2="FIGURE 3. H" captionStart-3="FIGURE 4. H" captionStart-4="FIGURE 5. H" captionStartId-0="1.[151,255,1751,1775]" captionStartId-1="2.[151,255,2003,2027]" captionStartId-2="3.[151,255,887,911]" captionStartId-3="3.[151,255,2001,2025]" captionStartId-4="4.[151,258,1615,1639]" captionTargetBox-0="[197,1402,815,1731]" captionTargetBox-1="[190,1394,290,1982]" captionTargetBox-2="[151,1434,181,865]" captionTargetBox-3="[185,1409,948,1979]" captionTargetBox-4="[171,1398,198,1614]" captionTargetId-0="figure@1.[197,1402,814,1731]" captionTargetId-1="figure@2.[190,1396,290,1982]" captionTargetId-2="figure@3.[151,1436,181,866]" captionTargetId-3="figure@3.[185,1410,930,1980]" captionTargetId-4="figure@4.[151,1436,176,1614]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionText-0="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." captionText-1="FIGURE 2. H. perijaensis sp. n. A) head and pronotum (dorsal); B) elytron (dorsal); C) elytron (ventral)." captionText-2="FIGURE 3. H. perijaensis sp. n. A) head (ventral); B) antenna; C) mesosternum and metaventrite (ventral)." captionText-3="FIGURE 4. H. perijaensis sp. n. A) terminal abdominal segments; female; B) terminal abdominal segments, male." captionText-4="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri-0="https://zenodo.org/record/275691/files/figure.png" httpUri-1="https://zenodo.org/record/275692/files/figure.png" httpUri-2="https://zenodo.org/record/275693/files/figure.png" httpUri-3="https://zenodo.org/record/275694/files/figure.png" httpUri-4="https://zenodo.org/record/275695/files/figure.png" pageId="1" pageNumber="50">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3A856573DF40831DA5D8FEC8CE28CEB3" pageId="1" pageNumber="50" type="materials_examined">
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<paragraph id="722036F8DF40831DA5D8FEC8CE28CEB3" blockId="1.[151,1437,263,775]" pageId="1" pageNumber="50">
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<emphasis id="40EBEAEADF40831DA5D8FEC8CD1FCD2E" bold="true" box="[151,338,263,289]" pageId="1" pageNumber="50">
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<typeStatus id="AD24885ADF40831DA5D8FEC8CC9BCD2E" box="[151,214,263,289]" pageId="1" pageNumber="50">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="AD24885ADF40831DA414FEC7CD9DCD2D" box="[347,464,264,290]" pageId="1" pageNumber="50" type="holotype">Holotype</typeStatus>
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: 3: “
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<collectingCountry id="0A887668DF40831DA743FEC7CE88CD2D" box="[524,709,264,290]" name="Venezuela" pageId="1" pageNumber="50">VENEZUELA</collectingCountry>
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: Zulia State / Perija National Park [at Toromo];
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<quantity id="B5679B1DDF40831DA040FEC7C916CD2D" box="[1295,1371,264,290]" metricMagnitude="2" metricUnit="m" metricValue="3.95" pageId="1" pageNumber="50" unit="m" value="395.0">395 m</quantity>
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/
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<geoCoordinate id="17AB503FDF40831DA03BFEC7CCB0CD48" direction="north" orientation="latitude" pageId="1" pageNumber="50" precision="1" value="10.049367">10º 2.962' N</geoCoordinate>
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,
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<geoCoordinate id="17AB503FDF40831DA446FEE2CDFDCD48" box="[265,432,301,327]" direction="west" orientation="longitude" pageId="1" pageNumber="50" precision="1" value="-72.71025">72º 42.615' W</geoCoordinate>
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/
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<date id="06211038DF40831DA489FEE2CE0BCD48" box="[454,582,301,327]" pageId="1" pageNumber="50" value="2007-09-23">23.ix.2007</date>
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; leg. A.E.Z. Short / AS-07-023”. Deposited in the Museo del Instituto de Zoología Agrícola (
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<collectionCode id="148EAE3DDF40831DA432FE9DCD83CD63" LSID="urn:lsid:biocol.org:col:33834" box="[381,462,338,364]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="50">MIZA</collectionCode>
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), Universidad Central de
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<collectingCountry id="0A887668DF40831DA7B7FE9DCF3DCD63" box="[760,880,338,364]" name="Venezuela" pageId="1" pageNumber="50">Venezuela</collectingCountry>
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(Maracay,
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<collectingCountry id="0A887668DF40831DA6BBFE9DC83FCD63" box="[1012,1138,338,364]" name="Venezuela" pageId="1" pageNumber="50">Venezuela</collectingCountry>
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). Allotype: Ƥ, (same data as
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<typeStatus id="AD24885ADF40831DA5F6FEB7CD64CD9D" box="[185,297,376,402]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
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;
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<collectionCode id="148EAE3DDF40831DA479FEB7CDC4CD9D" LSID="urn:lsid:biocol.org:col:33834" box="[310,393,376,402]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="50">MIZA</collectionCode>
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).
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<typeStatus id="AD24885ADF40831DA4D1FEB7CE5BCD9D" box="[414,534,376,402]" pageId="1" pageNumber="50" type="paratype">Paratypes</typeStatus>
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(142 exs.): same data as
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<typeStatus id="AD24885ADF40831DA61EFEB7CFF0CD9D" box="[849,957,376,402]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
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(35 exs.); same locality but
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<date id="06211038DF40831DA06FFEB7C9D5CD9D" box="[1312,1432,376,402]" pageId="1" pageNumber="50" value="2009-01-28">28.i.2009</date>
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, VZ
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<date id="06211038DF40831DA5F2FE52CD7ECDB8" box="[189,307,413,439]" pageId="1" pageNumber="50" value="2802-09-01">09012802</date>
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A (72 exs.).
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<collectingCountry id="0A887668DF40831DA48CFE52CE3BCDB8" box="[451,630,413,439]" name="Venezuela" pageId="1" pageNumber="50">VENEZUELA</collectingCountry>
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: Zulia State, Perija National Park,
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<quantity id="B5679B1DDF40831DA149FE52C81FCDB8" box="[1030,1106,413,439]" metricMagnitude="2" metricUnit="m" metricValue="4.35" pageId="1" pageNumber="50" unit="m" value="435.0">435 m</quantity>
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,
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<date id="06211038DF40831DA112FE52C883CDB8" box="[1117,1230,413,439]" pageId="1" pageNumber="50" value="2009-01-28">28.i.2009</date>
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,
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<geoCoordinate id="17AB503FDF40831DA195FE52C926CDB8" box="[1242,1387,413,439]" direction="north" orientation="latitude" pageId="1" pageNumber="50" precision="1" value="10.050966">10º 3.058’N</geoCoordinate>
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,
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<geoCoordinate id="17AB503FDF40831DA039FE52CD5DCDD3" direction="west" orientation="longitude" pageId="1" pageNumber="50" precision="1" value="-72.71623">72º 42.974’W</geoCoordinate>
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, leg. A. Short, M. Garcia, & J. Camacho, stream & seep, VZ
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<date id="06211038DF40831DA6BCFE0DC821CDD3" box="[1011,1132,450,476]" pageId="1" pageNumber="50" value="2801-09-01">09012801</date>
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B (35 exs.) In addition to
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<collectionCode id="148EAE3DDF40831DA5D8FE27CCAACE0D" LSID="urn:lsid:biocol.org:col:33834" box="[151,231,488,514]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="50">MIZA</collectionCode>
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,
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<typeStatus id="AD24885ADF40831DA5BBFE27CD24CE0D" box="[244,361,488,514]" pageId="1" pageNumber="50" type="paratype">paratypes</typeStatus>
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will be distributed in the following institutions: Division of Entomology, University of Kansas (Lawrence, KS,
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<collectingCountry id="0A887668DF40831DA4F1FDC2CDB2CE28" box="[446,511,525,551]" name="United States of America" pageId="1" pageNumber="50">USA</collectingCountry>
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), Department of Entomology, University of Arizona (Tucson AZ,
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<collectingCountry id="0A887668DF40831DA06AFDC2C92BCE28" box="[1317,1382,525,551]" name="United States of America" pageId="1" pageNumber="50">USA</collectingCountry>
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), W. Eugene Hall Collection (Tucson, AZ,
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<collectingCountry id="0A887668DF40831DA73CFDFDCEF8CE43" box="[627,693,562,588]" name="United States of America" pageId="1" pageNumber="50">USA</collectingCountry>
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), Museo de Artrópodos, Universidad del Zulia (Maracaibo,
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<collectingCountry id="0A887668DF40831DA5D8FD97CD57CE7D" box="[151,282,600,626]" name="Venezuela" pageId="1" pageNumber="50">Venezuela</collectingCountry>
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),
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<collectingCountry id="0A887668DF40831DA460FD97CD99CE7D" box="[303,468,600,626]" name="United States of America" pageId="1" pageNumber="50">United States</collectingCountry>
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National Museum of Natural History (Washington DC,
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<collectingCountry id="0A887668DF40831DA1C5FD97C886CE7D" box="[1162,1227,600,626]" name="United States of America" pageId="1" pageNumber="50">USA</collectingCountry>
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), Vienna Natural History Museum (Vienna,
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<collectingCountry id="0A887668DF40831DA482FDB2CE65CE98" box="[461,552,637,663]" name="Austria" pageId="1" pageNumber="50">Austria</collectingCountry>
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), Czech National Museum (Prague,
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<collectingCountry id="0A887668DF40831DA682FDB2C8C4CE98" box="[973,1161,637,663]" name="Czech Republic" pageId="1" pageNumber="50">Czech Republic</collectingCountry>
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) and the Field Museum of Natural History (Chicago,
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<collectingCountry id="0A887668DF40831DA4A0FD6DCE40CEB3" box="[495,525,674,700]" name="Iceland" pageId="1" pageNumber="50">IL</collectingCountry>
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,
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<collectingCountry id="0A887668DF40831DA756FD6DCE14CEB3" box="[537,601,674,700]" name="United States of America" pageId="1" pageNumber="50">USA</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3A856573DF408319A58BFD08CDCACEB3" lastPageId="5" lastPageNumber="54" pageId="1" pageNumber="50" type="description">
|
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<paragraph id="722036F8DF40831DA58BFD08C828CF08" blockId="1.[151,1437,263,775]" pageId="1" pageNumber="50">
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<emphasis id="40EBEAEADF40831DA58BFD08CD14CEEE" bold="true" box="[196,345,711,737]" pageId="1" pageNumber="50">Description</emphasis>
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. Body of adults fusiform, elongate, somewhat depressed (
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<figureCitation id="EAA42A7DDF40831DA150FD07C825CEED" box="[1055,1128,712,738]" captionStart="FIGURE 1" captionStartId="1.[151,255,1751,1775]" captionTargetBox="[197,1402,815,1731]" captionTargetId="figure@1.[197,1402,814,1731]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="1" pageNumber="50">Figs 1</figureCitation>
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A–1C); length
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<quantity id="B5679B1DDF40831DA06CFD07CC85CF08" metricMagnitude="-3" metricUnit="m" metricValue="1.16" metricValueMax="1.17" metricValueMin="1.15" pageId="1" pageNumber="50" unit="mm" value="1.16" valueMax="1.17" valueMin="1.15">1.15–1.17 mm</quantity>
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; color generally light brown to dark brown; pubescence golden to light yellow.
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</paragraph>
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<caption id="26E06670DF40831DA5D8F918C8FDCAE0" box="[151,1200,1751,1775]" httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="1" pageNumber="50" targetBox="[197,1402,815,1731]" targetPageId="1">
|
||
<paragraph id="722036F8DF40831DA5D8F918C8FDCAE0" blockId="1.[151,1200,1751,1775]" box="[151,1200,1751,1775]" pageId="1" pageNumber="50">
|
||
<emphasis id="40EBEAEADF40831DA5D8F918CD54CAE0" bold="true" box="[151,281,1751,1775]" pageId="1" pageNumber="50">FIGURE 1.</emphasis>
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Habitus of
|
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<taxonomicName id="B59F4D7BDF40831DA4D9F917CE61CAE0" box="[406,556,1752,1775]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
|
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<emphasis id="40EBEAEADF40831DA4D9F917CE61CAE0" box="[406,556,1752,1775]" italics="true" pageId="1" pageNumber="50">H. perijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40EBEAEADF40831DA77DF918CE23CAE0" bold="true" box="[562,622,1751,1775]" pageId="1" pageNumber="50">
|
||
<taxonomicNameLabel id="5BD85791DF40831DA77DF918CE23CAE0" box="[562,622,1751,1775]" pageId="1" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm.
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</paragraph>
|
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</caption>
|
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<paragraph id="722036F8DF40831EA58BF8D2C926CCF3" blockId="1.[151,1436,1821,2034]" lastBlockId="2.[151,1437,152,252]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
|
||
<emphasis id="40EBEAEADF40831DA58BF8D2CD5CCB38" bold="true" box="[196,273,1821,1847]" pageId="1" pageNumber="50">Head.</emphasis>
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Moderate, wider than long (
|
||
<figureCitation id="EAA42A7DDF40831DA73BF8D2CEF3CB38" box="[628,702,1821,1847]" captionStart="FIGURE 3. H" captionStartId="3.[151,255,887,911]" captionTargetBox="[151,1434,181,865]" captionTargetId="figure@3.[151,1436,181,866]" captionTargetPageId="3" captionText="FIGURE 3. H. perijaensis sp. n. A) head (ventral); B) antenna; C) mesosternum and metaventrite (ventral)." httpUri="https://zenodo.org/record/275693/files/figure.png" pageId="1" pageNumber="50">Fig. 3</figureCitation>
|
||
A), widest behind eyes, prognathus, anteriorly arcuate with pronounced frontal ridge; surface with moderately shallow punctures. Antennae sub-clavate, cylindrical (
|
||
<figureCitation id="EAA42A7DDF40831DA022F88DCCEBCB8D" captionStart="FIGURE 3. H" captionStartId="3.[151,255,887,911]" captionTargetBox="[151,1434,181,865]" captionTargetId="figure@3.[151,1436,181,866]" captionTargetPageId="3" captionText="FIGURE 3. H. perijaensis sp. n. A) head (ventral); B) antenna; C) mesosternum and metaventrite (ventral)." httpUri="https://zenodo.org/record/275693/files/figure.png" pageId="1" pageNumber="50">Fig. 3</figureCitation>
|
||
B), extended beyond anterolateral margins of head, possessing 8 antennomeres, segment VIII enlarged and elongate, longer than antennomeres V-VII combined, antennae inserted beneath the frontal ridge, between eyes and base of mandibles. Labrum well-developed, fused to clypeus, apically arcuate. Mandibles small, apices blunt, bidentate apically, Maxillary palpi prominent, possessing four palpomeres, apical one small, no longer than wide, apex acute; penultimate enlarged and three times as long as apical. Labial palpi possessing three palpomeres, small and obscure. Gular sutures present, widely separated then converging anteriorly. Tentorium well-developed. Eyes small, oval, widely separated, not protuberant, interfacetal setae present.
|
||
</paragraph>
|
||
<caption id="26E06670DF43831EA5D8F81CC94FCBE4" box="[151,1282,2003,2027]" httpUri="https://zenodo.org/record/275692/files/figure.png" pageId="2" pageNumber="51" targetBox="[190,1394,290,1982]" targetPageId="2">
|
||
<paragraph id="722036F8DF43831EA5D8F81CC94FCBE4" blockId="2.[151,1282,2003,2027]" box="[151,1282,2003,2027]" pageId="2" pageNumber="51">
|
||
<emphasis id="40EBEAEADF43831EA5D8F81CCD54CBE4" bold="true" box="[151,281,2003,2027]" pageId="2" pageNumber="51">FIGURE 2.</emphasis>
|
||
<taxonomicName id="B59F4D7BDF43831EA46FF81BCDF8CBE4" box="[288,437,2004,2027]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
|
||
<emphasis id="40EBEAEADF43831EA46FF81BCDF8CBE4" box="[288,437,2004,2027]" italics="true" pageId="2" pageNumber="51">H. perijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40EBEAEADF43831EA4F3F81CCDB5CBE4" bold="true" box="[444,504,2003,2027]" pageId="2" pageNumber="51">
|
||
<taxonomicNameLabel id="5BD85791DF43831EA4F3F81CCDB5CBE4" box="[444,504,2003,2027]" pageId="2" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) head and pronotum (dorsal); B) elytron (dorsal); C) elytron (ventral).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26E06670DF42831FA5D8FCB8C954CF81" box="[151,1305,887,911]" httpUri="https://zenodo.org/record/275693/files/figure.png" pageId="3" pageNumber="52" targetBox="[151,1434,181,865]" targetPageId="3">
|
||
<paragraph id="722036F8DF42831FA5D8FCB8C954CF81" blockId="3.[151,1305,887,911]" box="[151,1305,887,911]" pageId="3" pageNumber="52">
|
||
<emphasis id="40EBEAEADF42831FA5D8FCB8CD54CF80" bold="true" box="[151,281,887,911]" pageId="3" pageNumber="52">FIGURE 3.</emphasis>
|
||
<taxonomicName id="B59F4D7BDF42831FA46FFCB8CDF8CF81" box="[288,437,887,910]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
|
||
<emphasis id="40EBEAEADF42831FA46FFCB8CDF8CF81" box="[288,437,887,910]" italics="true" pageId="3" pageNumber="52">H. perijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40EBEAEADF42831FA4F3FCB8CDB5CF80" bold="true" box="[444,504,887,911]" pageId="3" pageNumber="52">
|
||
<taxonomicNameLabel id="5BD85791DF42831FA4F3FCB8CDB5CF80" box="[444,504,887,911]" pageId="3" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) head (ventral); B) antenna; C) mesosternum and metaventrite (ventral).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26E06670DF42831FA5D8F81EC929CBE7" box="[151,1380,2001,2025]" httpUri="https://zenodo.org/record/275694/files/figure.png" pageId="3" pageNumber="52" targetBox="[185,1409,948,1979]" targetPageId="3">
|
||
<paragraph id="722036F8DF42831FA5D8F81EC929CBE7" blockId="3.[151,1380,2001,2025]" box="[151,1380,2001,2025]" pageId="3" pageNumber="52">
|
||
<emphasis id="40EBEAEADF42831FA5D8F81ECD54CBE6" bold="true" box="[151,281,2001,2025]" pageId="3" pageNumber="52">FIGURE 4.</emphasis>
|
||
<taxonomicName id="B59F4D7BDF42831FA46FF81DCDF8CBE6" box="[288,437,2002,2025]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
|
||
<emphasis id="40EBEAEADF42831FA46FF81DCDF8CBE6" box="[288,437,2002,2025]" italics="true" pageId="3" pageNumber="52">H. perijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40EBEAEADF42831FA4F3F81ECDB5CBE6" bold="true" box="[444,504,2001,2025]" pageId="3" pageNumber="52">
|
||
<taxonomicNameLabel id="5BD85791DF42831FA4F3F81ECDB5CBE6" box="[444,504,2001,2025]" pageId="3" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) terminal abdominal segments; female; B) terminal abdominal segments, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26E06670DF458318A5D8F980CF30CA88" httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="4" pageNumber="53" targetBox="[171,1398,198,1614]" targetPageId="4">
|
||
<paragraph id="722036F8DF458318A5D8F980CF30CA88" blockId="4.[151,1436,1615,1671]" pageId="4" pageNumber="53">
|
||
<emphasis id="40EBEAEADF458318A5D8F980CD52CA68" bold="true" box="[151,287,1615,1639]" pageId="4" pageNumber="53">FIGURE 5.</emphasis>
|
||
<taxonomicName id="B59F4D7BDF458318A468F99FCD88CA68" box="[295,453,1616,1639]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="perijaensis" status="sp. nov.">
|
||
<emphasis id="40EBEAEADF458318A468F99FCD88CA68" box="[295,453,1616,1639]" italics="true" pageId="4" pageNumber="53">H. perijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40EBEAEADF458318A483F980CE40CA68" bold="true" box="[460,525,1615,1639]" pageId="4" pageNumber="53">
|
||
<taxonomicNameLabel id="5BD85791DF458318A483F980CE40CA68" box="[460,525,1615,1639]" pageId="4" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="722036F8DF458318A58BF960C930CB35" blockId="4.[151,1436,1711,2036]" pageId="4" pageNumber="53">
|
||
<emphasis id="40EBEAEADF458318A58BF960CD07CAC6" bold="true" box="[196,330,1711,1737]" pageId="4" pageNumber="53">Pronotum.</emphasis>
|
||
Broader than head (
|
||
<figureCitation id="EAA42A7DDF458318A776F97FCECDCAC5" box="[569,640,1712,1738]" captionStart="FIGURE 1" captionStartId="1.[151,255,1751,1775]" captionTargetBox="[197,1402,815,1731]" captionTargetId="figure@1.[197,1402,814,1731]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="4" pageNumber="53">Figs 1</figureCitation>
|
||
A–1B, 2A), widest at hind angles, nearly as wide as elytra; anterior margin excavated to receive head; lateral margins arcuate, sides deeply explanate, posterior margin arcuate, distance between hind angles twice the length of pronotum anterior to hind margins medially. Prosternum short, spinasternum depressed, obscure. Procoxal cavities opened posteriorly, procoxae conical, prominent.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF458318A589F88ACF5BCB8B" blockId="4.[151,1436,1711,2036]" pageId="4" pageNumber="53">
|
||
<emphasis id="40EBEAEADF458318A589F88ACD35CB50" bold="true" box="[198,376,1861,1887]" pageId="4" pageNumber="53">Mesosternum.</emphasis>
|
||
Short, mesocoxal cavities widely separated, mesocoxae small, ovate; mesosternal disc wider than long, separating mesocoxae (
|
||
<figureCitation id="EAA42A7DDF458318A720F8A5CEF0CB8B" box="[623,701,1898,1924]" captionStart="FIGURE 1" captionStartId="1.[151,255,1751,1775]" captionTargetBox="[197,1402,815,1731]" captionTargetId="figure@1.[197,1402,814,1731]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="4" pageNumber="53">Figs. 1</figureCitation>
|
||
B, 3C).
|
||
</paragraph>
|
||
<paragraph id="722036F8DF458318A58BF85FCE06CBFB" blockId="4.[151,1436,1711,2036]" pageId="4" pageNumber="53">
|
||
Metaventrite long (
|
||
<figureCitation id="EAA42A7DDF458318A4E6F85FCDBCCBA5" box="[425,497,1936,1962]" captionStart="FIGURE 1" captionStartId="1.[151,255,1751,1775]" captionTargetBox="[197,1402,815,1731]" captionTargetId="figure@1.[197,1402,814,1731]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="4" pageNumber="53">Figs 1</figureCitation>
|
||
B, 3C), widest at hind angles, metacoxal cavities widely separated, metacoxae transverse, posteriorly enlarged and forming a coxal plate partially covering hind femora. Trochanters small, triangular. Femora and tibiae normal.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFF58CEA5CCBD" blockId="5.[151,1437,151,1298]" box="[196,744,151,178]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFF58CD04CCBE" bold="true" box="[196,329,151,177]" pageId="5" pageNumber="54">Scutellum.</emphasis>
|
||
Acute posteriorly, wider than long.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFF72CCB6CD2D" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFF72CD56CCD8" bold="true" box="[196,283,189,215]" pageId="5" pageNumber="54">Elytra.</emphasis>
|
||
Truncate, exposing abdominal tergites, elytron longer (
|
||
<quantity id="B5679B1DDF448319A6E2FF72C857CCD8" box="[941,1050,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="54" unit="mm" value="0.6">0.60 mm</quantity>
|
||
) than wide (
|
||
<quantity id="B5679B1DDF448319A1E2FF72C950CCD8" box="[1197,1309,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="54" unit="mm" value="0.37">0.37 mm</quantity>
|
||
), widest at anterior half; setae moderately flat, setal length two-three times distance between elytral punctures (
|
||
<figureCitation id="EAA42A7DDF448319A07BFF2DC936CCF3" box="[1332,1403,226,252]" captionStart="FIGURE 1" captionStartId="1.[151,255,1751,1775]" captionTargetBox="[197,1402,815,1731]" captionTargetId="figure@1.[197,1402,814,1731]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of H. perijaensis sp. n.. A) dorsal; B) ventral; C) lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/275691/files/figure.png" pageId="5" pageNumber="54">Figs 1</figureCitation>
|
||
A, 2B–2C).
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFEE2C84FCD63" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFEE2CD12CD48" bold="true" box="[196,351,301,327]" pageId="5" pageNumber="54">
|
||
<collectingCountry id="0A887668DF448319A58BFEE2CD48CD48" box="[196,261,301,327]" name="India" pageId="5" pageNumber="54">Hind</collectingCountry>
|
||
wings.
|
||
</emphasis>
|
||
Present, fringe of setae along margin of wing membrane, setae longest (
|
||
<quantity id="B5679B1DDF448319A197FEE2C904CD48" box="[1240,1353,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="54" unit="mm" value="0.22">0.22 mm</quantity>
|
||
) along distal margin of wing, width of wing membrane widest (
|
||
<quantity id="B5679B1DDF448319A664FE9DCFDACD63" box="[811,919,338,364]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="54" unit="mm" value="0.41">0.41 mm</quantity>
|
||
) distally.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFEB8CE7DCDD3" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFEB8CD46CD9E" bold="true" box="[196,267,375,401]" pageId="5" pageNumber="54">Tarsi.</emphasis>
|
||
3–3–3; claws moderate, wider at base than distally, males possessing an oblong disc at base of protarsal claws; meso- and metatarsal claws of male subequal, the outer claw possessing an invagination medially, anterioly to base of claw.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFE28CDCACEB3" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFE28CD04CE0E" bold="true" box="[196,329,487,513]" pageId="5" pageNumber="54">Abdomen.</emphasis>
|
||
Narrowed posteriorly, possessing a dense mat of recumbent setae on tergite IV, forming a plastron beneath elytra; tergite VIII of female rounded apically (
|
||
<figureCitation id="EAA42A7DDF448319A69FFDC2C850CE28" box="[976,1053,525,551]" captionStart="FIGURE 5. H" captionStartId="4.[151,258,1615,1639]" captionTargetBox="[171,1398,198,1614]" captionTargetId="figure@4.[151,1436,176,1614]" captionTargetPageId="4" captionText="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="5" pageNumber="54">Fig. 5</figureCitation>
|
||
A), terminal segments tapered posteriorly, acute, bifid (
|
||
<figureCitation id="EAA42A7DDF448319A49AFDFDCE6FCE43" box="[469,546,562,588]" captionStart="FIGURE 5. H" captionStartId="4.[151,258,1615,1639]" captionTargetBox="[171,1398,198,1614]" captionTargetId="figure@4.[151,1436,176,1614]" captionTargetPageId="4" captionText="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="5" pageNumber="54">Fig. 5</figureCitation>
|
||
C); abdominal sternite VIII of male possessing two widely separated, moderately acute lobes along posterior margin (
|
||
<figureCitation id="EAA42A7DDF448319A78CFD97CF45CE7D" box="[707,776,600,626]" captionStart="FIGURE 5. H" captionStartId="4.[151,258,1615,1639]" captionTargetBox="[171,1398,198,1614]" captionTargetId="figure@4.[151,1436,176,1614]" captionTargetPageId="4" captionText="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="5" pageNumber="54">Fig. 5</figureCitation>
|
||
B), sternite IX not tapered, hind margin arcuate medially with hind angles widely separated (
|
||
<figureCitation id="EAA42A7DDF448319A773FDB2CECECE98" box="[572,643,637,663]" captionStart="FIGURE 5. H" captionStartId="4.[151,258,1615,1639]" captionTargetBox="[171,1398,198,1614]" captionTargetId="figure@4.[151,1436,176,1614]" captionTargetPageId="4" captionText="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="5" pageNumber="54">Fig. 5</figureCitation>
|
||
D). Aedeagus
|
||
<quantity id="B5679B1DDF448319A662FDB2CFDACE98" box="[813,919,637,663]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="54" unit="mm" value="0.3">0.30 mm</quantity>
|
||
long, acute posteriorly, widest anteriorly at basal bulb (
|
||
<figureCitation id="EAA42A7DDF448319A451FD6DCD29CEB3" box="[286,356,674,700]" captionStart="FIGURE 5. H" captionStartId="4.[151,258,1615,1639]" captionTargetBox="[171,1398,198,1614]" captionTargetId="figure@4.[151,1436,176,1614]" captionTargetPageId="4" captionText="FIGURE 5. H. perijaensis sp. n. A) tergite VIII, female; B) sternite VIII, male; C) terminal abdominal segments, female; D) terminal abdominal segments, male; E) aedeagus (lateral)." httpUri="https://zenodo.org/record/275695/files/figure.png" pageId="5" pageNumber="54">Fig. 5</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A856573DF448319A58BFD08CFFFCF08" pageId="5" pageNumber="54" type="distribution">
|
||
<paragraph id="722036F8DF448319A58BFD08CFFFCF08" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFD08CD29CEEE" bold="true" box="[196,356,711,737]" pageId="5" pageNumber="54">Distribution.</emphasis>
|
||
Currently known from two sites located less than
|
||
<quantity id="B5679B1DDF448319A6F9FD07CFBCCEED" box="[950,1009,712,738]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="54" unit="km" value="1.0">1 km</quantity>
|
||
apart in the foothills of the Serrania de Perija near Toromo, along the northwestern border of
|
||
<collectingCountry id="0A887668DF448319A67EFD22CFE0CF08" box="[817,941,749,775]" name="Venezuela" pageId="5" pageNumber="54">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A856573DF448319A58BFCDDC86BC91D" pageId="5" pageNumber="54" type="etymology">
|
||
<paragraph id="722036F8DF448319A58BFCDDCF8ECF23" blockId="5.[151,1437,151,1298]" box="[196,963,786,812]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFCDDCD1CCF23" bold="true" box="[196,337,786,812]" pageId="5" pageNumber="54">Etymology.</emphasis>
|
||
The specific epithet is derived from the
|
||
<typeStatus id="AD24885ADF448319A663FCDDCF13CF23" box="[812,862,786,812]" pageId="5" pageNumber="54">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFCF8CE2CC8C8" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">
|
||
<emphasis id="40EBEAEADF448319A58BFCF8CD0BCF5E" bold="true" box="[196,326,823,849]" pageId="5" pageNumber="54">Bionomics</emphasis>
|
||
. The species was first encountered in the wet season (September) living in a seepage flowing over exposed, sunlit rock adjacent to a small stream (
|
||
<figureCitation id="EAA42A7DDF448319A64BFC92CF01CF78" box="[772,844,861,887]" captionStart="FIGURE 6" captionStartId="6.[151,256,1906,1930]" captionTargetBox="[151,1436,264,1883]" captionTargetId="figure@6.[151,1436,264,1884]" captionTargetPageId="6" captionText="FIGURE 6. Type locality of H. perijaensis sp. n., Venezuela: Zulia State, edge of Perija National Park at Toromo. A) Seepage habitat at type locality on 23 September 2007 (wet season); B) overview of the type locality on 28 January 2009 (dry season), and C) 23 September 2007." httpUri="https://zenodo.org/record/275696/files/figure.png" pageId="5" pageNumber="54">Fig. 6</figureCitation>
|
||
). The same site was visited in the early dry season (January) at which time the seepages were completely dry. However, the small stream was now reduced to a small trickle with areas mimicking the now-dry side seepage in which many specimens of the new species were found, particularly in clumps of moss. The species was also found in a similar exposed seepage habitat in a second stream less than
|
||
<quantity id="B5679B1DDF448319A4B4FC3DCE77C803" box="[507,570,1010,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="54" unit="km" value="1.0">1 km</quantity>
|
||
west of the first site. No specimens were found in other co-occurring habitats such as sandy stream margins or leaf packs. Over the last several years a number of Venezuelan rock seepage habitats have been examined by AEZS and several Venezuelan colleagues including sites in the primary Venezuelan Andes, the Cordillera de la Costa, and in
|
||
<collectingCountry id="0A887668DF448319A6C3FBADCFAEC873" box="[908,995,1122,1148]" name="Guyana" pageId="5" pageNumber="54">Guiana</collectingCountry>
|
||
Shield-associated outcrops. Despite these efforts no other
|
||
<taxonomicName id="B59F4D7BDF448319A4DAFB47CE7FC8AE" box="[405,562,1160,1185]" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40EBEAEADF448319A4DAFB47CE7FC8AE" box="[405,562,1160,1185]" italics="true" pageId="5" pageNumber="54">Hydroscapha</emphasis>
|
||
</taxonomicName>
|
||
specimens have yet been found, suggesting the genus is not widespread or at least highly localized in distribution.
|
||
</paragraph>
|
||
<paragraph id="722036F8DF448319A58BFB1DC86BC91D" blockId="5.[151,1437,151,1298]" pageId="5" pageNumber="54">Numerous females were found to contain one to two nearly-fully developed eggs in specimens collected in the wet season (dry season specimens were not examined for this purpose).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |