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<document ID-DOI="10.11646/zootaxa.5277.1.6" ID-GBIF-Dataset="11ad2e19-0175-4b89-8b57-0192c1437803" ID-ISSN="1175-5326" ID-Zenodo-Dep="7893034" ID-ZooBank="CD726173-9C2E-4E33-91BF-DD3D47A791D6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682931241685" checkinUser="plazi" docAuthor="Tan, Ming Kai, Muhammad, Amira Aqilah &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2023" docId="FA3587B43C3CFFC8FF32F9549190FE2F" docLanguage="en" docName="zootaxa.5277.1.6.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pendleburyella eirmosa Tan &amp; Muhammad &amp; Wahab 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="144" masterDocId="060CFFCC3C37FFC5FFA5FF989365FFFE" masterDocTitle="The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae)" masterLastPageNumber="148" masterPageNumber="131" pageNumber="142" updateTime="1683132538267" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The taxonomy and bioacoustics of the elusive crickets from the genus Pendleburyella Chopard, 1969 (Gryllidae: Pentacentrinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tan, Ming Kai</mods:namePart>
<mods:affiliation>Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Muhammad, Amira Aqilah</mods:namePart>
<mods:affiliation>Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur 50603, Malaysia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation>Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
<mods:nameIdentifier type="email">rodzay.wahab@ubd.edu.bn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5277</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>148</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.1.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5277.1.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">11ad2e19-0175-4b89-8b57-0192c1437803</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7893034</mods:identifier>
<mods:identifier type="ZooBank">CD726173-9C2E-4E33-91BF-DD3D47A791D6</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7893058" ID-Zenodo-Dep="7893058" LSID="urn:lsid:plazi:treatment:FA3587B43C3CFFC8FF32F9549190FE2F" httpUri="http://treatment.plazi.org/id/FA3587B43C3CFFC8FF32F9549190FE2F" lastPageId="13" lastPageNumber="144" pageId="11" pageNumber="142">
<subSubSection box="[151,538,1740,1767]" pageId="11" pageNumber="142" type="nomenclature">
<paragraph blockId="11.[151,603,1740,1803]" box="[151,538,1740,1767]" pageId="11" pageNumber="142">
<heading bold="true" box="[151,538,1740,1767]" fontSize="11" level="1" pageId="11" pageNumber="142" reason="1">
<emphasis bold="true" box="[151,538,1740,1767]" pageId="11" pageNumber="142">
<taxonomicName authority="Tan &amp; Muhammad &amp; Wahab, 2023" authorityName="Tan &amp; Muhammad &amp; Wahab" authorityYear="2023" box="[151,437,1740,1767]" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="142" phylum="Arthropoda" rank="species" species="eirmosa" status="sp. nov.">
<emphasis bold="true" box="[151,437,1740,1767]" italics="true" pageId="11" pageNumber="142">Pendleburyella eirmosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[444,538,1740,1766]" pageId="11" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,603,1776,1803]" pageId="11" pageNumber="142" type="description">
<paragraph blockId="11.[151,603,1740,1803]" box="[151,603,1776,1803]" pageId="11" pageNumber="142">
(
<figureCitation box="[159,237,1776,1802]" captionStart="FIGURE 1" captionStartId="1.[151,250,1061,1085]" captionTargetBox="[152,1436,181,1037]" captionTargetId="figure-19@1.[152,1436,181,1037]" captionTargetPageId="1" captionText="FIGURE 1. Pendleburyella J adult among the leaf litter in Kuala Belalong Field Studies Centre, Brunei Darussalam. The specimen was uncollected but identified as presumably Pendleburyella eirmosa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7893038" httpUri="https://zenodo.org/record/7893038/files/figure.png" pageId="11" pageNumber="142">Figs. 1</figureCitation>
,
<figureCitation box="[251,285,1776,1803]" captionStart="FIGURE 5" captionStartId="5.[151,250,2007,2031]" captionTargetBox="[256,1320,866,1970]" captionTargetId="figure-36@5.[203,1384,824,1983]" captionTargetPageId="5" captionText="FIGURE 5. Pendleburyella pimela sp. nov. (A) and Pendleburyella eirmosa sp. nov. (B) J dorsal habitus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7893054" httpUri="https://zenodo.org/record/7893054/files/figure.png" pageId="11" pageNumber="142">5B</figureCitation>
,
<figureCitation box="[297,423,1776,1802]" captionStart="FIGURE 6" captionStartId="6.[151,250,1905,1929]" captionTargetBox="[170,1389,723,1865]" captionTargetId="figure-238@6.[157,1430,705,1882]" captionTargetPageId="6" captionText="FIGURE 6. Pendleburyella J morphology: head and pronotum in dorsal view (A, B), face in anterior view (C, D), body in lateral view (E, F). Pendleburyella pimela sp. nov. (A, C, E); Pendleburyella eirmosa sp. nov. (B, D, F). Scale bars: 1 mm (A–D), 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.7893056" httpUri="https://zenodo.org/record/7893056/files/figure.png" pageId="11" pageNumber="142">6B, 6D, 6F</figureCitation>
,
<figureCitation box="[434,470,1777,1803]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[161,1427,181,1918]" captionTargetId="figure-23@8.[161,1427,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Pendleburyella J head and pronotum in lateral view: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella sp. 1 (E), Pendleburyella sp. 2 (F). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893060" httpUri="https://zenodo.org/record/7893060/files/figure.png" pageId="11" pageNumber="142">7D</figureCitation>
,
<figureCitation box="[482,518,1776,1802]" captionStart="FIGURE 8" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[212,1359,543,1876]" captionTargetId="figure-196@9.[195,1393,524,1910]" captionTargetPageId="9" captionText="FIGURE 8. Pendleburyella J FWs: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella “intermedia” (E), Pendleburyella sp. 1 (F), Pendleburyella sp. 2 (G). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893064" httpUri="https://zenodo.org/record/7893064/files/figure.png" pageId="11" pageNumber="142">8D</figureCitation>
,
<figureCitation box="[530,595,1776,1802]" captionStart="FIGURE 9" captionStartId="10.[151,250,1721,1745]" captionTargetBox="[151,1436,181,1697]" captionTargetId="figure-23@10.[151,1436,181,1697]" captionTargetPageId="10" captionText="FIGURE 9. Pendleburyella J genitalia: Pendleburyella pimela sp. nov. (A–C) and Pendleburyella eirmosa sp. nov. (D–F) in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893066" httpUri="https://zenodo.org/record/7893066/files/figure.png" pageId="11" pageNumber="142">9D–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="142" type="diagnosis">
<paragraph blockId="11.[151,1436,1841,2012]" pageId="11" pageNumber="142">
<emphasis bold="true" box="[151,274,1841,1867]" pageId="11" pageNumber="142">Diagnosis.</emphasis>
This species is characterised by a medium-sized habitus (similar to
<taxonomicName authorityName="Chopard" authorityYear="1969" box="[1098,1373,1842,1868]" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="142" phylum="Arthropoda" rank="species" species="testacea">
<emphasis box="[1098,1373,1842,1868]" italics="true" pageId="11" pageNumber="142">Pendleburyella testacea</emphasis>
</taxonomicName>
), the maxillary palps with the apical segment also somewhat rectangular (similar to that of
<taxonomicName authorityName="Chopard" authorityYear="1969" box="[1142,1391,1878,1904]" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="142" phylum="Arthropoda" rank="species" species="vicina">
<emphasis box="[1142,1391,1878,1904]" italics="true" pageId="11" pageNumber="142">Pendleburyella vicina</emphasis>
</taxonomicName>
but stouter); the mirror longer than broad, the dividing vein of the mirror broadly and roundly curved at the basal end; the apical part of FW longer relative to the FWL. The genitalia also differs from that of
<taxonomicName authority="Tan &amp; Muhammad &amp; Wahab, 2023" authorityName="Tan &amp; Muhammad &amp; Wahab" authorityYear="2023" box="[1141,1395,1950,1976]" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="142" phylum="Arthropoda" rank="species" species="pimela" status="sp. nov.">
<emphasis box="[1141,1395,1950,1976]" italics="true" pageId="11" pageNumber="142">Pendleburyella pimela</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="11" pageNumber="142">
<taxonomicNameLabel pageId="11" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see the diagnosis for the latter species).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7893072" ID-Zenodo-Dep="7893072" httpUri="https://zenodo.org/record/7893072/files/figure.png" pageId="12" pageNumber="143" startId="12.[151,250,1098,1122]" targetBox="[156,1436,188,1073]" targetPageId="12" targetType="figure">
<paragraph blockId="12.[151,1436,1098,1159]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[151,290,1098,1122]" pageId="12" pageNumber="143">FIGURE 11.</emphasis>
<taxonomicName authority="Tan &amp; Muhammad &amp; Wahab, 2023" authorityName="Tan &amp; Muhammad &amp; Wahab" authorityYear="2023" box="[299,533,1098,1122]" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="143" phylum="Arthropoda" rank="species" species="pimela" status="sp. nov.">
<emphasis box="[299,533,1098,1122]" italics="true" pageId="12" pageNumber="143">Pendleburyella pimela</emphasis>
</taxonomicName>
<emphasis bold="true" box="[542,656,1098,1122]" pageId="12" pageNumber="143">
<taxonomicNameLabel box="[542,626,1098,1122]" pageId="12" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
♂
</emphasis>
calling song: oscillograms showing a continuous trill (A) and a sequence showing ten complete syllables (B); spectrogram of the same sequence (C).
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="143" type="etymology">
<paragraph blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[199,334,1205,1231]" pageId="12" pageNumber="143">Etymology.</emphasis>
The species name refers to the broadly rounded emargination between the dorsal and ventral processes of the pseudepiphallic paramere (distinguished from the narrowly angular emargination in
<taxonomicName authority="Tan &amp; Muhammad &amp; Wahab, 2023" authorityName="Tan &amp; Muhammad &amp; Wahab" authorityYear="2023" class="Insecta" family="Gryllidae" genus="Pendleburyella" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="143" phylum="Arthropoda" rank="species" species="pimela" status="sp. nov.">
<emphasis italics="true" pageId="12" pageNumber="143">Pendleburyella pimela</emphasis>
</taxonomicName>
<emphasis bold="true" box="[234,324,1277,1303]" pageId="12" pageNumber="143">
<taxonomicNameLabel box="[234,324,1277,1303]" pageId="12" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); eirmós = continuity in Ancient Greek.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="143" type="materials_examined">
<paragraph blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[199,430,1312,1338]" pageId="12" pageNumber="143">Material examined.</emphasis>
<materialsCitation collectingDate="2019-03-01" collectionCode="UBDM" collectorName="M. K. Tan &amp; H. Yeo" country="Brunei" elevation="5" latitude="4.34186" location="Wasai Wong Kadir Recreational Park" longLatPrecision="1" longitude="114.44611" pageId="12" pageNumber="143" specimenCount="1" stateProvince="Belait" typeStatus="holotype">
<typeStatus box="[434,543,1312,1338]" pageId="12" pageNumber="143">Holotype</typeStatus>
:
<collectingCountry box="[551,852,1313,1339]" name="Brunei" pageId="12" pageNumber="143">BRUNEI DARUSSALAM</collectingCountry>
• 1♁;
<collectingRegion box="[918,1076,1312,1338]" country="Brunei" name="Belait" pageId="12" pageNumber="143">Belait District</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:FA3587B43C3CFFC8FF32F9549190FE2F:774360793C3BFFC9FB99FAB893ABFAA0" country="Brunei" latitude="4.34186" longLatPrecision="1" longitude="114.44611" name="Wasai Wong Kadir Recreational Park" pageId="12" pageNumber="143" stateProvince="Belait">Wasai Wong Kadir Recreational Park</location>
;
<geoCoordinate box="[218,332,1348,1374]" degrees="4.34186" direction="north" orientation="latitude" pageId="12" pageNumber="143" precision="1" value="4.34186">N4.34186</geoCoordinate>
,
<geoCoordinate box="[344,481,1348,1374]" degrees="114.44611" direction="east" orientation="longitude" pageId="12" pageNumber="143" precision="1" value="114.44611">E114.44611</geoCoordinate>
, 35.8±
<quantity box="[556,628,1348,1375]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="143" unit="m" value="5.2">
<elevation box="[556,628,1348,1375]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="143" unit="m" value="5.2">5.2 m</elevation>
</quantity>
.a.s.l.;
<date box="[695,856,1348,1375]" pageId="12" pageNumber="143" value="2019-03-01">
<collectingDate box="[695,856,1348,1375]" pageId="12" pageNumber="143" value="2019-03-01">1 March 2019</collectingDate>
</date>
, 18h54; calling on a leaf; coll.
<collectorName box="[1206,1314,1349,1375]" pageId="12" pageNumber="143">M.K. Tan</collectorName>
&amp;
<collectorName box="[1351,1433,1349,1375]" pageId="12" pageNumber="143">H. Yeo</collectorName>
; BRU.19.47 (
<collectionCode box="[293,380,1385,1411]" pageId="12" pageNumber="143">UBDM</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[199,472,1420,1447]" pageId="12" pageNumber="143">Photograph examined.</emphasis>
<materialsCitation collectingDate="2017-07-26" collectorName="M. K. Tan" country="Brunei" elevation="7" latitude="4.54619" location="Ashton Trail" longLatPrecision="1" longitude="115.1569" municipality="Kuala Belalong Field Studies Centre" pageId="12" pageNumber="143" specimenCount="1" stateProvince="Temburong">
<collectingCountry box="[480,786,1421,1447]" name="Brunei" pageId="12" pageNumber="143">BRUNEI DARUSSALAM</collectingCountry>
• 1♁;
<collectingRegion box="[865,1089,1420,1446]" country="Brunei" name="Temburong" pageId="12" pageNumber="143">Temburong District</collectingRegion>
,
<collectingMunicipality pageId="12" pageNumber="143">Kuala Belalong Field Studies Centre</collectingMunicipality>
, along
<location LSID="urn:lsid:plazi:treatment:FA3587B43C3CFFC8FF32F9549190FE2F:774360793C3BFFC9FE88FA2892DFFA34" box="[301,442,1456,1482]" country="Brunei" latitude="4.54619" longLatPrecision="1" longitude="115.1569" municipality="Kuala Belalong Field Studies Centre" name="Ashton Trail" pageId="12" pageNumber="143" stateProvince="Temburong">Ashton Trail</location>
;
<geoCoordinate box="[450,565,1456,1483]" degrees="4.54619" direction="north" orientation="latitude" pageId="12" pageNumber="143" precision="1" value="4.54619">N4.54619</geoCoordinate>
,
<geoCoordinate box="[572,710,1456,1483]" degrees="115.15690" direction="east" orientation="longitude" pageId="12" pageNumber="143" precision="1" value="115.1569">E115.15690</geoCoordinate>
, 110.8±
<quantity box="[795,863,1456,1482]" metricMagnitude="0" metricUnit="m" metricValue="6.9" pageId="12" pageNumber="143" unit="m" value="6.9">
<elevation box="[795,863,1456,1482]" metricMagnitude="0" metricUnit="m" metricValue="6.9" pageId="12" pageNumber="143" unit="m" value="6.9">6.9 m</elevation>
</quantity>
.a.s.l.;
<date box="[927,1068,1456,1483]" pageId="12" pageNumber="143" value="2017-07-26">
<collectingDate box="[927,1068,1456,1483]" pageId="12" pageNumber="143" value="2017-07-26">26 July2017</collectingDate>
</date>
, dusk; on leaf litter; photographed
<collectorName box="[151,259,1493,1519]" pageId="12" pageNumber="143">M.K. Tan</collectorName>
(
<figureCitation box="[274,339,1492,1519]" captionStart="FIGURE 1" captionStartId="1.[151,250,1061,1085]" captionTargetBox="[152,1436,181,1037]" captionTargetId="figure-19@1.[152,1436,181,1037]" captionTargetPageId="1" captionText="FIGURE 1. Pendleburyella J adult among the leaf litter in Kuala Belalong Field Studies Centre, Brunei Darussalam. The specimen was uncollected but identified as presumably Pendleburyella eirmosa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7893038" httpUri="https://zenodo.org/record/7893038/files/figure.png" pageId="12" pageNumber="143">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="144" pageId="12" pageNumber="143" type="description">
<paragraph blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[199,345,1528,1554]" pageId="12" pageNumber="143">Description.</emphasis>
Medium-sized species for this genus. Maxillary palpi with apical segment stout and cylindrical to somewhat rectangular; with subapical segment very stout, distinctly shorter than apical and third segments (
<figureCitation captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[161,1427,181,1918]" captionTargetId="figure-23@8.[161,1427,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Pendleburyella J head and pronotum in lateral view: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella sp. 1 (E), Pendleburyella sp. 2 (F). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893060" httpUri="https://zenodo.org/record/7893060/files/figure.png" pageId="12" pageNumber="143">Fig. 7D</figureCitation>
). Pronotal disc brown, about 1.7 times as wide as long, widening posteriorly (posterior margin about 1.6 times as wide as anterior margin); anterior margin of disc somewhat straight; posterior margin of disc slightly convex (
<figureCitation captionStart="FIGURE 6" captionStartId="6.[151,250,1905,1929]" captionTargetBox="[170,1389,723,1865]" captionTargetId="figure-238@6.[157,1430,705,1882]" captionTargetPageId="6" captionText="FIGURE 6. Pendleburyella J morphology: head and pronotum in dorsal view (A, B), face in anterior view (C, D), body in lateral view (E, F). Pendleburyella pimela sp. nov. (A, C, E); Pendleburyella eirmosa sp. nov. (B, D, F). Scale bars: 1 mm (A–D), 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.7893056" httpUri="https://zenodo.org/record/7893056/files/figure.png" pageId="12" pageNumber="143">Fig. 6B</figureCitation>
). Pronotal lateral lobe 2.0 times as long as high (
<figureCitation box="[733,824,1672,1699]" captionStart="FIGURE 7" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[161,1427,181,1918]" captionTargetId="figure-23@8.[161,1427,181,1919]" captionTargetPageId="8" captionText="FIGURE 7. Pendleburyella J head and pronotum in lateral view: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella sp. 1 (E), Pendleburyella sp. 2 (F). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893060" httpUri="https://zenodo.org/record/7893060/files/figure.png" pageId="12" pageNumber="143">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="143">
<emphasis bold="true" box="[199,267,1709,1735]" pageId="12" pageNumber="143">Male.</emphasis>
FW 2.6 times as long as broad (
<figureCitation box="[616,704,1708,1735]" captionStart="FIGURE 8" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[212,1359,543,1876]" captionTargetId="figure-196@9.[195,1393,524,1910]" captionTargetPageId="9" captionText="FIGURE 8. Pendleburyella J FWs: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella “intermedia” (E), Pendleburyella sp. 1 (F), Pendleburyella sp. 2 (G). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893064" httpUri="https://zenodo.org/record/7893064/files/figure.png" pageId="12" pageNumber="143">Fig. 8D</figureCitation>
), covering abdomen and slightly surpassing apex of FIII. Venation (
<figureCitation box="[159,253,1744,1771]" captionStart="FIGURE 8" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[212,1359,543,1876]" captionTargetId="figure-196@9.[195,1393,524,1910]" captionTargetPageId="9" captionText="FIGURE 8. Pendleburyella J FWs: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella “intermedia” (E), Pendleburyella sp. 1 (F), Pendleburyella sp. 2 (G). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893064" httpUri="https://zenodo.org/record/7893064/files/figure.png" pageId="12" pageNumber="143">Fig. 8D</figureCitation>
): 1A vein transverse, faintly sinuous; diagonal faintly bent, with 3 distinct oblique veins in harp area; posterior two oblique veins parallel to each other, anterior most vein distinctly shorter and running nearly along length of FW. Mirror about 1.2 times as long as wide, dividing vein broadly curved at basal end. Lateral field around 7 branches on Sc (
<figureCitation box="[357,442,1852,1878]" captionStart="FIGURE 6" captionStartId="6.[151,250,1905,1929]" captionTargetBox="[170,1389,723,1865]" captionTargetId="figure-238@6.[157,1430,705,1882]" captionTargetPageId="6" captionText="FIGURE 6. Pendleburyella J morphology: head and pronotum in dorsal view (A, B), face in anterior view (C, D), body in lateral view (E, F). Pendleburyella pimela sp. nov. (A, C, E); Pendleburyella eirmosa sp. nov. (B, D, F). Scale bars: 1 mm (A–D), 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.7893056" httpUri="https://zenodo.org/record/7893056/files/figure.png" pageId="12" pageNumber="143">Fig. 6F</figureCitation>
). Apical field 0.11 times as long as FWL (
<figureCitation box="[914,1004,1852,1879]" captionStart="FIGURE 8" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[212,1359,543,1876]" captionTargetId="figure-196@9.[195,1393,524,1910]" captionTargetPageId="9" captionText="FIGURE 8. Pendleburyella J FWs: Pendleburyella testacea (A), Pendleburyella vicina (B), Pendleburyella pimela sp. nov. (C), Pendleburyella eirmosa sp. nov. (D), Pendleburyella “intermedia” (E), Pendleburyella sp. 1 (F), Pendleburyella sp. 2 (G). Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893064" httpUri="https://zenodo.org/record/7893064/files/figure.png" pageId="12" pageNumber="143">Fig. 8D</figureCitation>
). Hind wings clearly surpassing FWs.
</paragraph>
<paragraph blockId="12.[151,1437,1204,2023]" lastBlockId="13.[151,1437,151,465]" lastPageId="13" lastPageNumber="144" pageId="12" pageNumber="143">
♁ genitalia (
<figureCitation box="[349,485,1888,1915]" captionStart="FIGURE 9" captionStartId="10.[151,250,1721,1745]" captionTargetBox="[151,1436,181,1697]" captionTargetId="figure-23@10.[151,1436,181,1697]" captionTargetPageId="10" captionText="FIGURE 9. Pendleburyella J genitalia: Pendleburyella pimela sp. nov. (A–C) and Pendleburyella eirmosa sp. nov. (D–F) in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7893066" httpUri="https://zenodo.org/record/7893066/files/figure.png" pageId="12" pageNumber="143">Figs. 9D–F</figureCitation>
). Pseudepiphallus [epiphallus] typical of genus. Posterior end of pseudepiphallus [epiphallus] produced into two lobular pseudepiphallic lophi [posterolateral epiphallic lobe], in dorsal view tongue-shaped with apex obtuse; more broadly emarginated between latero-apical lobules; in profile distinctly flattened. Pseudepiphallic paramere [ectoparamere] not surpassing pseudepiphallic lophi, sclerotized, also slender and tapering into subacute apex. Dorsal process of pseudepiphallic paramere not surpassing ventral process. Ventral process of pseudepiphallic paramere obtuse at apex in ventral view. In profile, emargination between dorsal and ventral processes of pseudepiphallic paramere broadly rounded. Ectophallic fold [rachis] typical of genus; in profile slenderer.
</paragraph>
<paragraph blockId="13.[151,1437,151,465]" pageId="13" pageNumber="144">
<emphasis bold="true" box="[199,518,294,321]" pageId="13" pageNumber="144">Measurements (♂, in mm).</emphasis>
BL = 6.5; BWL = 10.1; HL = 0.9; PronL = 1.2; PronW = 2.0; FWL = 6.2; FWW = 2.4; HWT = 2.1; FIIIL = 4.0; TIIIL = 3.1; TaIIIL = 1.6.
</paragraph>
</subSubSection>
<subSubSection box="[199,1404,367,393]" pageId="13" pageNumber="144" type="biology_ecology">
<paragraph blockId="13.[151,1437,151,465]" box="[199,1404,367,393]" pageId="13" pageNumber="144">
<emphasis bold="true" box="[199,300,367,393]" pageId="13" pageNumber="144">Ecology.</emphasis>
The species was found dwelling among the leaf litter as well as on the leaves of shrub/small trees.
</paragraph>
</subSubSection>
<subSubSection box="[199,860,402,429]" pageId="13" pageNumber="144" type="distribution">
<paragraph blockId="13.[151,1437,151,465]" box="[199,860,402,429]" pageId="13" pageNumber="144">
<emphasis bold="true" box="[199,353,402,428]" pageId="13" pageNumber="144">Distribution.</emphasis>
Borneo (
<collectingCountry box="[456,765,403,429]" name="Brunei" pageId="13" pageNumber="144">BRUNEI DARUSSALAM</collectingCountry>
,
<collectingRegion box="[776,848,403,429]" country="Brunei" name="Belait" pageId="13" pageNumber="144">Belait</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,757,438,465]" pageId="13" pageNumber="144" type="materials_examined">
<paragraph blockId="13.[151,1437,151,465]" box="[199,757,438,465]" pageId="13" pageNumber="144">
<materialsCitation box="[199,753,438,465]" country="Brunei" location="Belait" pageId="13" pageNumber="144" specimenCount="1" stateProvince="Belait" typeStatus="holotype">
<emphasis bold="true" box="[199,358,438,465]" pageId="13" pageNumber="144">
<typeStatus box="[199,258,439,465]" pageId="13" pageNumber="144">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[364,672,439,465]" name="Brunei" pageId="13" pageNumber="144">BRUNEI DARUSSALAM</collectingCountry>
,
<collectingRegion box="[682,753,439,465]" country="Brunei" name="Belait" pageId="13" pageNumber="144">Belait</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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