845 lines
94 KiB
XML
845 lines
94 KiB
XML
<document id="C07A1E879AE0A8C4CF6D2865ECB7C56B" ID-DOI="10.5281/zenodo.200605" ID-GBIF-Dataset="c4f6386e-38aa-49c4-bf75-211356305e33" ID-ISSN="1175-5326" ID-Zenodo-Dep="200605" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460329340757" checkinUser="plazi" docAuthor="Říčan, Oldřich, Piálek, Lubomír, Almirón, Adriana & Casciotta, Jorge" docDate="2011" docId="FA2F69118609FFCA22A4FDCDFC3AF89A" docLanguage="en" docName="zt02982p026.pdf" docOrigin="Zootaxa 2982" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Australoheros angiru Říčan, Piálek, Almirón & Casciotta, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="061611698607FFDB2233FF8AFFC2FFBD" masterDocTitle="Two new species of Australoheros (Teleostei: Cichlidae), with notes on diversity of the genus and biogeography of the Río de la Plata basin" masterLastPageNumber="26" masterPageNumber="1" pageNumber="15" updateTime="1698258417310" updateUser="plazi">
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<mods:mods id="02C997DBBE3034A24A14A385F3BF8F4F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1C446F430C04F78C36E5ADE8C597D210">
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<mods:title id="4870BD653C19507501198DA206FDEA10">Two new species of Australoheros (Teleostei: Cichlidae), with notes on diversity of the genus and biogeography of the Río de la Plata basin</mods:title>
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</mods:titleInfo>
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<mods:name id="F228A8F7E5C46F76226F4B9C9E8A5E02" type="personal">
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<mods:roleTerm id="625723CD1DFE96E779463B43D48A8217">Author</mods:roleTerm>
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<mods:namePart id="B67AAEFB6DBCE6CEE8C193B555E4CD3C">Říčan, Oldřich</mods:namePart>
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</mods:name>
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<mods:roleTerm id="AF3B1FA6326608D027A724287E929E09">Author</mods:roleTerm>
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<mods:namePart id="2591629AA42D924285CDB79F1EFA2C1A">Piálek, Lubomír</mods:namePart>
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<mods:roleTerm id="24F7DD9D63E0FA77F94DFD10F4ED9FAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C277BF9A9BB2D740DE2F080284DDE4B">Almirón, Adriana</mods:namePart>
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</mods:name>
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<mods:name id="4B49C760718B0FD8EA9638847C254E26" type="personal">
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<mods:roleTerm id="A21ADD562084D1C2C5BBAAF1B6247DA8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A2B9E17F0E07482965D0751D22BEBB42">Casciotta, Jorge</mods:namePart>
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<mods:typeOfResource id="B949E41258ADE3C05E63063B155405A4">text</mods:typeOfResource>
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<mods:title id="EAE8FE5473B0F4ADA047CD0EE84A9BC6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AA5EEC27040354E766D58D271503D4A0">
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<mods:date id="46C90DF671624515ACA1E5867B1272D9">2011</mods:date>
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<mods:detail id="73B9A4A71051602DCC2AEAADCCE08607" type="volume">
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<mods:number id="0B957574FAFBEE38619462F5A7544479">2982</mods:number>
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<mods:start id="07E7D3727F89E1666C7898F021290F68">1</mods:start>
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<mods:identifier id="A8961B786CD210351B2FF4EFEE84DF95" type="DOI">10.5281/zenodo.200605</mods:identifier>
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<mods:identifier id="1EFA915B780D0965A15D49FB1119DA2E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A7C35F12AECE57127B50567EF02B7A2A" type="Zenodo-Dep">200605</mods:identifier>
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<treatment id="FA2F69118609FFCA22A4FDCDFC3AF89A" ID-DOI="http://doi.org/10.5281/zenodo.6192230" ID-GBIF-Taxon="119404273" ID-Zenodo-Dep="6192230" LSID="urn:lsid:plazi:treatment:FA2F69118609FFCA22A4FDCDFC3AF89A" httpUri="http://treatment.plazi.org/id/FA2F69118609FFCA22A4FDCDFC3AF89A" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
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<subSubSection id="3A9C8B8C8609FFD522A4FDCDFEECFD3F" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="7239D8078609FFD522A4FDCDFE3DFDDC" blockId="14.[151,511,583,642]" box="[151,511,583,609]" pageId="14" pageNumber="15">
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<heading id="29716F6B8609FFD522A4FDCDFE3DFDDC" bold="true" box="[151,511,583,609]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="40F204158609FFD522A4FDCDFE3DFDDC" bold="true" box="[151,511,583,609]" pageId="14" pageNumber="15">
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<taxonomicName id="B586A3848609FFD522A4FDCDFE59FDDC" ID-CoL="689B7" box="[151,411,583,609]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angiru" status="sp. nov.">
|
||
<emphasis id="40F204158609FFD522A4FDCDFE59FDDC" bold="true" box="[151,411,583,609]" italics="true" pageId="14" pageNumber="15">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BC1B96E8609FFD52391FDCDFE3DFDDC" box="[418,511,583,609]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
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</emphasis>
|
||
</heading>
|
||
</paragraph>
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||
<paragraph id="7239D8078609FFD522A4FDE0FEECFD3F" blockId="14.[151,511,583,642]" box="[151,302,618,642]" pageId="14" pageNumber="15">
|
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(
|
||
<figureCitation id="EABDC4828609FFD522ACFDE0FF25FD3F" box="[159,231,618,642]" captionStart="FIGURE 7" captionStartId="13.[151,250,1505,1528]" captionTargetBox="[161,1428,253,1489]" captionTargetId="figure@13.[151,1436,243,1499]" captionTargetPageId="13" captionText="FIGURE 7. Color plate. Horizontaly from upper left to lower right. Australoheros forquilha, rio Forquilha, rio Uruguai drainage, Rio Grande do Sul, Brazil (not preserved). Australoheros ykeregua (MACN-ict 9467, holotype), río Uruguay drainage, arroyo Paraiso (or Canal Muerto), Misiones province, Argentina. Australoheros kaaygua (MACN-ict 9473), río Iguazú drainage, small stream 7 km SW from Andresito, Misiones province, Argentina. Australoheros angiru, male in neutral colors, rio Chopim, rio Iguaçu drainage, Paraná, Brazil (not preserved). A. angiru, male and female in breeding colors guarding fry, same locality (not preserved). All A. angiru photographs courtesy of Wolfgang Staeck." httpUri="https://zenodo.org/record/200612/files/figure.png" pageId="14" pageNumber="15">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="EABDC4828609FFD522C0FDE0FEC1FD3F" box="[243,259,618,642]" captionStart="FIGURE 8" captionStartId="15.[151,250,864,887]" captionTargetBox="[151,1436,195,842]" captionTargetId="figure@15.[151,1436,195,842]" captionTargetPageId="15" captionText="FIGURE 8. Australoheros angiru. Holotype, MCP 13937, 73.2 mm SL, rio Jacutinga, rio Uruguai drainage, Brazil." httpUri="https://zenodo.org/record/200613/files/figure.png" pageId="14" pageNumber="15">8</figureCitation>
|
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,
|
||
<figureCitation id="EABDC4828609FFD5233CFDE0FEDDFD3F" box="[271,287,618,642]" captionStart="FIGURE 9" captionStartId="15.[151,250,1537,1560]" captionTargetBox="[151,1435,926,1526]" captionTargetId="figure@15.[151,1435,926,1526]" captionTargetPageId="15" captionText="FIGURE 9. Australoheros angiru. Paratype, MCP 13011, 48.1 mm SL, rio Jacutinga, rio Uruguai drainage, Brazil." httpUri="https://zenodo.org/record/200614/files/figure.png" pageId="14" pageNumber="15">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A9C8B8C8609FFD522A4FD38FCE3FCFE" pageId="14" pageNumber="15" type="reference_group">
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<paragraph id="7239D8078609FFD522A4FD38FDBBFD5A" blockId="14.[151,801,690,835]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="5296FF298609FFD522A4FD38FDBBFD5A" pageId="14" pageNumber="15">
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<emphasis id="40F204158609FFD522A4FD38FE5AFD75" box="[151,408,690,712]" italics="true" pageId="14" pageNumber="15">
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||
“
|
||
<taxonomicName id="B586A3848609FFD522ADFD38FEE6FD75" ID-CoL="3PTG" box="[158,292,690,712]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
|
||
” facetum—
|
||
</emphasis>
|
||
<treatmentCitation id="F327FE168609FFD523AAFD38FD8AFD75" author="Staeck" box="[409,584,690,712]" page="62" pageId="14" pageNumber="15" year="1998">
|
||
<bibRefCitation id="1617A5F68609FFD523AAFD38FDE6FD75" author="Staeck" box="[409,548,690,712]" pageId="14" pageNumber="15" refString="Staeck, W. (1998 a) Neuer Chanchito. DCG informationen, 29, 62 - 63." type="journal article" year="1998" yearSuffix="a">Staeck 1998a</bibRefCitation>
|
||
: 62
|
||
</treatmentCitation>
|
||
–63; 1998b: 81–85
|
||
<emphasis id="40F204158609FFD522A4FD5BFEE9FD5A" box="[151,299,721,743]" italics="true" pageId="14" pageNumber="15">
|
||
“
|
||
<taxonomicName id="B586A3848609FFD522ADFD5BFEE6FD5A" ID-CoL="3PTG" box="[158,292,721,743]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
sp. Iguaçu
|
||
<emphasis id="40F204158609FFD523A9FD58FE72FD5A" box="[410,432,722,743]" italics="true" pageId="14" pageNumber="15">—</emphasis>
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||
<treatmentCitation id="F327FE168609FFD52382FD5BFD91FD5A" author="Staeck" box="[433,595,721,743]" page="64" pageId="14" pageNumber="15" year="2003">
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<bibRefCitation id="1617A5F68609FFD52382FD5BFDF3FD5A" author="Staeck" box="[433,561,721,743]" pageId="14" pageNumber="15" refString="Staeck, W. (2003) " Cichlasoma " facetum komplex: Ein ruckblick auf die Anfange der Aquaristik. Die Aquarien- und Terrarienzeitschrift, 9, 60 - 65." type="journal article" year="2003">Staeck 2003</bibRefCitation>
|
||
: 64
|
||
</treatmentCitation>
|
||
–65
|
||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7239D8078609FFD522A4FD7AFCE3FCFE" blockId="14.[151,801,690,835]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="5296FF298609FFD522A4FD7AFCE3FCFE" pageId="14" pageNumber="15">
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<emphasis id="40F204158609FFD522A4FD7AFEE9FCBB" box="[151,299,752,774]" italics="true" pageId="14" pageNumber="15">
|
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“
|
||
<taxonomicName id="B586A3848609FFD522ADFD7AFEE6FCBB" ID-CoL="3PTG" box="[158,292,752,774]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
sp. Iguaçu
|
||
<emphasis id="40F204158609FFD523A9FD7BFE72FCBB" box="[410,432,753,774]" italics="true" pageId="14" pageNumber="15">—</emphasis>
|
||
<treatmentCitation id="F327FE168609FFD52382FD7AFCD8FCBB" author="Stawikowski" box="[433,794,752,774]" page="455" pageId="14" pageNumber="15" year="2004">
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<bibRefCitation id="1617A5F68609FFD52382FD7AFD29FCB8" author="Stawikowski" box="[433,747,752,774]" pageId="14" pageNumber="15" refString="Stawikowski, R. & Werner, U. (2004) Die Buntbarsche Amerikas. Band 3: Erdfresser, Hecht- un Kammbuntbarsche. Ulmer Verlag, Stuttgart, 478 pp." type="book" year="2004">Stawikowski and Werner 2004</bibRefCitation>
|
||
: 455
|
||
</treatmentCitation>
|
||
<taxonomicName id="B586A3848609FFD522A4FC85FEE4FC98" ID-CoL="3634" box="[151,294,783,805]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="40F204158609FFD522A4FC85FEE4FC98" box="[151,294,783,805]" italics="true" pageId="14" pageNumber="15">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
sp. jacutinga
|
||
<emphasis id="40F204158609FFD5239FFC9AFE00FC98" box="[428,450,784,805]" italics="true" pageId="14" pageNumber="15">—</emphasis>
|
||
<treatmentCitation id="F327FE168609FFD523F1FC80FD25FC99" author="Rican" box="[450,743,778,805]" page="6" pageId="14" pageNumber="15" year="2006">
|
||
<bibRefCitation id="1617A5F68609FFD523F1FC80FD12FC99" author="Rican" box="[450,720,778,805]" pageId="14" pageNumber="15" refString="Rican, O. & Kullander, S. O. (2006) Character- and Tree-based delimitation of species in the´Cichlasoma´facetum group (Teleostei, Cichlidae) with the description of a new genus. Journal of Zoological Systematics and Evolutionary Research, 44, 136 - 152." type="journal article" year="2006">Říčan and Kullander 2006</bibRefCitation>
|
||
: 6
|
||
</treatmentCitation>
|
||
<emphasis id="40F204158609FFD522A4FCA7FE5AFCFE" box="[151,408,813,835]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="B586A3848609FFD522A4FCA7FEE4FCFE" ID-CoL="3634" box="[151,294,813,835]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">Australoheros</taxonomicName>
|
||
kaaygua—
|
||
</emphasis>
|
||
<treatmentCitation id="F327FE168609FFD523ABFCA2FD0AFCFF" author="Rican" box="[408,712,808,835]" page="28" pageId="14" pageNumber="15" year="2008">
|
||
<bibRefCitation id="1617A5F68609FFD523ABFCA2FD67FCFF" author="Rican" box="[408,677,808,835]" pageId="14" pageNumber="15" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan and Kullander 2008</bibRefCitation>
|
||
: 28
|
||
</treatmentCitation>
|
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(in part)
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="3A9C8B8C8609FFD522A4FCFAFBF0FAE1" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="7239D8078609FFD522A4FCFAFCBDFC11" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<emphasis id="40F204158609FFD522A4FCFAFEC9FC35" bold="true" box="[151,267,880,904]" pageId="14" pageNumber="15">
|
||
<typeStatus id="AD3D66A58609FFD522A4FCFAFEC4FC35" box="[151,262,880,904]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="149740C28609FFD52327FCE5FE8DFC3A" box="[276,335,879,903]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="14" pageNumber="15">MCP</collectionCode>
|
||
13937,
|
||
<quantity id="B57E75E28609FFD5239CFCE5FDD7FC3A" box="[431,533,879,903]" metricMagnitude="-2" metricUnit="m" metricValue="7.32" pageId="14" pageNumber="15" unit="mm" value="73.2">73.2 mm</quantity>
|
||
SL,
|
||
<collectingCountry id="0A9198978609FFD5207CFCE5FD54FC3A" box="[591,662,879,903]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
|
||
, Santa Catarina State, rio
|
||
<collectingCountry id="0A9198978609FFD521F0FCE5FBDDFC3A" box="[963,1055,879,903]" name="Uruguay" pageId="14" pageNumber="15">Uruguai</collectingCountry>
|
||
drainage, rio Jacutinga, road BR 283 from Ceará to Concordia, col: Bergmann
|
||
<emphasis id="40F204158609FFD520AAFC1FFD13FC11" box="[665,721,916,940]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
,
|
||
<date id="0638FEC78609FFD520ECFC1EFCB9FC11" box="[735,891,916,940]" pageId="14" pageNumber="15" value="1988-10">October 1988</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7239D8078609FFD522F4FC32FC79FB81" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<emphasis id="40F204158609FFD522F4FC32FE84FC6D" bold="true" box="[199,326,952,976]" pageId="14" pageNumber="15">
|
||
<typeStatus id="AD3D66A58609FFD522F4FC32FE83FC6D" box="[199,321,952,976]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="6480138E8609FFD5237FFC3DFE25FC72" box="[332,487,951,975]" pageId="14" pageNumber="15" type="generic">13 specimens</specimenCount>
|
||
, 24.6–77.0 mm SL, all from
|
||
<collectingCountry id="0A9198978609FFD5211BFC3DFCB2FC72" box="[808,880,951,975]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
|
||
. Santa Catarina State, rio
|
||
<collectingCountry id="0A9198978609FFD526A3FC3DFB2EFC72" box="[1168,1260,951,975]" name="Uruguay" pageId="14" pageNumber="15">Uruguai</collectingCountry>
|
||
drainage:
|
||
<collectionCode id="149740C28609FFD52752FC3DFA5EFC72" box="[1377,1436,951,975]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="14" pageNumber="15">MCP</collectionCode>
|
||
13383, 6 ex., 24.6–77.0 mm SL, rio Jacutinga, road BR 283 from Ceará to Concordia, col: Reis
|
||
<emphasis id="40F204158609FFD526DEFC57FAE5FC49" box="[1261,1319,988,1012]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
,
|
||
<date id="0638FEC78609FFD52704FC56FF10FBAA" pageId="14" pageNumber="15" value="1989-02">February 1989</date>
|
||
.
|
||
<collectionCode id="149740C28609FFD522EEFC75FEDAFBAA" box="[221,280,1023,1047]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="14" pageNumber="15">MCP</collectionCode>
|
||
12509, 1 ex., 75.0 mm SL, same data as
|
||
<typeStatus id="AD3D66A58609FFD520D6FC75FC88FBAA" box="[741,842,1023,1047]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
.
|
||
<collectionCode id="149740C28609FFD52167FC75FC4DFBAA" box="[852,911,1023,1047]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="14" pageNumber="15">MCP</collectionCode>
|
||
13011, 6 ex.,
|
||
<quantity id="B57E75E28609FFD5261EFC75FB12FBAA" box="[1069,1232,1023,1047]" metricMagnitude="-2" metricUnit="m" metricValue="5.279999999999999" metricValueMax="6.14" metricValueMin="4.42" pageId="14" pageNumber="15" unit="mm" value="52.8" valueMax="61.4" valueMin="44.2">44.2–61.4 mm</quantity>
|
||
SL, rio Jacutinga, road BR 283 from Ceará to Concordia, col: Reis
|
||
<emphasis id="40F204158609FFD5208FFBAFFD36FB81" box="[700,756,1060,1084]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
,
|
||
<date id="0638FEC78609FFD52131FBAEFC75FB81" box="[770,951,1060,1084]" pageId="14" pageNumber="15" value="1988-12">December 1988</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7239D8078609FFD522F4FBCDFBF0FAE1" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<emphasis id="40F204158609FFD522F4FBCDFDE8FBDD" bold="true" box="[199,554,1095,1120]" pageId="14" pageNumber="15">Additional non-type material.</emphasis>
|
||
Paraná State, rio Iguaçu drainage:
|
||
<collectionCode id="149740C28609FFD52182FBC3FC2BFBDD" box="[945,1001,1097,1120]" pageId="14" pageNumber="15">NUP</collectionCode>
|
||
3913, 2 ex., rio São Pedro, tributary to rio Iguaçu, Pinhão county,
|
||
<geoCoordinate id="17B2BEC08609FFD523D3FBE6FDFFFB39" box="[480,573,1132,1156]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-26.083334">26º05´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="17B2BEC08609FFD5207CFBE6FD72FB39" box="[591,688,1132,1156]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-51.75">51º45'W</geoCoordinate>
|
||
, col: Nupélia staff,
|
||
<date id="0638FEC78609FFD52194FBE6FBFBFB39" box="[935,1081,1132,1156]" pageId="14" pageNumber="15" value="1993-03">March 1993</date>
|
||
.
|
||
<collectionCode id="149740C28609FFD52678FBE7FB41FB39" box="[1099,1155,1133,1156]" pageId="14" pageNumber="15">NUP</collectionCode>
|
||
3914, 1 ex, rio Iratim (Linígrafo), tributary to rio Iguaçu, Palmas county, boundary with Pìnhão-PR,
|
||
<geoCoordinate id="17B2BEC08609FFD5261CFB05FB49FB1A" box="[1071,1163,1167,1191]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-26.083334">26º05´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="17B2BEC08609FFD526A9FB05FB3DFB1A" box="[1178,1279,1167,1191]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-51.75">51º45´W</geoCoordinate>
|
||
, col: Nupélia staff,
|
||
<date id="0638FEC78609FFD522EBFB3EFE95FB71" box="[216,343,1204,1228]" pageId="14" pageNumber="15" value="1993-04">April 1993</date>
|
||
.
|
||
<collectionCode id="149740C28609FFD52356FB3FFE5FFB71" box="[357,413,1205,1228]" pageId="14" pageNumber="15">NUP</collectionCode>
|
||
3915, 1 ex, rio São Pedro, tributary to rio Iguaçu, Pinhão county,
|
||
<geoCoordinate id="17B2BEC08609FFD526A8FB3EFB35FB71" box="[1179,1271,1204,1228]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-26.083334">26º05´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="17B2BEC08609FFD52737FB3EFAA7FB71" box="[1284,1381,1204,1228]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-51.75">51º45'W</geoCoordinate>
|
||
, col: Nupélia staff,
|
||
<date id="0638FEC78609FFD5230BFB5DFE06FB52" box="[312,452,1239,1263]" pageId="14" pageNumber="15" value="1993-03">March 1993</date>
|
||
. Rio Grande do Sul State, rio
|
||
<collectingCountry id="0A9198978609FFD52129FB5DFCB4FB52" box="[794,886,1239,1263]" name="Uruguay" pageId="14" pageNumber="15">Uruguai</collectingCountry>
|
||
drainage:
|
||
<collectionCode id="149740C28609FFD521DDFB5DFBEBFB52" box="[1006,1065,1239,1263]" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="14" pageNumber="15">MCP</collectionCode>
|
||
46328, 13 ex., Sanga das Aguas Frias, Irai, col: Malabarba
|
||
<emphasis id="40F204158609FFD523F3FB77FE3AFAA9" box="[448,504,1276,1300]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
, 1985.
|
||
<collectingCountry id="0A9198978609FFD5207FFB76FD7CFAA9" box="[588,702,1276,1300]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
|
||
, Misiones province, río
|
||
<collectingCountry id="0A9198978609FFD521FFFB77FBECFAA8" box="[972,1070,1277,1301]" name="Uruguay" pageId="14" pageNumber="15">Uruguay</collectingCountry>
|
||
drainage:
|
||
<collectionCode id="149740C28609FFD52697FB76FB1FFAA9" box="[1188,1245,1276,1300]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="15">ZSM</collectionCode>
|
||
23482a, 1 ex., P, río Soberbio, El Soberbio, col: J. Foerster, 1966.
|
||
<collectionCode id="149740C28609FFD520F8FAAAFCC7FA85" box="[715,773,1312,1336]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="15">ZSM</collectionCode>
|
||
23060a, 4 ex., río Soberbio, El Soberbio, col: J. Foerster, 1966.
|
||
<collectionCode id="149740C28609FFD522EEFACEFED5FAE1" box="[221,279,1348,1372]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="15">ZSM</collectionCode>
|
||
23060c, 2 ex. (C&S), río Soberbio, El Soberbio, col: J. Foerster, 1966.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A9C8B8C8609FFD522F4FAEDFB43F8C5" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="7239D8078609FFD522F4FAEDFD79FA75" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<emphasis id="40F204158609FFD522F4FAEDFE83FA3D" bold="true" box="[199,321,1383,1408]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
|
||
<taxonomicName id="B586A3848609FFD52379FAE2FDF9FA3D" box="[330,571,1384,1408]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158609FFD52379FAE2FDF9FA3D" box="[330,571,1384,1408]" italics="true" pageId="14" pageNumber="15">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is one of the most deep-bodied species of
|
||
<taxonomicName id="B586A3848609FFD52615FAE2FB06FA3D" box="[1062,1220,1384,1408]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="40F204158609FFD52615FAE2FB06FA3D" box="[1062,1220,1384,1408]" italics="true" pageId="14" pageNumber="15">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
(body depth in SL>49%; shared with
|
||
<taxonomicName id="B586A3848609FFD52342FA06FE2BFA19" box="[369,489,1420,1444]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="guarani">
|
||
<emphasis id="40F204158609FFD52342FA06FE2BFA19" box="[369,489,1420,1444]" italics="true" pageId="14" pageNumber="15">A. guarani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848609FFD5202DFA06FD4EFA19" box="[542,652,1420,1444]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="facetus">
|
||
<emphasis id="40F204158609FFD5202DFA06FD4EFA19" box="[542,652,1420,1444]" italics="true" pageId="14" pageNumber="15">A. facetus</emphasis>
|
||
</taxonomicName>
|
||
). It has been previously associated with
|
||
<taxonomicName id="B586A3848609FFD52661FA06FB12FA19" box="[1106,1232,1420,1444]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158609FFD52661FA06FB12FA19" box="[1106,1232,1420,1444]" italics="true" pageId="14" pageNumber="15">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
, but it is the sister species of
|
||
<taxonomicName id="B586A3848609FFD5233EFA3AFE4CFA75" box="[269,398,1456,1480]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="40F204158609FFD5233EFA3AFE4CFA75" box="[269,398,1456,1480]" italics="true" pageId="14" pageNumber="15">A. minuano</emphasis>
|
||
</taxonomicName>
|
||
based on DNA characters.
|
||
</paragraph>
|
||
<paragraph id="7239D8078609FFD522F6FA5EFAC7F91D" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="B586A3848609FFD522F6FA5EFE76FA51" box="[197,436,1492,1516]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158609FFD522F6FA5EFE76FA51" box="[197,436,1492,1516]" italics="true" pageId="14" pageNumber="15">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="B586A3848609FFD52082FA5EFCEDFA51" box="[689,815,1492,1516]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158609FFD52082FA5EFCEDFA51" box="[689,815,1492,1516]" italics="true" pageId="14" pageNumber="15">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
by having less scale rows between anterior end of dorsal fin and upper lateral line (ch4 states 1–2
|
||
<emphasis id="40F204158609FFD520BAFA73FD6AF9AD" box="[649,680,1529,1552]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
0), by a very narrow or missing caudal base spot, by a pure yellow ground color (
|
||
<emphasis id="40F204158609FFD5230BF997FE94F989" box="[312,342,1565,1588]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
yellowish-green), by yellow eyes (
|
||
<emphasis id="40F204158609FFD520D0F997FCC3F989" box="[739,769,1565,1588]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
dark green), by more scales between anterior end of dorsal fin and upper lateral line (5
|
||
<emphasis id="40F204158609FFD523E5F9CBFE36F9E5" box="[470,500,1601,1624]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
4), more anal fin spines (7
|
||
<emphasis id="40F204158609FFD52103F9CBFC8CF9E5" box="[816,846,1601,1624]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
6), more anal fin rays (> 7
|
||
<emphasis id="40F204158609FFD526B9F9CBFB6AF9E5" box="[1162,1192,1601,1624]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
<7), more dorsal fin rays (9
|
||
<emphasis id="40F204158609FFD522D9F9EFFECAF9C1" box="[234,264,1637,1660]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
8), less E0 scales (24
|
||
<emphasis id="40F204158609FFD523CDF9EFFDDEF9C1" box="[510,540,1637,1660]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
> 25), more L1 scales (> 17–18
|
||
<emphasis id="40F204158609FFD521B5F9EFFC66F9C1" box="[902,932,1637,1660]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
16), less L2 scales (8
|
||
<emphasis id="40F204158609FFD526A9F9EFFB7AF9C1" box="[1178,1208,1637,1660]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
> 9), and by a being more deep-bodied (49.6%
|
||
<emphasis id="40F204158609FFD523F2F903FE22F91D" box="[449,480,1673,1696]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
43.8% SL), and having a shorter caudal peduncle (7.4%
|
||
<emphasis id="40F204158609FFD5266CF903FBBFF91D" box="[1119,1149,1673,1696]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
10.4% SL).
|
||
</paragraph>
|
||
<paragraph id="7239D8078609FFD522F6F926FD0FF8E9" blockId="14.[151,1437,879,1912]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="B586A3848609FFD522F6F926FE76F979" box="[197,436,1708,1732]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158609FFD522F6F926FE76F979" box="[197,436,1708,1732]" italics="true" pageId="14" pageNumber="15">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="B586A3848609FFD5209EF926FCECF979" box="[685,814,1708,1732]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="40F204158609FFD5209EF926FCECF979" box="[685,814,1708,1732]" italics="true" pageId="14" pageNumber="15">A. minuano</emphasis>
|
||
</taxonomicName>
|
||
by a large and dominant midlateral blotch, very narrow or missing caudal base spot, by lacking a pinkish body coloration, by a small terminal or subterminal mouth (
|
||
<emphasis id="40F204158609FFD5274EF95BFA56F955" box="[1405,1428,1745,1768]" italics="true" pageId="14" pageNumber="15">vs</emphasis>
|
||
. large supraterminal), by more scales between the anterior end of the dorsal fin and the upper lateral line (5
|
||
<emphasis id="40F204158609FFD52764F97FFAB7F8B1" box="[1367,1397,1781,1804]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
4), less anal fin rays (7
|
||
<emphasis id="40F204158609FFD5234AF893FE5AF88D" box="[377,408,1817,1840]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
8), less dorsal fin rays (9
|
||
<emphasis id="40F204158609FFD5208EF893FD1EF88D" box="[701,732,1817,1840]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
10), and by slight differences in body depth (49.6%
|
||
<emphasis id="40F204158609FFD5271DF893FA8EF88D" box="[1326,1356,1817,1840]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
46.9% SL) and in preorbital distance (7.3%
|
||
<emphasis id="40F204158609FFD52006F8B7FD96F8E9" box="[565,596,1853,1876]" italics="true" pageId="14" pageNumber="15">vs.</emphasis>
|
||
6.0% SL).
|
||
</paragraph>
|
||
<paragraph id="7239D8078609FFD522F6F8EBFB43F8C5" blockId="14.[151,1437,879,1912]" box="[197,1153,1888,1912]" pageId="14" pageNumber="15">
|
||
For distinguishing characters to all other
|
||
<taxonomicName id="B586A3848609FFD520A2F8EAFCECF8C5" box="[657,814,1888,1912]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="40F204158609FFD520A2F8EAFCECF8C5" box="[657,814,1888,1912]" italics="true" pageId="14" pageNumber="15">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
species see the Notes section.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="26F9888F8608FFD422A4FCEAFAEDFCCB" box="[151,1327,864,887]" httpUri="https://zenodo.org/record/200613/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1436,195,842]" targetPageId="15">
|
||
<paragraph id="7239D8078608FFD422A4FCEAFAEDFCCB" blockId="15.[151,1327,864,887]" box="[151,1327,864,887]" pageId="15" pageNumber="16">
|
||
<emphasis id="40F204158608FFD422A4FCEAFED0FCC8" bold="true" box="[151,274,864,887]" pageId="15" pageNumber="16">FIGURE 8.</emphasis>
|
||
<taxonomicName id="B586A3848608FFD4232AFCEAFE33FCCB" box="[281,497,864,886]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158608FFD4232AFCEAFE33FCCB" box="[281,497,864,886]" italics="true" pageId="15" pageNumber="16">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
. Holotype, MCP 13937, 73.2 mm SL, rio Jacutinga, rio Uruguai drainage, Brazil.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26F9888F8608FFD422A4F98BFAE5F9A5" box="[151,1319,1537,1560]" httpUri="https://zenodo.org/record/200614/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1435,926,1526]" targetPageId="15">
|
||
<paragraph id="7239D8078608FFD422A4F98BFAE5F9A5" blockId="15.[151,1319,1537,1560]" box="[151,1319,1537,1560]" pageId="15" pageNumber="16">
|
||
<emphasis id="40F204158608FFD422A4F98BFED0F9AA" bold="true" box="[151,274,1537,1560]" pageId="15" pageNumber="16">FIGURE 9.</emphasis>
|
||
<taxonomicName id="B586A3848608FFD4232AF988FE33F9A5" box="[281,497,1538,1560]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158608FFD4232AF988FE33F9A5" box="[281,497,1538,1560]" italics="true" pageId="15" pageNumber="16">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
. Paratype, MCP 13011, 48.1 mm SL, rio Jacutinga, rio Uruguai drainage, Brazil.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3A9C8B8C8608FFCB22F6F9CDFE09FA21" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="7239D8078608FFD422F6F9CDFC9CF939" blockId="15.[151,1437,1607,2031]" pageId="15" pageNumber="16">
|
||
<emphasis id="40F204158608FFD422F6F9CDFE94F9DD" bold="true" box="[197,342,1607,1632]" pageId="15" pageNumber="16">Description.</emphasis>
|
||
Based on specimens over
|
||
<quantity id="B57E75E28608FFD420B3F9C2FD0DF9DD" box="[640,719,1608,1632]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="60.0">60 mm</quantity>
|
||
SL, with notes on smaller specimens. Meristic data are summarized in Table 2, morphometric data are summarized in
|
||
<tableCitation id="3F04EDBC8608FFD42137F9E6FC9BF939" box="[772,857,1644,1668]" captionStart="TABLE 3" captionStartId="9.[151,235,152,173]" captionText="TABLE 3. Proportional measurements in percents of standard length (SL) of the two new species (A. ykeregua, A. angiru) and the two species with which they have been previously associated (A. forquilha, A. kaaygua). SD = standard deviation. A. forquilha A. ykeregua" pageId="15" pageNumber="16">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7239D8078608FFD422F6F91AFEB5F971" blockId="15.[151,1437,1607,2031]" pageId="15" pageNumber="16">Comparatively deep bodied (mean body depth 49.6% SL). Snout short, straight in lateral view. Jaws isognathous. Mouth small.</paragraph>
|
||
<paragraph id="7239D8078608FFD422F6F952FD40F852" blockId="15.[151,1437,1607,2031]" pageId="15" pageNumber="16">Scales on head and chest not distinctly smaller than on flanks. Scales in E0 row 23(3), 24(16*), 25(4). Upper lateral line scales 16(1), 17(6*), 18(8). Lower lateral line scales 7(4), 8(7*), 9(4). Scales between upper lateral line and dorsal fin 4 anteriorly, 1 large plus 1 small posteriorly. Cheek scale rows 3(14*), 4(2). About 8 scale rows between the opercular flap and the anterior insertion of the pelvic fin. Dorsal fin with one basal scale row, starting from the 7th or 8th spine and running posteriad; interradial scales appear from 14th or 15th spine membrane, in single rows. Anal fin with one basal scale row; interradial scales in single rows, from penultimate spine. Caudal fin densely scaled, scales ctenoid; interradial scales in single rows; hind margin of scaly area concave, extending to between one-third and middle of caudal fin.</paragraph>
|
||
<paragraph id="7239D8078617FFCB22F6FF1DFDDEFE82" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">Soft dorsal fin pointed, extending beyond middle of caudal fin. D. XVI,9(16*), XVI,10(13), XVII,8(2). Soft anal fin pointed, of about the same length as dorsal fin. A. VI,7(2), VI,8(3), VII,7(17*), VII,8(8), VIII,6(1). Anal fin pterygiophores 11(2), 12(22*), 13(7). First pelvic fin ray longest, extending up to the second anal fin spine. Pectoral fin with a rounded tip, third and fourth rays longest, extending just to the midlateral blotch. P. 12(11*), 13(5). Caudal fin rounded to subtruncate.</paragraph>
|
||
<paragraph id="7239D8078617FFCB22F6FEC6FB37FE72" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
All teeth caniniform, slightly curved. Outer row teeth increasing in size symphysiad, upper jaw anterior teeth longest, lower jaw anterior teeth subequal. Number of lower jaw teeth up to
|
||
<quantity id="B57E75E28617FFCB21C7FEE5FBECFE3A" box="[1012,1070,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="16" pageNumber="17" unit="in" value="16.0">16 in</quantity>
|
||
one outer hemiseries, upper jaw tooth row much shorter, with about 7 or 8 teeth in one outer hemiseries. Lower pharyngeal tooth plate not studied. Gill rakers externally on first gill arch, 2 epibranchial,
|
||
<quantity id="B57E75E28617FFCB20C9FE3DFCE7FE72" box="[762,805,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0">1 in</quantity>
|
||
angle, 5(4), 6(11*), 7(1) ceratobranchial.
|
||
</paragraph>
|
||
<paragraph id="7239D8078617FFCB22A4FE56FE93FDAA" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">Vertebrae 13+13=26(29*), 13+14=27(2). Caudal peduncle with no vertebrae (10) or containing 0.5(4), 1(14*), 1.5(1) vertebrae.</paragraph>
|
||
<paragraph id="7239D8078617FFCB22F4FDA9FF14FB89" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
<emphasis id="40F204158617FFCB22F4FDA9FE31FD81" bold="true" box="[199,499,547,572]" pageId="16" pageNumber="17">Color pattern in alcohol.</emphasis>
|
||
Six to seven vertical flank bars, a caudal peduncle bar confluent with the caudalbase bar, and a midlateral stripe bearing the midlateral blotch in the fourth flank bar (sensu
|
||
<bibRefCitation id="1617A5F68617FFCB26A2FDC8FA92FDE2" author="Rican" box="[1169,1360,578,608]" pageId="16" pageNumber="17" refString="Rican, O., Musilova, Z., Muska, M. & Novak, J. (2005) Development of coloration patterns in Neotropical cichlids (Perciformes: Cichlidae: Cichlasomatinae). Folia Zoologica, 54, 1 - 46." type="journal article" year="2005">
|
||
Říčan
|
||
<emphasis id="40F204158617FFCB26EAFDC3FAD2FDDD" box="[1241,1296,584,608]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
) make up the principal markings. All fins and body are without conspicuous spots or blotches. The midlateral stripe is more distinct anteriorly from the midlateral blotch than posteriorly, and the midlateral blotch itself is a dominant coloration pattern element. Vertical bars are relatively wide, faint, indistinct in their ventral parts. The midlateral stripe posteriorly from the midlateral blotch does not align with the lower lateral line and aligns with the E1 scale row and does not continue in the E0 scale row. Posteriorly from the midlateral blotch, the stripe is slightly decomposed into two blotches in the respective vertical flank bars. The blotch posterior from the midlateral blotch is centered in the same scale row as the midlateral blotch (
|
||
<emphasis id="40F204158617FFCB20D7FCCFFCC5FCE1" box="[740,775,837,860]" italics="true" pageId="16" pageNumber="17">i.e.</emphasis>
|
||
E1 scale row), whereas the second blotch is more elongate along the vertical axis and centered in the E2 scale row, making the impression that the midlateral stripe makes a dorsally directed turn at its posterior end. The arrangement of the bars on the body in essentially the same as described for
|
||
<taxonomicName id="B586A3848617FFCB2300FC3AFD1FFC7A" authority="Rican & Kullander 2003" authorityName="Rican & Kullander" authorityYear="2003" box="[307,733,938,968]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="scitulus">
|
||
<emphasis id="40F204158617FFCB2300FC3AFE6BFC75" box="[307,425,944,968]" italics="true" pageId="16" pageNumber="17">A. scitulus</emphasis>
|
||
(
|
||
<bibRefCitation id="1617A5F68617FFCB2389FC20FD14FC7A" author="Rican" box="[442,726,938,968]" pageId="16" pageNumber="17" refString="Rican, O. & Kullander, S. O. (2003) ' Cichlasoma' scitulum: a new species of cichlid fish from the Rio de La Plata Region in Argentina, Brazil, and Uruguay. Copeia, 2003, 794 - 802." type="journal article" year="2003">Říčan & Kullander 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Very small spots present on the bases of some body scales in adult specimens. In juveniles the spotted pattern of the body is much more pronounced, with virtually every scale on the body having a spot at its base, including those in the anterior part of the E4 scale row (
|
||
<emphasis id="40F204158617FFCB2699FC73FB0EFBAD" box="[1194,1228,1017,1040]" italics="true" pageId="16" pageNumber="17">i.e.</emphasis>
|
||
as in adult
|
||
<taxonomicName id="B586A3848617FFCB277EFC72FF04FB89" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="scitulus">
|
||
<emphasis id="40F204158617FFCB277EFC72FF04FB89" italics="true" pageId="16" pageNumber="17">A. scitulus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7239D8078617FFCB22F4FBB5FE09FA21" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
<emphasis id="40F204158617FFCB22F4FBB5FEBDFBE5" bold="true" box="[199,383,1087,1112]" pageId="16" pageNumber="17">Life coloration.</emphasis>
|
||
Coloration of life specimens from the rio
|
||
<collectingCountry id="0A9198978617FFCB216FFBB5FC7AFBEA" box="[860,952,1087,1111]" name="Uruguay" pageId="16" pageNumber="17">Uruguai</collectingCountry>
|
||
drainage is unknown to us.
|
||
<bibRefCitation id="1617A5F68617FFCB26C6FBB5FA55FBEA" author="Staeck" box="[1269,1431,1087,1111]" pageId="16" pageNumber="17" refString="Staeck, W. (1998 a) Neuer Chanchito. DCG informationen, 29, 62 - 63." type="journal article" year="1998" yearSuffix="a">Staeck (1998a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1617A5F68617FFCB22A4FBEEFF22FBC1" author="Staeck" box="[151,224,1124,1148]" pageId="16" pageNumber="17" refString="Staeck, W. (1998 b) Ein neuer cichlide aus dem Cichlasoma facetum komplex. DCG informationen, 29, 81 - 85." type="journal article" year="1998" yearSuffix="b">1998b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1617A5F68617FFCB22D9FBEEFEE4FBC1" author="Staeck" box="[234,294,1124,1148]" pageId="16" pageNumber="17" refString="Staeck, W. (2003) " Cichlasoma " facetum komplex: Ein ruckblick auf die Anfange der Aquaristik. Die Aquarien- und Terrarienzeitschrift, 9, 60 - 65." type="journal article" year="2003">2003</bibRefCitation>
|
||
: p. 64) photographed specimens from the rio Iguaçu drainage (
|
||
<figureCitation id="EABDC4828617FFCB21D0FBEEFBE5FBC1" box="[995,1063,1124,1148]" captionStart="FIGURE 7" captionStartId="13.[151,250,1505,1528]" captionTargetBox="[161,1428,253,1489]" captionTargetId="figure@13.[151,1436,243,1499]" captionTargetPageId="13" captionText="FIGURE 7. Color plate. Horizontaly from upper left to lower right. Australoheros forquilha, rio Forquilha, rio Uruguai drainage, Rio Grande do Sul, Brazil (not preserved). Australoheros ykeregua (MACN-ict 9467, holotype), río Uruguay drainage, arroyo Paraiso (or Canal Muerto), Misiones province, Argentina. Australoheros kaaygua (MACN-ict 9473), río Iguazú drainage, small stream 7 km SW from Andresito, Misiones province, Argentina. Australoheros angiru, male in neutral colors, rio Chopim, rio Iguaçu drainage, Paraná, Brazil (not preserved). A. angiru, male and female in breeding colors guarding fry, same locality (not preserved). All A. angiru photographs courtesy of Wolfgang Staeck." httpUri="https://zenodo.org/record/200612/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
|
||
). These specimens have a yellow ground coloration with dark vertical bars and a dark horizontal stripe. Several other species of
|
||
<taxonomicName id="B586A3848617FFCB26F1FB02FA9DFB1D" box="[1218,1375,1160,1184]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis id="40F204158617FFCB26F1FB02FA9DFB1D" box="[1218,1375,1160,1184]" italics="true" pageId="16" pageNumber="17">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
have a yellowish ground color, but it is best developed in
|
||
<taxonomicName id="B586A3848617FFCB20ECFB26FC8BFB79" box="[735,841,1196,1220]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB20ECFB26FC8BFB79" box="[735,841,1196,1220]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
. The iris is also yellow. The caudal fin has red dorsal and ventral margins and corners. This character is not unique for
|
||
<taxonomicName id="B586A3848617FFCB2143FB5AFC1BFB55" box="[880,985,1232,1256]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB2143FB5AFC1BFB55" box="[880,985,1232,1256]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
, and can also be seen in
|
||
<taxonomicName id="B586A3848617FFCB26DCFB5AFAAFFB55" box="[1263,1389,1232,1256]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB26DCFB5AFAAFFB55" box="[1263,1389,1232,1256]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
and in populations of
|
||
<taxonomicName id="B586A3848617FFCB236CFB7EFE0CFAB1" box="[351,462,1268,1292]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="facetus">
|
||
<emphasis id="40F204158617FFCB236CFB7EFE0CFAB1" box="[351,462,1268,1292]" italics="true" pageId="16" pageNumber="17">A. facetus</emphasis>
|
||
</taxonomicName>
|
||
from the state of
|
||
<collectingCountry id="0A9198978617FFCB20A8FB7FFD3DFAB0" box="[667,767,1269,1293]" name="Uruguay" pageId="16" pageNumber="17">Uruguay</collectingCountry>
|
||
. Breeding animals have the typical
|
||
<taxonomicName id="B586A3848617FFCB26A7FB7EFAF3FAB1" box="[1172,1329,1268,1292]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis id="40F204158617FFCB26A7FB7EFAF3FAB1" box="[1172,1329,1268,1292]" italics="true" pageId="16" pageNumber="17">Australoheros</emphasis>
|
||
</taxonomicName>
|
||
breeding coloration with the horizontal interruption of the black vertical bars in their dorsal portion between the opercle and the midlateral blotch (
|
||
<bibRefCitation id="1617A5F68617FFCB23A3FABCFD6AFAE9" author="Rican" box="[400,680,1334,1364]" pageId="16" pageNumber="17" refString="Rican, O. & Kullander, S. O. (2003) ' Cichlasoma' scitulum: a new species of cichlid fish from the Rio de La Plata Region in Argentina, Brazil, and Uruguay. Copeia, 2003, 794 - 802." type="journal article" year="2003">Říčan & Kullander 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1617A5F68617FFCB2087FAB6FC8CFAE9" author="Staeck" box="[692,846,1340,1364]" pageId="16" pageNumber="17" refString="Staeck, W. (1998 a) Neuer Chanchito. DCG informationen, 29, 62 - 63." type="journal article" year="1998" yearSuffix="a">Staeck 1998a</bibRefCitation>
|
||
: p. 82, 1998b: p. 62, 2003: p. 65). Females in breeding coloration develop a black blotch in the dorsal fin.
|
||
<bibRefCitation id="1617A5F68617FFCB212DFAD5FC06FACA" author="Staeck" box="[798,964,1375,1399]" pageId="16" pageNumber="17" refString="Staeck, W. (1998 b) Ein neuer cichlide aus dem Cichlasoma facetum komplex. DCG informationen, 29, 81 - 85." type="journal article" year="1998" yearSuffix="b">Staeck (1998b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1617A5F68617FFCB21E0FAD5FBCFFACA" author="Staeck" box="[979,1037,1375,1399]" pageId="16" pageNumber="17" refString="Staeck, W. (2003) " Cichlasoma " facetum komplex: Ein ruckblick auf die Anfange der Aquaristik. Die Aquarien- und Terrarienzeitschrift, 9, 60 - 65." type="journal article" year="2003">2003</bibRefCitation>
|
||
) describes behavior and spawning under aquarium conditions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A9C8B8C8617FFCB22F4FA2DFB3DFA59" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="7239D8078617FFCB22F4FA2DFB3DFA59" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
<emphasis id="40F204158617FFCB22F4FA2DFEA2FA7D" bold="true" box="[199,352,1447,1472]" pageId="16" pageNumber="17">Distribution.</emphasis>
|
||
<taxonomicName id="B586A3848617FFCB235BFA22FD9AFA7D" box="[360,600,1448,1472]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB235BFA22FD9AFA7D" box="[360,600,1448,1472]" italics="true" pageId="16" pageNumber="17">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
has a disjunct distribution in the rio Iguaçu and in the upper rio
|
||
<collectingCountry id="0A9198978617FFCB270AFA2DFA5AFA02" box="[1337,1432,1447,1471]" name="Uruguay" pageId="16" pageNumber="17">Uruguai</collectingCountry>
|
||
. One locality is so far known from the middle río
|
||
<collectingCountry id="0A9198978617FFCB208FFA47FCDCFA58" box="[700,798,1485,1509]" name="Uruguay" pageId="16" pageNumber="17">Uruguay</collectingCountry>
|
||
in Misiones province,
|
||
<collectingCountry id="0A9198978617FFCB262CFA46FB4DFA59" box="[1055,1167,1484,1508]" name="Argentina" pageId="16" pageNumber="17">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="EABDC4828617FFCB26ADFA46FB33FA59" box="[1182,1265,1484,1508]" captionStart="FIGURE 10" captionStartId="18.[151,250,894,917]" captionTargetBox="[151,1434,195,872]" captionTargetId="figure@18.[151,1436,195,872]" captionTargetPageId="18" captionText="FIGURE 10. Map of the middle Río de la Plata basin. Distributions of the two new species (A. angiru and A. ykeregua) and their relatives, as well as five areas of endemism are shown. Percent values and corresponding arrows demonstrate sequence divergences in the cyt b gene (see Fig. 2) between the species and areas of endemism in the río Iguazú and río Uruguay river drainages (plus the arroyo Urugua – í). Divergence of A. ykeregua from its sister species A. forquilha is 2.3 %. This divergence probably represents the minimum age of the Salto Moconá. Divergence of A. kaaygua from its sister species A. tembe is 3.8 %, and of A. angiru from A. tembe is similarly 3.6 – 3.7 %. This probably represents the age of the division of the arroyo Urugua – í from the río Iguazú. Divergence of A. angiru from its sister species A. minuano is 4.2 %. This is likely a divergence of the rio Iguaçu and río Uruguay drainages. Divergence of A. angiru from A. kaaygua (4.8 %), two unrelated species endemic to the río Iguazú river drainage, demonstrates an old divergence within the Iguazú drainage basin itself. See Discussion for more detailed description of the biogeography." httpUri="https://zenodo.org/record/200615/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A9C8B8C8617FFCA22F4FA7AFC3AF89A" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="7239D8078617FFCB22F4FA7AFD8AF9C9" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
<emphasis id="40F204158617FFCB22F4FA7AFE8CF9B5" bold="true" box="[199,334,1520,1544]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
The
|
||
<taxonomicName id="B586A3848617FFCB23B5FA65FE22F9BA" box="[390,480,1519,1543]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="21" phylum="Chordata" rank="species" species="guarani">Guaraní</taxonomicName>
|
||
word
|
||
<taxonomicName id="B586A3848617FFCB201BFA7AFDB1F9B5" box="[552,627,1520,1544]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB201BFA7AFDB1F9B5" box="[552,627,1520,1544]" italics="true" pageId="16" pageNumber="17">angirû</emphasis>
|
||
</taxonomicName>
|
||
means friend, partner (amigo or compañero in Spanish). The etymology is based on the fact that
|
||
<taxonomicName id="B586A3848617FFCB239EF99EFDDBF991" box="[429,537,1556,1580]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB239EF99EFDDBF991" box="[429,537,1556,1580]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848617FFCB2062F99EFD13F991" box="[593,721,1556,1580]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB2062F99EFD13F991" box="[593,721,1556,1580]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
have been confused as one species (
|
||
<bibRefCitation id="1617A5F68617FFCB2647F984FA53F991" author="Rican" box="[1140,1425,1550,1580]" pageId="16" pageNumber="17" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan & Kullander 2008</bibRefCitation>
|
||
). New data have however demonstrate that they are two non-sister group species living in the same river drainage (río Iguazú), though not sympatrically.
|
||
</paragraph>
|
||
<paragraph id="7239D8078617FFCB22F4F90AFBDFF861" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
|
||
<emphasis id="40F204158617FFCB22F4F90AFED3F925" bold="true" box="[199,273,1664,1688]" pageId="16" pageNumber="17">Notes.</emphasis>
|
||
Part of
|
||
<taxonomicName id="B586A3848617FFCB235FF90AFD9EF925" box="[364,604,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB235FF90AFD9EF925" box="[364,604,1664,1688]" italics="true" pageId="16" pageNumber="17">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
material (MCP 6262) has been previously considered conspecific with
|
||
<taxonomicName id="B586A3848617FFCB27B7F90AFDF1F901" authority="Rican & Kullander 2008" authorityName="Rican & Kullander" authorityYear="2008" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB27B7F90AFF34F901" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
(
|
||
<bibRefCitation id="1617A5F68617FFCB233AF914FDE9F901" author="Rican" box="[265,555,1694,1724]" pageId="16" pageNumber="17" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan & Kullander 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The authors were aware of the morphological variation within
|
||
<taxonomicName id="B586A3848617FFCB2728F92EFA5EF901" box="[1307,1436,1700,1724]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB2728F92EFA5EF901" box="[1307,1436,1700,1724]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="1617A5F68617FFCB22DAF948FDC9F962" author="Rican" box="[233,523,1730,1760]" pageId="16" pageNumber="17" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan & Kullander 2008</bibRefCitation>
|
||
), but lack of DNA data and of first hand examination of the
|
||
<typeStatus id="AD3D66A58617FFCB26EEF942FACFF95D" box="[1245,1293,1736,1760]" pageId="16" pageNumber="17">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="B586A3848617FFCB27B7F942FF34F8B9" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB27B7F942FF34F8B9" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
made them sceptical about describing a new species with an additionally unusual distribution (occuring in the same river basin, río Iguazú as
|
||
<taxonomicName id="B586A3848617FFCB2012F89AFD62F895" box="[545,672,1808,1832]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB2012F89AFD62F895" box="[545,672,1808,1832]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
, but not in sympatry, and at the same time also in the río
|
||
<collectingCountry id="0A9198978617FFCB2719F885FA53F89A" box="[1322,1425,1807,1831]" name="Uruguay" pageId="16" pageNumber="17">Uruguay</collectingCountry>
|
||
). DNA data from the rio Iguaçu populations in
|
||
<collectingCountry id="0A9198978617FFCB20A5F8BEFD18F8F1" box="[662,730,1844,1868]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
|
||
(
|
||
<taxonomicName id="B586A3848617FFCB20D9F8BEFC96F8F1" box="[746,852,1844,1868]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB20D9F8BEFC96F8F1" box="[746,852,1844,1868]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
) however show no relationship to
|
||
<taxonomicName id="B586A3848617FFCB26EBF8BEFA94F8F1" box="[1240,1366,1844,1868]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB26EBF8BEFA94F8F1" box="[1240,1366,1844,1868]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
in the río Iguazú in
|
||
<collectingCountry id="0A9198978617FFCB231FF8DDFE59F8D2" box="[300,411,1879,1903]" name="Argentina" pageId="16" pageNumber="17">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="EABDC4828617FFCB2399F8DDFE2DF8D2" box="[426,495,1879,1903]" captionStart="FIGURE 2" captionStartId="6.[151,250,1938,1961]" captionTargetBox="[167,1388,208,1889]" captionTargetId="figure@6.[151,1436,161,1917]" captionTargetPageId="6" captionText="FIGURE 2. Molecular phylogeny of the Río de la Plata basin Australoheros species using BI. Node support values shown for MP / BI analyses. The alternative dotted topology represents neighbor-joining (NJ) analysis. Asterisk denotes posterior probability of 1.00." httpUri="https://zenodo.org/record/200607/files/figure.png" pageId="16" pageNumber="17">Fig. 2</figureCitation>
|
||
). DNA data from the río
|
||
<collectingCountry id="0A9198978617FFCB2134F8D3FCABF8CC" box="[775,873,1881,1905]" name="Uruguay" pageId="16" pageNumber="17">Uruguay</collectingCountry>
|
||
are so far lacking. A more detailed morphological analysis (
|
||
<figureCitation id="EABDC4828617FFCB2332F8F6FE80F829" box="[257,322,1916,1940]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1960]" captionTargetBox="[165,1418,1057,1919]" captionTargetId="figure@5.[151,1436,1028,1930]" captionTargetPageId="5" captionText="FIGURE 1. Tree-based delimitation using MP phylogenetic analysis of morphological data. The tree shown is one of two MP trees (L = 693; N = 2; CI = 0.51; RI = 0.66), which differ only in the internal topology of A. angiru. Branch lengths represent morphological divergences." httpUri="https://zenodo.org/record/200606/files/figure.png" pageId="16" pageNumber="17">Fig. 1</figureCitation>
|
||
) also supports the notion of two unrelated species, with populations of
|
||
<taxonomicName id="B586A3848617FFCB2659F8F6FB17F829" box="[1130,1237,1916,1940]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB2659F8F6FB17F829" box="[1130,1237,1916,1940]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
from both the rio Iguaçu and from the río
|
||
<collectingCountry id="0A9198978617FFCB239EF82BFDCDF804" box="[429,527,1953,1977]" name="Uruguay" pageId="16" pageNumber="17">Uruguay</collectingCountry>
|
||
forming a homogenous clade with short intraspecific branch lengths. The sister species of
|
||
<taxonomicName id="B586A3848617FFCB233EF84EFEB5F861" box="[269,375,1988,2012]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158617FFCB233EF84EFEB5F861" box="[269,375,1988,2012]" italics="true" pageId="16" pageNumber="17">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="B586A3848617FFCB23A4F84EFDDAF861" box="[407,536,1988,2012]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="40F204158617FFCB23A4F84EFDDAF861" box="[407,536,1988,2012]" italics="true" pageId="16" pageNumber="17">A. minuano</emphasis>
|
||
</taxonomicName>
|
||
, while that of
|
||
<taxonomicName id="B586A3848617FFCB2088F84EFCFBF861" box="[699,825,1988,2012]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158617FFCB2088F84EFCFBF861" box="[699,825,1988,2012]" italics="true" pageId="16" pageNumber="17">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="B586A3848617FFCB2169F84EFC7EF861" box="[858,956,1988,2012]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="tembe">
|
||
<emphasis id="40F204158617FFCB2169F84EFC7EF861" box="[858,956,1988,2012]" italics="true" pageId="16" pageNumber="17">A. tembe</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EABDC4828617FFCB21F8F84EFBCDF861" box="[971,1039,1988,2012]" captionStart="FIGURE 3" captionStartId="7.[151,250,1124,1147]" captionTargetBox="[166,1418,214,1073]" captionTargetId="figure@7.[151,1436,193,1103]" captionTargetPageId="7" captionText="FIGURE 3. Combined MP morphological-molecular phylogeny with between-state scaling internal weighting between morphological and molecular data (L = 2457; N = 1; CI = 0.58; RI = 0.49). Node support values show MP bootstrap for two types of analyses (left: between-state scaling internal weighting structure / right: all characters weighted equally)." httpUri="https://zenodo.org/record/200608/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6FF1DFE43FEA1" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
The MCP 6262 lot additionally included two species (
|
||
<bibRefCitation id="1617A5F68616FFCA211CFF18FBA6FF12" author="Rican" box="[815,1124,146,176]" pageId="17" pageNumber="18" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan and Kullander, 2008</bibRefCitation>
|
||
). Nine specimens from this lot are
|
||
<typeStatus id="AD3D66A58616FFCA22D7FF37FE8DFF68" box="[228,335,189,213]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="B586A3848616FFCA2346FF36FC7AFF69" authority="Rican and Kullander, 2008" authorityName="Rican and Kullander" authorityYear="2008" box="[373,952,182,212]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="forquilha">
|
||
<emphasis id="40F204158616FFCA2346FF36FD43FF69" box="[373,641,188,212]" italics="true" pageId="17" pageNumber="18">Australoheros forquilha</emphasis>
|
||
<bibRefCitation id="1617A5F68616FFCA20BBFF3CFC7AFF69" author="Rican" box="[648,952,182,212]" pageId="17" pageNumber="18" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan and Kullander, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
. Thirteen specimens from this lot represent
|
||
<taxonomicName id="B586A3848616FFCA22A4FF6AFEC0FF45" box="[151,258,224,248]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA22A4FF6AFEC0FF45" box="[151,258,224,248]" italics="true" pageId="17" pageNumber="18">A. angiru</emphasis>
|
||
</taxonomicName>
|
||
(previously erroneously treated as
|
||
<taxonomicName id="B586A3848616FFCA20A0FF6AFCD0FF45" box="[659,786,224,248]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158616FFCA20A0FF6AFCD0FF45" box="[659,786,224,248]" italics="true" pageId="17" pageNumber="18">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="1617A5F68616FFCA210BFF50FBA9FF4A" author="Rican" box="[824,1131,218,248]" pageId="17" pageNumber="18" refString="Rican, O. & Kullander, S. O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724, 1 - 51." type="journal article" year="2008">Říčan and Kullander, 2008</bibRefCitation>
|
||
) and were separated into a new lot MCP 46328.
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6FEADFCECFDE2" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
Additional diagnostic characters of
|
||
<taxonomicName id="B586A3848616FFCA205BFEA2FC9FFEFD" box="[616,861,296,320]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA205BFEA2FC9FFEFD" box="[616,861,296,320]" italics="true" pageId="17" pageNumber="18">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
that separate it from all other species except
|
||
<taxonomicName id="B586A3848616FFCA27B7FEA2FF34FED9" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kaaygua">
|
||
<emphasis id="40F204158616FFCA27B7FEA2FF34FED9" italics="true" pageId="17" pageNumber="18">A. kaaygua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848616FFCA2318FEC6FE69FED9" box="[299,427,332,356]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="minuano">
|
||
<emphasis id="40F204158616FFCA2318FEC6FE69FED9" box="[299,427,332,356]" italics="true" pageId="17" pageNumber="18">A. minuano</emphasis>
|
||
</taxonomicName>
|
||
are as follows. It is distinguished (in decreasing order of overall similarity; except for species from coastal drainages treated as last) from
|
||
<taxonomicName id="B586A3848616FFCA2085FEFAFCEDFE35" box="[694,815,368,392]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="charrua">
|
||
<emphasis id="40F204158616FFCA2085FEFAFCEDFE35" box="[694,815,368,392]" italics="true" pageId="17" pageNumber="18">A. charrua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848616FFCA2155FEFAFC19FE35" box="[870,987,368,392]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="scitulus">
|
||
<emphasis id="40F204158616FFCA2155FEFAFC19FE35" box="[870,987,368,392]" italics="true" pageId="17" pageNumber="18">A. scitulus</emphasis>
|
||
</taxonomicName>
|
||
by having less scale rows between posterior end of upper lateral line and dorsal fin (ch3 state 2
|
||
<emphasis id="40F204158616FFCA213DFE1FFCEFFE11" box="[782,813,405,428]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0
|
||
<emphasis id="40F204158616FFCA2174FE1FFCA4FE11" box="[839,870,405,428]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
1), less caudal vertebrae (13
|
||
<emphasis id="40F204158616FFCA2699FE1FFB03FE11" box="[1194,1217,405,428]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 14), in being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA2365FE33FEAFFE6D" box="[342,365,441,464]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 45% SL), and in having less E0 scales (24
|
||
<emphasis id="40F204158616FFCA2157FE33FCB9FE6D" box="[868,891,441,464]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
.>25). Additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2773FE32FF7CFE49" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="charrua">
|
||
<emphasis id="40F204158616FFCA2773FE32FF7CFE49" italics="true" pageId="17" pageNumber="18">A. charrua</emphasis>
|
||
</taxonomicName>
|
||
by details in the shape of the midlatral stripe (see description) and by lacking a pinkish body coloration. Additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA23FEFD8AFD83FDA5" box="[461,577,512,536]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="scitulus">
|
||
<emphasis id="40F204158616FFCA23FEFD8AFD83FDA5" box="[461,577,512,536]" italics="true" pageId="17" pageNumber="18">A. scitulus</emphasis>
|
||
</taxonomicName>
|
||
by lacking black blotches on the opercular series, having less anal fin spines (7
|
||
<emphasis id="40F204158616FFCA2286FDAFFF0EFD81" box="[181,204,549,572]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 8), less dorsal fin spines (16
|
||
<emphasis id="40F204158616FFCA202AFDAFFDF2FD81" box="[537,560,549,572]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 17), less caudal vertebrae (13
|
||
<emphasis id="40F204158616FFCA21BFFDAFFC61FD81" box="[908,931,549,572]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 14), in being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA2777FDAFFA99FD81" box="[1348,1371,549,572]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 45% SL), and in having less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA2081FDC3FD0BFDDD" box="[690,713,585,608]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14).
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6FDE6FD1CFCE1" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B586A3848616FFCA22F6FDE6FE77FD39" box="[197,437,620,644]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA22F6FDE6FE77FD39" box="[197,437,620,644]" italics="true" pageId="17" pageNumber="18">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2081FDE6FCD7FD39" box="[690,789,620,644]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tembe">
|
||
<emphasis id="40F204158616FFCA2081FDE6FCD7FD39" box="[690,789,620,644]" italics="true" pageId="17" pageNumber="18">A. tembe</emphasis>
|
||
</taxonomicName>
|
||
by having less scale rows between anterior end of dorsal fin and upper lateral line (ch4 states 1–2
|
||
<emphasis id="40F204158616FFCA205BFD1BFD45FD15" box="[616,647,657,680]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), by a very narrow or missing caudal base spot, a shorter dorsal fin scale cover (ch1 state 1
|
||
<emphasis id="40F204158616FFCA2391FD3FFE02FD71" box="[418,448,693,716]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), less scale rows between the posterior end of the upper lateral line and dorsal fin (ch3 state 2
|
||
<emphasis id="40F204158616FFCA22DBFD53FEC4FD4D" box="[232,262,729,752]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), by lacking thick lips, by having more anal fin spines (7
|
||
<emphasis id="40F204158616FFCA219EFD53FC06FD4D" box="[941,964,729,752]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 6), less caudal vertebrae (13
|
||
<emphasis id="40F204158616FFCA272AFD53FAF2FD4D" box="[1305,1328,729,752]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 14), and less caudal peduncle vertebrae (0
|
||
<emphasis id="40F204158616FFCA2025FD77FDEFFCA9" box="[534,557,765,788]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 3). It is distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2140FD76FC2EFCA9" box="[883,1004,764,788]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="guarani">
|
||
<emphasis id="40F204158616FFCA2140FD76FC2EFCA9" box="[883,1004,764,788]" italics="true" pageId="17" pageNumber="18">A. guarani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA21C8FD76FBA8FCA9" box="[1019,1130,764,788]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="facetus">
|
||
<emphasis id="40F204158616FFCA21C8FD76FBA8FCA9" box="[1019,1130,764,788]" italics="true" pageId="17" pageNumber="18">A. facetus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA264AFD76FAC5FCA9" box="[1145,1287,764,788]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="acaroides">
|
||
<emphasis id="40F204158616FFCA264AFD76FAC5FCA9" box="[1145,1287,764,788]" italics="true" pageId="17" pageNumber="18">A. acaroides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848616FFCA270CFD76FA5EFCA9" box="[1343,1436,764,788]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="taura">
|
||
<emphasis id="40F204158616FFCA270CFD76FA5EFCA9" box="[1343,1436,764,788]" italics="true" pageId="17" pageNumber="18">A. taura</emphasis>
|
||
</taxonomicName>
|
||
by a large and dominant midlateral blotch (except
|
||
<taxonomicName id="B586A3848616FFCA20FFFCAAFCF8FC85" box="[716,826,800,824]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="facetus">
|
||
<emphasis id="40F204158616FFCA20FFFCAAFCF8FC85" box="[716,826,800,824]" italics="true" pageId="17" pageNumber="18">A. facetus</emphasis>
|
||
</taxonomicName>
|
||
), very narrow or missing caudal base spot, and details in the shape of the midlatral stripe (see description).
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6FCE2FB9AFB79" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B586A3848616FFCA22F6FCE2FE74FC3D" box="[197,438,872,896]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA22F6FCE2FE74FC3D" box="[197,438,872,896]" italics="true" pageId="17" pageNumber="18">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2170FCE2FC7FFC3D" box="[835,957,872,896]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="guarani">
|
||
<emphasis id="40F204158616FFCA2170FCE2FC7FFC3D" box="[835,957,872,896]" italics="true" pageId="17" pageNumber="18">A. guarani</emphasis>
|
||
</taxonomicName>
|
||
by a small terminal or subterminal mouth (
|
||
<emphasis id="40F204158616FFCA2293FC07FF75FC19" box="[160,183,909,932]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. large supraterminal), more anal fin spines (7
|
||
<emphasis id="40F204158616FFCA208BFC07FD0DFC19" box="[696,719,909,932]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 6), shorter preorbital distance (21
|
||
<emphasis id="40F204158616FFCA2665FC07FBB7FC19" box="[1110,1141,909,932]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
25% HL), and less C1 gill rakers (6
|
||
<emphasis id="40F204158616FFCA2337FC3BFED9FC75" box="[260,283,945,968]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 7). Additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA20F4FC3AFCF5FC75" box="[711,823,944,968]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="facetus">
|
||
<emphasis id="40F204158616FFCA20F4FC3AFCF5FC75" box="[711,823,944,968]" italics="true" pageId="17" pageNumber="18">A. facetus</emphasis>
|
||
</taxonomicName>
|
||
by a longer dorsal fin scale cover (ch1 state 1
|
||
<emphasis id="40F204158616FFCA2765FC3BFAB7FC75" box="[1366,1397,945,968]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
2), more anal fin spines (7
|
||
<emphasis id="40F204158616FFCA2391FC5FFE7BFC51" box="[418,441,981,1004]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 6), less anal fin rays (7
|
||
<emphasis id="40F204158616FFCA20E0FC5FFD28FC51" box="[723,746,981,1004]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 8), less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA265BFC5FFBBDFC51" box="[1128,1151,981,1004]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14), and less C1 gill rakers (6
|
||
<emphasis id="40F204158616FFCA2331FC73FEDBFBAD" box="[258,281,1017,1040]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 7–8). It is additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2136FC72FC50FBAD" box="[773,914,1016,1040]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="acaroides">
|
||
<emphasis id="40F204158616FFCA2136FC72FC50FBAD" box="[773,914,1016,1040]" italics="true" pageId="17" pageNumber="18">A. acaroides</emphasis>
|
||
</taxonomicName>
|
||
by a longer dorsal fin scale cover (ch1 state 1
|
||
<emphasis id="40F204158616FFCA22A4FB97FF77FB89" box="[151,181,1053,1076]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
2), shorter caudal peduncle (40% CPD
|
||
<emphasis id="40F204158616FFCA20B7FB97FD59FB89" box="[644,667,1053,1076]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 50–60% CPD), by being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA26E6FB97FB2EFB89" box="[1237,1260,1053,1076]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 45% SL), and having a narrower interorbital distance (35
|
||
<emphasis id="40F204158616FFCA20B8FBCBFD60FBE5" box="[651,674,1089,1112]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 40–45% HL). It is distinguished from
|
||
<taxonomicName id="B586A3848616FFCA265CFBCAFB0CFBE5" box="[1135,1230,1088,1112]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="taura">
|
||
<emphasis id="40F204158616FFCA265CFBCAFB0CFBE5" box="[1135,1230,1088,1112]" italics="true" pageId="17" pageNumber="18">A. taura</emphasis>
|
||
</taxonomicName>
|
||
by also lacking a pinkish body coloration, by a small terminal or subterminal mouth (
|
||
<emphasis id="40F204158616FFCA21A1FBEFFC6BFBC1" box="[914,937,1125,1148]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. large supraterminal), shorter caudal peduncle (40% CPD
|
||
<emphasis id="40F204158616FFCA2373FB03FE95FB1D" box="[320,343,1161,1184]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 50% CPD), by being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA2166FB03FCAEFB1D" box="[853,876,1161,1184]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 40% SL), by a narrower interorbital distance (35
|
||
<emphasis id="40F204158616FFCA22A4FB27FF6CFB79" box="[151,174,1197,1220]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 40% HL), less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA2044FB27FD4CFB79" box="[631,654,1197,1220]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14), and less E0 scales (24
|
||
<emphasis id="40F204158616FFCA21C5FB27FBCFFB79" box="[1014,1037,1197,1220]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
.>25).
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6FB5AFCE9F9ED" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B586A3848616FFCA22F6FB5AFE75FB55" box="[197,439,1232,1256]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA22F6FB5AFE75FB55" box="[197,439,1232,1256]" italics="true" pageId="17" pageNumber="18">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="B586A3848616FFCA208FFB5AFC86FB55" box="[700,836,1232,1256]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ykeregua">
|
||
<emphasis id="40F204158616FFCA208FFB5AFC86FB55" box="[700,836,1232,1256]" italics="true" pageId="17" pageNumber="18">A. ykeregua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848616FFCA214CFB5AFBCAFB55" box="[895,1032,1232,1256]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="forquilha">
|
||
<emphasis id="40F204158616FFCA214CFB5AFBCAFB55" box="[895,1032,1232,1256]" italics="true" pageId="17" pageNumber="18">A. forquilha</emphasis>
|
||
</taxonomicName>
|
||
by a shorter dorsal fin scale cover (ch1 state 1
|
||
<emphasis id="40F204158616FFCA2317FB7FFE81FAB1" box="[292,323,1269,1292]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), a different scale pattern along anterior border of dorsal fin (ch2 state 0
|
||
<emphasis id="40F204158616FFCA2697FB7FFB00FAB1" box="[1188,1218,1269,1292]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
1), less scale rows between posterior end of upper lateral line and dorsal fin (ch3 state 2
|
||
<emphasis id="40F204158616FFCA218BFA93FC14FA8D" box="[952,982,1305,1328]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), very narrow or missing caudal base spot, absence of opalescent spots below orbit, unpaired fins without checker-board spotted pattern, absence of red colored lower head area and opercular membrane, by a small terminal or subterminal mouth (
|
||
<emphasis id="40F204158616FFCA2685FAEBFB0FFAC5" box="[1206,1229,1377,1400]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. large supraterminal), less dorsal fin rays (9
|
||
<emphasis id="40F204158616FFCA23FCFA0FFE24FA21" box="[463,486,1413,1436]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 10), less caudal peduncle vertebrae (0
|
||
<emphasis id="40F204158616FFCA2198FA0FFC00FA21" box="[939,962,1413,1436]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 2
|
||
<emphasis id="40F204158616FFCA21DBFA0FFC3DFA21" box="[1000,1023,1413,1436]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 2.5), shorter caudal peduncle (40% CPD
|
||
<emphasis id="40F204158616FFCA22E7FA23FF29FA7D" box="[212,235,1449,1472]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 60% CPD), by being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA20D9FA23FCC3FA7D" box="[746,769,1449,1472]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 45
|
||
<emphasis id="40F204158616FFCA2100FA23FC88FA7D" box="[819,842,1449,1472]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 40% SL), with a wider head (55
|
||
<emphasis id="40F204158616FFCA26F4FA23FB1CFA7D" box="[1223,1246,1449,1472]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. <50% HL), and in having less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA2074FA47FD9CFA59" box="[583,606,1485,1508]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14). Additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA2607FA46FB7BFA59" box="[1076,1209,1484,1508]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ykeregua">
|
||
<emphasis id="40F204158616FFCA2607FA46FB7BFA59" box="[1076,1209,1484,1508]" italics="true" pageId="17" pageNumber="18">A. ykeregua</emphasis>
|
||
</taxonomicName>
|
||
by a large and dominant midlateral blotch, and more anal fin spines (7
|
||
<emphasis id="40F204158616FFCA20C1FA7BFCCBF9B5" box="[754,777,1521,1544]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 6). Additionally distinguished from
|
||
<taxonomicName id="B586A3848616FFCA268AFA7AFA80F9B5" box="[1209,1346,1520,1544]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="forquilha">
|
||
<emphasis id="40F204158616FFCA268AFA7AFA80F9B5" box="[1209,1346,1520,1544]" italics="true" pageId="17" pageNumber="18">A. forquilha</emphasis>
|
||
</taxonomicName>
|
||
by less scale rows between anterior end of dorsal fin and upper lateral line (ch4 state 1
|
||
<emphasis id="40F204158616FFCA2628F99FFBFBF991" box="[1051,1081,1557,1580]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
0), absence of opalescent scale rows on body, and less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA209CF9B3FD04F9ED" box="[687,710,1593,1616]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14).
|
||
</paragraph>
|
||
<paragraph id="7239D8078616FFCA22F6F9D6FC3AF89A" blockId="17.[151,1437,146,1833]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B586A3848616FFCA22F6F9D6FE76F9C9" box="[197,436,1628,1652]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="angiru">
|
||
<emphasis id="40F204158616FFCA22F6F9D6FE76F9C9" box="[197,436,1628,1652]" italics="true" pageId="17" pageNumber="18">Australoheros angiru</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all the Atlantic coast species north of
|
||
<taxonomicName id="B586A3848616FFCA2666F9D6FB23F9C9" box="[1109,1249,1628,1652]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="acaroides">
|
||
<emphasis id="40F204158616FFCA2666F9D6FB23F9C9" box="[1109,1249,1628,1652]" italics="true" pageId="17" pageNumber="18">A. acaroides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B586A3848616FFCA272BF9D6FAB6F9C9" box="[1304,1396,1628,1652]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="taura">
|
||
<emphasis id="40F204158616FFCA272BF9D6FAB6F9C9" box="[1304,1396,1628,1652]" italics="true" pageId="17" pageNumber="18">A. taura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="B586A3848616FFCA27B7F9D6FF28F925" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="autrani">
|
||
<emphasis id="40F204158616FFCA27B7F9D6FF28F925" italics="true" pageId="17" pageNumber="18">A. autrani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA22C8F90AFE47F925" box="[251,389,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="barbosae">
|
||
<emphasis id="40F204158616FFCA22C8F90AFE47F925" box="[251,389,1664,1688]" italics="true" pageId="17" pageNumber="18">A. barbosae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA23A4F90AFDE3F925" box="[407,545,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="capixaba">
|
||
<emphasis id="40F204158616FFCA23A4F90AFDE3F925" box="[407,545,1664,1688]" italics="true" pageId="17" pageNumber="18">A. capixaba</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA2001F90AFD2BF925" box="[562,745,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ipatinguensis">
|
||
<emphasis id="40F204158616FFCA2001F90AFD2BF925" box="[562,745,1664,1688]" italics="true" pageId="17" pageNumber="18">A. ipatinguensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA20C8F90AFC6FF925" box="[763,941,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="macacuensis">
|
||
<emphasis id="40F204158616FFCA20C8F90AFC6FF925" box="[763,941,1664,1688]" italics="true" pageId="17" pageNumber="18">A. macacuensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA218DF90AFB97F925" box="[958,1109,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="macaensis">
|
||
<emphasis id="40F204158616FFCA218DF90AFB97F925" box="[958,1109,1664,1688]" italics="true" pageId="17" pageNumber="18">A. macaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA2654F90AFB1BF925" box="[1127,1241,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="muriae">
|
||
<emphasis id="40F204158616FFCA2654F90AFB1BF925" box="[1127,1241,1664,1688]" italics="true" pageId="17" pageNumber="18">A. muriae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA26D8F90AFAB0F925" box="[1259,1394,1664,1688]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="paraibae">
|
||
<emphasis id="40F204158616FFCA26D8F90AFAB0F925" box="[1259,1394,1664,1688]" italics="true" pageId="17" pageNumber="18">A. paraibae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA27B7F90AFF33F901" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ribeirae">
|
||
<emphasis id="40F204158616FFCA27B7F90AFF33F901" italics="true" pageId="17" pageNumber="18">A. ribeirae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA2332F92EFE43F901" box="[257,385,1700,1724]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="40F204158616FFCA2332F92EFE43F901" box="[257,385,1700,1724]" italics="true" pageId="17" pageNumber="18">A. robustus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B586A3848616FFCA23A1F92EFDEDF901" box="[402,559,1700,1724]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="saquarema">
|
||
<emphasis id="40F204158616FFCA23A1F92EFDEDF901" box="[402,559,1700,1724]" italics="true" pageId="17" pageNumber="18">A. saquarema</emphasis>
|
||
</taxonomicName>
|
||
) by a longer dorsal fin scale cover (ch1 state 1
|
||
<emphasis id="40F204158616FFCA2668F92FFBBBF901" box="[1115,1145,1701,1724]" italics="true" pageId="17" pageNumber="18">vs.</emphasis>
|
||
2), a large and dominant midlateral blotch, details in the shape of the midlatral stripe (see description), shorter caudal peduncle (40% CPD
|
||
<emphasis id="40F204158616FFCA22A4F967FF6CF8B9" box="[151,174,1773,1796]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
.>50% CPD), in being more deep-bodied (50
|
||
<emphasis id="40F204158616FFCA20FBF967FD1DF8B9" box="[712,735,1773,1796]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 45% SL), with a narrower interorbital distance (35
|
||
<emphasis id="40F204158616FFCA2772F967FA9AF8B9" box="[1345,1368,1773,1796]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 40% HL), less pectoral fin rays (12–13
|
||
<emphasis id="40F204158616FFCA202BF89BFDEDF895" box="[536,559,1809,1832]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
. 13–14), and less E0 scales (24
|
||
<emphasis id="40F204158616FFCA21A4F89BFC6CF895" box="[919,942,1809,1832]" italics="true" pageId="17" pageNumber="18">vs</emphasis>
|
||
.>25).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |