462 lines
65 KiB
XML
462 lines
65 KiB
XML
<document id="59828CE458A2383E707FA1DE1CC7DFAC" ID-DOI="10.12782/specdiv.25.49" ID-GBIF-Dataset="7259cd46-a1db-4d03-bdd6-5993474ded22" ID-ISSN="2189-7301" ID-Zenodo-Dep="3751644" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586877699773" checkinUser="felipe" docAuthor="Fujiwara, Kyoji & Motomura, Hiroyuki" docDate="2020" docId="FA1F87F3FFDAFF865CE2F8C28716DA14" docLanguage="en" docName="SpeciesDiversity.25.49-54.pdf.imf" docOrigin="Species Diversity 25" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Kopua nuimata Hardy 1984" docType="treatment" docVersion="5" lastPageNumber="53" masterDocId="0626FF8BFFDAFF825F37FF86862EDD0F" masterDocTitle="Revised Diagnosis of the Rare Clingfish Kopua nuimata (Gobiesocidae) with Notes on Fresh Coloration and First Australian Record" masterLastPageNumber="54" masterPageNumber="49" pageNumber="49" updateTime="1698737539097" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3631613C84A5024E36D311902D1A6057">Revised Diagnosis of the Rare Clingfish Kopua nuimata (Gobiesocidae) with Notes on Fresh Coloration and First Australian Record</mods:title>
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<mods:namePart id="EA006D39A26BB2B1BA13B2ECA5B0342E">Fujiwara, Kyoji</mods:namePart>
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<mods:affiliation id="920841258BA14D3A90E39BBB52B4F667">The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 - 0065, Japan</mods:affiliation>
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<mods:namePart id="F1AE5B74D498F28FDCC4829CFB489344">Motomura, Hiroyuki</mods:namePart>
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<mods:affiliation id="A101910883B6FC9B3B08DB010CE4F44D">The Kagoshima University Museum, 1 - 21 - 30 Korimoto, Kagoshima 890 - 0065, Japan</mods:affiliation>
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<mods:title id="8711F5C9303F65E5008183229CBCC81E">Species Diversity</mods:title>
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<mods:date id="B9E04470D3786B474C2A1ED579F32807">2020</mods:date>
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<taxonomicName id="B5B64D66FFDAFF825CE2F8C282D6DA53" ID-CoL="3RD48" authority="Hardy, 1984" authorityName="Hardy" authorityYear="1984" box="[981,1272,1860,1884]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="nuimata">
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<emphasis id="40C2EAF7FFDAFF825CE2F8C28259DA53" bold="true" box="[981,1143,1860,1884]" italics="true" pageId="0" pageNumber="49">Kopua nuimata</emphasis>
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<bibRefCitation id="16274B14FFDAFF825B4AF8C282D6DA53" author="Hardy, G. S." box="[1149,1272,1860,1884]" pageId="0" pageNumber="49" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy, 1984</bibRefCitation>
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</paragraph>
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<subSubSection id="3AAC656EFFDAFF825CF6F8E28322DA73" box="[961,1292,1892,1916]" pageId="0" pageNumber="49" type="vernacular_names">
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<paragraph id="720936E5FFDAFF825CF6F8E28322DA73" blockId="0.[961,1292,1892,1916]" box="[961,1292,1892,1916]" pageId="0" pageNumber="49">[English name: Bigeye Clingfish]</paragraph>
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</subSubSection>
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<subSubSection id="3AAC656EFFDAFF825B04F80382B7DA92" box="[1075,1177,1925,1949]" pageId="0" pageNumber="49" type="description">
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<paragraph id="720936E5FFDAFF825B04F80382B7DA92" blockId="0.[1075,1177,1925,1949]" box="[1075,1177,1925,1949]" pageId="0" pageNumber="49">
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(
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<figureCitation id="EA8D2A60FFDAFF825B0DF80382BFDA92" box="[1082,1169,1925,1949]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[156,187,1373,1395]" captionStartId-1="2.[156,187,1211,1233]" captionStartId-2="3.[156,187,1946,1968]" captionStartId-3="3.[821,852,2002,2024]" captionTargetBox-0="[188,1398,206,1331]" captionTargetBox-1="[189,1368,210,1135]" captionTargetBox-2="[160,764,772,1904]" captionTargetBox-3="[824,1428,1355,1959]" captionTargetId-0="figure@1.[187,1399,205,1332]" captionTargetId-1="figure@2.[187,1400,205,1170]" captionTargetId-2="figure@3.[158,766,770,1906]" captionTargetId-3="figure@3.[822,1430,1353,1961]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionText-0="Fig. 1. Preserved specimen of Kopua nuimata from Norfolk Ridge, south of Norfolk Island, Australia (CSIRO H6007-17, 29.0mm SL). A, lateral view; B, dorsal view; C, ventral view; D, radiograph." captionText-1="Fig. 2. Kopua nuimata (CSIRO H6007-17, 29.0mm SL), Norfolk Ridge, south of Norfolk Island, Australia. Photographed subsequent to thawing. Photographs provided by CSIRO." captionText-2="Fig. 3. Illustration of head sensory canal pores of Kopua nuimata, based on CSIRO H6007-17, 29.0mm SL. AN and PN, anterior and posterior nostrils, respectively; LC1–2, lacrimal canal pores; NC1–2, nasal canal pores; PO1, postorbital canal pore; PR2–3, preopercular canal pores. Bar indicates 1mm." captionText-3="Fig. 4. Distributional records of Kopua nuimata. Star and circles indicate new and previous records, respectively. Arrowhead indicates type locality." figureDoi-0="http://doi.org/10.5281/zenodo.3751646" figureDoi-1="http://doi.org/10.5281/zenodo.3751648" figureDoi-2="http://doi.org/10.5281/zenodo.3751650" figureDoi-3="http://doi.org/10.5281/zenodo.3751652" httpUri-0="https://zenodo.org/record/3751646/files/figure.png" httpUri-1="https://zenodo.org/record/3751648/files/figure.png" httpUri-2="https://zenodo.org/record/3751650/files/figure.png" httpUri-3="https://zenodo.org/record/3751652/files/figure.png" pageId="0" pageNumber="49">Figs 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3AAC656EFFDAFF835C02F841844BDB96" lastPageId="1" lastPageNumber="50" pageId="0" pageNumber="49" type="reference_group">
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<paragraph id="720936E5FFDAFF835C02F841844BDB96" blockId="0.[821,1431,1990,2079]" lastBlockId="1.[181,768,1469,1689]" lastPageId="1" lastPageNumber="50" pageId="0" pageNumber="49">
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<treatmentCitation id="F31710F4FFDAFF825C02F84182B1DAD1" author="Hardy, G. S." box="[821,1183,1990,2014]" page="246" pageId="0" pageNumber="49" year="1984">
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<taxonomicName id="B5B64D66FFDAFF825C02F84182B1DAD1" ID-CoL="3RD48" authority="Hardy, 1984: 246" authorityName="Hardy" authorityPageNumber="246" authorityYear="1984" box="[821,1183,1990,2014]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="0" pageNumber="49" phylum="Chordata" rank="species" species="nuimata">
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<emphasis id="40C2EAF7FFDAFF825C02F84185F9DAD1" box="[821,983,1991,2014]" italics="true" pageId="0" pageNumber="49">Kopua nuimata</emphasis>
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<bibRefCitation id="16274B14FFDAFF825CD3F84082B1DAD1" author="Hardy, G. S." box="[996,1183,1990,2014]" pageId="0" pageNumber="49" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy, 1984: 246</bibRefCitation>
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, figs 1–5 (original description; type locality: Rangatira Knoll, northwest of White Island,
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);
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<treatmentCitation id="F31710F4FFDAFF825BB2F781835ED510" author="Paulin, C. D. & Stewart, A. L. & Roberts, C. D. & McMillan, P. J." box="[1157,1392,2055,2079]" page="141" pageId="0" pageNumber="49" year="1989">
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<bibRefCitation id="16274B14FFDAFF825BB2F781835ED510" author="Paulin, C. D. & Stewart, A. L. & Roberts, C. D. & McMillan, P. J." box="[1157,1392,2055,2079]" pageId="0" pageNumber="49" pagination="1 - 279" refId="ref3837" refString="Paulin, C. D., Stewart, A. L., Roberts, C. D., and McMillan, P. J. 1989. New Zealand fish a complete guide. National Museum of New Zealand Miscellaneous Series 19: i-xiv + 1 - 279." type="book chapter" year="1989">
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Paulin
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<emphasis id="40C2EAF7FFDAFF825BF8F78E832AD510" box="[1231,1284,2055,2079]" italics="true" pageId="0" pageNumber="49">et al.</emphasis>
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1989: 141
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</bibRefCitation>
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</treatmentCitation>
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(in key; northern
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<collectingCountry id="0AA17675FFDBFF835E7DFA3887F9D8DA" box="[330,471,1469,1493]" name="New Zealand" pageId="1" pageNumber="50">New Zealand</collectingCountry>
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);
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<treatmentCitation id="F31710F4FFDBFF835EDEFA3B849AD8DA" author="Hutchins, J. B." box="[489,692,1469,1494]" page="464" pageId="1" pageNumber="50" year="1991">
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<bibRefCitation id="16274B14FFDBFF835EDEFA3B849AD8DA" author="Hutchins, J. B." box="[489,692,1469,1494]" pageId="1" pageNumber="50" pagination="463 - 468" refId="ref3692" refString="Hutchins, J. B. 1991. Description of a new deepwater clingfish (Gobiesocidae) from New South Wales. Records of the Western Australian Museum 15: 463 - 468." type="journal article" year="1991">Hutchins 1991: 464</bibRefCitation>
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</treatmentCitation>
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, fig. 1b [sketches of head sensory canal pores; same as type locality: based on NMNZ P. 13110 (
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<typeStatus id="AD0D8847FFDBFF835D2AFA798453DB18" box="[541,637,1535,1559]" pageId="1" pageNumber="50" type="paratype">paratype</typeStatus>
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)];
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<treatmentCitation id="F31710F4FFDBFF835DA0FA788783DB38" author="Shinohara, G. & Katayama, E." page="436" pageId="1" pageNumber="50" year="2015">
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<bibRefCitation id="16274B14FFDBFF835DA0FA788783DB38" author="Shinohara, G. & Katayama, E." pageId="1" pageNumber="50" pagination="431 - 438" refId="ref4097" refString="Shinohara, G. and Katayama, E. 2015. A new species of the clingfish genus Kopua (Gobiesociformes: Gobiesocidae) from Japan. Ichthyological Research doi: 10.1007 / s 10228 - 015 - 0456 - 9 (31 January 2015), 62: 431 - 438 (November 2015)." type="journal article" year="2015">Shinohara and Katayama 2015: 436</bibRefCitation>
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</treatmentCitation>
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(in key; northern
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<collectingCountry id="0AA17675FFDBFF835D5EF9A684DADB38" box="[617,756,1567,1591]" name="New Zealand" pageId="1" pageNumber="50">New Zealand</collectingCountry>
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);
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<treatmentCitation id="F31710F4FFDBFF835F82F9C68754DB58" author="Stewart, A. L." box="[181,378,1599,1624]" page="1551" pageId="1" pageNumber="50" year="2015">
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<bibRefCitation id="16274B14FFDBFF835F82F9C68754DB58" author="Stewart, A. L." box="[181,378,1599,1624]" pageId="1" pageNumber="50" pagination="1539 - 1555" refId="ref4188" refString="Stewart, A. L. 2015. 218 Family Gobiesocidae. Pp. 1539 - 1555. In: Roberts, C. D., Stewart, A. L., and Struthers, C. D. (Eds) The Fishes of New Zealand. Volume Four. Systematic Accounts. Te Papa Press, Wellington." type="book chapter" year="2015">Stewart 2015: 1551</bibRefCitation>
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</treatmentCitation>
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, fig. 218.8 (brief description; same as type locality: based on NMNZ P. 13110);
|
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<treatmentCitation id="F31710F4FFDBFF835D5CF9E6873BDB96" author="Fujiwara, K. & Okamoto, M. & Motomura, H." page="449" pageId="1" pageNumber="50" year="2018">
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<bibRefCitation id="16274B14FFDBFF835D5CF9E6873BDB96" author="Fujiwara, K. & Okamoto, M. & Motomura, H." pageId="1" pageNumber="50" pagination="433 - 453" refId="ref3591" refString="Fujiwara, K., Okamoto, M., and Motomura, H. 2018. Review of the clingfish genus Kopua (Gobiesocidae: Trachelochisminae) in Japan, with description of a new species. Ichthyological Research doi: 10.1007 / s 10228 - 0180633 - 8 (22 May 2018), 65: 433 - 453 (6 November 2018)." type="journal article" year="2018">
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Fujiwara
|
||
<emphasis id="40C2EAF7FFDBFF835DFBF9E7852EDB77" box="[716,768,1632,1656]" italics="true" pageId="1" pageNumber="50">et al.</emphasis>
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2018: 449
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</bibRefCitation>
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</treatmentCitation>
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(in key; northern
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<collectingCountry id="0AA17675FFDBFF835EF8F9078475DB96" box="[463,603,1665,1689]" name="New Zealand" pageId="1" pageNumber="50">New Zealand</collectingCountry>
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).
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</treatmentCitationGroup>
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</paragraph>
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<caption id="26C9666DFFDBFF835FABFADB84E7D880" ID-DOI="http://doi.org/10.5281/zenodo.3751646" ID-Zenodo-Dep="3751646" httpUri="https://zenodo.org/record/3751646/files/figure.png" pageId="1" pageNumber="50" startId="1.[156,187,1373,1395]" targetBox="[188,1398,206,1331]" targetPageId="1">
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<paragraph id="720936E5FFDBFF835FABFADB84E7D880" blockId="1.[156,1433,1372,1423]" pageId="1" pageNumber="50">
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Fig. 1. Preserved specimen of
|
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<taxonomicName id="B5B64D66FFDBFF835EFEFADB8479D87D" authorityName="Hardy" authorityYear="1984" box="[457,599,1373,1394]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="1" pageNumber="50" phylum="Chordata" rank="species" species="nuimata">
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<emphasis id="40C2EAF7FFDBFF835EFEFADB8479D87D" box="[457,599,1373,1394]" italics="true" pageId="1" pageNumber="50">Kopua nuimata</emphasis>
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</taxonomicName>
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from Norfolk Ridge, south of Norfolk Island, Australia (CSIRO H6007-17, 29.0 mm SL). A, lateral view; B, dorsal view; C, ventral view; D, radiograph.
|
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</paragraph>
|
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</caption>
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<subSubSection id="3AAC656EFFDBFF835F82F9448443DA14" pageId="1" pageNumber="50" type="materials_examined">
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<paragraph id="720936E5FFDBFF835F82F9448443DA14" blockId="1.[156,767,1730,2079]" pageId="1" pageNumber="50">
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<emphasis id="40C2EAF7FFDBFF835F82F94487ADDBD5" bold="true" box="[181,387,1730,1754]" pageId="1" pageNumber="50">Material examined.</emphasis>
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<materialsCitation id="C2DE3CB8FFDBFF835EBDF9448443DA14" ID-GBIF-Occurrence="2597620215" collectingDate="2013-05-14" collectionCode="CSIRO" country="Norfolk Island" elevation="322" latitude="-29.7" location="Norfolk Ridge" longLatPrecision="1225" longitude="168.01666" pageId="1" pageNumber="50" specimenCode="H6007-17" specimenCount="1" stateProvince="New South Wales">
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<collectionCode id="14A7AE20FFDBFF835EBDF94487FCDBD5" box="[394,466,1730,1754]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="1" pageNumber="50" type="Museum">CSIRO</collectionCode>
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<specimenCode id="22109E9EFFDBFF835EEEF944846EDBD5" box="[473,576,1730,1754]" collectionCode="CSIRO" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="1" pageNumber="50" type="Museum">H6007-17</specimenCode>
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, 29.0 mm SL,
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<location id="7769603EFFDBFF835DF9F9448728DBF5" LSID="urn:lsid:plazi:treatment:FA1F87F3FFDAFF865CE2F8C28716DA14:7769603EFFDBFF835DF9F9448728DBF5" country="Norfolk Island" latitude="-29.7" longLatPrecision="1225" longitude="168.01666" name="Norfolk Ridge" pageId="1" pageNumber="50" stateProvince="New South Wales">Norfolk Ridge</location>
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, south of
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<collectingCountry id="0AA17675FFDBFF835E5AF964842DDBF5" box="[365,515,1762,1786]" name="Norfolk Island" pageId="1" pageNumber="50">Norfolk Island</collectingCountry>
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,
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<collectingRegion id="B072F807FFDBFF835D39F96584EADBF5" box="[526,708,1762,1786]" country="Australia" name="New South Wales" pageId="1" pageNumber="50">New South Wales</collectingRegion>
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,
|
||
<collectingCountry id="0AA17675FFDBFF835DF8F96486FDDA14" name="Australia" pageId="1" pageNumber="50">Australia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="17825022FFDBFF835FEBF8858702DA14" box="[220,300,1791,1819]" degrees="29" direction="south" minutes="42" orientation="latitude" pageId="1" pageNumber="50" precision="925" value="-29.7">29°42′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="17825022FFDBFF835E01F88587BADA15" box="[310,404,1791,1819]" degrees="168" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="50" precision="925" value="168.01666">168°01′E</geoCoordinate>
|
||
,
|
||
<quantity id="B54E9B00FFDBFF835EA9F88587F5DA15" box="[414,475,1795,1819]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="1" pageNumber="50" unit="m" value="322.0">
|
||
<elevation id="F99BD1D6FFDBFF835EA9F88587F5DA15" box="[414,475,1795,1819]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="1" pageNumber="50" unit="m" value="322.0">322 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="06081025FFDBFF835ED2F8858447DA14" box="[485,617,1795,1819]" pageId="1" pageNumber="50" value="2013-05-14">
|
||
<collectingDate id="164CE9CDFFDBFF835ED2F8858447DA14" box="[485,617,1795,1819]" pageId="1" pageNumber="50" value="2013-05-14">14 May 2013</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AAC656EFFDBFF835F82F8A5835ADA14" pageId="1" pageNumber="50" type="diagnosis">
|
||
<paragraph id="720936E5FFDBFF835F82F8A5835ADA14" blockId="1.[156,767,1730,2079]" lastBlockId="1.[820,1432,1469,2079]" pageId="1" pageNumber="50">
|
||
<emphasis id="40C2EAF7FFDBFF835F82F8A5870BDA34" bold="true" box="[181,293,1827,1851]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="B5B64D66FFDBFF835E83F8A287DBDA34" authorityName="Hardy" authorityYear="1984" box="[436,501,1828,1851]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="1" pageNumber="50" phylum="Chordata" rank="genus">
|
||
<emphasis id="40C2EAF7FFDBFF835E83F8A287DBDA34" box="[436,501,1828,1851]" italics="true" pageId="1" pageNumber="50">Kopua</emphasis>
|
||
</taxonomicName>
|
||
distinguished from other members of the genus by the following combination of characters: 10 or 11 dorsal-fin rays; 8–10 anal-fin rays; 23–25 pectoral-fin rays; 7 or 8 gill rakers on 2nd arch; 34–36 vertebrae; 2 pores in nasal, lacrimal, and preopercular canals; snout short, its length 7.5–10.4% SL; gill opening wide, its depth 7.5% SL, upper end of gill membrane level with 2nd to 4th pectoral-fin ray base in lateral view; anterior, poste- rior and least interorbital widths 7.9, 15.0, and 1.2% SL, respectively; disc large, its length 19.6–22.7% SL, disc region D with
|
||
<emphasis id="40C2EAF7FFDBFF835C8AFA7985F4DB19" box="[957,986,1535,1558]" italics="true" pageId="1" pageNumber="50">ca.</emphasis>
|
||
6 rows of papillae; caudal-peduncle depth 8.2–11.2% SL; anus slightly closer to posterior margin of disc than to anal-fin origin, disc and disc C to anus lengths 14.3 and 27.4% SL, respectively; dorsal and anal-fin origins located posteriorly on body, pre-dorsal- and anal-fin lengths 66.7–71.4 and 71.4–74.1% SL, respectively, disc and disc C to anal-fin origin 31.9 and 43.8% SL, respectively; post-dorsal-caudal length 14.4% SL; arch-shaped blotches on lateral aspect of body; and two reddish-orange stripes on cheek.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AAC656EFFDBFF815C79F8A58223DE05" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50" type="description">
|
||
<paragraph id="720936E5FFDBFF805C79F8A587CFDB57" blockId="1.[820,1432,1469,2079]" lastBlockId="2.[156,768,1307,2079]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
|
||
<emphasis id="40C2EAF7FFDBFF835C79F8A58573DA53" bold="true" pageId="1" pageNumber="50">Description of Australian specimen (CSIRO H6007- 17).</emphasis>
|
||
Dorsal-fin rays 10; anal-fin rays 10; pectoral-fin rays 23; caudal-fin rays 6–5+5–6; gill rakers 7 on each arch; vertebrae 34. The following morphometrics are expressed as percentage of SL: head length 34.2; postorbital length 16.4; head depth 10.8; head width 19.8; body depth 12.1; body width 17.2; gill-opening depth 7.5; snout length 7.5; snout depth 6.9; upper-jaw length 11.9; orbit diameter 10.9; anterior in- terorbital width 7.9; posterior interorbital width 15.0; least interorbital width 1.2; disc length 22.4; caudal-peduncle length 10.1; caudal-peduncle depth 8.2; pre-disc length 20.7; pre-anus length 57.8; disc to anal-fin origin length 31.9; disc to anus length 14.3; disc C to anal-fin origin length 43.8; disc C to anus length 27.4; pre-dorsal-fin length 68.5; preanal-fin length 74.1; dorsal-caudal length 32.6; post-dorsalcaudal length 14.4; anal-caudal length 25.8; dorsal-fin base length 18.8; anal-fin base length 17.4; pectoral-fin length 16.8; and caudal-fin length 20.3.
|
||
</paragraph>
|
||
<caption id="26C9666DFFD8FF805FABFB3D8430D9E3" ID-DOI="http://doi.org/10.5281/zenodo.3751648" ID-Zenodo-Dep="3751648" httpUri="https://zenodo.org/record/3751648/files/figure.png" pageId="2" pageNumber="51" startId="2.[156,187,1211,1233]" targetBox="[189,1368,210,1135]" targetPageId="2">
|
||
<paragraph id="720936E5FFD8FF805FABFB3D8430D9E3" blockId="2.[156,1431,1211,1261]" pageId="2" pageNumber="51">
|
||
Fig. 2.
|
||
<taxonomicName id="B5B64D66FFD8FF805FC7FB3A8753D9DE" authorityName="Hardy" authorityYear="1984" box="[240,381,1212,1233]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFD8FF805FC7FB3A8753D9DE" box="[240,381,1212,1233]" italics="true" pageId="2" pageNumber="51">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
(CSIRO H6007-17, 29.0mm SL), Norfolk Ridge, south of Norfolk Island, Australia. Photographed subsequent to thawing. Photographs provided by CSIRO.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="720936E5FFD8FF805F82F9E68285D85C" blockId="2.[156,768,1307,2079]" lastBlockId="2.[820,1433,1307,2079]" pageId="2" pageNumber="51">
|
||
Body slender, cylindrical, compressed at caudal peduncle (
|
||
<figureCitation id="EA8D2A60FFD8FF805F94F90786CADB96" box="[163,228,1665,1689]" captionStart="Fig" captionStartId="1.[156,187,1373,1395]" captionTargetBox="[188,1398,206,1331]" captionTargetId="figure@1.[187,1399,205,1332]" captionTargetPageId="1" captionText="Fig. 1. Preserved specimen of Kopua nuimata from Norfolk Ridge, south of Norfolk Island, Australia (CSIRO H6007-17, 29.0mm SL). A, lateral view; B, dorsal view; C, ventral view; D, radiograph." figureDoi="http://doi.org/10.5281/zenodo.3751646" httpUri="https://zenodo.org/record/3751646/files/figure.png" pageId="2" pageNumber="51">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="EA8D2A60FFD8FF805FC2F907872ADB96" box="[245,260,1665,1689]" captionStart="Fig" captionStartId="2.[156,187,1211,1233]" captionTargetBox="[189,1368,210,1135]" captionTargetId="figure@2.[187,1400,205,1170]" captionTargetPageId="2" captionText="Fig. 2. Kopua nuimata (CSIRO H6007-17, 29.0mm SL), Norfolk Ridge, south of Norfolk Island, Australia. Photographed subsequent to thawing. Photographs provided by CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3751648" httpUri="https://zenodo.org/record/3751648/files/figure.png" pageId="2" pageNumber="51">2</figureCitation>
|
||
). Head large, depressed anteriorly. Snout somewhat rounded in lateral view, triangular in dorsal view; dorsal profile of snout slightly concave. Anterior and posterior nostrils with long membranous tube. Posterior nostril located in front of anterodorsal margin of orbit; anterior nostril located between posterior nostril and nasal canal pore (NC1). Eye very large. Interorbital region very narrow, flattened, its width 10% of orbit diameter. Mouth terminal. Upper jaw slightly longer than lower jaw, posterior margin of former reaching to anterior margin of eye lens. Upper lip thickened. Single row of thin, broad incisiviform teeth in both jaws; tips of upper-jaw teeth slightly pointed; posterior part of upper symphysis with much smaller, villiform tooth patch; lower-jaw teeth somewhat larger than upper-jaw teeth, tips slightly curved posteriorly.
|
||
</paragraph>
|
||
<paragraph id="720936E5FFD8FF805C79FADA823EDA73" blockId="2.[820,1433,1307,2079]" pageId="2" pageNumber="51">
|
||
First gill arch with one row of gill filaments, 2nd to 4th arches with 2 rows of gill filaments. Gill rakers very short, somewhat pointed. Each side of gill membrane united on underside of head, free from isthmus. Upper attachment of gill membrane level with 2nd pectoral-fin ray in lateral view. All soft-fin rays unbranched. Dorsal and anal fins well separated from caudal fin. Origin of anal fin located just below 3rd dorsal-fin ray. First dorsal- and anal-fin rays very short. Last dorsal- and anal-fin rays connected to caudal peduncle by membrane. Pectoral- and caudal-fin margins rounded. Uppermost pectoral-fin ray minute. Pelvic fins and pectoralgirdle elements forming circular “double”
|
||
<typeStatus id="AD0D8847FFD8FF805BD7F9448322DBD5" box="[1248,1292,1730,1754]" pageId="2" pageNumber="51">type</typeStatus>
|
||
disc. Disc region B squarish, larger than disc region A. Posterior margin of disc with long fringe. Lower 10th pectoral-fin ray base attached to 4th pelvic-fin ray by membrane. Disc with small papillae (in very poor condition, only disc region C with 1 or 2 rows of papillae).
|
||
</paragraph>
|
||
<paragraph id="720936E5FFD8FF815C79F8038787DCAB" blockId="2.[820,1433,1307,2079]" lastBlockId="3.[156,767,201,713]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
|
||
Head sensory canal pores well-developed, comprising 2 nasal, 2 lacrimal and 2 preopercular pores, and 1 postorbital pore (
|
||
<figureCitation id="EA8D2A60FFD8FF805C43F840859DDAD1" box="[884,947,1990,2014]" captionStart="Fig" captionStartId="3.[156,187,1946,1968]" captionTargetBox="[160,764,772,1904]" captionTargetId="figure@3.[158,766,770,1906]" captionTargetPageId="3" captionText="Fig. 3. Illustration of head sensory canal pores of Kopua nuimata, based on CSIRO H6007-17, 29.0mm SL. AN and PN, anterior and posterior nostrils, respectively; LC1–2, lacrimal canal pores; NC1–2, nasal canal pores; PO1, postorbital canal pore; PR2–3, preopercular canal pores. Bar indicates 1mm." figureDoi="http://doi.org/10.5281/zenodo.3751650" httpUri="https://zenodo.org/record/3751650/files/figure.png" pageId="2" pageNumber="51">Fig. 3</figureCitation>
|
||
); mandibular canal pores absent. Nasal canal pore smaller than nostrils, NC1 and NC2 located slightly before anterior margin of anterior and posterior nostrils, re- spectively, in dorsal view; lacrimal canal pore same size as nasal canal pore, LC1 located in front of anterior margin of eye, LC2 located posteroventrally below LC1; 1 postocular canal pore (largest of all pores) located just behind posterior margin of orbit; PR2 and PR3 located on ventral surface and lateral to operculum, respectively; all pores with distinct membranous tube (
|
||
<figureCitation id="EA8D2A60FFD9FF815E56FE0A87B2DCAB" box="[353,412,396,420]" captionStart="Fig" captionStartId="3.[156,187,1946,1968]" captionTargetBox="[160,764,772,1904]" captionTargetId="figure@3.[158,766,770,1906]" captionTargetPageId="3" captionText="Fig. 3. Illustration of head sensory canal pores of Kopua nuimata, based on CSIRO H6007-17, 29.0mm SL. AN and PN, anterior and posterior nostrils, respectively; LC1–2, lacrimal canal pores; NC1–2, nasal canal pores; PO1, postorbital canal pore; PR2–3, preopercular canal pores. Bar indicates 1mm." figureDoi="http://doi.org/10.5281/zenodo.3751650" httpUri="https://zenodo.org/record/3751650/files/figure.png" pageId="3" pageNumber="52">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="720936E5FFD9FF815F82FE2A8223DE05" blockId="3.[156,767,201,713]" lastBlockId="3.[821,1432,201,1298]" pageId="3" pageNumber="52">
|
||
<emphasis id="40C2EAF7FFD9FF815F82FE2A871EDCCB" bold="true" box="[181,304,428,452]" pageId="3" pageNumber="52">Coloration.</emphasis>
|
||
Based on
|
||
<figureCitation id="EA8D2A60FFD9FF815EA8FE2A87F2DCCB" box="[415,476,428,452]" captionStart="Fig" captionStartId="2.[156,187,1211,1233]" captionTargetBox="[189,1368,210,1135]" captionTargetId="figure@2.[187,1400,205,1170]" captionTargetPageId="2" captionText="Fig. 2. Kopua nuimata (CSIRO H6007-17, 29.0mm SL), Norfolk Ridge, south of Norfolk Island, Australia. Photographed subsequent to thawing. Photographs provided by CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3751648" httpUri="https://zenodo.org/record/3751648/files/figure.png" pageId="3" pageNumber="52">Fig. 2</figureCitation>
|
||
. Body ground color whitish to pale pink.
|
||
<emphasis id="40C2EAF7FFD9FF815E13FE488714DCEA" box="[292,314,462,485]" italics="true" pageId="3" pageNumber="52">ca</emphasis>
|
||
. 9 reddish-orange irregular bands on lateral aspect of body (upper part of some bands disrupted), each band interconnected at mid-line of body and forming archshaped blotches (4th and 9th bands distinct and isolated); 1st to 3rd band located above pelvic disc, merging with ground color dorsally; 4th and 5th band located around anus, somewhat broad, connected with antimere dorsally; 6–8th band (7th and 8th rudimentary) located below dor- sal-fin base; posteriormost band located just anterior to caudal-fin base, slightly broadening ventrally. 4th and 9th band extended further ventrally than other bands; 9th band connected with antimere. Dorsal surface of anterior body with irregular faint yellowish bands and dots. Head ground color similar to body color. Snout with somewhat broad yellowish-orange stripe laterally, faint yellow “V” shaped tinge dorsally. Mouth faint yellowish-orange. Eye red, transitioning to brownish-green dorsally, a pale ring encircling black pupil. Two reddish-orange stripes from eye to cheek; upper stripe narrow, horizontally level with upper end of pupil, its tip close to 1st band; lower stripe directed diagonally downward, broad but narrowing to tip, extending to underside of head. Dorsal, anal and caudal fins reddish to translucent whitish. Center of caudal fin with faint red tinge, fin base with broad white band. Pectoral fin translucent white. Disc whitish to faint reddish. Preserved specimen uniformly yellowish-white (
|
||
<figureCitation id="EA8D2A60FFD9FF815CF3FD7485D3DE05" box="[964,1021,754,778]" captionStart="Fig" captionStartId="1.[156,187,1373,1395]" captionTargetBox="[188,1398,206,1331]" captionTargetId="figure@1.[187,1399,205,1332]" captionTargetPageId="1" captionText="Fig. 1. Preserved specimen of Kopua nuimata from Norfolk Ridge, south of Norfolk Island, Australia (CSIRO H6007-17, 29.0mm SL). A, lateral view; B, dorsal view; C, ventral view; D, radiograph." figureDoi="http://doi.org/10.5281/zenodo.3751646" httpUri="https://zenodo.org/record/3751646/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AAC656EFFD9FF815C79FC948364DE84" pageId="3" pageNumber="52" type="distribution">
|
||
<paragraph id="720936E5FFD9FF815C79FC948364DE84" blockId="3.[821,1432,201,1298]" pageId="3" pageNumber="52">
|
||
<emphasis id="40C2EAF7FFD9FF815C79FC9485F7DE25" bold="true" box="[846,985,786,810]" pageId="3" pageNumber="52">Distribution.</emphasis>
|
||
Currently recorded only from
|
||
<collectingCountry id="0AA17675FFD9FF815A7FFC94855BDE44" name="Norfolk Island" pageId="3" pageNumber="52">Norfolk Island</collectingCountry>
|
||
,
|
||
<collectingCountry id="0AA17675FFD9FF815CB6FCB585F3DE44" box="[897,989,819,843]" name="Australia" pageId="3" pageNumber="52">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="EA8D2A60FFD9FF815CDAFCB58202DE44" box="[1005,1068,819,843]" captionStart="Fig" captionStartId="3.[821,852,2002,2024]" captionTargetBox="[824,1428,1355,1959]" captionTargetId="figure@3.[822,1430,1353,1961]" captionTargetPageId="3" captionText="Fig. 4. Distributional records of Kopua nuimata. Star and circles indicate new and previous records, respectively. Arrowhead indicates type locality." figureDoi="http://doi.org/10.5281/zenodo.3751652" httpUri="https://zenodo.org/record/3751652/files/figure.png" pageId="3" pageNumber="52">Fig. 4</figureCitation>
|
||
), and the Three Kings Islands and White Island,
|
||
<collectingCountry id="0AA17675FFD9FF815CFDFCD2827BDE64" box="[970,1109,851,875]" name="New Zealand" pageId="3" pageNumber="52">New Zealand</collectingCountry>
|
||
(
|
||
<bibRefCitation id="16274B14FFD9FF815B51FCD582C8DE64" author="Hardy, G. S." box="[1126,1254,851,875]" pageId="3" pageNumber="52" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy 1984</bibRefCitation>
|
||
). These localities are on the Zealandia Continent (
|
||
<bibRefCitation id="16274B14FFD9FF815B4AFCF28311DE84" author="Roberts, C. D. & Stewart A. L. & Struthers, C. D." box="[1149,1343,883,908]" pageId="3" pageNumber="52" refId="ref3889" refString="Roberts, C. D., Stewart A. L., and Struthers, C. D. 2015. The Fishes of New Zealand. 4 Volumes. Te Papa Press, Wellington, 2008 pp." type="book" year="2015">
|
||
Roberts
|
||
<emphasis id="40C2EAF7FFD9FF815BE3FCF2832BDE83" box="[1236,1285,884,908]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AAC656EFFD9FF865C79FC128716DA14" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="discussion">
|
||
<paragraph id="720936E5FFD9FF865C79FC128769DF68" blockId="3.[821,1432,201,1298]" lastBlockId="4.[156,768,201,1819]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
|
||
<emphasis id="40C2EAF7FFD9FF815C79FC12859CDEA3" bold="true" box="[846,946,916,940]" pageId="3" pageNumber="52">Remarks.</emphasis>
|
||
The present specimen agrees closely with the original description of
|
||
<taxonomicName id="B5B64D66FFD9FF815B1BFC3082E3DEC3" authorityName="Hardy" authorityYear="1984" box="[1068,1229,949,973]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFD9FF815B1BFC3082E3DEC3" box="[1068,1229,949,973]" italics="true" pageId="3" pageNumber="52">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
in having the following combination of characters: 10 dorsal-fin rays; 23 pectoral-fin rays; 6–5+5–6 caudal-fin rays; 7 gill rakers on each arch; very large eyes and narrow interorbital width, orbit diameter 10.9% SL and
|
||
<quantity id="B54E9B00FFD9FF815B52FBB1828ED940" box="[1125,1184,1079,1103]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="52" unit="in" value="0.1">0.1 in</quantity>
|
||
least interorbital width; “double”
|
||
<typeStatus id="AD0D8847FFD9FF815CA4FBDE8591D97F" box="[915,959,1112,1136]" pageId="3" pageNumber="52">type</typeStatus>
|
||
disc, with disc region B squarish; 4 gill arches with filaments; and gill membrane on each side united on underside of head and free from isthmus (
|
||
<bibRefCitation id="16274B14FFD9FF815A38FB1E83A3D9BE" author="Hardy, G. S." box="[1295,1421,1176,1201]" pageId="3" pageNumber="52" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy 1984</bibRefCitation>
|
||
). In addition, the head sensory canal pore characters of the present specimen matched those of
|
||
<taxonomicName id="B5B64D66FFD9FF815B8CFB5C831FD9FE" authorityName="Hardy" authorityYear="1984" box="[1211,1329,1242,1265]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFD9FF815B8CFB5C831FD9FE" box="[1211,1329,1242,1265]" italics="true" pageId="3" pageNumber="52">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
provided by
|
||
<bibRefCitation id="16274B14FFD9FF815C63FB7C85D3D81D" author="Hutchins, J. B." box="[852,1021,1274,1298]" pageId="3" pageNumber="52" pagination="463 - 468" refId="ref3692" refString="Hutchins, J. B. 1991. Description of a new deepwater clingfish (Gobiesocidae) from New South Wales. Records of the Western Australian Museum 15: 463 - 468." type="journal article" year="1991">Hutchins (1991)</bibRefCitation>
|
||
. However, the location of the gill mem- brane and counts of anal-fin rays and vertebrae of the present specimen differed slightly from those given by
|
||
<bibRefCitation id="16274B14FFDEFF865D88FF6F86F0DC2D" author="Hardy, G. S." pageId="4" pageNumber="53" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy (1984)</bibRefCitation>
|
||
(
|
||
<emphasis id="40C2EAF7FFDEFF865FD8FE8C873CDC2E" box="[239,274,266,289]" italics="true" pageId="4" pageNumber="53">viz.</emphasis>
|
||
, upper attachment of gill membrane same level as 2nd pectoral-fin ray, 10 anal-fin rays, and 34 vertebrae in the former
|
||
<emphasis id="40C2EAF7FFDEFF865E05FECA8762DC6C" box="[306,332,332,355]" italics="true" pageId="4" pageNumber="53">vs.</emphasis>
|
||
4th ray, 8 or 9, and 35 or 36, respectively, in the latter). These minor differences were regarded as intraspecific variations in this study because similar variations exist in other species of
|
||
<taxonomicName id="B5B64D66FFDEFF865EFCFE2B8422DCCB" authorityName="Hardy" authorityYear="1984" box="[459,524,429,452]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="40C2EAF7FFDEFF865EFCFE2B8422DCCB" box="[459,524,429,452]" italics="true" pageId="4" pageNumber="53">Kopua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="40C2EAF7FFDEFF865D2BFE2B8413DCCB" box="[540,573,429,452]" italics="true" pageId="4" pageNumber="53">e.g.</emphasis>
|
||
, see
|
||
<bibRefCitation id="16274B14FFDEFF865D58FE2A86FFDCEA" author="Fujiwara, K. & Okamoto, M. & Motomura, H." pageId="4" pageNumber="53" pagination="433 - 453" refId="ref3591" refString="Fujiwara, K., Okamoto, M., and Motomura, H. 2018. Review of the clingfish genus Kopua (Gobiesocidae: Trachelochisminae) in Japan, with description of a new species. Ichthyological Research doi: 10.1007 / s 10228 - 0180633 - 8 (22 May 2018), 65: 433 - 453 (6 November 2018)." type="journal article" year="2018">
|
||
Fujiwara
|
||
<emphasis id="40C2EAF7FFDEFF865DF9FE2B852EDCCB" box="[718,768,428,452]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Although
|
||
<bibRefCitation id="16274B14FFDEFF865E78FE4B87F4DCEA" author="Hardy, G. S." box="[335,474,461,485]" pageId="4" pageNumber="53" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy (1984)</bibRefCitation>
|
||
provided a detailed pattern of disc papillae (
|
||
<bibRefCitation id="16274B14FFDEFF865E75FE6B8792DF09" author="Hardy, G. S." box="[322,444,493,518]" pageId="4" pageNumber="53" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy 1984</bibRefCitation>
|
||
: fig. 3), this character could not be examined completely here (only papillae on disc region C confirmed) because of the poor condition of the
|
||
<collectingCountry id="0AA17675FFDEFF865D98FDA886F4DF68" name="Norfolk Island" pageId="4" pageNumber="53">Norfolk Island</collectingCountry>
|
||
specimen.
|
||
</paragraph>
|
||
<caption id="26C9666DFFD9FF815FABF81C8432D510" ID-DOI="http://doi.org/10.5281/zenodo.3751650" ID-Zenodo-Dep="3751650" httpUri="https://zenodo.org/record/3751650/files/figure.png" pageId="3" pageNumber="52" startId="3.[156,187,1946,1968]" targetBox="[160,764,772,1904]" targetPageId="3">
|
||
<paragraph id="720936E5FFD9FF815FABF81C8432D510" blockId="3.[156,768,1946,2080]" pageId="3" pageNumber="52">
|
||
Fig. 3. Illustration of head sensory canal pores of
|
||
<taxonomicName id="B5B64D66FFD9FF815D4AF81D8680DAC3" authorityName="Hardy" authorityYear="1984" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFD9FF815D4AF81D8680DAC3" italics="true" pageId="3" pageNumber="52">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
, based on CSIRO H6007-17, 29.0 mm SL. AN and PN, anterior and posterior nostrils, respectively; LC1–2, lacrimal canal pores; NC1–2, nasal canal pores; PO1, postorbital canal pore; PR2–3, preopercular canal pores. Bar indicates 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26C9666DFFD9FF815C02F85485F7D52F" ID-DOI="http://doi.org/10.5281/zenodo.3751652" ID-Zenodo-Dep="3751652" httpUri="https://zenodo.org/record/3751652/files/figure.png" pageId="3" pageNumber="52" startId="3.[821,852,2002,2024]" targetBox="[824,1428,1355,1959]" targetPageId="3">
|
||
<paragraph id="720936E5FFD9FF815C02F85485F7D52F" blockId="3.[821,1431,2002,2080]" pageId="3" pageNumber="52">
|
||
Fig. 4. Distributional records of
|
||
<taxonomicName id="B5B64D66FFD9FF815B45F85582D1DAE7" authorityName="Hardy" authorityYear="1984" box="[1138,1279,2003,2024]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFD9FF815B45F85582D1DAE7" box="[1138,1279,2003,2024]" italics="true" pageId="3" pageNumber="52">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
. Star and circles indicate new and previous records, respectively. Arrowhead indicates type locality.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="720936E5FFDEFF865F82FDF68736DB58" blockId="4.[156,768,201,1819]" pageId="4" pageNumber="53">
|
||
To date, six nominal species of
|
||
<taxonomicName id="B5B64D66FFDEFF865D20FDF68476DF88" authorityName="Hardy" authorityYear="1984" box="[535,600,624,647]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="40C2EAF7FFDEFF865D20FDF68476DF88" box="[535,600,624,647]" italics="true" pageId="4" pageNumber="53">Kopua</emphasis>
|
||
</taxonomicName>
|
||
have been described, five being regarded as valid,
|
||
<emphasis id="40C2EAF7FFDEFF865D25FD17841BDFA7" box="[530,565,657,680]" italics="true" pageId="4" pageNumber="53">viz.</emphasis>
|
||
<taxonomicName id="B5B64D66FFDEFF865D0CFD178722DFC6" authority="(Doderlein, 1887)" baseAuthorityName="Doderlein" baseAuthorityYear="1887" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="40C2EAF7FFDEFF865D0CFD17848BDFA7" box="[571,677,657,680]" italics="true" pageId="4" pageNumber="53">K. minima</emphasis>
|
||
(Döderlein, 1887)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865E2CFD3787A1DFC7" authorityName="Hardy" authorityYear="1984" box="[283,399,689,712]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865E2CFD3787A1DFC7" box="[283,399,689,712]" italics="true" pageId="4" pageNumber="53">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865EA8FD378487DFC6" authority="Hutchins, 1991" authorityName="Hutchins" authorityYear="1991" box="[415,681,689,713]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="kuiteri">
|
||
<emphasis id="40C2EAF7FFDEFF865EA8FD37842FDFC6" box="[415,513,689,713]" italics="true" pageId="4" pageNumber="53">K. kuiteri</emphasis>
|
||
<bibRefCitation id="16274B14FFDEFF865D3CFD378487DFC6" author="Hutchins, J. B." box="[523,681,689,713]" pageId="4" pageNumber="53" pagination="463 - 468" refId="ref3692" refString="Hutchins, J. B. 1991. Description of a new deepwater clingfish (Gobiesocidae) from New South Wales. Records of the Western Australian Museum 15: 463 - 468." type="journal article" year="1991">Hutchins, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865D8FFD378411DFE6" authority="Shinohara and Katayama, 2015" authorityName="Shinohara and Katayama" authorityYear="2015" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="vermiculata">
|
||
<emphasis id="40C2EAF7FFDEFF865D8FFD3786DDDFE6" italics="true" pageId="4" pageNumber="53">K. vermiculata</emphasis>
|
||
<bibRefCitation id="16274B14FFDEFF865FCBFD578411DFE6" author="Shinohara, G. & Katayama, E." box="[252,575,721,746]" pageId="4" pageNumber="53" pagination="431 - 438" refId="ref4097" refString="Shinohara, G. and Katayama, E. 2015. A new species of the clingfish genus Kopua (Gobiesociformes: Gobiesocidae) from Japan. Ichthyological Research doi: 10.1007 / s 10228 - 015 - 0456 - 9 (31 January 2015), 62: 431 - 438 (November 2015)." type="journal article" year="2015">Shinohara and Katayama, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B5B64D66FFDEFF865D4EFD54840EDE05" authority="Fujiwara, Okamoto, and Motomura, 2018" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865D4EFD5484E8DFE6" box="[633,710,721,745]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
<bibRefCitation id="16274B14FFDEFF865DF8FD57840EDE05" author="Fujiwara, K. & Okamoto, M. & Motomura, H." pageId="4" pageNumber="53" pagination="433 - 453" refId="ref3591" refString="Fujiwara, K., Okamoto, M., and Motomura, H. 2018. Review of the clingfish genus Kopua (Gobiesocidae: Trachelochisminae) in Japan, with description of a new species. Ichthyological Research doi: 10.1007 / s 10228 - 0180633 - 8 (22 May 2018), 65: 433 - 453 (6 November 2018)." type="journal article" year="2018">Fujiwara, Okamoto, and Motomura, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="16274B14FFDEFF865D1AFD7484D5DE05" author="Fujiwara, K. & Okamoto, M. & Motomura, H." box="[557,763,754,778]" pageId="4" pageNumber="53" pagination="433 - 453" refId="ref3591" refString="Fujiwara, K., Okamoto, M., and Motomura, H. 2018. Review of the clingfish genus Kopua (Gobiesocidae: Trachelochisminae) in Japan, with description of a new species. Ichthyological Research doi: 10.1007 / s 10228 - 0180633 - 8 (22 May 2018), 65: 433 - 453 (6 November 2018)." type="journal article" year="2018">
|
||
Fujiwara
|
||
<emphasis id="40C2EAF7FFDEFF865DB8FD7484ECDE05" box="[655,706,754,778]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="16274B14FFDEFF865FABFC9487E0DE25" author="Fujiwara, K. & Motomura, H." box="[156,462,786,810]" pageId="4" pageNumber="53" pagination="50 - 67" refId="ref3516" refString="Fujiwara, K. and Motomura, H. 2019. Kopua minima (Doderlein 1887), a senior synonym of K. japonica Moore, Hutchins and Okamoto 2012, and description of a new species of Aspasma (Gobiesocidae). Ichthyological Research doi: 10.1007 / s 10228 - 019 - 00701 - 0 (25 June 2019), 67: 50 - 67 (17 January 2020)." type="journal article" year="2019">Fujiwara and Motomura 2019</bibRefCitation>
|
||
).
|
||
<taxonomicName id="B5B64D66FFDEFF865EE9FC958457DE25" authorityName="Hardy" authorityYear="1984" box="[478,633,787,810]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865EE9FC958457DE25" box="[478,633,787,810]" italics="true" pageId="4" pageNumber="53">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from other congeners by the following features: 10 or 11 dorsal-fin rays (
|
||
<emphasis id="40C2EAF7FFDEFF865ED0FCD2842CDE64" box="[487,514,852,875]" italics="true" pageId="4" pageNumber="53">vs.</emphasis>
|
||
<quantity id="B54E9B00FFDEFF865D3CFCD58460DE64" box="[523,590,851,875]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="4" pageNumber="53" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
|
||
the latter), 8–10 anal-fin rays (4–8), 23–25 pectoral-fin rays (21–23), 2 nasal canal pores (1; very rarely
|
||
<quantity id="B54E9B00FFDEFF865E82FC1287F1DEA3" box="[437,479,916,940]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="53" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="B5B64D66FFDEFF865ED0FC13841ADEA3" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" box="[487,564,916,940]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865ED0FC13841ADEA3" box="[487,564,916,940]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
[left side only in 1 of
|
||
<specimenCount id="64B0FD6CFFDEFF865F8EFC338764DEC2" box="[185,330,949,973]" pageId="4" pageNumber="53" type="generic">14 specimens</specimenCount>
|
||
]), and papillae on disc region D. In other meristic characters,
|
||
<taxonomicName id="B5B64D66FFDEFF865E5AFC5087CEDEE2" authorityName="Hardy" authorityYear="1984" box="[365,480,982,1005]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865E5AFC5087CEDEE2" box="[365,480,982,1005]" italics="true" pageId="4" pageNumber="53">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="B5B64D66FFDEFF865D5AFC5084F7DEE2" baseAuthorityName="Doderlein" baseAuthorityYear="1887" box="[621,729,982,1005]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="40C2EAF7FFDEFF865D5AFC5084F7DEE2" box="[621,729,982,1005]" italics="true" pageId="4" pageNumber="53">K. minima</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865DDFFC50873CD901" authorityName="Shinohara and Katayama" authorityYear="2015" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="vermiculata">
|
||
<emphasis id="40C2EAF7FFDEFF865DDFFC50873CD901" italics="true" pageId="4" pageNumber="53">K. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B5B64D66FFDEFF865E7AFC7187B4D901" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" box="[333,410,1014,1038]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865E7AFC7187B4D901" box="[333,410,1014,1038]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
: 7 or 8 gill rakers on the 2nd arch (
|
||
<emphasis id="40C2EAF7FFDEFF865F92FB918691D921" box="[165,191,1047,1070]" italics="true" pageId="4" pageNumber="53">vs.</emphasis>
|
||
<quantity id="B54E9B00FFDEFF865FF3FB9086C5D921" box="[196,235,1046,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="53" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="B5B64D66FFDEFF865FC6FB9187ADD921" authorityName="Shinohara and Katayama" authorityYear="2015" box="[241,387,1046,1070]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="vermiculata">
|
||
<emphasis id="40C2EAF7FFDEFF865FC6FB9187ADD921" box="[241,387,1046,1070]" italics="true" pageId="4" pageNumber="53">K. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="B54E9B00FFDEFF865E82FB9087DBD921" box="[437,501,1046,1070]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="4" pageNumber="53" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
|
||
<taxonomicName id="B5B64D66FFDEFF865ECCFB91846BD921" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" box="[507,581,1046,1070]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865ECCFB91846BD921" box="[507,581,1046,1070]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
) and 34–36 vertebrae (
|
||
<quantity id="B54E9B00FFDEFF865FE0FBB1871CD940" box="[215,306,1079,1103]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" metricValueMax="8.382" metricValueMin="7.874" pageId="4" pageNumber="53" unit="in" value="32.0" valueMax="33.0" valueMin="31.0">31–33 in</quantity>
|
||
<taxonomicName id="B5B64D66FFDEFF865E0EFBBE878AD940" baseAuthorityName="Doderlein" baseAuthorityYear="1887" box="[313,420,1080,1103]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="40C2EAF7FFDEFF865E0EFBBE878AD940" box="[313,420,1080,1103]" italics="true" pageId="4" pageNumber="53">K. minima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5B64D66FFDEFF865EEDFBBE8408D940" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" box="[474,550,1079,1103]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865EEDFBBE8408D940" box="[474,550,1079,1103]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
). Some morphometrics (see above) can also be used for identification purposes (
|
||
<bibRefCitation id="16274B14FFDEFF865F95FBFE8758D99F" author="Fujiwara, K. & Okamoto, M. & Motomura, H." box="[162,374,1144,1168]" pageId="4" pageNumber="53" pagination="433 - 453" refId="ref3591" refString="Fujiwara, K., Okamoto, M., and Motomura, H. 2018. Review of the clingfish genus Kopua (Gobiesocidae: Trachelochisminae) in Japan, with description of a new species. Ichthyological Research doi: 10.1007 / s 10228 - 0180633 - 8 (22 May 2018), 65: 433 - 453 (6 November 2018)." type="journal article" year="2018">
|
||
Fujiwara
|
||
<emphasis id="40C2EAF7FFDEFF865E32FBFF8717D99F" box="[261,313,1144,1168]" italics="true" pageId="4" pageNumber="53">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). The head sensory canal pore condition of
|
||
<taxonomicName id="B5B64D66FFDEFF865FDCFB1F8771D9BF" authorityName="Hardy" authorityYear="1984" box="[235,351,1177,1200]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865FDCFB1F8771D9BF" box="[235,351,1177,1200]" italics="true" pageId="4" pageNumber="53">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
is the most developed in the genus (
|
||
<taxonomicName id="B5B64D66FFDEFF865DDFFB1F86E4D9DE" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865DDFFB1F86E4D9DE" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
being most similar) and distinctly different from
|
||
<taxonomicName id="B5B64D66FFDEFF865DDFFB3C86C7D9FE" baseAuthorityName="Doderlein" baseAuthorityYear="1887" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="40C2EAF7FFDEFF865DDFFB3C86C7D9FE" italics="true" pageId="4" pageNumber="53">K. minima</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865FCDFB5C8772D9FD" authorityName="Hutchins" authorityYear="1991" box="[250,348,1242,1266]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="kuiteri">
|
||
<emphasis id="40C2EAF7FFDEFF865FCDFB5C8772D9FD" box="[250,348,1242,1266]" italics="true" pageId="4" pageNumber="53">K. kuiteri</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B5B64D66FFDEFF865EAAFB5C841AD9FD" authorityName="Shinohara and Katayama" authorityYear="2015" box="[413,564,1242,1266]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="vermiculata">
|
||
<emphasis id="40C2EAF7FFDEFF865EAAFB5C841AD9FD" box="[413,564,1242,1266]" italics="true" pageId="4" pageNumber="53">K. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
(but see above) in having 2 lacrimal and preopercular canal pores (
|
||
<emphasis id="40C2EAF7FFDEFF865DBFFB7D848DD81D" box="[648,675,1275,1298]" italics="true" pageId="4" pageNumber="53">vs.</emphasis>
|
||
no lacrimal or preopercular canal pores). The fresh coloration of
|
||
<taxonomicName id="B5B64D66FFDEFF865DDFFA9D86DED85C" authorityName="Hardy" authorityYear="1984" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865DDFFA9D86DED85C" italics="true" pageId="4" pageNumber="53">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
, detailed here for the first time, also provides very good field characters for species identification. Although the arch-shaped blotches on the lateral aspect of the body and two reddish-orange stripes on the cheek in
|
||
<taxonomicName id="B5B64D66FFDEFF865DBBFA1886E1D8DA" authorityName="Hardy" authorityYear="1984" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865DBBFA1886E1D8DA" italics="true" pageId="4" pageNumber="53">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
are shared with
|
||
<taxonomicName id="B5B64D66FFDEFF865E4CFA3887CBD8DA" baseAuthorityName="Doderlein" baseAuthorityYear="1887" box="[379,485,1470,1493]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="minima">
|
||
<emphasis id="40C2EAF7FFDEFF865E4CFA3887CBD8DA" box="[379,485,1470,1493]" italics="true" pageId="4" pageNumber="53">K. minima</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5B64D66FFDEFF865D2DFA388448D8DA" authorityName="Fujiwara, Okamoto, and Motomura" authorityYear="2018" box="[538,614,1469,1493]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="yoko">
|
||
<emphasis id="40C2EAF7FFDEFF865D2DFA388448D8DA" box="[538,614,1469,1493]" italics="true" pageId="4" pageNumber="53">K. yoko</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5B64D66FFDEFF865D44FA3884FFD8DA" authorityName="Hutchins" authorityYear="1991" box="[627,721,1469,1493]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="kuiteri">
|
||
<emphasis id="40C2EAF7FFDEFF865D44FA3884FFD8DA" box="[627,721,1469,1493]" italics="true" pageId="4" pageNumber="53">K. kuiteri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5B64D66FFDEFF865FABFA59871ED8F9" authorityName="Shinohara and Katayama" authorityYear="2015" box="[156,304,1502,1526]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="vermiculata">
|
||
<emphasis id="40C2EAF7FFDEFF865FABFA59871ED8F9" box="[156,304,1502,1526]" italics="true" pageId="4" pageNumber="53">K. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
differ significantly, both lacking arch-shaped blotches on the body, and having four or five reddish-orange stripes and a triangular reddish-orange blotch on the cheek, respectively.
|
||
</paragraph>
|
||
<paragraph id="720936E5FFDEFF865F82F9E78716DA14" blockId="4.[156,768,201,1819]" pageId="4" pageNumber="53">
|
||
<taxonomicName id="B5B64D66FFDEFF865F82F9E7877ADB77" authorityName="Hardy" authorityYear="1984" box="[181,340,1633,1656]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865F82F9E7877ADB77" box="[181,340,1633,1656]" italics="true" pageId="4" pageNumber="53">Kopua nuimata</emphasis>
|
||
</taxonomicName>
|
||
has previously been recorded only from northern
|
||
<collectingCountry id="0AA17675FFDEFF865E37F90787A7DB97" box="[256,393,1664,1688]" name="New Zealand" pageId="4" pageNumber="53">New Zealand</collectingCountry>
|
||
(Three Kings Islands and White Island;
|
||
<bibRefCitation id="16274B14FFDEFF865FEDF9278774DBB6" author="Hardy, G. S." box="[218,346,1697,1721]" pageId="4" pageNumber="53" pagination="244 - 247" refId="ref3658" refString="Hardy, G. S. 1984. A new genus and species of deepwater clingfish (family Gobiesocidae) from New Zealand. Bulletin of Marine Science 34: 244 - 247." type="journal article" year="1984">Hardy 1984</bibRefCitation>
|
||
). The present specimen, from south of
|
||
<collectingCountry id="0AA17675FFDEFF865FABF9478719DBD6" box="[156,311,1729,1753]" name="Norfolk Island" pageId="4" pageNumber="53">Norfolk Island</collectingCountry>
|
||
, represents the first record of
|
||
<taxonomicName id="B5B64D66FFDEFF865DBFF94484D0DBD6" authorityName="Hardy" authorityYear="1984" box="[648,766,1730,1753]" class="Actinopterygii" family="Gobiesocidae" genus="Kopua" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="nuimata">
|
||
<emphasis id="40C2EAF7FFDEFF865DBFF94484D0DBD6" box="[648,766,1730,1753]" italics="true" pageId="4" pageNumber="53">K. nuimata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="0AA17675FFDEFF865FEFF964871ADBF5" box="[216,308,1762,1786]" name="Australia" pageId="4" pageNumber="53">Australia</collectingCountry>
|
||
as well as the northernmost record for the species (
|
||
<figureCitation id="EA8D2A60FFDEFF865FC7F8848702DA14" box="[240,300,1794,1819]" captionStart="Fig" captionStartId="3.[821,852,2002,2024]" captionTargetBox="[824,1428,1355,1959]" captionTargetId="figure@3.[822,1430,1353,1961]" captionTargetPageId="3" captionText="Fig. 4. Distributional records of Kopua nuimata. Star and circles indicate new and previous records, respectively. Arrowhead indicates type locality." figureDoi="http://doi.org/10.5281/zenodo.3751652" httpUri="https://zenodo.org/record/3751652/files/figure.png" pageId="4" pageNumber="53">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |