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<document id="2DBD83D6080338B4EC0468BCED937E7E" ID-DOI="10.11646/zootaxa.5503.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745712" ID-ZooBank="7385E4AA-D7AD-4E79-A05C-684319E0843B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051594773" checkinUser="plazi" docAuthor="Hotèkpo, Sourou Joseph, Schön, Isa &amp; Martens, Koen" docDate="2024" docId="FA1F073A453FFFAFFF411BD9FA1E2AD6" docLanguage="en" docName="zootaxa.5503.1.1.pdf" docOrigin="Zootaxa 5503 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5503.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Benincandona martini Hotèkpo, Schön &amp; Martens, 2024, gen. et spec. nov." docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="06267F424535FFBCFFD61B4EFF8E2D35" masterDocTitle="An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="11" updateTime="1726052453190" updateUser="ExternalLinkService">
<mods:mods id="DC0F39AFDA7D3A8C7BEE279C4C542874" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48F8E230D5D7C9D3F6BDF36CCF6E1F4C">
<mods:title id="23E750C991F959EA557B4A8F167B67FE">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
</mods:titleInfo>
<mods:name id="4BE08F8468BA7A8BCA0ABD5D49523324" type="personal">
<mods:role id="69342259D1736C97B812337797E20EC6">
<mods:roleTerm id="76726A4B2F9F164A6B52EE4D55E59E7E">Author</mods:roleTerm>
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<mods:namePart id="EE79A571823AF7937D551FAF094C7D69">Hotèkpo, Sourou Joseph</mods:namePart>
<mods:nameIdentifier id="6FC4B670A5D84E9D5AD42C860A65307B" type="ORCID">0000-0001-9091-433X</mods:nameIdentifier>
<mods:affiliation id="C75BF32B6798D3D25F3DDD09651C7B31">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université d’Abomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="800AB7867B7253E18B2ECD0B82E4FBC4" type="email">jshotekpo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="44B85E94848DF59AF0D66841C512AF47" type="personal">
<mods:role id="9F08919AC28D28998F8F9F919FCDE15B">
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<mods:namePart id="37E03D996ABEC481F184F3506DAFB263">Schön, Isa</mods:namePart>
<mods:nameIdentifier id="965F1C3F65C6D13BE41309D2393579EC" type="ORCID">0000-0001-9269-6487</mods:nameIdentifier>
<mods:affiliation id="C3479E4693735C4FDF74B1AC50498DD4">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="25F722758D59958FB1C3E4D9704F2D85" type="email">ischoen@naturalsciences.be</mods:nameIdentifier>
</mods:name>
<mods:name id="992002C37E576333A4B3ABAA49D85F2C" type="personal">
<mods:role id="5C1716985A32EC9123B1095026499819">
<mods:roleTerm id="9A77A3B82AC11AF99C45773DF7AEA30F">Author</mods:roleTerm>
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<mods:namePart id="8784AC9633CC3F07CA3ACFBE34F14D59">Martens, Koen</mods:namePart>
<mods:nameIdentifier id="94E4E7FEE322C7F28F4279671D0CD6DF" type="ORCID">0000-0001-8680-973X</mods:nameIdentifier>
<mods:affiliation id="DE967356FE87619EEC2B7E29FC19BCA3">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="804E7DEF457ADD85AFD5FA30D5393841" type="email">kmartens@naturalsciences.be</mods:nameIdentifier>
</mods:name>
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<mods:title id="90A4CD5F39A3BBE04281EB59EADB508D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C4E8A86D67331E9AF8FCC51318D1464B">
<mods:date id="EA5490A02381E5F11E959C11D3CB0348">2024</mods:date>
<mods:detail id="06DEF2A7007FE5C34354C7AD58A96187" type="pubDate">
<mods:number id="A2D20BAAE571313FCABC08CC6D9259F4">2024-09-04</mods:number>
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<mods:number id="85CDEBB8058B57EC8C5C59FBAFA418F9">5503</mods:number>
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<mods:number id="9E1885591F7DCD6C058AB935C6B7693D">1</mods:number>
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<mods:url id="4C661997A1C882ED955AC6FAC145BC69">http://dx.doi.org/10.11646/zootaxa.5503.1.1</mods:url>
</mods:location>
<mods:classification id="1C65BBB7BFF012B808B279CEE0C80631">journal article</mods:classification>
<mods:identifier id="148BD1C7859CCE3388F8C99CFDA7475E" type="DOI">10.11646/zootaxa.5503.1.1</mods:identifier>
<mods:identifier id="A5AF98F29AD5C2E119E1CFBAA667C0E4" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="E182AC3E52A4E0C940AA4D2E37D57254" type="Zenodo-Dep">13745712</mods:identifier>
<mods:identifier id="BDBBA7DDFB04513051856C2825360492" type="ZooBank">7385E4AA-D7AD-4E79-A05C-684319E0843B</mods:identifier>
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<treatment id="FA1F073A453FFFAFFF411BD9FA1E2AD6" LSID="urn:lsid:plazi:treatment:FA1F073A453FFFAFFF411BD9FA1E2AD6" httpUri="http://treatment.plazi.org/id/FA1F073A453FFFAFFF411BD9FA1E2AD6" lastPageId="19" lastPageNumber="20" pageId="10" pageNumber="11">
<subSubSection id="3AACE5A7453FFFB6FF411BD9FC4E2DE3" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="7209B62C453FFFB6FF411BD9FC4E2DE3" blockId="10.[151,960,150,249]" pageId="10" pageNumber="11">
<heading id="29410140453FFFB6FF411BD9FC082D84" bold="true" box="[151,902,150,178]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="3">
<emphasis id="40C26A3E453FFFB6FF411BD9FC082D84" bold="true" box="[151,902,150,178]" pageId="10" pageNumber="11">
<taxonomicName id="B5B6CDAF453FFFB6FF411BD9FE242D84" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,426,151,178]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453FFFB6FF411BD9FE242D84" bold="true" box="[151,426,151,178]" italics="true" pageId="10" pageNumber="11">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453FFFB6FE671BD9FD0C2D84" box="[433,642,151,177]" pageId="10" pageNumber="11" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
</heading>
<uri id="0627BA2E453FFFB6FF411BF5FC4E2DE3" box="[151,960,187,214]" pageId="10" pageNumber="11">
urn:lsid:zoobank.org:act:
<uuid id="06108CF9453FFFB6FE7F1BF5FC4E2DE3" box="[425,960,187,214]" pageId="10" pageNumber="11">CBC82271-797B-4368-8958-DD94C3584F1D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411B91FE012DCC" box="[151,399,223,249]" pageId="10" pageNumber="11" type="description">
<paragraph id="7209B62C453FFFB6FF411B91FE012DCC" blockId="10.[151,960,150,249]" box="[151,399,223,249]" pageId="10" pageNumber="11">
(
<figureCitation id="EA8DAAA9453FFFB6FF491B91FEA92DCC" box="[159,295,223,249]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStartId-0="11.[151,250,1933,1957]" captionStartId-1="12.[151,250,1944,1968]" captionStartId-2="13.[151,250,1943,1967]" captionStartId-3="14.[151,250,1981,2005]" captionStartId-4="16.[151,250,1981,2005]" captionStartId-5="17.[151,250,1943,1967]" captionStartId-6="18.[151,250,1943,1967]" captionTargetBox-0="[172,1415,181,1910]" captionTargetBox-1="[157,1430,181,1920]" captionTargetBox-2="[170,1415,185,1915]" captionTargetBox-3="[169,1422,182,1942]" captionTargetBox-4="[186,1404,198,1941]" captionTargetBox-5="[196,1391,191,1914]" captionTargetBox-6="[172,1415,185,1915]" captionTargetId-0="figure-17@11.[172,1415,178,1910]" captionTargetId-1="figure-31@12.[157,1430,181,1920]" captionTargetId-2="figure-17@13.[167,1419,181,1919]" captionTargetId-3="figure-31@14.[166,1422,181,1957]" captionTargetId-4="figure-31@16.[166,1422,181,1957]" captionTargetId-5="figure-17@17.[186,1401,181,1919]" captionTargetId-6="figure-31@18.[168,1419,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." captionText-1="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." captionText-2="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200– INV.159203). Scale bar = 50 µm." captionText-3="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-4="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." captionText-5="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). B–D. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." captionText-6="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13745720" figureDoi-1="http://doi.org/10.5281/zenodo.13745722" figureDoi-2="http://doi.org/10.5281/zenodo.13745726" figureDoi-3="http://doi.org/10.5281/zenodo.13745728" figureDoi-4="http://doi.org/10.5281/zenodo.13745730" figureDoi-5="http://doi.org/10.5281/zenodo.13745732" figureDoi-6="http://doi.org/10.5281/zenodo.13745734" httpUri-0="https://zenodo.org/record/13745720/files/figure.png" httpUri-1="https://zenodo.org/record/13745722/files/figure.png" httpUri-2="https://zenodo.org/record/13745726/files/figure.png" httpUri-3="https://zenodo.org/record/13745728/files/figure.png" httpUri-4="https://zenodo.org/record/13745730/files/figure.png" httpUri-5="https://zenodo.org/record/13745732/files/figure.png" httpUri-6="https://zenodo.org/record/13745734/files/figure.png" pageId="10" pageNumber="11">Figures 3–9</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FEE21B91FE092DCC" box="[308,391,223,249]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). A–D. B. martini. E–F. B. ibikounlei. G–J. B. sakitii. K–N. B. cebios. O–R. B. trapezoides. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745806" httpUri="https://zenodo.org/record/13745806/files/figure.png" pageId="10" pageNumber="11">41A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A69FBD22C98" pageId="10" pageNumber="11" type="etymology">
<paragraph id="7209B62C453FFFB6FF411A69FE962C74" blockId="10.[151,1436,295,430]" box="[151,280,295,321]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A69FE962C74" bold="true" box="[151,280,295,321]" pageId="10" pageNumber="11">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411A05FBD22C98" blockId="10.[151,1436,295,430]" pageId="10" pageNumber="11">
The species is named after Dr Patrick Martin (RBINS,
<collectingRegion id="B07278CE453FFFB6FCDD1A05FCE32C50" box="[779,877,331,357]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="10" pageNumber="11">Brussels</collectingRegion>
,
<collectingCountry id="0AA1F6BC453FFFB6FCAF1A05FC552C50" box="[889,987,331,357]" name="Belgium" pageId="10" pageNumber="11">Belgium</collectingCountry>
) in acknowledgement of his significant contributions to groundwater biodiversity research in
<collectingCountry id="0AA1F6BC453FFFB6FD2A1A21FCCC2CBC" box="[764,834,367,393]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
, and with gratitude for his continuous support of the career of SJH and in recognition of decades of collegiality with IS and KM at RBINS.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF411A9AFE552FF2" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="7209B62C453FFFB6FF411A9AFE842CDB" blockId="10.[151,1437,468,711]" box="[151,266,468,494]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411A9AFE842CDB" bold="true" box="[151,266,468,494]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411AB6FE552FF2" blockId="10.[151,1437,468,711]" pageId="10" pageNumber="11">CpRl elongated (L/H = 2.0), sub-triangular, with greatest height in the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, surface with ridges and polygonal ornamentation at the mid-ventral side. CpD and CpV narrow, with greatest width in the middle, lateral sides evenly rounded, anterior and posterior extremities pointed. A2 with seta X long, reaching beyond tip of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with short seta d2. Hp with ls and ms with diverging positions, ls relatively short, reaching slightly beyond ms.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFB6FF4119A0FF742864" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="7209B62C453FFFB6FF4119A0FEB02E32" blockId="10.[151,318,749,776]" box="[151,318,749,776]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4119A0FEB02E32" bold="true" box="[151,318,749,776]" pageId="10" pageNumber="11">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411861FE8A2E7C" blockId="10.[151,1282,815,878]" box="[151,260,815,841]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411861FE8A2E7C" bold="true" box="[151,260,815,841]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF411861FE8A2E7C" box="[151,260,815,841]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF41181DFA8C2E5B" blockId="10.[151,1282,815,878]" box="[151,1282,851,878]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF41181DFA8C2E5B" box="[151,1282,851,878]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA5453FFFB6FF41181DFF4D2E5B" box="[151,195,851,878]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
, with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE9453FFFB6FBB0181DFB182E58" box="[1126,1174,851,877]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159200).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118DAFF722E9B" blockId="10.[151,780,916,979]" box="[151,252,916,942]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118DAFF722E9B" bold="true" box="[151,252,916,942]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118DAFF722E9B" box="[151,252,916,942]" pageId="10" pageNumber="11" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118F7FC822EE6" blockId="10.[151,780,916,979]" box="[151,780,952,979]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF4118F7FC822EE6" box="[151,780,952,979]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA5453FFFB6FF4118F7FF4F2EE6" box="[151,193,953,979]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
, dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FE2918F6FDEF2EE7" box="[511,609,952,978]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FDA618F6FD282EE6" box="[624,678,952,979]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159201).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C453FFFB6FF4118B4FE812921" blockId="10.[151,1436,1018,1189]" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF4118B4FE812921" bold="true" box="[151,271,1018,1044]" pageId="10" pageNumber="11">
<typeStatus id="AD0D088E453FFFB6FF4118B4FE812921" box="[151,271,1018,1044]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F50FDF92990" blockId="10.[151,1436,1018,1189]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411F50FCC1290C" box="[151,847,1054,1081]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FF411F50FF45290D" box="[151,203,1054,1080]" count="1" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
dissected and stored as the
<typeStatus id="AD0D088E453FFFB6FDEB1F50FD11290D" box="[573,671,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FD611F50FD62290C" box="[695,748,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159202)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FCB21F50FEA32968" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FCB21F50FC23290D" box="[868,941,1054,1080]" count="2" pageId="10" pageNumber="11" type="male">2 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E453FFFB6FB461F50FB7C290D" box="[1168,1266,1054,1080]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE9453FFFB6FADC1F50FAB0290C" box="[1290,1342,1054,1081]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159203, INV.159204)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FEE91F0CFBF72968" box="[319,1145,1090,1117]" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FEE91F0CFE0A2969" box="[319,388,1090,1116]" count="3" pageId="10" pageNumber="11" type="female">3 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE9453FFFB6FD701F0CFD542968" box="[678,730,1090,1117]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159205, INV.159206, INV.159207)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB5D1F0CFBBE29B4" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB5D1F0CFB5E2969" box="[1163,1232,1090,1116]" count="3" pageId="10" pageNumber="11" type="male">3 ♂♂</specimenCount>
carapaces stored dry in micropalaeontological slides (
<collectionCode id="14A72EE9453FFFB6FD811F28FD0529B4" box="[599,651,1126,1153]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159208, INV.159209, INV.159210)
</materialsCitation>
;
<materialsCitation id="C2DEBC71453FFFB6FB931F28FDFD2990" collectionCode="INV" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="64B07DA5453FFFB6FB931F28FBF629B5" box="[1093,1144,1126,1152]" count="1" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
carapace stored dry in micropalaeontological slide (
<collectionCode id="14A72EE9453FFFB6FE0A1FC4FD9F2990" box="[476,529,1162,1189]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="10" pageNumber="11" type="Herbarium">INV</collectionCode>
.159211)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C453FFFB6FF411F85FEBE29D0" blockId="10.[151,1437,1227,1361]" box="[151,304,1227,1253]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411F85FEBE29D0" bold="true" box="[151,304,1227,1253]" pageId="10" pageNumber="11">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411FA1FF742864" blockId="10.[151,1437,1227,1361]" pageId="10" pageNumber="11">
<materialsCitation id="C2DEBC71453FFFB6FF411FA1FB89283C" box="[151,1031,1263,1290]" country="Benin" location="Adjohoun District" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingCountry id="0AA1F6BC453FFFB6FF411FA1FF54283C" box="[151,218,1263,1289]" name="Benin" pageId="10" pageNumber="11">Benin</collectingCountry>
,
<collectingRegion id="B07278CE453FFFB6FF301FA1FE4C283F" box="[230,450,1263,1290]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé Department</collectingRegion>
,
<location id="7769E0F7453FFFB6FE1F1FA1FD1B283C" LSID="urn:lsid:plazi:treatment:FA1F073A453FFFAFFF411BD9FA1E2AD6:7769E0F7453FFFB6FE1F1FA1FD1B283C" box="[457,661,1263,1289]" country="Benin" name="Adjohoun District" pageId="10" pageNumber="11" stateProvince="Oueme">Adjohoun District</location>
, town of Azowlissé, located in the
</materialsCitation>
<materialsCitation id="C2DEBC71453FFFB6FBDA1FA1FF782864" collectingDate="2016-08-27" collectingDateMax="2016-08-29" collectingDateMin="2016-08-27" collectorName="Leg. &amp; S. J. Hotekpo" country="Benin" latitude="6.66605" location="Oueme" longLatPrecision="1" longitude="2.4920833" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Oueme">
<collectingRegion id="B07278CE453FFFB6FBDA1FA1FBD3283C" box="[1036,1117,1263,1289]" country="Benin" name="Oueme" pageId="10" pageNumber="11">Ouémé</collectingRegion>
catchment area, coordinates:
<geoCoordinate id="1782D0EB453FFFB6FF411E5DFEAA2818" box="[151,292,1299,1325]" degrees="6" direction="north" minutes="39.963" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="6.66605">6° 39.963’N</geoCoordinate>
,
<geoCoordinate id="1782D0EB453FFFB6FEF91E5DFE36281B" box="[303,440,1299,1326]" degrees="2" direction="east" minutes="29.525" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="2.4920833">2° 29.525’E</geoCoordinate>
,
<collectorName id="DF43D3FA453FFFB6FE151E5AFE74281B" box="[451,506,1300,1326]" pageId="10" pageNumber="11">Leg.</collectorName>
:
<collectorName id="DF43D3FA453FFFB6FDD21E5DFD182818" box="[516,662,1299,1325]" pageId="10" pageNumber="11">S.J. Hotèkpo</collectorName>
, collected on
<date id="060890EC453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">
<collectingDate id="164C6904453FFFB6FCF91E5DFBBE2818" box="[815,1072,1299,1325]" pageId="10" pageNumber="11" value="2016-08-27" valueMax="2016-08-29" valueMin="2016-08-27">27 and 29 August 2016</collectingDate>
</date>
(Locality code BVS
<quantity id="B54E1BC9453FFFB6FAC11E5DFAC02818" box="[1303,1358,1299,1325]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="10" pageNumber="11" unit="in" value="19.0">19 in</quantity>
<tableCitation id="3F348397453FFFB6FA851E5DFF2B2864" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="10" pageNumber="11">Tables 1</tableCitation>
and
<tableCitation id="3F348397453FFFB6FF0A1E79FF632864" box="[220,237,1335,1361]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="10" pageNumber="11">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A7453FFFAFFF411E36FD8A28A8" lastPageId="19" lastPageNumber="20" pageId="10" pageNumber="11" type="description">
<paragraph id="7209B62C453FFFB6FF411E36FE0F28A6" blockId="10.[151,1437,1400,2039]" box="[151,385,1400,1427]" pageId="10" pageNumber="11">
<emphasis id="40C26A3E453FFFB6FF411E36FE0F28A6" bold="true" box="[151,385,1400,1427]" pageId="10" pageNumber="11">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C453FFFB6FF411ED2FB592BBA" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
CpRl (
<figureCitation id="EA8DAAA9453FFFB6FF091ED2FEFC2882" box="[223,370,1436,1463]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figure 3A, C</figureCitation>
) elongated and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; external ornamentation consisting of linear ridges (
<figureCitation id="EA8DAAA9453FFFB6FE061D62FDE42B72" box="[464,618,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figure 3A–D</figureCitation>
) and polygonal structures (
<figureCitation id="EA8DAAA9453FFFB6FC741D62FB802B72" box="[930,1038,1580,1607]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figure 3I</figureCitation>
) in mid-ventral position of valves. CpD (
<figureCitation id="EA8DAAA9453FFFB6FF0C1D1EFEC52B5E" box="[218,331,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figure 3E</figureCitation>
) and CpV (
<figureCitation id="EA8DAAA9453FFFB6FE1C1D1EFDB42B5E" box="[458,570,1616,1643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figure 3F</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins evenly rounded; LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111DD6FB8C2A77" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
LVi (
<figureCitation id="EA8DAAA9453FFFB6FED71DD6FE0D2B86" box="[257,387,1688,1715]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figures 3G</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE421DD7FE552B86" box="[404,475,1689,1715]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="10" pageNumber="11">4A, C</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle, overlap present at that site, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; CMS (
<figureCitation id="EA8DAAA9453FFFB6FC261C4AFBED2A2A" box="[1008,1123,1796,1823]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="10" pageNumber="11">Figure 4C</figureCitation>
) consisting of six scars: one large dorsal, three anterior and two posterior ones; one short antero-ventral il.
</paragraph>
<paragraph id="7209B62C453FFFB6FF111C02FCDE2AC3" blockId="10.[151,1437,1400,2039]" pageId="10" pageNumber="11">
RVi (
<figureCitation id="EA8DAAA9453FFFB6FF291C02FE182A52" box="[255,406,1868,1895]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figures 3H, J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FE771C03FE6D2A52" box="[417,483,1869,1895]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="10" pageNumber="11">4B, D</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest with height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin but with a blunt angle in the middle, posterior part sloping almost straight to posterior margin, ventral margin almost straight. CMS (
<figureCitation id="EA8DAAA9453FFFB6FE441CF6FD862AE6" box="[402,520,1976,2003]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="10" pageNumber="11">Figures 3J</figureCitation>
,
<figureCitation id="EA8DAAA9453FFFB6FDC51CF7FDB92AE6" box="[531,567,1977,2003]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="10" pageNumber="11">4D</figureCitation>
) consisting of six scars: one large dorsal (1), three anterior (2–4) and two (5,6) posterior ones; il running submarginally along ventral margin.
</paragraph>
<caption id="26C9E6A4453EFFB7FF411CC3FC722ADB" ID-DOI="http://doi.org/10.5281/zenodo.13745720" ID-Zenodo-Dep="13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1933,1957]" targetBox="[172,1415,181,1910]" targetPageId="11" targetType="figure">
<paragraph id="7209B62C453EFFB7FF411CC3FC722ADB" blockId="11.[151,1437,1933,2030]" pageId="11" pageNumber="12">
<emphasis id="40C26A3E453EFFB7FF411CC3FE802A90" bold="true" box="[151,270,1933,1957]" pageId="11" pageNumber="12">FIGURE 3</emphasis>
.
<taxonomicName id="B5B6CDAF453EFFB7FECD1CC3FD8E2A90" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[283,512,1933,1957]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453EFFB7FECD1CC3FD8E2A90" box="[283,512,1933,1957]" italics="true" pageId="11" pageNumber="12">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453EFFB7FDD11CC0FD352A90" box="[519,699,1934,1958]" pageId="11" pageNumber="12" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (A, C, E–H, J ♂; B, D, I ♀).
<emphasis id="40C26A3E453EFFB7FBEF1CC3FB0C2A93" bold="true" box="[1081,1154,1933,1958]" pageId="11" pageNumber="12">A–D, I</emphasis>
. CpRl, (A, C. INV.159208; B, D, I. INV.159211).
<emphasis id="40C26A3E453EFFB7FEAB1CFCFE002AFF" bold="true" box="[381,398,1970,1994]" pageId="11" pageNumber="12">E</emphasis>
. INV.159209. CpD.
<emphasis id="40C26A3E453EFFB7FDB41CFCFDFF2AFF" bold="true" box="[610,625,1970,1994]" pageId="11" pageNumber="12">F</emphasis>
. INV.159210. CpV.
<emphasis id="40C26A3E453EFFB7FC951CFFFCD92AFC" bold="true" box="[835,855,1969,1993]" pageId="11" pageNumber="12">G</emphasis>
. INV.159200. LVi.
<emphasis id="40C26A3E453EFFB7FBF61CFCFBBA2AFF" bold="true" box="[1056,1076,1970,1994]" pageId="11" pageNumber="12">H</emphasis>
. INV.159200. RVi.
<emphasis id="40C26A3E453EFFB7FAD61CFCFA832AFF" bold="true" box="[1280,1293,1970,1994]" pageId="11" pageNumber="12">J</emphasis>
. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44539FFB0FF411CD6FC792AE0" ID-DOI="http://doi.org/10.5281/zenodo.13745722" ID-Zenodo-Dep="13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1944,1968]" targetBox="[157,1430,181,1920]" targetPageId="12" targetType="figure">
<paragraph id="7209B62C4539FFB0FF411CD6FC792AE0" blockId="12.[151,1436,1944,2005]" pageId="12" pageNumber="13">
<emphasis id="40C26A3E4539FFB0FF411CD6FE9D2A85" bold="true" box="[151,275,1944,1968]" pageId="12" pageNumber="13">FIGURE 4.</emphasis>
<taxonomicName id="B5B6CDAF4539FFB0FECC1CD6FE702A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[282,510,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4539FFB0FECC1CD6FE702A85" box="[282,510,1944,1968]" italics="true" pageId="12" pageNumber="13">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454539FFB0FDD31CD7FD392A85" box="[517,695,1945,1969]" pageId="12" pageNumber="13" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀)
<emphasis id="40C26A3E4539FFB0FB2F1CD6FAA72A85" bold="true" box="[1273,1321,1944,1968]" pageId="12" pageNumber="13">A, E</emphasis>
. LVi.
<emphasis id="40C26A3E4539FFB0FABE1CD6FA1B2A85" bold="true" box="[1384,1429,1944,1968]" pageId="12" pageNumber="13">B, F</emphasis>
. RVi,
<emphasis id="40C26A3E4539FFB0FF1D1CF2FEB32AE1" bold="true" box="[203,317,1980,2004]" pageId="12" pageNumber="13">C, D, G, H</emphasis>
. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44538FFB1FF411CD9FD0F2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745726" ID-Zenodo-Dep="13745726" httpUri="https://zenodo.org/record/13745726/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1943,1967]" targetBox="[170,1415,185,1915]" targetPageId="13" targetType="figure">
<paragraph id="7209B62C4538FFB1FF411CD9FD0F2AE1" blockId="13.[151,1436,1943,2004]" pageId="13" pageNumber="14">
<emphasis id="40C26A3E4538FFB1FF411CD9FE9A2A85" bold="true" box="[151,276,1943,1968]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
<taxonomicName id="B5B6CDAF4538FFB1FECA1CD6FD8F2A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[284,513,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4538FFB1FECA1CD6FD8F2A9A" box="[284,513,1944,1968]" italics="true" pageId="13" pageNumber="14">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454538FFB1FDDF1CD6FD312A9A" box="[521,703,1944,1968]" pageId="13" pageNumber="14" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4538FFB1FCEA1CD9FCC02A9A" bold="true" box="[828,846,1943,1967]" pageId="13" pageNumber="14">A</emphasis>
. A1 (INV.159203).
<emphasis id="40C26A3E4538FFB1FBCB1CD6FBC02A9A" bold="true" box="[1053,1102,1943,1968]" pageId="13" pageNumber="14">B, C</emphasis>
. MdCoxa, (INV.159202).
<emphasis id="40C26A3E4538FFB1FA881CD6FAFE2A85" bold="true" box="[1374,1392,1944,1968]" pageId="13" pageNumber="14">D</emphasis>
. A2 (INV.159200– INV.159203). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4453BFFB2FF411CF3FA8E2ACF" ID-DOI="http://doi.org/10.5281/zenodo.13745728" ID-Zenodo-Dep="13745728" httpUri="https://zenodo.org/record/13745728/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1981,2005]" targetBox="[169,1422,182,1942]" targetPageId="14" targetType="figure">
<paragraph id="7209B62C453BFFB2FF411CF3FA8E2ACF" blockId="14.[151,1436,1981,2042]" pageId="14" pageNumber="15">
<emphasis id="40C26A3E453BFFB2FF411CF3FE982AE0" bold="true" box="[151,278,1981,2005]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
<taxonomicName id="B5B6CDAF453BFFB2FEC91CF3FD8B2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[287,517,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E453BFFB2FEC91CF3FD8B2AE0" box="[287,517,1981,2005]" italics="true" pageId="14" pageNumber="15">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745453BFFB2FDD81CF0FD492AE0" box="[526,711,1982,2006]" pageId="14" pageNumber="15" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E453BFFB2FC9E1CF3FCD42AE0" bold="true" box="[840,858,1981,2005]" pageId="14" pageNumber="15">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159200).
<emphasis id="40C26A3E453BFFB2FA901CF0FAD92AE3" bold="true" box="[1350,1367,1982,2006]" pageId="14" pageNumber="15">B</emphasis>
. Mx1, respiratory plate (INV.159202).
<emphasis id="40C26A3E453BFFB2FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="14" pageNumber="15">C</emphasis>
. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C453AFFB3FF111BD9FCD62F2C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A1 (
<figureCitation id="EA8DAAA9453AFFB3FF2E1BD9FEE32D84" box="[248,365,150,177]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200– INV.159203). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745726" httpUri="https://zenodo.org/record/13745726/files/figure.png" pageId="15" pageNumber="16">Figure 5A</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment
<emphasis id="40C26A3E453AFFB3FAD01BD9FA942D85" box="[1286,1306,151,176]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
three times as long as basal width, with two sub-apical, ventral setae, one long and one shorter, the latter about half of the length of the longer seta; and two shorter setae on the dorsal side, situated some distance from each other (one sub-apical and one more proximal). First distal segment slightly longer than basal width, with one short dorso-apical seta. Second distal segment about twice as long as wide, set with one short dorso-apical seta. Third distal segment with about the same length/width ratio as the second segment, carrying one short apical seta on each side. Fourth distal segment about three times as long as wide, with one ventro-apical seta, and one long dorso-apical seta, reaching beyond tip of sixth segment with almost one third of total length. Fifth distal segment four times as long as wide, set with two long dorso-apical setae and one short ventro-apical seta. Sixth distal segment narrow,
<emphasis id="40C26A3E453AFFB3FB261AF9FA8D2CE5" box="[1264,1283,439,464]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
four times as long as wide, with four apical structures: one aesthetasc Ya of intermediate length, two long setae more than twice the length of Ya and one short seta, about half the length of Ya.
</paragraph>
<paragraph id="7209B62C453AFFB3FF11196DFCE5296C" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
A2 (
<figureCitation id="EA8DAAA9453AFFB3FF20196DFEE72F08" box="[246,361,546,573]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200– INV.159203). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745726" httpUri="https://zenodo.org/record/13745726/files/figure.png" pageId="15" pageNumber="16">Figure 5D</figureCitation>
) with basal segment (first protopodite) broad and rounded, mid-ventrally with one seta of medium length, subapically with two setae of unequal length, proximal one
<emphasis id="40C26A3E453AFFB3FCAC1909FC032F55" box="[890,909,583,608]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E453AFFB3FD8A1925FDE12FB1" box="[604,623,619,644]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as basal width, set with one long ventro-subapical seta X, reaching tip of next segment (first endopodal segment). No remnant of exopodite visible. Endopodite consisting of four segments (penultimate segment divided). First endopodal segment large and stout,
<emphasis id="40C26A3E453AFFB3FBB319FDFBFF2FF9" box="[1125,1137,691,716]" italics="true" pageId="15" pageNumber="16">c</emphasis>
four times as long as basal width, carrying one long and narrow mid-ventral aesthetasc Y (reaching tip of segment) and two ventro-subapical setae, one short and one large and stout, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E453AFFB3FAFF19B5FAB22E21" box="[1321,1340,763,788]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 and t3 hyper-developed into male bristles, t2 ventro-apical, t3 centro-apical. Third endopodal segment (segment 2B) smaller than previous segment and about as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9453AFFB3FBD518C5FBF82E90" box="[1027,1142,907,933]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="15" pageNumber="16">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of equal length; centrally with one short seta z3; dorso-apically with the seta z1 transformed into claw, length about twice as long as penultimate segment. Claw G1 slightly shorter than G2; seta z2 transformed into long claw, slightly shorter than G1. Fourth, terminal segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and a long seta g slightly shorter than claw Gm.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F2DFBDC29F0" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdCoxa (
<figureCitation id="EA8DAAA9453AFFB3FEEF1F2DFE5F2948" box="[313,465,1122,1149]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200– INV.159203). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745726" httpUri="https://zenodo.org/record/13745726/files/figure.png" pageId="15" pageNumber="16">Figure 5B, C</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (6–7) of strong teeth, interspaced with setae; subapically with a short seta.
<emphasis id="40C26A3E453AFFB3FAEB1FC9FA1A2994" box="[1341,1428,1159,1185]" italics="true" pageId="15" pageNumber="16">Remark</emphasis>
: asymmetry illustrated in
<figureCitation id="EA8DAAA9453AFFB3FE781FE5FDAE29F0" box="[430,544,1194,1221]" captionStart="FIGURE 5" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[170,1415,185,1915]" captionTargetId="figure-17@13.[167,1419,181,1919]" captionTargetPageId="13" captionText="FIGURE 5. Benincandona martini gen. et spec. nov., limbs (♂). A. A1 (INV.159203). B, C. MdCoxa, (INV.159202). D. A2 (INV.159200– INV.159203). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745726" httpUri="https://zenodo.org/record/13745726/files/figure.png" pageId="15" pageNumber="16">Figure 5C</figureCitation>
possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111F81FE092B64" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
MdPalp (
<figureCitation id="EA8DAAA9453AFFB3FEF81F81FE2F29DC" box="[302,417,1231,1257]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745728" httpUri="https://zenodo.org/record/13745728/files/figure.png" pageId="15" pageNumber="16">Figure 6C</figureCitation>
) set at about one third of the tip of the coxa and consisting of four segments. Basal segment dorsally carrying the respiratory plate, set with six distal rays (four long, two short), subapically with one long and two short rays, proximally with one long ray (this chaetotaxy approximate); ventro-subapically with one long hirsute seta (S1) and more apically S2 hirsute seta, shorter and stout, length about one third of the length of S1; in between with the short flagellum-like alpha seta, and one smooth and short, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with two setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, length of dorsal one (slightly shorter than the fused claw) about 1.5 times length of ventral one.
</paragraph>
<paragraph id="7209B62C453AFFB3FF111D15FCB12A78" blockId="15.[151,1437,150,1869]" pageId="15" pageNumber="16">
Mx1 (
<figureCitation id="EA8DAAA9453AFFB3FEDD1D15FE2E2B40" box="[267,416,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745728" httpUri="https://zenodo.org/record/13745728/files/figure.png" pageId="15" pageNumber="16">Figure 6A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Palp (
<figureCitation id="EA8DAAA9453AFFB3FD771D31FC992BAC" box="[673,791,1663,1689]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745728" httpUri="https://zenodo.org/record/13745728/files/figure.png" pageId="15" pageNumber="16">Figure 6A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying four distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E453AFFB3FB701D89FAB22BD4" box="[1190,1340,1735,1761]" italics="true" pageId="15" pageNumber="16">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E453AFFB3FB141DA5FB5B2A31" box="[1218,1237,1771,1796]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
half the length of the long one. Respiratory plate (
<figureCitation id="EA8DAAA9453AFFB3FE2F1C41FDE22A1C" box="[505,620,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[169,1422,182,1942]" captionTargetId="figure-31@14.[166,1422,181,1957]" captionTargetPageId="14" captionText="FIGURE 6. Benincandona martini gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159200). B. Mx1, respiratory plate (INV.159202). C. MdPalp (INV.159204– INV.159203). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745728" httpUri="https://zenodo.org/record/13745728/files/figure.png" pageId="15" pageNumber="16">Figure 6B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E453AFFB3FC8E1C41FCE52A1D" box="[856,875,1807,1832]" italics="true" pageId="15" pageNumber="16">c.</emphasis>
15 plumose respiratory rays, two short distal setae and a group of five shorter proximal setae of unequal length.
</paragraph>
<caption id="26C9E6A44525FFACFF411CF3FD432ACF" ID-DOI="http://doi.org/10.5281/zenodo.13745730" ID-Zenodo-Dep="13745730" httpUri="https://zenodo.org/record/13745730/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1981,2005]" targetBox="[186,1404,198,1941]" targetPageId="16" targetType="figure">
<paragraph id="7209B62C4525FFACFF411CF3FD432ACF" blockId="16.[151,1436,1981,2042]" pageId="16" pageNumber="17">
<emphasis id="40C26A3E4525FFACFF411CF3FE962AE3" bold="true" box="[151,280,1981,2006]" pageId="16" pageNumber="17">FIGURE 7.</emphasis>
<taxonomicName id="B5B6CDAF4525FFACFEF21CF3FD832AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[292,525,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4525FFACFEF21CF3FD832AE0" box="[292,525,1981,2005]" italics="true" pageId="16" pageNumber="17">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454525FFACFDCE1CF0FD542AE0" box="[536,730,1982,2006]" pageId="16" pageNumber="17" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4525FFACFCB51CF3FCFB2AE0" bold="true" box="[867,885,1981,2005]" pageId="16" pageNumber="17">A</emphasis>
. T2 (INV.159200).
<emphasis id="40C26A3E4525FFACFB9B1CF0FBD02AE3" bold="true" box="[1101,1118,1982,2006]" pageId="16" pageNumber="17">B</emphasis>
. CR (INV.159200).
<emphasis id="40C26A3E4525FFACFAEB1CF3FAC12AE0" bold="true" box="[1341,1359,1981,2005]" pageId="16" pageNumber="17">C</emphasis>
. CRatt (INV.159200).
<emphasis id="40C26A3E4525FFACFEE71CACFECD2ACF" bold="true" box="[305,323,2018,2042]" pageId="16" pageNumber="17">D</emphasis>
. T3 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44524FFADFF411CD9FDB32ACD" ID-DOI="http://doi.org/10.5281/zenodo.13745732" ID-Zenodo-Dep="13745732" httpUri="https://zenodo.org/record/13745732/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1943,1967]" targetBox="[196,1391,191,1914]" targetPageId="17" targetType="figure">
<paragraph id="7209B62C4524FFADFF411CD9FDB32ACD" blockId="17.[151,1437,1943,2040]" pageId="17" pageNumber="18">
<emphasis id="40C26A3E4524FFADFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="17" pageNumber="18">FIGURE 8.</emphasis>
<taxonomicName id="B5B6CDAF4524FFADFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4524FFADFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="17" pageNumber="18">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454524FFADFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="17" pageNumber="18" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4524FFADFC9B1CD9FCD12A9A" bold="true" box="[845,863,1943,1967]" pageId="17" pageNumber="18">A</emphasis>
. Zenker organ (INV.159200).
<emphasis id="40C26A3E4524FFADFB771CD6FB5E2A85" bold="true" box="[1185,1232,1944,1968]" pageId="17" pageNumber="18">B–D</emphasis>
. Hp (INV.159200).
<emphasis id="40C26A3E4524FFADFF411CF2FF262AE1" bold="true" box="[151,168,1980,2004]" pageId="17" pageNumber="18">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E4524FFADFE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="17" pageNumber="18">C</emphasis>
. Internal view (photograph with interference contrast).
<emphasis id="40C26A3E4524FFADFBCE1CF2FBA42AE1" bold="true" box="[1048,1066,1980,2004]" pageId="17" pageNumber="18">D</emphasis>
. Internal view (photograph without interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44527FFAEFF411CD9FA9F2AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745734" ID-Zenodo-Dep="13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1943,1967]" targetBox="[172,1415,185,1915]" targetPageId="18" targetType="figure">
<paragraph id="7209B62C4527FFAEFF411CD9FA9F2AE1" blockId="18.[151,1436,1943,2004]" pageId="18" pageNumber="19">
<emphasis id="40C26A3E4527FFAEFF411CD9FE982A9A" bold="true" box="[151,278,1943,1967]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
<taxonomicName id="B5B6CDAF4527FFAEFEF61CD6FD892A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[288,519,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4527FFAEFEF61CD6FD892A9A" box="[288,519,1944,1968]" italics="true" pageId="18" pageNumber="19">Benincandona martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454527FFAEFDC61CD6FD452A9A" box="[528,715,1944,1968]" pageId="18" pageNumber="19" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (A–B ♂ and C–E ♀).
<emphasis id="40C26A3E4527FFAEFBDE1CD9FB942A9A" bold="true" box="[1032,1050,1943,1967]" pageId="18" pageNumber="19">A</emphasis>
. T1, Lpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFAF01CD6FAB92A85" bold="true" box="[1318,1335,1944,1968]" pageId="18" pageNumber="19">B</emphasis>
. T1, Rpp (INV.159202).
<emphasis id="40C26A3E4527FFAEFEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="18" pageNumber="19">C</emphasis>
. A2 (INV.159205, INV.159207).
<emphasis id="40C26A3E4527FFAEFD401CF2FD262AE1" bold="true" box="[662,680,1980,2004]" pageId="18" pageNumber="19">D</emphasis>
. CR (INV.159205).
<emphasis id="40C26A3E4527FFAEFCA11CF2FC062AE1" bold="true" box="[887,904,1980,2004]" pageId="18" pageNumber="19">E</emphasis>
. T1 (INV.159200). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFF111BD9FD062C50" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF201BD9FE012D84" box="[246,399,151,177]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) consisting of basal part (protopodite), carrying respiratory plates (exopodites) represented here by one long and one short plumose setae, and slightly asymmetrical prehensile palps (endopodites). Protopodite (
<figureCitation id="EA8DAAA94526FFAFFF491B91FE9E2DCC" box="[159,272,223,249]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9B</figureCitation>
) distally set with 5+2 setae of different morphology and length (not illustrated); more proximally carrying d, d’ and b setae of medium length. Prehensile palps (
<figureCitation id="EA8DAAA94526FFAFFD341A4DFCFB2C28" box="[738,885,259,285]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps apically set with a short sensory organ, proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA94526FFAFFA871A69FF352C50" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA94526FFAFFDD31A05FDF52C50" box="[517,635,331,357]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111A21FCD12F08" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T2 (
<figureCitation id="EA8DAAA94526FFAFFF201A21FEE32CBC" box="[246,365,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745730" httpUri="https://zenodo.org/record/13745730/files/figure.png" pageId="19" pageNumber="20">Figure 7A</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), generally elongated. First basal segment and knee-segment without setae d1 and d2. First segment of endopodite elongated (length at least 4x the basal width), without setae. Second endopodal segment with length
<emphasis id="40C26A3E4526FFAFFC451AF9FC292CE5" box="[915,935,439,464]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
3x basal width, with one short ventro-apical seta f. Third endopodal segment of about the same length/width ratio as the previous segment and with one ventro-subapical seta g. Terminal segment about 1.5 times as long as basal width, apically with two short setae (h1 and h3) and a long and slender claw (h2), distally set with short setulae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111909FBDB2FC4" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
T3 (
<figureCitation id="EA8DAAA94526FFAFFF211909FEE12F54" box="[247,367,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745730" httpUri="https://zenodo.org/record/13745730/files/figure.png" pageId="19" pageNumber="20">Figure 7D</figureCitation>
) with first segment carrying only one long subapical seta d1. Second (knee-) segment with a short seta d2. Third segment about 5x as long as wide, not carrying any setae. Fourth segment narrower, but still
<emphasis id="40C26A3E4526FFAFFA5F1925FA122FB1" box="[1417,1436,619,644]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
5x basal width and carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4526FFAFFF7919FDFF4C2FF9" box="[175,194,691,716]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), almost reaching knee-segment and two ventrally directed and curved setae h1 &amp; h2, the former
<emphasis id="40C26A3E4526FFAFFCDD1999FC902FC5" box="[779,798,727,752]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4526FFAFFF1119B5FC852E0C" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
CR (
<figureCitation id="EA8DAAA94526FFAFFF2819B5FEFD2E20" box="[254,371,763,789]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745730" httpUri="https://zenodo.org/record/13745730/files/figure.png" pageId="19" pageNumber="20">Figure 7B</figureCitation>
) with ramus stout; claw Ga well-developed, stout and reaching widely beyond half L of the ramus, setae Sp, Sa and claw Gp reduced to small setae.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11180DFCA92E68" blockId="19.[151,1437,151,1005]" box="[199,807,835,861]" pageId="19" pageNumber="20">
CRAtt (
<figureCitation id="EA8DAAA94526FFAFFEC9180DFE1D2E68" box="[287,403,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1981,2005]" captionTargetBox="[186,1404,198,1941]" captionTargetId="figure-31@16.[166,1422,181,1957]" captionTargetPageId="16" captionText="FIGURE 7. Benincandona martini gen. et spec. nov., limbs (♂). A. T2 (INV.159200). B. CR (INV.159200). C. CRatt (INV.159200). D. T3 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745730" httpUri="https://zenodo.org/record/13745730/files/figure.png" pageId="19" pageNumber="20">Figure 7C</figureCitation>
) short and stout, distally bifurcated.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111829FD4E2EFC" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
Hp (
<figureCitation id="EA8DAAA94526FFAFFF2F1829FE1E2EB4" box="[249,400,871,897]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). B–D. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745732" httpUri="https://zenodo.org/record/13745732/files/figure.png" pageId="19" pageNumber="20">Figure 8B–D</figureCitation>
) relatively small, with bluntly pointed ls, reaching slightly beyond asymmetrically rounded ms; lobe ‘h’ shorter and sub-rectangular, ls and ms with diverging positions; internal anatomy (
<figureCitation id="EA8DAAA94526FFAFFB1218C5FAD52E90" box="[1220,1371,907,933]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). B–D. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745732" httpUri="https://zenodo.org/record/13745732/files/figure.png" pageId="19" pageNumber="20">Figure 8C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4526FFAFFF11189DFB892ED8" blockId="19.[151,1437,151,1005]" box="[199,1031,979,1005]" pageId="19" pageNumber="20">
Zenker organ (
<figureCitation id="EA8DAAA94526FFAFFEBD189DFE6F2ED8" box="[363,481,979,1005]" captionStart="FIGURE 8" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[196,1391,191,1914]" captionTargetId="figure-17@17.[186,1401,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. Benincandona martini gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159200). B–D. Hp (INV.159200). B. External view (drawing). C. Internal view (photograph with interference contrast). D. Internal view (photograph without interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745732" httpUri="https://zenodo.org/record/13745732/files/figure.png" pageId="19" pageNumber="20">Figure 8A</figureCitation>
) short and broad, with eight-nine spinous whirls.
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F54FE182901" blockId="19.[151,1436,1050,1545]" box="[151,406,1050,1076]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF411F54FE182901" bold="true" box="[151,406,1050,1076]" pageId="19" pageNumber="20">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF411F71FB10296C" blockId="19.[151,1436,1050,1545]" box="[151,1182,1087,1113]" pageId="19" pageNumber="20">
Cp and valves as in the male, without obvious sexual dimorphism (
<figureCitation id="EA8DAAA94526FFAFFC501F71FBCE296C" box="[902,1088,1087,1113]" captionStart="FIGURE 3" captionStartId="11.[151,250,1933,1957]" captionTargetBox="[172,1415,181,1910]" captionTargetId="figure-17@11.[172,1415,178,1910]" captionTargetPageId="11" captionText="FIGURE 3. Benincandona martini gen. et spec. nov., valves (A, C, E–H, J ♂; B, D, I ♀). A–D, I. CpRl, (A, C. INV.159208; B, D, I. INV.159211). E. INV.159209. CpD. F. INV.159210. CpV. G. INV.159200. LVi. H. INV.159200. RVi. J. INV.159200. CMS. Scale bars. 1 (A, B, E, F, G, H) = 500 µm; 2 (C, D) = 200 µm; 3 (I, J) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745720" httpUri="https://zenodo.org/record/13745720/files/figure.png" pageId="19" pageNumber="20">Figures 3B, D, I</figureCitation>
;
<figureCitation id="EA8DAAA94526FFAFFB9A1F71FB1C296C" box="[1100,1170,1087,1113]" captionStart="FIGURE 4" captionStartId="12.[151,250,1944,1968]" captionTargetBox="[157,1430,181,1920]" captionTargetId="figure-31@12.[157,1430,181,1920]" captionTargetPageId="12" captionText="FIGURE 4. Benincandona martini gen. et spec. nov., valves (A–D: INV.159202 ♂ and E–H. INV.159201 ♀) A, E. LVi. B, F. RVi, C, D, G, H. CMS. Scale bars. 1 (A, B, E, F) = 500 µm; 2 (C, D, G, H) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745722" httpUri="https://zenodo.org/record/13745722/files/figure.png" pageId="19" pageNumber="20">4E–H</figureCitation>
).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111F2DFBF52948" blockId="19.[151,1436,1050,1545]" box="[199,1147,1123,1149]" pageId="19" pageNumber="20">
Limbs (
<figureCitation id="EA8DAAA94526FFAFFECB1F2DFE3E2948" box="[285,432,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9C–E</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111FC9FC232838" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
A2 (
<figureCitation id="EA8DAAA94526FFAFFF2F1FC9FEE12994" box="[249,367,1159,1185]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9C</figureCitation>
) with second endopodal segment undivided, with two t-setae (one long and plumose (t1), one smooth short and thin (t2)); apical chaetotaxy (
<figureCitation id="EA8DAAA94526FFAFFD771FE5FC9E29F0" box="[673,784,1195,1221]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="19" pageNumber="20">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4526FFAFFF111E59FC31284C" blockId="19.[151,1436,1050,1545]" pageId="19" pageNumber="20">
T1 (
<figureCitation id="EA8DAAA94526FFAFFF2F1E59FEE12804" box="[249,367,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[172,1415,185,1915]" captionTargetId="figure-31@18.[168,1419,181,1919]" captionTargetPageId="18" captionText="FIGURE 9. Benincandona martini gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159202). B. T1, Rpp (INV.159202). C. A2 (INV.159205, INV.159207). D. CR (INV.159205). E. T1 (INV.159200). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745734" httpUri="https://zenodo.org/record/13745734/files/figure.png" pageId="19" pageNumber="20">Figure 9E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4526FFAFFDA01E59FD072805" box="[630,649,1303,1328]" italics="true" pageId="19" pageNumber="20">c.</emphasis>
10 distal setae of unequal length, setae d and b present, with long additional seta (d’) present close to seta d; respiratory plate reduced to about three short rays. Endopodite set with a broad and short palp, with three subequal but short apical setae (h1–h3).
</paragraph>
<paragraph id="7209B62C4526FFAFFF111ECDFD8A28A8" blockId="19.[151,1436,1050,1545]" box="[199,516,1411,1437]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111ECDFE0E28A8" bold="true" box="[199,384,1411,1437]" pageId="19" pageNumber="20">Measurements:</emphasis>
see
<tableCitation id="3F3483974526FFAFFE671ECDFD8A28A8" box="[433,516,1411,1437]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="19" pageNumber="20">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111EE8FDD328F4" box="[199,605,1446,1473]" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="7209B62C4526FFAFFF111EE8FDD328F4" blockId="19.[151,1436,1050,1545]" box="[199,605,1446,1473]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111EE8FE5B28F5" bold="true" box="[199,469,1446,1472]" pageId="19" pageNumber="20">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974526FFAFFDDC1EE9FDD328F4" box="[522,605,1447,1473]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="19" pageNumber="20">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74526FFAFFF111E85FA1E2AD6" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="7209B62C4526FFAFFF111E85FE6128D1" blockId="19.[151,1436,1050,1545]" box="[199,495,1482,1509]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF111E85FE6128D1" bold="true" box="[199,495,1482,1509]" pageId="19" pageNumber="20">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4526FFAFFF111EA1FBD02B3C" blockId="19.[151,1436,1050,1545]" box="[199,1118,1519,1545]" pageId="19" pageNumber="20">
This subterranean species is presently only known from its
<typeStatus id="AD0D088E4526FFAFFCB61EA1FC1E2B3C" box="[864,912,1519,1545]" pageId="19" pageNumber="20">type</typeStatus>
locality (
<tableCitation id="3F3483974526FFAFFC2C1EA1FBC32B3C" box="[1018,1101,1519,1545]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="19" pageNumber="20">Table 1</tableCitation>
).
</paragraph>
<caption id="26C9E6A44526FFAFFF4E1D76FB332B64" box="[152,1213,1592,1617]" pageId="19" pageNumber="20" startId="19.[152,244,1592,1616]" targetType="table">
<paragraph id="7209B62C4526FFAFFF4E1D76FB332B64" blockId="19.[152,1213,1592,1617]" box="[152,1213,1592,1617]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFF4E1D76FE812B65" bold="true" box="[152,271,1592,1616]" pageId="19" pageNumber="20">TABLE 3.</emphasis>
Measurements of illustrated specimens of
<taxonomicName id="B5B6CDAF4526FFAFFD0F1D76FCFD2B65" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[729,883,1592,1616]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFD0F1D76FCFD2B65" box="[729,883,1592,1616]" italics="true" pageId="19" pageNumber="20">Benincandona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFCAF1D77FBBA2B65" box="[889,1076,1593,1617]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, using SEM.
</paragraph>
</caption>
<paragraph id="7209B62C4526FFAFFE8C1D2EFAF62AA7" pageId="19" pageNumber="20">
<table id="00B6448C45260043FF751D2EFA0B2AA7" box="[163,1413,1632,1938]" gridcols="7" gridrows="8" pageId="19" pageNumber="20">
<tr id="CC86B46E45260043FF751D2EFA0B2B4D" box="[163,1413,1632,1656]" gridrow="0" pageId="19" pageNumber="20">
<th id="8F57DD1245260043FF751D2EFE312B4D" box="[163,447,1632,1656]" gridcol="0" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFE8C1D2EFE272B4D" bold="true" box="[346,425,1632,1656]" pageId="19" pageNumber="20">Species</emphasis>
</th>
<th id="8F57DD1245260043FD4D1D2EFD4F2B4D" box="[667,705,1632,1656]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFD4D1D2EFD4F2B4D" bold="true" box="[667,705,1632,1656]" pageId="19" pageNumber="20">Sex</emphasis>
</th>
<th id="8F57DD1245260043FCD51D2EFCF12B4D" box="[771,895,1632,1656]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFCD91D2EFCFD2B4D" bold="true" box="[783,883,1632,1656]" pageId="19" pageNumber="20">specimen</emphasis>
</th>
<th id="8F57DD1245260043FC611D2EFC7E2B4D" box="[951,1008,1632,1656]" gridcol="3" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFC611D2EFC7E2B4D" bold="true" box="[951,1008,1632,1656]" pageId="19" pageNumber="20">V/Cp</emphasis>
</th>
<th id="8F57DD1245260043FBE71D2EFBF52B4D" box="[1073,1147,1632,1656]" gridcol="4" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFBE71D2EFBF52B4D" bold="true" box="[1073,1147,1632,1656]" pageId="19" pageNumber="20">L (µm)</emphasis>
</th>
<th id="8F57DD1245260043FB641D2EFA8F2B4D" box="[1202,1281,1632,1656]" gridcol="5" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB641D2EFA8F2B4D" bold="true" box="[1202,1281,1632,1656]" pageId="19" pageNumber="20">H (µm)</emphasis>
</th>
<th id="8F57DD1245260043FAE41D2EFA0B2B4D" box="[1330,1413,1632,1656]" gridcol="6" gridrow="0" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFAE41D2EFA0B2B4D" bold="true" box="[1330,1413,1632,1656]" pageId="19" pageNumber="20">W (µm)</emphasis>
</th>
</tr>
<tr id="CC86B46E45260043FF751DC6FA0B2B94" box="[163,1413,1672,1697]" gridrow="1" pageId="19" pageNumber="20" rowspan-6="1">
<th id="8F57DD1245260043FF751DC6FE312B94" box="[163,447,1672,1697]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">
<taxonomicName id="B5B6CDAF4526FFAFFF751DC7FE842B95" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[163,266,1673,1696]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="martini" status="gen. et spec. nov.">
<emphasis id="40C26A3E4526FFAFFF751DC7FE842B95" box="[163,266,1673,1696]" italics="true" pageId="19" pageNumber="20">B. martini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454526FFAFFEC61DC7FE312B95" box="[272,447,1673,1697]" pageId="19" pageNumber="20" rank="species">gen. et spec. nov.</taxonomicNameLabel>
</th>
<td id="8F57DD1245260043FD4D1DC6FD4F2B94" box="[667,705,1672,1697]" gridcol="1" gridrow="1" pageId="19" pageNumber="20">♂</td>
<td id="8F57DD1245260043FCD51DC6FCF12B94" box="[771,895,1672,1697]" gridcol="2" gridrow="1" pageId="19" pageNumber="20">INV.159200</td>
<td id="8F57DD1245260043FC611DC6FC7E2B94" box="[951,1008,1672,1697]" gridcol="3" gridrow="1" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71DC6FBF52B94" box="[1073,1147,1672,1697]" gridcol="4" gridrow="1" pageId="19" pageNumber="20">706.7</td>
<td id="8F57DD1245260043FB641DC6FA8F2B94" box="[1202,1281,1672,1697]" gridcol="5" gridrow="1" pageId="19" pageNumber="20">331.7</td>
</tr>
<tr id="CC86B46E45260043FF751DFFFA0B2BFC" box="[163,1413,1713,1737]" gridrow="2" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611DFFFC7E2BFC" box="[951,1008,1713,1737]" gridcol="3" gridrow="2" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71DFFFBF52BFC" box="[1073,1147,1713,1737]" gridcol="4" gridrow="2" pageId="19" pageNumber="20">697.9</td>
<td id="8F57DD1245260043FB641DFFFA8F2BFC" box="[1202,1281,1713,1737]" gridcol="5" gridrow="2" pageId="19" pageNumber="20">310.7</td>
</tr>
<tr id="CC86B46E45260043FF751D97FA0B2BC4" box="[163,1413,1753,1777]" gridrow="3" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51D97FCF12BC4" box="[771,895,1753,1777]" gridcol="2" gridrow="3" pageId="19" pageNumber="20">INV.159202</td>
<td id="8F57DD1245260043FC611D97FC7E2BC4" box="[951,1008,1753,1777]" gridcol="3" gridrow="3" pageId="19" pageNumber="20">LVi</td>
<td id="8F57DD1245260043FBE71D97FBF52BC4" box="[1073,1147,1753,1777]" gridcol="4" gridrow="3" pageId="19" pageNumber="20">667.5</td>
<td id="8F57DD1245260043FB641D97FA8F2BC4" box="[1202,1281,1753,1777]" gridcol="5" gridrow="3" pageId="19" pageNumber="20">317.5</td>
</tr>
<tr id="CC86B46E45260043FF751C4FFA0B2A2C" box="[163,1413,1793,1817]" gridrow="4" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1">
<td id="8F57DD1245260043FC611C4FFC7E2A2C" box="[951,1008,1793,1817]" gridcol="3" gridrow="4" pageId="19" pageNumber="20">RVi</td>
<td id="8F57DD1245260043FBE71C4FFBF52A2C" box="[1073,1147,1793,1817]" gridcol="4" gridrow="4" pageId="19" pageNumber="20">656.7</td>
<td id="8F57DD1245260043FB641C4FFA8F2A2C" box="[1202,1281,1793,1817]" gridcol="5" gridrow="4" pageId="19" pageNumber="20">295.0</td>
</tr>
<tr id="CC86B46E45260043FF751C67FA0B2A77" box="[163,1413,1833,1858]" gridrow="5" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-6="1">
<td id="8F57DD1245260043FCD51C67FCF12A77" box="[771,895,1833,1858]" gridcol="2" gridrow="5" pageId="19" pageNumber="20">INV.159208</td>
<td id="8F57DD1245260043FC611C67FC7E2A77" box="[951,1008,1833,1858]" gridcol="3" gridrow="5" pageId="19" pageNumber="20">CpRl</td>
<td id="8F57DD1245260043FBE71C67FBF52A77" box="[1073,1147,1833,1858]" gridcol="4" gridrow="5" pageId="19" pageNumber="20">692.9</td>
<td id="8F57DD1245260043FB641C67FA8F2A77" box="[1202,1281,1833,1858]" gridcol="5" gridrow="5" pageId="19" pageNumber="20">334.3</td>
</tr>
<tr id="CC86B46E45260043FF751C1CFA0B2A5F" box="[163,1413,1874,1898]" gridrow="6" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C1CFCF12A5F" box="[771,895,1874,1898]" gridcol="2" gridrow="6" pageId="19" pageNumber="20">INV.159209</td>
<td id="8F57DD1245260043FC611C1CFC7E2A5F" box="[951,1008,1874,1898]" gridcol="3" gridrow="6" pageId="19" pageNumber="20">CpD</td>
<td id="8F57DD1245260043FBE71C1CFBF52A5F" box="[1073,1147,1874,1898]" gridcol="4" gridrow="6" pageId="19" pageNumber="20">698.6</td>
<td id="8F57DD1245260043FAE41C1CFA0B2A5F" box="[1330,1413,1874,1898]" gridcol="6" gridrow="6" pageId="19" pageNumber="20">185.0</td>
</tr>
<tr id="CC86B46E45260043FF751C34FA0B2AA7" box="[163,1413,1914,1938]" gridrow="7" pageId="19" pageNumber="20" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="8F57DD1245260043FCD51C34FCF12AA7" box="[771,895,1914,1938]" gridcol="2" gridrow="7" pageId="19" pageNumber="20">INV.159210</td>
<td id="8F57DD1245260043FC611C34FC7E2AA7" box="[951,1008,1914,1938]" gridcol="3" gridrow="7" pageId="19" pageNumber="20">CpV</td>
<td id="8F57DD1245260043FBE71C34FBF52AA7" box="[1073,1147,1914,1938]" gridcol="4" gridrow="7" pageId="19" pageNumber="20">679.3</td>
<td id="8F57DD1245260043FAE41C34FA0B2AA7" box="[1330,1413,1914,1938]" gridcol="6" gridrow="7" pageId="19" pageNumber="20">190.7</td>
</tr>
</table>
</paragraph>
<paragraph id="7209B62C4526FFAFFB8F1C85FA1E2AD6" blockId="19.[1113,1424,1995,2019]" box="[1113,1424,1995,2019]" pageId="19" pageNumber="20">
<emphasis id="40C26A3E4526FFAFFB8F1C85FA1E2AD6" box="[1113,1424,1995,2019]" italics="true" pageId="19" pageNumber="20">......continued on the next page</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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