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<document id="101A23A926569C24A2287AD2556C500B" ID-DOI="10.11646/zootaxa.5503.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745712" ID-ZooBank="7385E4AA-D7AD-4E79-A05C-684319E0843B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726051594773" checkinUser="plazi" docAuthor="Hotèkpo, Sourou Joseph, Schön, Isa &amp; Martens, Koen" docDate="2024" docId="FA1F073A4520FFA1FF41190AFB072994" docLanguage="en" docName="zootaxa.5503.1.1.pdf" docOrigin="Zootaxa 5503 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5503.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Benincandona ibikounlei Hotèkpo, Schön &amp; Martens, 2024, gen. et spec. nov." docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="06267F424535FFBCFFD61B4EFF8E2D35" masterDocTitle="An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="22" updateTime="1726052453190" updateUser="ExternalLinkService">
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<mods:title id="B762EBF795BB46933969A6E4C1F58383">An endemic species flock of Candonidae Kaufmann, 1900 (Crustacea, Ostracoda) from wells in Benin (Africa), with the description of a new subfamily, a new genus and five new species</mods:title>
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<mods:namePart id="BE5F64B0B72DCE97C25EA72D8BC33362">Hotèkpo, Sourou Joseph</mods:namePart>
<mods:nameIdentifier id="CD263E789CC767D640BF2782A90FD431" type="ORCID">0000-0001-9091-433X</mods:nameIdentifier>
<mods:affiliation id="20782F70B62D11CB55EA0210E79107F9">Centre de Recherche pour la lutte contre les Maladies Infectieuses Tropicales (CReMIT / TIDRC), Université d’Abomey-Calavi, Bénin jshotekpo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9091 - 433 X &amp; Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="CFFDBB1E3B0B9B3311034352B50D99A0" type="email">jshotekpo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A5AEDE1FF072A00C373B8DBE912188CF">Schön, Isa</mods:namePart>
<mods:nameIdentifier id="17A131761FB7A4159104647655EFE548" type="ORCID">0000-0001-9269-6487</mods:nameIdentifier>
<mods:affiliation id="178857D617E7E6B21DAA4F53A2223791">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; University of Hasselt, Research Group Zoology, Agoralaan Building D, B- 3590 Diepenbeek, Belgium ischoen @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 9269 - 6487</mods:affiliation>
<mods:nameIdentifier id="3D123578075046DBA224CF1680F758D5" type="email">ischoen@naturalsciences.be</mods:nameIdentifier>
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<mods:role id="03DFD55C99530A6E71FF8BAA8CD129DD">
<mods:roleTerm id="B46B3F3E70505C6F1C3C87869911D8CA">Author</mods:roleTerm>
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<mods:namePart id="812A52E8997C5D490AEA8BF8E9961E06">Martens, Koen</mods:namePart>
<mods:nameIdentifier id="67BC7D514920E1CADEE33F18D9C75CBB" type="ORCID">0000-0001-8680-973X</mods:nameIdentifier>
<mods:affiliation id="193BDE6777F5C84F024E791122E5A62D">Royal Belgian Institute of Natural Sciences, Freshwater Biology, Vautierstraat 29, B- 1000 Brussels, Belgium &amp; Ghent University, Dept. Biology, K. L. Ledeganckstraat 35, B- 9000 Ghent, Belgium kmartens @ naturalsciences. be; https: // orcid. org / 0000 - 0001 - 8680 - 973 X * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="3402F4BC6419A5E4C39A987DB40C9FB4" type="email">kmartens@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="7C3E53A3EEC45F70C8E7C2AD0D2371FA">Zootaxa</mods:title>
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<mods:part id="86ACC9D8CD6630F7893681ECEFB50877">
<mods:date id="43DE7D38B90A36EBDAAB922C38E1A452">2024</mods:date>
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<mods:number id="0755C09C052D8D313328B9C4AC2756EA">2024-09-04</mods:number>
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<mods:number id="73AA554C7EABD5B29A5EBE73B821B473">5503</mods:number>
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<mods:identifier id="95920F5F89F754D51D9687A57DE65907" type="DOI">10.11646/zootaxa.5503.1.1</mods:identifier>
<mods:identifier id="595182C6A965937F5A4717C556A2D4C2" type="ISSN">1175-5334</mods:identifier>
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<treatment id="FA1F073A4520FFA1FF41190AFB072994" LSID="urn:lsid:plazi:treatment:FA1F073A4520FFA1FF41190AFB072994" httpUri="http://treatment.plazi.org/id/FA1F073A4520FFA1FF41190AFB072994" lastPageId="29" lastPageNumber="30" pageId="21" pageNumber="22">
<subSubSection id="3AACE5A74520FFA9FF41190AFC5A2FB1" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="7209B62C4520FFA9FF41190AFC5A2FB1" blockId="21.[151,980,580,680]" pageId="21" pageNumber="22">
<heading id="294101404520FFA9FF41190AFC282F6A" bold="true" box="[151,934,580,607]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="40C26A3E4520FFA9FF41190AFC282F6A" bold="true" box="[151,934,580,607]" pageId="21" pageNumber="22">
<taxonomicName id="B5B6CDAF4520FFA9FF41190AFE442F6A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[151,458,580,607]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FF41190AFE442F6A" bold="true" box="[151,458,580,607]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454520FFA9FE04190BFD2C2F6A" box="[466,674,581,607]" pageId="21" pageNumber="22" rank="species">gen. et spec. nov.</taxonomicNameLabel>
Hotèkpo &amp; Martens
</emphasis>
</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="3AACE5A74520FFA9FF4119C3FE2D2F92" box="[151,419,653,680]" pageId="21" pageNumber="22" type="description">
<paragraph id="7209B62C4520FFA9FF4119C3FE2D2F92" blockId="21.[151,980,580,680]" box="[151,419,653,680]" pageId="21" pageNumber="22">
(
<figureCitation id="EA8DAAA94520FFA9FF4919C3FECA2F9D" box="[159,324,653,680]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStartId-0="21.[151,250,1855,1879]" captionStartId-1="22.[151,250,1943,1967]" captionStartId-2="25.[151,250,1981,2005]" captionStartId-3="26.[151,250,1943,1967]" captionStartId-4="27.[151,250,1943,1967]" captionStartId-5="28.[151,250,1943,1967]" captionTargetBox-0="[151,1436,883,1832]" captionTargetBox-1="[183,1405,185,1915]" captionTargetBox-2="[166,1422,183,1955]" captionTargetBox-3="[191,1395,185,1916]" captionTargetBox-4="[229,1359,191,1909]" captionTargetBox-5="[176,1407,181,1915]" captionTargetId-0="figure-159@21.[151,1436,883,1832]" captionTargetId-1="figure-31@22.[179,1407,181,1919]" captionTargetId-2="figure-17@25.[166,1422,181,1957]" captionTargetId-3="figure-31@26.[187,1399,181,1919]" captionTargetId-4="figure-17@27.[219,1369,181,1919]" captionTargetId-5="figure-31@28.[176,1411,181,1919]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="25" captionTargetPageId-3="26" captionTargetPageId-4="27" captionTargetPageId-5="28" captionText-0="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." captionText-1="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." captionText-2="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." captionText-3="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." captionText-4="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). B–D. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." captionText-5="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.13745738" figureDoi-1="http://doi.org/10.5281/zenodo.13745742" figureDoi-2="http://doi.org/10.5281/zenodo.13745746" figureDoi-3="http://doi.org/10.5281/zenodo.13745748" figureDoi-4="http://doi.org/10.5281/zenodo.13745750" figureDoi-5="http://doi.org/10.5281/zenodo.13745752" httpUri-0="https://zenodo.org/record/13745738/files/figure.png" httpUri-1="https://zenodo.org/record/13745742/files/figure.png" httpUri-2="https://zenodo.org/record/13745746/files/figure.png" httpUri-3="https://zenodo.org/record/13745748/files/figure.png" httpUri-4="https://zenodo.org/record/13745750/files/figure.png" httpUri-5="https://zenodo.org/record/13745752/files/figure.png" pageId="21" pageNumber="22">
Figures 10–
<geoCoordinate id="1782D0EB4520FFA9FEFF19C3FECA2F9D" box="[297,324,653,680]" degrees="15, 41" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="15.41">15</geoCoordinate>
</figureCitation>
,
<figureCitation id="EA8DAAA94520FFA9FE8619C3FE122F92" box="[336,412,653,679]" captionStart="FIGURE 41" captionStartId="61.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-17@61.[151,1436,181,1089]" captionTargetPageId="61" captionText="FIGURE 41. Benincandona gen. et spec. nov., CpRl. (A, C, E, F, G, I, K, N, O, Q ♂ and B, D, H, J, L, M, P, R ♀). A–D. B. martini. E–F. B. ibikounlei. G–J. B. sakitii. K–N. B. cebios. O–R. B. trapezoides. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745806" httpUri="https://zenodo.org/record/13745806/files/figure.png" pageId="21" pageNumber="22">41E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74520FFA9FF41199BFEB92E6E" pageId="21" pageNumber="22" type="etymology">
<paragraph id="7209B62C4520FFA9FF41199BFE962FDA" blockId="21.[151,1436,725,859]" box="[151,280,725,751]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF41199BFE962FDA" bold="true" box="[151,280,725,751]" pageId="21" pageNumber="22">Etymology</emphasis>
</paragraph>
<paragraph id="7209B62C4520FFA9FF4119B7FEB92E6E" blockId="21.[151,1436,725,859]" pageId="21" pageNumber="22">
The species is named after Prof. Moudachirou Ibikounlé (Université d’Abomey Calavi,
<collectingCountry id="0AA1F6BC4520FFA9FB5E19B7FB432E26" box="[1160,1229,761,787]" name="Benin" pageId="21" pageNumber="22">Bénin</collectingCountry>
) in recognition of his contribution to the advancement of scientific research in
<collectingCountry id="0AA1F6BC4520FFA9FCEF1853FCF12E02" box="[825,895,797,823]" name="Benin" pageId="21" pageNumber="22">Benin</collectingCountry>
, and in gratitude for his continued support to the career of SJH.
</paragraph>
</subSubSection>
<caption id="26C9E6A44520FFA9FF411C71FCDB2A49" ID-DOI="http://doi.org/10.5281/zenodo.13745738" ID-Zenodo-Dep="13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1855,1879]" targetBox="[151,1436,883,1832]" targetPageId="21" targetType="figure">
<paragraph id="7209B62C4520FFA9FF411C71FCDB2A49" blockId="21.[151,1437,1855,1916]" pageId="21" pageNumber="22">
<emphasis id="40C26A3E4520FFA9FF411C71FEAE2A62" bold="true" box="[151,288,1855,1879]" pageId="21" pageNumber="22">FIGURE 10.</emphasis>
<taxonomicName id="B5B6CDAF4520FFA9FEF01C71FDAD2A62" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[294,547,1855,1879]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4520FFA9FEF01C71FDAD2A62" box="[294,547,1855,1879]" italics="true" pageId="21" pageNumber="22">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454520FFA9FDFC1C0EFD552A62" box="[554,731,1856,1880]" pageId="21" pageNumber="22" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, valves.
<emphasis id="40C26A3E4520FFA9FCE41C71FCCA2A62" bold="true" box="[818,836,1855,1879]" pageId="21" pageNumber="22">A</emphasis>
. INV.159222. CpRl.
<emphasis id="40C26A3E4520FFA9FBCA1C71FBA32A62" bold="true" box="[1052,1069,1855,1879]" pageId="21" pageNumber="22">B</emphasis>
. INV.159222. CpV.
<emphasis id="40C26A3E4520FFA9FB2D1C71FA832A62" bold="true" box="[1275,1293,1855,1879]" pageId="21" pageNumber="22">C</emphasis>
. INV.159223. CpD.
<emphasis id="40C26A3E4520FFA9FF021C2DFF682A4E" bold="true" box="[212,230,1891,1915]" pageId="21" pageNumber="22">D</emphasis>
. INV.159224. LVi.
<emphasis id="40C26A3E4520FFA9FE7B1C2DFE302A4E" bold="true" box="[429,446,1891,1915]" pageId="21" pageNumber="22">E</emphasis>
. INV.159224. RVi. Scale bar = 500 µm.
</paragraph>
</caption>
<caption id="26C9E6A44523FFAAFF411CD9FC362AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745742" ID-Zenodo-Dep="13745742" httpUri="https://zenodo.org/record/13745742/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1943,1967]" targetBox="[183,1405,185,1915]" targetPageId="22" targetType="figure">
<paragraph id="7209B62C4523FFAAFF411CD9FC362AE1" blockId="22.[151,1436,1943,2004]" pageId="22" pageNumber="23">
<emphasis id="40C26A3E4523FFAAFF411CD9FEAE2A9A" bold="true" box="[151,288,1943,1967]" pageId="22" pageNumber="23">FIGURE 11.</emphasis>
<taxonomicName id="B5B6CDAF4523FFAAFEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4523FFAAFEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="22" pageNumber="23">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454523FFAAFDE71CD6FD662A9A" box="[561,744,1944,1968]" pageId="22" pageNumber="23" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E4523FFAAFCB11CD9FCF72A9A" bold="true" box="[871,889,1943,1967]" pageId="22" pageNumber="23">A</emphasis>
. A2 (INV.159219, INV.159217, INV.159218).
<emphasis id="40C26A3E4523FFAAFA891CD6FAFE2A85" bold="true" box="[1375,1392,1944,1968]" pageId="22" pageNumber="23">B</emphasis>
. A1 (INV.159219, INV.159218).
<emphasis id="40C26A3E4523FFAAFE6C1CF5FE652AE1" bold="true" box="[442,491,1979,2004]" pageId="22" pageNumber="23">C, D</emphasis>
. MdCoxa, (INV.159218). Scale bar = 50 µm.
</paragraph>
</caption>
<subSubSection id="3AACE5A74522FFABFF411BD8FD132CBC" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="7209B62C4522FFABFF411BD8FE842D85" blockId="23.[151,1436,150,393]" box="[151,266,150,176]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411BD8FE842D85" bold="true" box="[151,266,150,176]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411BF5FD132CBC" blockId="23.[151,1436,150,393]" pageId="23" pageNumber="24">CpRl elongated (L/H = 2.4), sub-triangular, with greatest height slightly in front of the middle; LV dorsally extending beyond RV, anterior and posterior margins evenly rounded, valve surface smooth. CpD and CpV narrow, with greatest width in the middle and angular lateral sides, anterior and posterior extremities pointed. A2 with seta X short, not reaching base of aesthetasc Y. Second palp segment of Mx1 with basal W almost equalling L. T3 with seta d2 absent. Hp with ls and ms with slightly diverging positions, narrower lobe ls on Hp with skewed rectangular distal margin, extending well beyond lobe ms.</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74522FFABFF411AF9FC4E296C" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="7209B62C4522FFABFF411AF9FEB02CE5" blockId="23.[151,318,438,465]" box="[151,318,438,465]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AF9FEB02CE5" bold="true" box="[151,318,438,465]" pageId="23" pageNumber="24">Type material</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411AB1FE8A2F2C" blockId="23.[151,1394,511,573]" box="[151,260,511,537]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411AB1FE8A2F2C" bold="true" box="[151,260,511,537]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411AB1FE8A2F2C" box="[151,260,511,537]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF41196DFAFC2F08" blockId="23.[151,1394,511,573]" box="[151,1394,546,573]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF41196DFAFC2F08" box="[151,1394,546,573]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="64B07DA54522FFABFF41196DFF4D2F08" box="[151,195,547,573]" count="1" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, specimen with soft parts dissected in glycerine in a sealed slide, and with valves stored dry (
<collectionCode id="14A72EE94522FFABFB01196DFA892F08" box="[1239,1287,547,573]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159215).
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF411924FF722FB1" blockId="23.[151,1436,618,717]" box="[151,252,618,644]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411924FF722FB1" bold="true" box="[151,252,618,644]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF411924FF722FB1" box="[151,252,618,644]" pageId="23" pageNumber="24" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119C1FEE92FF8" blockId="23.[151,1436,618,717]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4119C1FEE92FF8" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="64B07DA54522FFABFF4119C1FF4F2F9C" box="[151,193,655,681]" count="1" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, soft part dissected as the male (
<collectionCode id="14A72EE94522FFABFDFA19C0FDEF2F9C" box="[556,609,654,681]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159216). Valves lost: the decalcified valves could not be preserved during the manipulations.
</materialsCitation>
</paragraph>
<paragraph id="7209B62C4522FFABFF4119B5FE812E20" blockId="23.[151,1436,763,933]" box="[151,271,763,789]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF4119B5FE812E20" bold="true" box="[151,271,763,789]" pageId="23" pageNumber="24">
<typeStatus id="AD0D088E4522FFABFF4119B5FE812E20" box="[151,271,763,789]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411851FEBF2E68" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF411851FC4A2E0C" box="[151,964,798,825]" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFF411851FF582E0C" box="[151,214,799,825]" count="3" pageId="23" pageNumber="24" type="male">3 ♂♂</specimenCount>
dissected as the
<typeStatus id="AD0D088E4522FFABFE591851FE7F2E0C" box="[399,497,799,825]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(
<collectionCode id="14A72EE94522FFABFE291850FDBD2E0C" box="[511,563,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159217, INV.159218, INV.159219)
</materialsCitation>
;
<materialsCitation id="C2DEBC714522FFABFC181851FEA32E68" collectionCode="INV" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="64B07DA54522FFABFC181851FB832E0C" box="[974,1037,799,825]" count="2" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
dissected as the male (
<collectionCode id="14A72EE94522FFABFADC1850FAB02E0C" box="[1290,1342,798,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13356" name="Inverness Museum and Art Gallery" pageId="23" pageNumber="24" type="Herbarium">INV</collectionCode>
.159220, INV.159221)
</materialsCitation>
.
</paragraph>
<paragraph id="7209B62C4522FFABFF111829FE872E90" blockId="23.[151,1436,763,933]" pageId="23" pageNumber="24">
The
<typeStatus id="AD0D088E4522FFABFF291829FEEF2EB4" box="[255,353,871,897]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
male comprises both valves and dissected soft parts, but no illustrations are based on this specimen.
</paragraph>
<paragraph id="7209B62C4522FFABFF41189DFEBE2ED9" blockId="23.[151,1436,978,1113]" box="[151,304,978,1005]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF41189DFEBE2ED9" bold="true" box="[151,304,978,1005]" pageId="23" pageNumber="24">Type locality</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF4118B9FC4E296C" blockId="23.[151,1436,978,1113]" pageId="23" pageNumber="24">
<materialsCitation id="C2DEBC714522FFABFF4118B9FC32296C" collectingDate="2016-08-09" collectorName="Leg. &amp; S. J. Hotekpo &amp; M. Lagnika &amp; P. Martin" country="Benin" county="Colline Department" latitude="7.8160834" location="Tchetti" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Zou">
<collectingCountry id="0AA1F6BC4522FFABFF4118B9FF542924" box="[151,218,1015,1041]" name="Benin" pageId="23" pageNumber="24">Benin</collectingCountry>
,
<collectingCounty id="9B68CEA04522FFABFF3C18B9FE422924" box="[234,460,1015,1041]" pageId="23" pageNumber="24">Colline Department</collectingCounty>
,
<collectingMunicipality id="926D2C564522FFABFE0F18B9FD1C2924" box="[473,658,1015,1041]" pageId="23" pageNumber="24">Savalou District</collectingMunicipality>
, town of
<location id="7769E0F74522FFABFD2918B9FCC12924" LSID="urn:lsid:plazi:treatment:FA1F073A4520FFA1FF41190AFB072994:7769E0F74522FFABFD2918B9FCC12924" box="[767,847,1015,1041]" country="Benin" county="Colline Department" latitude="7.8160834" longLatPrecision="1" longitude="1.67185" municipality="Savalou District" name="Tchetti" pageId="23" pageNumber="24" stateProvince="Zou">Tchetti</location>
, located in the
<collectingRegion id="B07278CE4522FFABFC2918B9FBA22924" box="[1023,1068,1015,1041]" country="Benin" name="Zou" pageId="23" pageNumber="24">Zou</collectingRegion>
catchment area (tributary of the
<collectingRegion id="B07278CE4522FFABFF411F55FF662900" box="[151,232,1051,1077]" country="Benin" name="Oueme" pageId="23" pageNumber="24">Ouémé</collectingRegion>
catchment area), coordinates:
<geoCoordinate id="1782D0EB4522FFABFD9A1F55FD532900" box="[588,733,1050,1077]" degrees="7" direction="north" minutes="48.965" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="7.8160834">7° 48.965’N</geoCoordinate>
,
<geoCoordinate id="1782D0EB4522FFABFD3D1F55FCF92900" box="[747,887,1051,1077]" degrees="1" direction="east" minutes="40.311" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="1.67185">1° 40.311’E</geoCoordinate>
,
<collectorName id="DF43D3FA4522FFABFC501F55FC332900" box="[902,957,1051,1077]" pageId="23" pageNumber="24">Leg.</collectorName>
:
<collectorName id="DF43D3FA4522FFABFC1D1F55FBEF2900" box="[971,1121,1051,1077]" pageId="23" pageNumber="24">S.J. Hotèkpo</collectorName>
,
<collectorName id="DF43D3FA4522FFABFBB91F55FB782900" box="[1135,1270,1051,1077]" pageId="23" pageNumber="24">M. Lagnika</collectorName>
and
<collectorName id="DF43D3FA4522FFABFAE41F55FA122900" box="[1330,1436,1051,1077]" pageId="23" pageNumber="24">P. Martin</collectorName>
collected on
<date id="060890EC4522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">
<collectingDate id="164C69044522FFABFEF01F71FE56296C" box="[294,472,1087,1113]" pageId="23" pageNumber="24" value="2016-08-09">09 August 2016</collectingDate>
</date>
(Locality code BEN072 in
<tableCitation id="3F3483974522FFABFCDB1F71FCE5296C" box="[781,875,1087,1113]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="23" pageNumber="24">Tables 1</tableCitation>
and
<tableCitation id="3F3483974522FFABFC771F71FC3C296C" box="[929,946,1087,1113]" captionStart="TABLE 2" captionStartId="6.[152,244,1634,1658]" captionText="TABLE 2. Measurements of physical and chemical water parameters and variables at the time of well fauna collection. WT= water temperature (°C), EC = electrical conductivity (µS cm-1), DO = dissolved oxygen (mg L-1). NM= no measurement." pageId="23" pageNumber="24">2</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74522FFA1FF411FC8FD9E2900" lastPageId="29" lastPageNumber="30" pageId="23" pageNumber="24" type="description">
<paragraph id="7209B62C4522FFABFF411FC8FE0F2994" blockId="23.[151,1437,1158,2013]" box="[151,385,1158,1185]" pageId="23" pageNumber="24">
<emphasis id="40C26A3E4522FFABFF411FC8FE0F2994" bold="true" box="[151,385,1158,1185]" pageId="23" pageNumber="24">Description of male</emphasis>
</paragraph>
<paragraph id="7209B62C4522FFABFF411FE5FC9F28A8" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
CpRl (
<figureCitation id="EA8DAAA94522FFABFF331FE5FEE229F0" box="[229,364,1195,1221]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="23" pageNumber="24">Figure 10A</figureCitation>
) elongated (L/H ratio = 2.4) and sub-triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height slightly in front of the middle, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; LV slightly overlapping RV along anterior, ventral and posterior sides, dorsally strongly overlapping RV; valves externally smooth. CpV (
<figureCitation id="EA8DAAA94522FFABFCF11E75FC272860" box="[807,937,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="23" pageNumber="24">Figure 10B</figureCitation>
) and CpD (
<figureCitation id="EA8DAAA94522FFABFBF81E75FB3F2860" box="[1070,1201,1339,1365]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="23" pageNumber="24">Figure 10C</figureCitation>
) narrow and pointed on both anterior and posterior sides, greatest width in the middle, lateral margins angular in the middle; CpV with LV forming a pronounced overlap over RV in the centre.
</paragraph>
<paragraph id="7209B62C4522FFABFF111EE9FBEA2B18" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
LVi (
<figureCitation id="EA8DAAA94522FFABFF291EE9FE0A28F4" box="[255,388,1447,1473]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="23" pageNumber="24">Figure 10D</figureCitation>
) elongated and triangular, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin bluntly pointed, with greatest height in the middle showing an overlap, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight, with a slight inward curl in the middle; one ventral il and antero-ventral tooth.
</paragraph>
<paragraph id="7209B62C4522FFABFF111D79FE062B88" blockId="23.[151,1437,1158,2013]" pageId="23" pageNumber="24">
RVi (
<figureCitation id="EA8DAAA94522FFABFED71D79FE0C2B64" box="[257,386,1591,1617]" captionStart="FIGURE 10" captionStartId="21.[151,250,1855,1879]" captionTargetBox="[151,1436,883,1832]" captionTargetId="figure-159@21.[151,1436,883,1832]" captionTargetPageId="21" captionText="FIGURE 10. Benincandona ibikounlei gen. et spec. nov., valves. A. INV.159222. CpRl. B. INV.159222. CpV. C. INV.159223. CpD. D. INV.159224. LVi. E. INV.159224. RVi. Scale bar = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.13745738" httpUri="https://zenodo.org/record/13745738/files/figure.png" pageId="23" pageNumber="24">Figure 10E</figureCitation>
) elongated, with anterior margin slightly more broadly rounded than posterior margin, dorsal margin greatest height situated about 1/3 from the rear, anterior part sloping almost straight to anterior margin, posterior part sloping almost straight to posterior margin, ventral margin almost straight; il running submarginally along ventral margin.
</paragraph>
<paragraph id="7209B62C4522FFA4FF111D89FE0D2C74" blockId="23.[151,1437,1158,2013]" lastBlockId="24.[151,1437,151,1905]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
A1 (
<figureCitation id="EA8DAAA94522FFABFF2E1D89FEF62BD4" box="[248,376,1735,1761]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745742" httpUri="https://zenodo.org/record/13745742/files/figure.png" pageId="23" pageNumber="24">Figure 11B</figureCitation>
) consisting of seven articulated segments, one basal and six distal. Basal segment large with
<emphasis id="40C26A3E4522FFABFA5F1D89FA122BD5" box="[1417,1436,1735,1760]" italics="true" pageId="23" pageNumber="24">c.</emphasis>
2.5 times as long as basal width, supporting two ventro-subapical setae, more proximal about three times as long as second one; and two setae on the dorsal side situated some distance from each other with the more proximal one slightly longer than the second one. First distal segment length about twice the basal width, with one short dorso-apical seta, no ventral apical seta present. Next (second distal segment) about twice as long as wide, with one dorso-apical seta. Third distal segment slightly longer than the previous one, with one short apical seta on each side. Fourth distal segment also slightly longer than the previous one (more than three time as long as wide) with one short ventro-apical seta and one long dorso-apical seta, reaching beyond tip of sixth distal segment with almost half the total length. Fifth distal segment more than four times as long as wide, with two long dorso-apical setae (almost five times longer than the length of the segment) and one short ventro-apical seta slightly exceeding the middle of the next segment. Sixth distal (terminal) segment, narrow,
<emphasis id="40C26A3E452DFFA4FCFC1B91FCB32DCD" box="[810,829,223,248]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as wide, bearing one short and two longer setae and one aesthetasc (Ya). Both long setae about 2.5x the length of Ya, length of the short one is about half the length of Ya.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111A05FBED2E68" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
A2 (
<figureCitation id="EA8DAAA9452DFFA4FF2E1A05FEF52C50" box="[248,379,331,357]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745742" httpUri="https://zenodo.org/record/13745742/files/figure.png" pageId="24" pageNumber="25">Figure 11A</figureCitation>
) with a basal segment broad and rounded, mid-ventrally with one seta of medium length, two subapical setae of unequal length, proximal one
<emphasis id="40C26A3E452DFFA4FD671A21FD4A2CBD" box="[689,708,367,392]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice the length of the more apical one. Second segment (second protopodite) large and
<emphasis id="40C26A3E452DFFA4FE4D1ADDFE202C99" box="[411,430,403,428]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as basal width, set with one short ventro-subapical seta X. No remnant of exopodite visible. Endopodite consisting of four segments with the penultimate segment divided. First endopodal segment long
<emphasis id="40C26A3E452DFFA4FEEF1A95FEC22CC1" box="[313,332,475,500]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width, carrying one long mid-ventral aesthetasc (Y) (reaching tip of segment) and two ventro-apical setae, one short and one long, hirsute in its distal 4/5. Second endopodal segment (segment 2A)
<emphasis id="40C26A3E452DFFA4FEEF196DFEC32F09" box="[313,333,547,572]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
twice as long as wide, carrying one stout dorso-apical seta and three t-setae; seta t1 long, narrow and plumose, accompanied by a long and slender, but well-developed aesthetasc y1; setae t2 (ventro-apical) and t3 (centro-apical) hyper-developed into male bristles. Third endopodal segment (segment 2B) smaller than previous segment and almost as long as wide, set with a complex (and sexually dimorphic) apical chaetotaxy (
<figureCitation id="EA8DAAA9452DFFA4FACA19C1FA1E2F9C" box="[1308,1424,655,681]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="24" pageNumber="25">Figure 40</figureCitation>
): ventro-apically with one short and slender aesthetasc y2 and the claw G3 transformed into short seta of almost equal length; centrally with one short seta z3; dorso-apically, with z1 claw-like, length twice as long as penultimate segment. Claw G2 about 2.5 times as long as z1; z2 seta transformed into long claw, slightly shorter than G1. Fourth endopodal, terminal, segment with one long claw (Gm), one short claw (GM), a long aesthetasc y3 with shorter accompanying seta, with fused part short, and seta g slightly longer than aesthetasc y3.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111829FC302EFC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdCoxa (
<figureCitation id="EA8DAAA9452DFFA4FEEE1829FE522EB4" box="[312,476,871,897]" captionStart="FIGURE 11" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[183,1405,185,1915]" captionTargetId="figure-31@22.[179,1407,181,1919]" captionTargetPageId="22" captionText="FIGURE 11. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. A2 (INV.159219, INV.159217, INV.159218). B. A1 (INV.159219, INV.159218). C, D. MdCoxa, (INV.159218). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745742" httpUri="https://zenodo.org/record/13745742/files/figure.png" pageId="24" pageNumber="25">Figure 11C–D</figureCitation>
) elongated, proximally sharply pointed, medially widened, distally with an obliquely widened end, set with a series (6–7) of strong teeth, interspaced with setae; subapically with a short seta. Remark: asymmetry illustrated possibly resulting from different position in slide.
</paragraph>
<paragraph id="7209B62C452DFFA4FF11189DFAEE29DC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
MdPalp (
<figureCitation id="EA8DAAA9452DFFA4FEE7189DFE3B2ED8" box="[305,437,979,1005]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745746" httpUri="https://zenodo.org/record/13745746/files/figure.png" pageId="24" pageNumber="25">Figure 12C</figureCitation>
) consisting of four segments. Basal segment ventro-apically with two hirsute setae, S1 (long, reaching tip of terminal segment) and S2 (short and stout, length about one third of the length of S1), in between with the short flagellum-like alpha seta (thin and about as long as S2), and one smooth and medium length, more proximal seta. Second segment of sub-triangular shape, dorso-apically with two setae of unequal length, ventro-apically with a group of four long setae, one of these being the beta-seta. Third segment with two unequal dorso-subapical setae, two mid-dorsal setae, one of which being the gamma seta and ventro-apically with 2 setae of sub-equal length. Terminal segment sub-rectangular, apically set with one large claw, fused with the segment over its entire distal margin, accompanied by one smooth lateral seta on each side, about as long as the fused claw.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111FBDFC2028D0" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
Mx1 (
<figureCitation id="EA8DAAA9452DFFA4FED91FBDFE322838" box="[271,444,1267,1293]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745746" httpUri="https://zenodo.org/record/13745746/files/figure.png" pageId="24" pageNumber="25">Figure 12A, B</figureCitation>
) with a basal (protopodite) part carrying a large respiratory plate (exopodite), three endites and a two-segmented palp (endopodite). Respiratory plate (
<figureCitation id="EA8DAAA9452DFFA4FC5D1E59FB802804" box="[907,1038,1303,1329]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745746" httpUri="https://zenodo.org/record/13745746/files/figure.png" pageId="24" pageNumber="25">Figure 12B</figureCitation>
) elongated, carrying
<emphasis id="40C26A3E452DFFA4FB281E59FA9C2805" box="[1278,1298,1303,1328]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
13 plumose respiratory rays, two short distal setae and a group of four shorter proximal setae of unequal length. Palp (
<figureCitation id="EA8DAAA9452DFFA4FAE31E75FA1A2860" box="[1333,1428,1339,1365]" captionStart="FIGURE 12" captionStartId="25.[151,250,1981,2005]" captionTargetBox="[166,1422,183,1955]" captionTargetId="figure-17@25.[166,1422,181,1957]" captionTargetPageId="25" captionText="FIGURE 12. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Mx1 (Chaetotaxy incomplete) (INV.159218). B. Mx1, respiratory plate (INV.159218). C. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745746" httpUri="https://zenodo.org/record/13745746/files/figure.png" pageId="24" pageNumber="25">Fig 12A</figureCitation>
) with first segment carrying four long dorso-apical setae; terminal segment short and rectangular, carrying also 4 distal setae, some slightly claw-like; chaetotaxy of three endites incompletely illustrated: third endite with two strong teeth carrying lateral spines (
<emphasis id="40C26A3E452DFFA4FDF11EE9FD3028F4" box="[551,702,1447,1473]" italics="true" pageId="24" pageNumber="25">Zahnborsten)</emphasis>
, basally with one lateral seta; sideways directed bristles near base of first endite one long and one short
<emphasis id="40C26A3E452DFFA4FDED1E85FDC028D1" box="[571,590,1483,1508]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
half the length of the long one.
</paragraph>
<paragraph id="7209B62C452DFFA4FF111EA1FF582BAC" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T1 (
<figureCitation id="EA8DAAA9452DFFA4FF251EA1FE1A2B3C" box="[243,404,1518,1545]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) basal part (protopodite) carrying respiratory plates (exopodites, not shown), distally set with 4+7 setae of different morphology and length; more proximally with setae d and b of medium length. Prehensile palps (
<figureCitation id="EA8DAAA9452DFFA4FF351D79FE052B64" box="[227,395,1590,1617]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="24" pageNumber="25">Figure 15A, B</figureCitation>
) one-segmented, distally sigmoid hook-like, both palps set with a short sensory organ and proximally with two subequal setae (also sensory organs); Lpp (
<figureCitation id="EA8DAAA9452DFFA4FCA11D15FC722B40" box="[887,1020,1626,1653]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="24" pageNumber="25">Figure 15A</figureCitation>
) slightly narrower than Rpp (
<figureCitation id="EA8DAAA9452DFFA4FA871D15FF472BAC" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="24" pageNumber="25">Figure 15B</figureCitation>
).
</paragraph>
<paragraph id="7209B62C452DFFA4FF111DEDFB5C2A44" blockId="24.[151,1437,151,1905]" pageId="24" pageNumber="25">
T2 (
<figureCitation id="EA8DAAA9452DFFA4FF231DEDFEF72B88" box="[245,377,1699,1725]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745748" httpUri="https://zenodo.org/record/13745748/files/figure.png" pageId="24" pageNumber="25">Figure 13D</figureCitation>
) with 4-segmented endopodite (penultimate segment divided), elongated, without basal setae. Setae d1 and d2 absent. First endopodal segment without any setae, length
<emphasis id="40C26A3E452DFFA4FC2B1D89FB9E2BD5" box="[1021,1040,1735,1760]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
five times as long as basal width. Second endopodal segment (2A) set with one short ventro-apical seta f, length
<emphasis id="40C26A3E452DFFA4FBC21DA5FBA92A31" box="[1044,1063,1771,1796]" italics="true" pageId="24" pageNumber="25">c.</emphasis>
3x basal width. Third endopodal segment (2B) of about the same length/width ratio as the previous segment and with one ventro-subapical seta g, reaching about 1/3 of following segment. Terminal segment about 1.5 times as long as basal width, apically two short setae, h1 ventral and h3 dorsal; and a long slender claw (h2), distally set with short setulae.
</paragraph>
<caption id="26C9E6A4452CFFA5FF411CF3FBFF2ACF" ID-DOI="http://doi.org/10.5281/zenodo.13745746" ID-Zenodo-Dep="13745746" httpUri="https://zenodo.org/record/13745746/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1981,2005]" targetBox="[166,1422,183,1955]" targetPageId="25" targetType="figure">
<paragraph id="7209B62C452CFFA5FF411CF3FBFF2ACF" blockId="25.[151,1436,1981,2042]" pageId="25" pageNumber="26">
<emphasis id="40C26A3E452CFFA5FF411CF3FEAE2AE0" bold="true" box="[151,288,1981,2005]" pageId="25" pageNumber="26">FIGURE 12.</emphasis>
<taxonomicName id="B5B6CDAF452CFFA5FEF11CF3FDAA2AE0" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[295,548,1981,2005]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452CFFA5FEF11CF3FDAA2AE0" box="[295,548,1981,2005]" italics="true" pageId="25" pageNumber="26">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452CFFA5FDFD1CF0FD532AE0" box="[555,733,1982,2006]" pageId="25" pageNumber="26" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452CFFA5FC811CF3FCE72AE0" bold="true" box="[855,873,1981,2005]" pageId="25" pageNumber="26">A</emphasis>
. Mx1 (Chaetotaxy incomplete) (INV.159218).
<emphasis id="40C26A3E452CFFA5FA9E1CF0FAD72AE3" bold="true" box="[1352,1369,1982,2006]" pageId="25" pageNumber="26">B</emphasis>
. Mx1, respiratory plate (INV.159218).
<emphasis id="40C26A3E452CFFA5FE0A1CAFFE602ACC" bold="true" box="[476,494,2017,2041]" pageId="25" pageNumber="26">C</emphasis>
. MdPalp (INV.159218). Scale bar. 1 (B), 2 (C), 3 (A) = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452FFFA6FF411CD9FD432AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745748" ID-Zenodo-Dep="13745748" httpUri="https://zenodo.org/record/13745748/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1943,1967]" targetBox="[191,1395,185,1916]" targetPageId="26" targetType="figure">
<paragraph id="7209B62C452FFFA6FF411CD9FD432AE1" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="27">
<emphasis id="40C26A3E452FFFA6FF411CD9FEAC2A9A" bold="true" box="[151,290,1943,1967]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
<taxonomicName id="B5B6CDAF452FFFA6FEFD1CD6FDA22A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[299,556,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452FFFA6FEFD1CD6FDA22A85" box="[299,556,1944,1968]" italics="true" pageId="26" pageNumber="27">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452FFFA6FDE31CD6FD602A9A" box="[565,750,1944,1968]" pageId="26" pageNumber="27" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452FFFA6FCB91CD9FC0F2A9A" bold="true" box="[879,897,1943,1967]" pageId="26" pageNumber="27">A</emphasis>
. CR (INV.159219).
<emphasis id="40C26A3E452FFFA6FB811CD6FBE62A85" bold="true" box="[1111,1128,1944,1968]" pageId="26" pageNumber="27">B</emphasis>
. CRAtt (INV.159219).
<emphasis id="40C26A3E452FFFA6FA891CD9FAFF2A9A" bold="true" box="[1375,1393,1943,1967]" pageId="26" pageNumber="27">C</emphasis>
. T3 (INV.159219).
<emphasis id="40C26A3E452FFFA6FEE71CF2FECD2AE1" bold="true" box="[305,323,1980,2004]" pageId="26" pageNumber="27">D</emphasis>
. T2 (INV.159217). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A4452EFFA7FF411CD9FDB32ACD" ID-DOI="http://doi.org/10.5281/zenodo.13745750" ID-Zenodo-Dep="13745750" httpUri="https://zenodo.org/record/13745750/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1943,1967]" targetBox="[229,1359,191,1909]" targetPageId="27" targetType="figure">
<paragraph id="7209B62C452EFFA7FF411CD9FDB32ACD" blockId="27.[151,1437,1943,2040]" pageId="27" pageNumber="28">
<emphasis id="40C26A3E452EFFA7FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="27" pageNumber="28">FIGURE 14.</emphasis>
<taxonomicName id="B5B6CDAF452EFFA7FEFE1CD9FDA82A9A" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[296,550,1943,1967]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E452EFFA7FEFE1CD9FDA82A9A" box="[296,550,1943,1967]" italics="true" pageId="27" pageNumber="28">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D745452EFFA7FDFB1CD6FD6F2A9A" box="[557,737,1944,1968]" pageId="27" pageNumber="28" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (♂).
<emphasis id="40C26A3E452EFFA7FC8A1CD9FCE02A9A" bold="true" box="[860,878,1943,1967]" pageId="27" pageNumber="28">A</emphasis>
. Zenker organ (INV.159219).
<emphasis id="40C26A3E452EFFA7FB701CD9FB5B2A9A" bold="true" box="[1190,1237,1943,1967]" pageId="27" pageNumber="28">B–D</emphasis>
. Hp (INV.159219).
<emphasis id="40C26A3E452EFFA7FF411CF5FF262AE6" bold="true" box="[151,168,1979,2003]" pageId="27" pageNumber="28">B</emphasis>
. External view (drawing).
<emphasis id="40C26A3E452EFFA7FE141CF5FE5A2AE6" bold="true" box="[450,468,1979,2003]" pageId="27" pageNumber="28">C</emphasis>
. Internal view (photograph without interference contrast).
<emphasis id="40C26A3E452EFFA7FBEE1CF5FBC42AE6" bold="true" box="[1080,1098,1979,2003]" pageId="27" pageNumber="28">D</emphasis>
. Internal view (photograph with interference contrast). Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="26C9E6A44529FFA0FF411CD9FB072AE1" ID-DOI="http://doi.org/10.5281/zenodo.13745752" ID-Zenodo-Dep="13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1943,1967]" targetBox="[176,1407,181,1915]" targetPageId="28" targetType="figure">
<paragraph id="7209B62C4529FFA0FF411CD9FB072AE1" blockId="28.[151,1436,1943,2004]" pageId="28" pageNumber="29">
<emphasis id="40C26A3E4529FFA0FF411CD9FEAF2A9A" bold="true" box="[151,289,1943,1967]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
<taxonomicName id="B5B6CDAF4529FFA0FEFF1CD6FDA62A85" authorityName="Hotèkpo &amp; Schön &amp; Martens" authorityYear="2024" box="[297,552,1944,1968]" class="Ostracoda" family="Candonidae" genus="Benincandona" kingdom="Animalia" order="Podocopida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ibikounlei" status="gen. et spec. nov.">
<emphasis id="40C26A3E4529FFA0FEFF1CD6FDA62A85" box="[297,552,1944,1968]" italics="true" pageId="28" pageNumber="29">Benincandona ibikounlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BF1D7454529FFA0FDE61CD6FD6B2A9A" box="[560,741,1944,1968]" pageId="28" pageNumber="29" rank="species">gen. et spec. nov.</taxonomicNameLabel>
, limbs (A–B ♂ and C–E ♀).
<emphasis id="40C26A3E4529FFA0FBC21CD9FBA82A9A" bold="true" box="[1044,1062,1943,1967]" pageId="28" pageNumber="29">A</emphasis>
. T1, Lpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FAFC1CD6FAB52A85" bold="true" box="[1322,1339,1944,1968]" pageId="28" pageNumber="29">B</emphasis>
. T1, Rpp (INV.159219).
<emphasis id="40C26A3E4529FFA0FEE71CF5FECD2AE6" bold="true" box="[305,323,1979,2003]" pageId="28" pageNumber="29">C</emphasis>
. A2 (INV.159216).
<emphasis id="40C26A3E4529FFA0FDDB1CF2FD912AE1" bold="true" box="[525,543,1980,2004]" pageId="28" pageNumber="29">D</emphasis>
. CR (INV.159216).
<emphasis id="40C26A3E4529FFA0FD381CF2FD712AE1" bold="true" box="[750,767,1980,2004]" pageId="28" pageNumber="29">E</emphasis>
. T1 (INV.159221). Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="7209B62C4528FFA1FF111BD9FE902C50" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
T3 (
<figureCitation id="EA8DAAA94528FFA1FF231BD9FEF82D84" box="[245,374,151,177]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745748" httpUri="https://zenodo.org/record/13745748/files/figure.png" pageId="29" pageNumber="30">Figure 13C</figureCitation>
) first segment with only one medium length seta d1. Second (knee-) segment without any seta. Third segment about 5x as long as wide, not carrying any seta. Fourth segment narrower,
<emphasis id="40C26A3E4528FFA1FB701BF5FB372DE1" box="[1190,1209,187,212]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
7x basal width and slightly longer than the previous segment, carrying one short subapical seta g. Terminal segment well-separated from penultimate segment, L
<emphasis id="40C26A3E4528FFA1FE241A4DFD8B2C29" box="[498,517,259,284]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
1.5x basal width and carrying three setae: one long and reflexed (h3), clearly exceeding half of the third segment and two ventrally directed short setae h1 and h2, the former
<emphasis id="40C26A3E4528FFA1FB0B1A69FB7E2C75" box="[1245,1264,295,320]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
half the length of the latter.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A21FB082C98" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
CR (
<figureCitation id="EA8DAAA94528FFA1FF2D1A21FEF32CBC" box="[251,381,367,393]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745748" httpUri="https://zenodo.org/record/13745748/files/figure.png" pageId="29" pageNumber="30">Figure 13A</figureCitation>
) with ramus stout; claw Ga well-developed, stout with length ratio ramus/claw Ga = 1.5, setae Sp (posterior seta), Sa (anterior seta) and claw Gp (posterior claw) reduced to small setae.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111AF9FABB2CE4" blockId="29.[151,1437,151,645]" box="[199,1333,439,465]" pageId="29" pageNumber="30">
CRAtt (
<figureCitation id="EA8DAAA94528FFA1FEC91AF9FE2F2CE4" box="[287,417,439,465]" captionStart="FIGURE 13" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[191,1395,185,1916]" captionTargetId="figure-31@26.[187,1399,181,1919]" captionTargetPageId="26" captionText="FIGURE 13. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. CR (INV.159219). B. CRAtt (INV.159219). C. T3 (INV.159219). D. T2 (INV.159217). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745748" httpUri="https://zenodo.org/record/13745748/files/figure.png" pageId="29" pageNumber="30">Figure 13B</figureCitation>
) long and stout, distally bifurcated and subapically with additional lateral branch.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111A95FD772F08" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Hp (
<figureCitation id="EA8DAAA94528FFA1FF2F1A95FE102CC0" box="[249,414,475,501]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). B–D. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745750" httpUri="https://zenodo.org/record/13745750/files/figure.png" pageId="29" pageNumber="30">Figure 14B–D</figureCitation>
) of medium size, with (ls) relatively narrow, with skewed rectangular distal part, reaching well beyond the asymmetrically rounded ms; lobe ‘h’ shorter and sub-rectangular; internal anatomy (
<figureCitation id="EA8DAAA94528FFA1FB241AB1FA1A2F2C" box="[1266,1428,511,537]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). B–D. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745750" httpUri="https://zenodo.org/record/13745750/files/figure.png" pageId="29" pageNumber="30">Figure 14C, D</figureCitation>
) with postlabyrinthal spermiduct with one circular coil.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111909FCDD2FB0" blockId="29.[151,1437,151,645]" pageId="29" pageNumber="30">
Zenker organ (
<figureCitation id="EA8DAAA94528FFA1FEA61909FE782F54" box="[368,502,583,609]" captionStart="FIGURE 14" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[229,1359,191,1909]" captionTargetId="figure-17@27.[219,1369,181,1919]" captionTargetPageId="27" captionText="FIGURE 14. Benincandona ibikounlei gen. et spec. nov., limbs (♂). A. Zenker organ (INV.159219). B–D. Hp (INV.159219). B. External view (drawing). C. Internal view (photograph without interference contrast). D. Internal view (photograph with interference contrast). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745750" httpUri="https://zenodo.org/record/13745750/files/figure.png" pageId="29" pageNumber="30">Figure 14A</figureCitation>
) with
<emphasis id="40C26A3E4528FFA1FD951909FDD92F55" box="[579,599,583,608]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
seven spinous whirls, relatively long and with an elongated central tube joining posterior funnel-shaped end plate to anterior end plate.
</paragraph>
<paragraph id="7209B62C4528FFA1FF4119FCFE182FF9" blockId="29.[151,1436,690,1185]" box="[151,406,690,716]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF4119FCFE182FF9" bold="true" box="[151,406,690,716]" pageId="29" pageNumber="30">Description of female</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF411999FB6F2FC4" blockId="29.[151,1436,690,1185]" box="[151,1249,727,753]" pageId="29" pageNumber="30">Cp and valves most likely as in the male, without obvious sexual dimorphism (see remark above).</paragraph>
<paragraph id="7209B62C4528FFA1FF1119B5FB072E20" blockId="29.[151,1436,690,1185]" box="[199,1161,762,789]" pageId="29" pageNumber="30">
Limbs (
<figureCitation id="EA8DAAA94528FFA1FECB19B5FE302E20" box="[285,446,762,789]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="29" pageNumber="30">Figure 15C–E</figureCitation>
) largely as in the male, but with sexually dimorphic A2 and T1.
</paragraph>
<paragraph id="7209B62C4528FFA1FF111851FC232E90" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
A2 (
<figureCitation id="EA8DAAA94528FFA1FF2F1851FEF22E0C" box="[249,380,798,825]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="29" pageNumber="30">Figure 15C</figureCitation>
) with second endopodal segment undivided, with two t-setae, t1 (long, narrow and plumose) and t2 (thin and smooth); apical chaetotaxy (
<figureCitation id="EA8DAAA94528FFA1FD47180DFC8F2E68" box="[657,769,835,861]" captionStart="FIGURE 40" captionStartId="60.[151,250,1516,1540]" captionTargetBox="[151,1435,181,1492]" captionTargetId="figure-31@60.[151,1435,181,1492]" captionTargetPageId="60" captionText="FIGURE 40. Diagrams of apical views of the A2 of Benincandona, showing position of (sub-) terminal claws, setae and aesthetascs of the terminal (small circle) and penultimate segment (large circle). For explanations of codes, see Material and Methods (Abbreviations used in text and figures)." figureDoi="http://doi.org/10.5281/zenodo.13745804" httpUri="https://zenodo.org/record/13745804/files/figure.png" pageId="29" pageNumber="30">Figure 40</figureCitation>
) with z1 claw-like, z2 and z3 being both setae, z1 twice as long as z2 and z3; claws G1, G2, G3 and GM all reaching to about the same point; seta g and aesthetasc y3 as in the male. Small claw Gm in female homologous to large claw in the male.
</paragraph>
<paragraph id="7209B62C4528FFA1FF1118E1FCFA2924" blockId="29.[151,1436,690,1185]" pageId="29" pageNumber="30">
T1 (
<figureCitation id="EA8DAAA94528FFA1FF2E18E1FEF52EFC" box="[248,379,942,969]" captionStart="FIGURE 15" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[176,1407,181,1915]" captionTargetId="figure-31@28.[176,1411,181,1919]" captionTargetPageId="28" captionText="FIGURE 15. Benincandona ibikounlei gen. et spec. nov., limbs (A–B ♂ and C–E ♀). A. T1, Lpp (INV.159219). B. T1, Rpp (INV.159219). C. A2 (INV.159216). D. CR (INV.159216). E. T1 (INV.159221). Scale bar = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.13745752" httpUri="https://zenodo.org/record/13745752/files/figure.png" pageId="29" pageNumber="30">Figure 15E</figureCitation>
) with basal part with
<emphasis id="40C26A3E4528FFA1FDA918E1FD1C2EFD" box="[639,658,943,968]" italics="true" pageId="29" pageNumber="30">c.</emphasis>
10 distal setae of unequal length, setae b and d present, with long additional seta (d’) present close to seta d; respiratory plate hidden on the dissected specimens. Endopodite a broad and short palp, with three subequal but short apical setae (h1-h3).
</paragraph>
<paragraph id="7209B62C4528FFA1FF111F55FD9E2900" blockId="29.[151,1436,690,1185]" box="[199,528,1051,1077]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F55FE0E2900" bold="true" box="[199,384,1051,1077]" pageId="29" pageNumber="30">Measurements:</emphasis>
See
<tableCitation id="3F3483974528FFA1FE601F55FD842900" box="[438,522,1051,1077]" captionStart="TABLE 3" captionStartId="19.[152,244,1592,1616]" captionText="TABLE 3. Measurements of illustrated specimens of Benincandona gen. et spec. nov., using SEM." pageId="29" pageNumber="30">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F70FDEA296C" box="[199,612,1086,1113]" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7209B62C4528FFA1FF111F70FDEA296C" blockId="29.[151,1436,690,1185]" box="[199,612,1086,1113]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F70FE5B296D" bold="true" box="[199,469,1086,1112]" pageId="29" pageNumber="30">Differential Diagnosis:</emphasis>
See
<tableCitation id="3F3483974528FFA1FDDC1F71FDEE296C" box="[522,608,1087,1113]" captionStart="TABLE 4" captionStartId="20.[152,245,1665,1689]" captionText="TABLE 4. Differential character states between the five new species of Benincandona gen. nov." pageId="29" pageNumber="30">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AACE5A74528FFA1FF111F2DFB072994" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="7209B62C4528FFA1FF111F2DFE612949" blockId="29.[151,1436,690,1185]" box="[199,495,1122,1149]" pageId="29" pageNumber="30">
<emphasis id="40C26A3E4528FFA1FF111F2DFE612949" bold="true" box="[199,495,1122,1149]" pageId="29" pageNumber="30">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="7209B62C4528FFA1FF111FC9FB072994" blockId="29.[151,1436,690,1185]" box="[199,1161,1159,1185]" pageId="29" pageNumber="30">
This subterranean species is presently known from its
<typeStatus id="AD0D088E4528FFA1FCF11FC9FCD92994" box="[807,855,1159,1185]" pageId="29" pageNumber="30">type</typeStatus>
locality only (see
<tableCitation id="3F3483974528FFA1FBF01FC9FBF62994" box="[1062,1144,1159,1185]" captionStart="TABLE 1" captionStartId="5.[152,244,929,955]" captionText="TABLE 1. Localities where Benincandona gen. nov. sp. nov. were collected, coordinates and hydro-morphological characteristics. In all stations, the water was clear and was used for both drinking water and household purposes by the riparian villages." pageId="29" pageNumber="30">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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