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<mods:title id="A318E84622BD5FE14459A5C905ECC03A">A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan</mods:title>
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<taxonomicName id="B5BD4D5BFF9E5F36D3B9572F29C40EC1" authority="(Seidlitz, 1895)" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" box="[151,737,366,393]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus" subGenus="Pelorinus">
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<emphasis id="40C9EACAFF9E5F36D3B9572F2A240EC0" bold="true" box="[151,257,366,393]" italics="true" pageId="2" pageNumber="453">Euboeus</emphasis>
(
<emphasis id="40C9EACAFF9E5F36D23C572F2AA20EC0" bold="true" box="[274,391,366,393]" italics="true" pageId="2" pageNumber="453">Pelorinus</emphasis>
)
<emphasis id="40C9EACAFF9E5F36D2B7572E29050EC0" bold="true" box="[409,544,367,393]" italics="true" pageId="2" pageNumber="453">corrugosus</emphasis>
(
<bibRefCitation id="162C4B29FF9E5F36D101572F29FC0EC1" author="Seidlitz, G. C. M. von" box="[559,729,366,392]" pageId="2" pageNumber="453" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895</bibRefCitation>
)
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<paragraph id="720236D8FF9E5F36D3B957D22A290EE4" blockId="2.[151,737,366,429]" box="[151,268,403,429]" pageId="2" pageNumber="453">
(
<figureCitation id="EA862A5DFF9E5F36D3B157D22BC30EE4" box="[159,230,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1845,1869]" captionTargetBox="[151,1436,1049,1822]" captionTargetId="figure-324@2.[151,1436,1049,1822]" captionTargetPageId="2" captionText="FIGURE 1. Euboeus corrugosus, lectotype, male, habitus.A = dorsally; B = laterally; C = labels." figureDoi="http://doi.org/10.5281/zenodo.6785913" httpUri="https://zenodo.org/record/6785913/files/figure.png" pageId="2" pageNumber="453">Figs 1</figureCitation>
,
<figureCitation id="EA862A5DFF9E5F36D3DA57D22A260EE4" box="[244,259,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[229,1350,530,1873]" captionTargetId="figure-156@3.[226,1361,525,1873]" captionTargetPageId="3" captionText="FIGURE 2. Euboeus corrugosus, types of E. parthorum, syn. nov., habitus, details of structure. A = male, holotype dorsally; B = label of holotype; C = male, paratype, ventrally; D = female, paratype, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = tegmen, ventrally; K = median lobe of aedeagus, ventrally; L = apical piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.6785917" httpUri="https://zenodo.org/record/6785917/files/figure.png" pageId="2" pageNumber="453">2</figureCitation>
)
</paragraph>
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<treatmentCitation id="F31C10C9FF9E5F36D3B9579629050EA6" author="Seidlitz, G. C. M. von" box="[151,544,471,495]" page="707" pageId="2" pageNumber="453" year="1895">
<taxonomicName id="B5BD4D5BFF9E5F36D3B9579629050EA6" authority="Seidlitz, 1895: 707" authorityName="Seidlitz" authorityPageNumber="707" authorityYear="1895" box="[151,544,471,495]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9E5F36D3B957962A720EA6" box="[151,343,471,495]" italics="true" pageId="2" pageNumber="453">Helops corrugosus</emphasis>
<bibRefCitation id="162C4B29FF9E5F36D270579629050EA6" author="Seidlitz, G. C. M. von" box="[350,544,471,495]" pageId="2" pageNumber="453" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895: 707</bibRefCitation>
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, 750 (part);
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</paragraph>
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<treatmentCitation id="F31C10C9FF9E5F36D3B957B728340D47" author="Leo, P. &amp; Ruzzante, G." box="[151,785,502,526]" page="85" pageId="2" pageNumber="453" year="2016">
<taxonomicName id="B5BD4D5BFF9E5F36D3B957B728340D47" authority=": Leo &amp; Ruzzante, 2016: 85" authorityName="Leo &amp; Ruzzante" authorityPageNumber="85" authorityYear="2016" box="[151,785,502,526]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus" subGenus="Pelorinus">
<emphasis id="40C9EACAFF9E5F36D3B957B72A200D47" box="[151,261,502,526]" italics="true" pageId="2" pageNumber="453">Probaticus</emphasis>
(
<emphasis id="40C9EACAFF9E5F36D23A57B72A520D47" box="[276,375,502,526]" italics="true" pageId="2" pageNumber="453">Pelorinus</emphasis>
)
<emphasis id="40C9EACAFF9E5F36D2AB57B62AD20D47" box="[389,503,503,526]" italics="true" pageId="2" pageNumber="453">corrugosus</emphasis>
:
<bibRefCitation id="162C4B29FF9E5F36D12B57B728340D47" author="Leo, P. &amp; Ruzzante, G." box="[517,785,502,526]" pageId="2" pageNumber="453" pagination="83 - 89" refId="ref22267" refString="Leo, P. &amp; Ruzzante, G. (2016) Dati nuovi e riassuntivi sui Tenebrionidi dell'isola di Lesbo (Grecia) (Coleoptera Tenebrionidae). Bollettino della Societa Entomologica Italiana, 148, 83 - 89." type="journal article" year="2016">Leo &amp; Ruzzante, 2016: 85</bibRefCitation>
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(part);
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<paragraph id="720236D8FF9E5F36D3B9545428570D64" blockId="2.[151,882,471,557]" box="[151,882,532,557]" pageId="2" pageNumber="453">
<treatmentCitationGroup id="52AD11F6FF9E5F36D3B9545428570D64" box="[151,882,532,557]" pageId="2" pageNumber="453">
=
<treatmentCitation id="F31C10C9FF9E5F36D3825455282F0D64" author="Medvedev, G. S." box="[172,778,532,557]" page="891" pageId="2" pageNumber="453" year="1976">
<taxonomicName id="B5BD4D5BFF9E5F36D3825455282F0D64" authority="G. Medvedev, 1976: 891" authorityName="G. Medvedev" authorityPageNumber="891" authorityYear="1976" box="[172,778,532,557]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="parthorum" subGenus="Pelorinus">
<emphasis id="40C9EACAFF9E5F36D38254552A3F0D65" box="[172,282,532,556]" italics="true" pageId="2" pageNumber="453">Probaticus</emphasis>
(
<emphasis id="40C9EACAFF9E5F36D20754552AA90D65" box="[297,396,532,556]" italics="true" pageId="2" pageNumber="453">Pelorinus</emphasis>
)
<emphasis id="40C9EACAFF9E5F36D2B4545529230D65" box="[410,518,532,556]" italics="true" pageId="2" pageNumber="453">parthorum</emphasis>
G.
<bibRefCitation id="162C4B29FF9E5F36D1025455282F0D64" author="Medvedev, G. S." box="[556,778,532,557]" pageId="2" pageNumber="453" pagination="889 - 902" refId="ref22349" refString="Medvedev, G. S. (1976) New species of darkling beetles (Coleoptera, Tenebrionidae) from Iran. Entomologicheskoe Obozrenie, 55, 889 - 902." type="journal article" year="1976">Medvedev, 1976: 891</bibRefCitation>
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,
<emphasis id="40C9EACAFF9E5F36D03A545428570D64" bold="true" box="[788,882,533,557]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="5BFA57B1FF9E5F36D03A545428570D64" box="[788,882,533,557]" pageId="2" pageNumber="453">syn. nov.</taxonomicNameLabel>
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<emphasis id="40C9EACAFF9E5F36D3B9541A2A630D3D" bold="true" box="[151,326,602,629]" pageId="2" pageNumber="453">Type material.</emphasis>
<typeStatus id="AD06887AFF9E5F36D263541A2A980D3C" box="[333,445,603,629]" pageId="2" pageNumber="453" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="B5BD4D5BFF9E5F36D2CA541A28790D3C" authority="Seidlitz, 1895" authorityName="Seidlitz" authorityYear="1895" box="[484,860,602,629]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9E5F36D2CA541A299C0D3D" box="[484,697,603,629]" italics="true" pageId="2" pageNumber="453">Helops corrugosus</emphasis>
<bibRefCitation id="162C4B29FF9E5F36D1EE541A28790D3C" author="Seidlitz, G. C. M. von" box="[704,860,602,629]" pageId="2" pageNumber="453" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895</bibRefCitation>
</taxonomicName>
(
<collectionCode id="14ACAE1DFF9E5F36D042541A289C0D3C" box="[876,953,603,629]" pageId="2" pageNumber="453">SNSD</collectionCode>
), designated here according to Article 61.1 and 74.1 of ICZN (1999), with labels: bronze square, “
<emphasis id="40C9EACAFF9E5F36D026543E28AD0DD0" box="[776,904,639,665]" italics="true" pageId="2" pageNumber="453">subrugosus</emphasis>
tut Allard”, “
<specimenCount id="64BBFD51FF9E5F36D70D543E2F1C0DD0" box="[1059,1081,639,665]" pageId="2" pageNumber="453" type="male"></specimenCount>
<emphasis id="40C9EACAFF9E5F36D76F543E2F800DD0" box="[1089,1189,639,665]" italics="true" pageId="2" pageNumber="453">astrabad</emphasis>
Christoph”, “13916”, “
<taxonomicName id="B5BD4D5BFF9E5F36D38A54E22A980DF4" authority="Seidl." authorityName="Seidl." box="[164,445,675,701]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugatus">
<emphasis id="40C9EACAFF9E5F36D38A54E22A510DF5" box="[164,372,675,701]" italics="true" pageId="2" pageNumber="453">Helops corrugatus</emphasis>
Seidl.
</taxonomicName>
” [handwritten by G. Seidlitz], “
<taxonomicName id="B5BD4D5BFF9E5F36D03154E228F80DF4" authority="Seid." authorityName="Seidlitz" authorityYear="1895" box="[799,989,675,701]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9E5F36D03154E228B80DF5" box="[799,925,675,700]" italics="true" pageId="2" pageNumber="453">corrugosus</emphasis>
Seid.
</taxonomicName>
nec
<taxonomicName id="B5BD4D5BFF9E5F36D73E54E22FAE0DF5" box="[1040,1163,675,700]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugatus">
<emphasis id="40C9EACAFF9E5F36D73E54E22FAE0DF5" box="[1040,1163,675,700]" italics="true" pageId="2" pageNumber="453">corrugatus</emphasis>
</taxonomicName>
det.Schuster”. The label “
<taxonomicName id="B5BD4D5BFF9E5F36D38A5486293E0DA8" authority="Seidlitz, 1895" authorityName="Seidlitz" authorityYear="1895" box="[164,539,710,737]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9E5F36D38A54862A5D0DA9" box="[164,376,711,737]" italics="true" pageId="2" pageNumber="453">Helops corrugosus</emphasis>
<bibRefCitation id="162C4B29FF9E5F36D2515486293E0DA8" author="Seidlitz, G. C. M. von" box="[383,539,710,737]" pageId="2" pageNumber="453" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895</bibRefCitation>
</taxonomicName>
des. M. Nabozhenko 2021” was added to the
<typeStatus id="AD06887AFF9E5F36D73054862FAD0DA8" box="[1054,1160,711,737]" pageId="2" pageNumber="453" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="720236D8FF9E5F36D3E954AA2F860B48" blockId="2.[151,1437,602,1025]" pageId="2" pageNumber="453">
<materialsCitation id="C2D53C85FF9E5F36D3E954AA2AA40C04" ID-GBIF-Occurrence="3828382319" collectingDate="1912-05-09" collectingDateMax="1912-05-12" collectingDateMin="1912-05-09" collectorName="Bil'kevich" country="Iran" location="Iran" pageId="2" pageNumber="453" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="AD06887AFF9E5F36D3E954AA2A0A0C4C" box="[199,303,747,773]" pageId="2" pageNumber="453" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="B5BD4D5BFF9E5F36D27D54AA29B10C4C" authority="(ZIN)" baseAuthorityName="ZIN" box="[339,660,747,773]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="parthorum">
<emphasis id="40C9EACAFF9E5F36D27D54AA296F0C4C" box="[339,586,747,773]" italics="true" pageId="2" pageNumber="453">Probaticus parthorum</emphasis>
(ZIN)
</taxonomicName>
,
<specimenCount id="64BBFD51FF9E5F36D1B054AA29960C4C" box="[670,691,747,773]" pageId="2" pageNumber="453" type="male"></specimenCount>
with the Cyrillic Russian handwritten label “Астрабад, Сев. Иран
<date id="06031018FF9E5F36D3B9554E2A000C60" box="[151,293,783,809]" pageId="2" pageNumber="453" value="1912-05-09" valueMax="1912-05-12" valueMin="1912-05-09">
<collectingDate id="1647E9F0FF9E5F36D3B9554E2A000C60" box="[151,293,783,809]" pageId="2" pageNumber="453" value="1912-05-09" valueMax="1912-05-12" valueMin="1912-05-09">912.V.1912</collectingDate>
</date>
Билькевич” (Astrabad, North
<collectingCountry id="0AAA7648FF9E5F36D1A9554E29920C60" box="[647,695,783,809]" name="Iran" pageId="2" pageNumber="453">Iran</collectingCountry>
, leg.
<collectorName id="DF48530EFF9E5F36D1DB554E284A0C60" box="[757,879,783,809]" pageId="2" pageNumber="453">Bilkevich</collectorName>
), “
<typeStatus id="AD06887AFF9E5F36D0A3554E2F210C60" box="[909,1028,783,809]" pageId="2" pageNumber="453" type="holotype">Holotypus</typeStatus>
<emphasis id="40C9EACAFF9E5F36D723554E2FB00C60" box="[1037,1173,783,809]" italics="true" pageId="2" pageNumber="453">Probathicus</emphasis>
<taxonomicName id="B5BD4D5BFF9E5F36D7B3554E2A530C04" authority="G. Medvedev, 1977" authorityName="G. Medvedev" authorityYear="1977" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="parthorum" subGenus="Pelorinus">
(
<emphasis id="40C9EACAFF9E5F36D788554E2E360C60" box="[1190,1299,783,809]" italics="true" pageId="2" pageNumber="453">Pelorinus</emphasis>
)
<emphasis id="40C9EACAFF9E5F36D60A554E2EB90C60" box="[1316,1436,783,809]" italics="true" pageId="2" pageNumber="453">parthorum</emphasis>
G. Medvedev, 1977
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="C2D53C85FF9E5F36D2A4557229260CDC" ID-GBIF-Occurrence="3828382333" collectorName="Philippovich" country="Iran" location="Cyrillic" pageId="2" pageNumber="453" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="AD06887AFF9E5F36D2A455722ADE0C04" box="[394,507,819,845]" pageId="2" pageNumber="453" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="64BBFD51FF9E5F36D12A557229020C04" box="[516,551,819,845]" pageId="2" pageNumber="453" type="male">1♂</specimenCount>
with labels: “Астрабад С. Персия [print] 6.III.904 [handwritten] Филиппович [print italic]” (
<location id="77626003FF9E5F36D26E55162ABF0C38" LSID="urn:lsid:plazi:treatment:FA1487CEFF9E5F37D3B9572F29A10EBC:77626003FF9E5F36D26E55162ABF0C38" box="[320,410,855,881]" country="Iran" name="Cyrillic" pageId="2" pageNumber="453">Cyrillic</location>
,
<collectorName id="DF48530EFF9E5F36D286551629200C38" box="[424,517,855,881]" pageId="2" pageNumber="453">Russian</collectorName>
:
<collectorName id="DF48530EFF9E5F36D13F551629B90C38" box="[529,668,855,881]" pageId="2" pageNumber="453">Astrabad, N</collectorName>
<collectingCountry id="0AAA7648FF9E5F36D188551629C80C38" box="[678,749,855,881]" name="Iran" pageId="2" pageNumber="453">Persia</collectingCountry>
, leg.
<collectorName id="DF48530EFF9E5F36D01E551628E40C38" box="[816,961,855,881]" pageId="2" pageNumber="453">Philippovich</collectorName>
) and “
<typeStatus id="AD06887AFF9E5F36D72155162FA50C38" box="[1039,1152,855,881]" pageId="2" pageNumber="453" type="paratype">Paratypus</typeStatus>
<emphasis id="40C9EACAFF9E5F36D7A455162E370C38" box="[1162,1298,855,881]" italics="true" pageId="2" pageNumber="453">Probathicus</emphasis>
<taxonomicName id="B5BD4D5BFF9E5F36D63355162ADC0CDC" authority="G. Medvedev, 1977" authorityName="G. Medvedev" authorityYear="1977" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="parthorum" subGenus="Pelorinus">
(
<emphasis id="40C9EACAFF9E5F36D60855162EB60C38" box="[1318,1427,855,881]" italics="true" pageId="2" pageNumber="453">Pelorinus</emphasis>
)
<emphasis id="40C9EACAFF9E5F36D3B9553A2A2A0CDC" box="[151,271,891,917]" italics="true" pageId="2" pageNumber="453">parthorum</emphasis>
G. Medvedev, 1977
</taxonomicName>
</materialsCitation>
;
<specimenCount id="64BBFD51FF9E5F36D121553A29100CDC" box="[527,565,891,917]" pageId="2" pageNumber="453" type="male">1♂</specimenCount>
: the same labels, but
<date id="06031018FF9E5F36D008553A28BB0CDC" box="[806,926,891,917]" pageId="2" pageNumber="453" value="1904-02-18">18.II.1904</date>
;
<specimenCount id="64BBFD51FF9E5F36D084553A28F50CDC" box="[938,976,891,917]" pageId="2" pageNumber="453" type="male">1♂</specimenCount>
: the same labels, but 5.V.[19]04;
<specimenCount id="64BBFD51FF9E5F36D666553A2E480CDC" box="[1352,1389,891,917]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
: the same labels, but 2324.IV.[1]905;
<specimenCount id="64BBFD51FF9E5F36D13155DE29610CF0" box="[543,580,927,953]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
: the same labels, but 29.V.[19]04 and additional labels
<emphasis id="40C9EACAFF9E5F36D79255DE2FE30CF0" box="[1212,1222,927,953]" italics="true" pageId="2" pageNumber="453"></emphasis>
<taxonomicName id="B5BD4D5BFF9E5F36D7E855DE2E4B0CF0" box="[1222,1390,927,953]" class="Insecta" family="Tenebrionidae" genus="Probaticus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="40C9EACAFF9E5F36D7E855DE2E600CF0" box="[1222,1349,927,953]" italics="true" pageId="2" pageNumber="453">Probaticus</emphasis>
sp.
</taxonomicName>
aff.
<emphasis id="40C9EACAFF9E5F36D3B955822BC90C94" box="[151,236,963,989]" italics="true" pageId="2" pageNumber="453">4-collis</emphasis>
Bdi Dr Z. Kaszab det., 1960”, “
<taxonomicName id="B5BD4D5BFF9E5F36D178558229D20C94" box="[598,759,963,989]" class="Insecta" family="Tenebrionidae" genus="Probaticus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="40C9EACAFF9E5F36D178558229F50C94" box="[598,720,963,989]" italics="true" pageId="2" pageNumber="453">Probaticus</emphasis>
sp.
</taxonomicName>
” Red labels with “
<typeStatus id="AD06887AFF9E5F36D0E455822F660C94" box="[970,1091,963,989]" pageId="2" pageNumber="453" type="holotype">Holotypus</typeStatus>
” and “
<typeStatus id="AD06887AFF9E5F36D7A055822E240C94" box="[1166,1281,963,989]" pageId="2" pageNumber="453" type="paratype">Paratypus</typeStatus>
” were pinned by a laboratory assistant (A.K. Chistaykova) with erroneous year and the name of the genus.
</paragraph>
<caption id="26C26650FF9E5F36D3B951742F520804" ID-DOI="http://doi.org/10.5281/zenodo.6785913" ID-Zenodo-Dep="6785913" box="[151,1143,1845,1869]" httpUri="https://zenodo.org/record/6785913/files/figure.png" pageId="2" pageNumber="453" startId="2.[151,250,1845,1869]" targetBox="[151,1436,1049,1822]" targetPageId="2">
<paragraph id="720236D8FF9E5F36D3B951742F520804" blockId="2.[151,1143,1845,1869]" box="[151,1143,1845,1869]" pageId="2" pageNumber="453">
<emphasis id="40C9EACAFF9E5F36D3B951742A360804" bold="true" box="[151,275,1845,1869]" pageId="2" pageNumber="453">FIGURE 1.</emphasis>
<taxonomicName id="B5BD4D5BFF9E5F36D23751742ACC0804" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" box="[281,489,1845,1869]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9E5F36D23751742ACC0804" box="[281,489,1845,1869]" italics="true" pageId="2" pageNumber="453">Euboeus corrugosus</emphasis>
</taxonomicName>
, lectotype, male, habitus. A = dorsally; B = laterally; C = labels.
</paragraph>
</caption>
<paragraph id="720236D8FF9E5F36D3E9513A2F490894" blockId="2.[151,1436,1915,2014]" pageId="2" pageNumber="453">
<emphasis id="40C9EACAFF9E5F36D3E9513A2A1308DC" bold="true" box="[199,310,1915,1941]" pageId="2" pageNumber="453">Material.</emphasis>
<materialsCitation id="C2D53C85FF9E5F36D211513A2E4908DC" ID-GBIF-Occurrence="3828382341" box="[319,1388,1915,1942]" collectingDate="1899-05" collectionCode="HNHM" collectorName="Coll. Hauser &amp; Gorgan" country="Iran" location="Golestan Prov." pageId="2" pageNumber="453" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Golestan">
<specimenCount id="64BBFD51FF9E5F36D211513A2A4608DC" box="[319,355,1915,1941]" pageId="2" pageNumber="453" type="male">1♂</specimenCount>
,
<specimenCount id="64BBFD51FF9E5F36D25F513A2AB108DC" box="[369,404,1915,1941]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
(
<collectionCode id="14ACAE1DFF9E5F36D28A513A292408DC" box="[420,513,1915,1941]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="453">HNHM</collectionCode>
): “Persia Astrabad 5.99
<collectorName id="DF48530EFF9E5F36D03D513A288008DC" box="[787,933,1915,1941]" pageId="2" pageNumber="453">Coll. Hauser</collectorName>
” (
<collectingCountry id="0AAA7648FF9E5F36D091513A28D508DC" box="[959,1008,1915,1941]" name="Iran" pageId="2" pageNumber="453">Iran</collectingCountry>
,
<collectingRegion id="B079F83AFF9E5F36D0D3513A2F8308DF" box="[1021,1190,1915,1942]" country="Iran" name="Golestan" pageId="2" pageNumber="453">Golestan Prov.</collectingRegion>
,
<collectorName id="DF48530EFF9E5F36D79C513A2E2D08DC" box="[1202,1288,1915,1941]" pageId="2" pageNumber="453">Gorgan</collectorName>
,
<date id="06031018FF9E5F36D63B513A2E4108DC" box="[1301,1380,1915,1941]" pageId="2" pageNumber="453" value="1899-05">
<collectingDate id="1647E9F0FF9E5F36D63B513A2E4108DC" box="[1301,1380,1915,1941]" pageId="2" pageNumber="453" value="1899-05">v.1899</collectingDate>
</date>
)
</materialsCitation>
;
<materialsCitation id="C2D53C85FF9E5F36D657513A2E1D08F0" ID-GBIF-Occurrence="3828382310" collectingDate="1999-04-07" collectionCode="SMNS" collectorName="J. Rejsek" country="Iran" location="Karim Ishan" pageId="2" pageNumber="453" specimenCount="1" specimenCount-male="1" stateProvince="Golestan">
<specimenCount id="64BBFD51FF9E5F36D657513A2EB908DC" box="[1401,1436,1915,1941]" pageId="2" pageNumber="453" type="male">1♂</specimenCount>
(
<collectionCode id="14ACAE1DFF9E5F36D3B151DE2BD708F0" box="[159,242,1951,1977]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="453">SMNS</collectionCode>
):
<collectingCountry id="0AAA7648FF9E5F36D22651E12A1D08F3" box="[264,312,1952,1978]" name="Iran" pageId="2" pageNumber="453">Iran</collectingCountry>
,
<collectingRegion id="B079F83AFF9E5F36D26851DE2AD408F3" box="[326,497,1951,1978]" country="Iran" name="Golestan" pageId="2" pageNumber="453">Golestan Prov.</collectingRegion>
,
<location id="77626003FF9E5F36D2D051DE29AA08F0" LSID="urn:lsid:plazi:treatment:FA1487CEFF9E5F37D3B9572F29A10EBC:77626003FF9E5F36D2D051DE29AA08F0" box="[510,655,1951,1977]" country="Iran" name="Karim Ishan" pageId="2" pageNumber="453" stateProvince="Golestan">Karim Ishan</location>
,
<locationDeviation id="63B02FFAFF9E5F36D1B351DE28C708F0" box="[669,994,1951,1978]" location="Gonbad-e-Kavus" pageId="2" pageNumber="453">
<quantity id="B5459B3DFF9E5F36D1B351DE29C208F0" box="[669,743,1951,1977]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="453" unit="km" value="70.0">70 km</quantity>
NE Gonbad-e-Kavus
</locationDeviation>
, 4
<date id="06031018FF9E5F36D72551DE2F5908F0" box="[1035,1148,1951,1977]" pageId="2" pageNumber="453" value="1999-04-07">
<collectingDate id="1647E9F0FF9E5F36D72551DE2F5908F0" box="[1035,1148,1951,1977]" pageId="2" pageNumber="453" value="1999-04-07">7.04.1999</collectingDate>
</date>
(leg.
<collectorName id="DF48530EFF9E5F36D7EC51E12E1508F0" box="[1218,1328,1951,1978]" pageId="2" pageNumber="453">J. Rejsek</collectorName>
)
</materialsCitation>
;
<materialsCitation id="C2D53C85FF9E5F36D66B51DE2F4D0894" ID-GBIF-Occurrence="3828382369" collectingDate="2001-06-07" collectionCode="SMNS" collectorName="G. Sama" country="Iran" location="Gaz" pageId="2" pageNumber="453" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Golestan">
<specimenCount id="64BBFD51FF9E5F36D66B51DE2E4E08F0" box="[1349,1387,1951,1977]" pageId="2" pageNumber="453" type="male">2♂</specimenCount>
,
<specimenCount id="64BBFD51FF9E5F36D65751DE2EB908F0" box="[1401,1436,1951,1977]" pageId="2" pageNumber="453" type="female">1♀</specimenCount>
(
<collectionCode id="14ACAE1DFF9E5F36D3B151822BD70894" box="[159,242,1987,2013]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="453">SMNS</collectionCode>
):
<collectingCountry id="0AAA7648FF9E5F36D22B51852A100897" box="[261,309,1988,2014]" name="Iran" pageId="2" pageNumber="453">Iran</collectingCountry>
,
<collectingRegion id="B079F83AFF9E5F36D26E51822ACD0897" box="[320,488,1987,2014]" country="Iran" name="Golestan" pageId="2" pageNumber="453">Golestan Prov.</collectingRegion>
,
<locationDeviation id="63B02FFAFF9E5F36D2DD5182298D0894" box="[499,680,1987,2013]" location="Bandar" pageId="2" pageNumber="453">
<quantity id="B5459B3DFF9E5F36D2DD5182291F0894" box="[499,570,1987,2013]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="453" unit="km" value="10.0">10 km</quantity>
S Bandar
</locationDeviation>
e
<location id="77626003FF9E5F36D1EC518229D70894" LSID="urn:lsid:plazi:treatment:FA1487CEFF9E5F37D3B9572F29A10EBC:77626003FF9E5F36D1EC518229D70894" box="[706,754,1987,2013]" country="Iran" name="Gaz" pageId="2" pageNumber="453" stateProvince="Golestan">Gaz</location>
, 27.05
<date id="06031018FF9E5F36D0645182289F0897" box="[842,954,1987,2014]" pageId="2" pageNumber="453" value="2001-06-07">
<collectingDate id="1647E9F0FF9E5F36D0645182289F0897" box="[842,954,1987,2014]" pageId="2" pageNumber="453" value="2001-06-07">7.06.2001</collectingDate>
</date>
(leg.
<collectorName id="DF48530EFF9E5F36D0D551822F450894" box="[1019,1120,1987,2013]" pageId="2" pageNumber="453">G. Sama</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="720236D8FF9F5F37D3E956D62AE20E98" blockId="3.[151,1437,150,501]" pageId="3" pageNumber="454">
<emphasis id="40C9EACAFF9F5F37D3E956D62A340FF8" bold="true" box="[199,273,151,177]" pageId="3" pageNumber="454">Notes.</emphasis>
This species was described from
<specimenCount id="64BBFD51FF9F5F37D1A756D629DF0FF8" box="[649,762,151,177]" pageId="3" pageNumber="454" type="male">two males</specimenCount>
from “Klein-Asien” and “Astrabad” (
<bibRefCitation id="162C4B29FF9F5F37D78D56D62E670FF8" author="Seidlitz, G. C. M. von" box="[1187,1346,150,177]" pageId="3" pageNumber="454" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895</bibRefCitation>
, p. 707) in the key.
<bibRefCitation id="162C4B29FF9F5F37D23F56FA29090F9C" author="Leo, P. &amp; Ruzzante, G." box="[273,556,187,213]" pageId="3" pageNumber="454" pagination="83 - 89" refId="ref22267" refString="Leo, P. &amp; Ruzzante, G. (2016) Dati nuovi e riassuntivi sui Tenebrionidi dell'isola di Lesbo (Grecia) (Coleoptera Tenebrionidae). Bollettino della Societa Entomologica Italiana, 148, 83 - 89." type="journal article" year="2016">Leo and Ruzzante (2016)</bibRefCitation>
mentioned that it is a “mysterious” species, described from these two different localities.
<bibRefCitation id="162C4B29FF9F5F37D225569E2AE60FB0" author="Sahlberg, J. R." box="[267,451,223,249]" pageId="3" pageNumber="454" pagination="1 - 282" refId="ref23309" refString="Sahlberg, J. R. (1913) Coleoptera mediterranea orientalis quae in aegypto, Palestina, Syria, Caramania collegerunt John Sahlberg et unio Saalas. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 55, 1 - 282." type="journal article" year="1913">Sahlberg (1913)</bibRefCitation>
,
<bibRefCitation id="162C4B29FF9F5F37D2FE569E294A0FB0" author="Reitter, E." box="[464,623,223,249]" pageId="3" pageNumber="454" pagination="1 - 44" refId="ref23264" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter (1922)</bibRefCitation>
and
<bibRefCitation id="162C4B29FF9F5F37D189569E28D00FB0" author="Nabozhenko, M. V. &amp; Lobl, I." box="[679,1013,223,249]" pageId="3" pageNumber="454" pagination="1802" refId="ref22757" refString="Nabozhenko, M. V. &amp; Lobl, I. (2008) Tribe Helopini Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo books, Stenstrup, pp. 241 - 257." type="book chapter" year="2008">Nabozhenko and Löbl (2008)</bibRefCitation>
mentioned different species from the
<collectingRegion id="B079F83AFF9F5F37D3B957422A650E54" box="[151,320,259,285]" country="Greece" name="Voreio Aigaio" pageId="3" pageNumber="454">Aegean region</collectingRegion>
.
<bibRefCitation id="162C4B29FF9F5F37D2625742291E0E54" author="Nabozhenko, M. V." box="[332,571,259,285]" pageId="3" pageNumber="454" pagination="1802" refId="ref22635" refString="Nabozhenko, M. V. (2020 b) Tribe Helopini Latreille, 1802. In: Iwan, D. &amp; Lobl, I. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 5. Tenebrionoidea. Leiden, Brill, pp. 314 - 339. https: // doi. org / 10.1163 / 9789004434998 _ 004" type="book chapter" year="2020">
Nabozhenko (2020
<emphasis id="40C9EACAFF9F5F37D10A574229170E54" box="[548,562,259,285]" italics="true" pageId="3" pageNumber="454">b</emphasis>
)
</bibRefCitation>
listed
<taxonomicName id="B5BD4D5BFF9F5F37D1A7574228020E55" baseAuthorityName="Seidlitz" baseAuthorityYear="1895" box="[649,807,259,284]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9F5F37D1A7574228020E55" box="[649,807,259,284]" italics="true" pageId="3" pageNumber="454">E. corrugosus</emphasis>
</taxonomicName>
for
<collectingCountry id="0AAA7648FF9F5F37D0775742288E0E54" box="[857,939,259,285]" name="Greece" pageId="3" pageNumber="454">Greece</collectingCountry>
,
<collectingCountry id="0AAA7648FF9F5F37D09957422F220E54" box="[951,1031,259,285]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
and
<collectingCountry id="0AAA7648FF9F5F37D76F57422F570E54" box="[1089,1138,259,285]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
. One
<typeStatus id="AD06887AFF9F5F37D79B57422E290E54" box="[1205,1292,259,285]" pageId="3" pageNumber="454" type="syntype">syntype</typeStatus>
(male) from “
<collectingCountry id="0AAA7648FF9F5F37D38F57662A0D0E08" box="[161,296,295,321]" name="Turkey" pageId="3" pageNumber="454">Asia Minor</collectingCountry>
” was originally mentioned from the collection of G. Kraatz, but it was not found in the Senckenberg Deutsche Entomologische Institut (Müncheberg,
<collectingCountry id="0AAA7648FF9F5F37D192570A280F0E2C" box="[700,810,331,357]" name="Germany" pageId="3" pageNumber="454">Germany</collectingCountry>
). I studied the second Iranian specimen (designated here as the
<typeStatus id="AD06887AFF9F5F37D3F1572E2A6D0EC0" box="[223,328,367,393]" pageId="3" pageNumber="454" type="lectotype">lectotype</typeStatus>
) (
<figureCitation id="EA862A5DFF9F5F37D271572E2A840EC0" box="[351,417,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1845,1869]" captionTargetBox="[151,1436,1049,1822]" captionTargetId="figure-324@2.[151,1436,1049,1822]" captionTargetPageId="2" captionText="FIGURE 1. Euboeus corrugosus, lectotype, male, habitus.A = dorsally; B = laterally; C = labels." figureDoi="http://doi.org/10.5281/zenodo.6785913" httpUri="https://zenodo.org/record/6785913/files/figure.png" pageId="3" pageNumber="454">Fig. 1</figureCitation>
), which is conspecific to Medvedevs
<taxonomicName id="B5BD4D5BFF9F5F37D07D572E28CF0EC0" authorityName="G. Medvedev" authorityYear="1977" box="[851,1002,367,393]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="parthorum">
<emphasis id="40C9EACAFF9F5F37D07D572E28CF0EC0" box="[851,1002,367,393]" italics="true" pageId="3" pageNumber="454">E. parthorum</emphasis>
</taxonomicName>
(
<figureCitation id="EA862A5DFF9F5F37D0D4572E2F180EC0" box="[1018,1085,367,393]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[229,1350,530,1873]" captionTargetId="figure-156@3.[226,1361,525,1873]" captionTargetPageId="3" captionText="FIGURE 2. Euboeus corrugosus, types of E. parthorum, syn. nov., habitus, details of structure. A = male, holotype dorsally; B = label of holotype; C = male, paratype, ventrally; D = female, paratype, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = tegmen, ventrally; K = median lobe of aedeagus, ventrally; L = apical piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.6785917" httpUri="https://zenodo.org/record/6785917/files/figure.png" pageId="3" pageNumber="454">Fig. 2</figureCitation>
) also described from Astrabad. Thus, the following new synonymy is proposed:
<taxonomicName id="B5BD4D5BFF9F5F37D1FA57D22F700EE4" authority="Seidlitz, 1895" authorityName="Seidlitz" authorityYear="1895" box="[724,1109,402,429]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="corrugosus">
<emphasis id="40C9EACAFF9F5F37D1FA57D228890EE5" box="[724,940,403,429]" italics="true" pageId="3" pageNumber="454">Helops corrugosus</emphasis>
<bibRefCitation id="162C4B29FF9F5F37D09857D22F700EE4" author="Seidlitz, G. C. M. von" box="[950,1109,402,429]" pageId="3" pageNumber="454" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von &amp; Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="B5BD4D5BFF9F5F37D75457D22A700E98" authority="G. Medvedev, 1976" authorityName="G. Medvedev" authorityYear="1976" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="parthorum">
<emphasis id="40C9EACAFF9F5F37D75457D22E520EE4" box="[1146,1399,403,429]" italics="true" pageId="3" pageNumber="454">Probaticus parthorum</emphasis>
G.
<bibRefCitation id="162C4B29FF9F5F37D3B957F62A700E98" author="Medvedev, G. S." box="[151,341,439,465]" pageId="3" pageNumber="454" pagination="889 - 902" refId="ref22349" refString="Medvedev, G. S. (1976) New species of darkling beetles (Coleoptera, Tenebrionidae) from Iran. Entomologicheskoe Obozrenie, 55, 889 - 902." type="journal article" year="1976">Medvedev, 1976</bibRefCitation>
</taxonomicName>
,
<emphasis id="40C9EACAFF9F5F37D24E57F62AE20E98" bold="true" box="[352,455,439,465]" pageId="3" pageNumber="454">
<taxonomicNameLabel id="5BFA57B1FF9F5F37D24E57F62AE20E98" box="[352,455,439,465]" pageId="3" pageNumber="454">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3AA76553FF9F5F37D3E9579B29A10EBC" box="[199,644,474,501]" pageId="3" pageNumber="454" type="distribution">
<paragraph id="720236D8FF9F5F37D3E9579B29A10EBC" blockId="3.[151,1437,150,501]" box="[199,644,474,501]" pageId="3" pageNumber="454">
<emphasis id="40C9EACAFF9F5F37D3E9579B2A440EBD" bold="true" box="[199,353,474,500]" pageId="3" pageNumber="454">Distribution.</emphasis>
<collectingCountry id="0AAA7648FF9F5F37D249579A2AB10EBC" box="[359,404,475,501]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
(
<collectingRegion id="B079F83AFF9F5F37D28D579A295D0EBC" box="[419,632,475,501]" country="Iran" name="Golestan" pageId="3" pageNumber="454">Golestan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>