309 lines
37 KiB
XML
309 lines
37 KiB
XML
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<mods:title id="230423D59BEA9A90C98504F65F57BC2B">A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan</mods:title>
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<treatment id="FA1487CEFF9B5F33D3B954CF28780988" ID-DOI="http://doi.org/10.5281/zenodo.6794266" ID-GBIF-Taxon="196377795" ID-Zenodo-Dep="6794266" LSID="urn:lsid:plazi:treatment:FA1487CEFF9B5F33D3B954CF28780988" httpUri="http://treatment.plazi.org/id/FA1487CEFF9B5F33D3B954CF28780988" lastPageNumber="458" pageId="7" pageNumber="458">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B954CF29800DE1" authority="(Allard, 1876)" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[151,677,654,681]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus" subGenus="Pelorinus">
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<emphasis id="40C9EACAFF9B5F33D3B954CF29800DE1" bold="true" box="[151,677,654,681]" pageId="7" pageNumber="458">
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<emphasis id="40C9EACAFF9B5F33D3B954CF2A240DE0" bold="true" box="[151,257,654,681]" italics="true" pageId="7" pageNumber="458">Euboeus</emphasis>
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(
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<emphasis id="40C9EACAFF9B5F33D23C54CF2AA20DE0" bold="true" box="[274,391,654,681]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
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)
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<emphasis id="40C9EACAFF9B5F33D2B754CE2ACA0DE0" bold="true" box="[409,495,655,681]" italics="true" pageId="7" pageNumber="458">vicinus</emphasis>
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(
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<bibRefCitation id="162C4B29FF9B5F33D2D154CF29B90DE1" author="Allard, E." box="[511,668,654,680]" pageId="7" pageNumber="458" pagination="1 - 80" refId="ref21895" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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<subSubSection id="3AA76553FF9B5F33D3B954F22BC90D84" box="[151,236,691,717]" pageId="7" pageNumber="458" type="description">
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<paragraph id="720236D8FF9B5F33D3B954F22BC90D84" blockId="7.[151,677,654,717]" box="[151,236,691,717]" pageId="7" pageNumber="458">
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(
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<figureCitation id="EA862A5DFF9B5F33D3B154F22BC60D84" box="[159,227,691,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1851,1875]" captionTargetBox="[213,1377,185,1824]" captionTargetId="figure-32@8.[208,1380,181,1827]" captionTargetPageId="8" captionText="FIGURE 6. Euboeus vicinus, habitus, labels, details of structure. A = male dorsally (Georgia, Asureti); B = female dorsally (Georgia, Asureti); C = female dorsally (Georgia, Asureti); D = female dorsally (Armenia, Alaverdi), holotype of Probaticus medvedevi; E = labels of non-type specimens from Georgia, Asureti (HNHM); F = labels of the holotype of Probaticus medvedevi; G = male inner sternite VIII, ventrally; H = spiculum gastrale; I = median lobe of aedeagus, ventrally; J = tegmen, ventrally; K = apical piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.6785931" httpUri="https://zenodo.org/record/6785931/files/figure.png" pageId="7" pageNumber="458">Fig. 6</figureCitation>
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)
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<paragraph id="720236D8FF9B5F33D3B954B629230C46" blockId="7.[151,1006,759,937]" box="[151,518,759,783]" pageId="7" pageNumber="458">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B954B62AFF0C46" authority="Allard, 1876: 14" authorityName="Allard" authorityPageNumber="14" authorityYear="1876" box="[151,474,759,783]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
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<emphasis id="40C9EACAFF9B5F33D3B954B62A080C46" box="[151,301,759,783]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
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<bibRefCitation id="162C4B29FF9B5F33D21C54B62AFF0C46" author="Allard, E." box="[306,474,759,783]" pageId="7" pageNumber="458" pagination="1 - 80" refId="ref21895" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876: 14</bibRefCitation>
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, 34;
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<paragraph id="720236D8FF9B5F33D3B9555729290C67" blockId="7.[151,1006,759,937]" box="[151,524,790,814]" pageId="7" pageNumber="458">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B955572AC70C67" authority=": Allard, 1877: 26" authorityName="Allard" authorityPageNumber="26" authorityYear="1877" box="[151,482,790,814]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
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<emphasis id="40C9EACAFF9B5F33D3B955572A080C64" box="[151,301,790,814]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
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:
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<bibRefCitation id="162C4B29FF9B5F33D21755572AC70C67" author="Allard, E." box="[313,482,790,814]" pageId="7" pageNumber="458" pagination="13 - 268" refId="ref21920" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">Allard, 1877: 26</bibRefCitation>
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, 91;
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<paragraph id="720236D8FF9B5F33D3B9557529270C05" blockId="7.[151,1006,759,937]" box="[151,514,820,844]" pageId="7" pageNumber="458">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B955752ADB0C05" authority=": Seidlitz, 1895: 706" authorityName=": Seidlitz" authorityPageNumber="706" authorityYear="1895" box="[151,510,820,844]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
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<emphasis id="40C9EACAFF9B5F33D3B955752A080C05" box="[151,301,820,844]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
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:
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<bibRefCitation id="162C4B29FF9B5F33D21455752ADB0C05" author="Seidlitz, G. C. M. von" box="[314,510,820,844]" pageId="7" pageNumber="458" pagination="609 - 800" refId="ref23469" refString="Seidlitz, G. C. M. von (1895) Tenebrionidae. In: Kiesenwetter, H. von & Seidlitz, G. C. M. von (Eds.), Naturgeschichte der Insecten Deutschlands. Begonnen von Dr. W. F. Erichson, fortgesetzt von Prof. Dr. H. Schaum, Dr. G. Kraatz, H. v. Kiesenwetter, Julius Weise, Edm. Reitter und Dr. G. Seidlitz. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800. https: // doi. org / 10.5962 / bhl. title. 8270" type="book chapter" year="1895">Seidlitz, 1895: 706</bibRefCitation>
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;
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<paragraph id="720236D8FF9B5F33D3B9551229CF0C22" blockId="7.[151,1006,759,937]" box="[151,746,851,875]" pageId="7" pageNumber="458">
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<treatmentCitation id="F31C10C9FF9B5F33D3B9551229A30C22" author="Reitter, E." box="[151,646,851,875]" page="33" pageId="7" pageNumber="458" year="1922">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B9551229A30C22" authority=": Reitter, 1922: 33" authorityName=": Reitter" authorityPageNumber="33" authorityYear="1922" box="[151,646,851,875]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus" subGenus="Pelorinus">
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<emphasis id="40C9EACAFF9B5F33D3B955122A200C22" box="[151,261,851,875]" italics="true" pageId="7" pageNumber="458">Probaticus</emphasis>
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(
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<emphasis id="40C9EACAFF9B5F33D23A55122A520C22" box="[276,375,851,875]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
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)
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<emphasis id="40C9EACAFF9B5F33D2AB55152AE80C22" box="[389,461,852,875]" italics="true" pageId="7" pageNumber="458">vicinus</emphasis>
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:
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<bibRefCitation id="162C4B29FF9B5F33D2F5551229A30C22" author="Reitter, E." box="[475,646,851,875]" pageId="7" pageNumber="458" pagination="1 - 44" refId="ref23264" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44 + 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922: 33</bibRefCitation>
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, nota 28.;
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<paragraph id="720236D8FF9B5F33D3B9553328CB0CC0" blockId="7.[151,1006,759,937]" box="[151,1006,882,906]" pageId="7" pageNumber="458">
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||
<taxonomicName id="B5BD4D5BFF9B5F33D3B95533285A0CC3" authority="Abdurakhmanov et Nabozhenko, 2011" authorityName="Abdurakhmanov et Nabozhenko" authorityYear="2011" box="[151,895,882,906]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi" subGenus="Pelorinus">
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<emphasis id="40C9EACAFF9B5F33D3B955332A200CC3" box="[151,261,882,906]" italics="true" pageId="7" pageNumber="458">Probaticus</emphasis>
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(
|
||
<emphasis id="40C9EACAFF9B5F33D23A55332A520CC3" box="[276,375,882,906]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
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)
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||
<emphasis id="40C9EACAFF9B5F33D2AB55332AD50CC3" box="[389,496,882,906]" italics="true" pageId="7" pageNumber="458">medvedevi</emphasis>
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<bibRefCitation id="162C4B29FF9B5F33D2DB5533285A0CC3" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[501,895,882,906]" pageId="7" pageNumber="458" refId="ref21743" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov et Nabozhenko, 2011</bibRefCitation>
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</taxonomicName>
|
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,
|
||
<emphasis id="40C9EACAFF9B5F33D0A4553328C20CC3" bold="true" box="[906,999,882,906]" pageId="7" pageNumber="458">
|
||
<taxonomicNameLabel id="5BFA57B1FF9B5F33D0A4553328C20CC3" box="[906,999,882,906]" pageId="7" pageNumber="458">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="720236D8FF9B5F33D3B955D1283A0CE0" blockId="7.[151,1006,759,937]" box="[151,799,912,937]" pageId="7" pageNumber="458">
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<treatmentCitationGroup id="52AD11F6FF9B5F33D3B955D1283A0CE0" box="[151,799,912,937]" pageId="7" pageNumber="458">
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<taxonomicName id="B5BD4D5BFF9B5F33D3B955D1283A0CE0" authority=": Nabozhenko et al., 2021 b: 441" authorityName="Nabozhenko" authorityPageNumber="441" authorityYear="2021" box="[151,799,912,937]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi" subGenus="Pelorinus">
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<emphasis id="40C9EACAFF9B5F33D3B955D12BCB0CE1" box="[151,238,912,936]" italics="true" pageId="7" pageNumber="458">Euboeus</emphasis>
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||
(
|
||
<emphasis id="40C9EACAFF9B5F33D3D355D12A450CE1" box="[253,352,912,936]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
|
||
)
|
||
<emphasis id="40C9EACAFF9B5F33D24055D12AFC0CE1" box="[366,473,912,936]" italics="true" pageId="7" pageNumber="458">medvedevi</emphasis>
|
||
:
|
||
<bibRefCitation id="162C4B29FF9B5F33D2C955D1283A0CE0" author="Nabozhenko, M. V. & Kalashian, M. Yu. & Mazmanyan, M. A." box="[487,799,912,937]" pageId="7" pageNumber="458" pagination="425 - 450" refId="ref22898" refString="Nabozhenko, M. V., Kalashian, M. Yu. & Mazmanyan, M. A. (2021 b) A faunistic review of darkling beetles (Coleoptera: Tenebrionidae; excluding Alleculinae) of Armenia and partly the Nakhichevan Autonomous Republic of Azerbaijan with new records and taxonomic notes. Caucasian Entomological Bulletin, 17, 425 - 450. https: // doi. org / 10.23885 / 181433262021172 - 425450" type="journal article" year="2021">
|
||
Nabozhenko
|
||
<emphasis id="40C9EACAFF9B5F33D14355D029BF0CE1" box="[621,666,912,936]" italics="true" pageId="7" pageNumber="458">et al</emphasis>
|
||
., 2021b: 441
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AA76553FF9B5F33D3B955962FC909D4" pageId="7" pageNumber="458" type="materials_examined">
|
||
<paragraph id="720236D8FF9B5F33D3B955962A9A0B14" blockId="7.[151,1437,982,1729]" pageId="7" pageNumber="458">
|
||
<emphasis id="40C9EACAFF9B5F33D3B955962A600CB9" bold="true" box="[151,325,982,1009]" pageId="7" pageNumber="458">Type material.</emphasis>
|
||
<typeStatus id="AD06887AFF9B5F33D26555962A970CB8" box="[331,434,983,1009]" pageId="7" pageNumber="458" type="syntype">Syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D2F85596295E0CB9" authorityName=": Seidlitz" authorityYear="1895" box="[470,635,983,1009]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
|
||
<emphasis id="40C9EACAFF9B5F33D2F85596295E0CB9" box="[470,635,983,1009]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
|
||
</taxonomicName>
|
||
were not found by curators in the Muséum national d’Histoire naturelle Paris.
|
||
<materialsCitation id="C2D53C85FF9B5F33D3CE55BA2A9E0B14" ID-GBIF-Occurrence="3828382367" collectingDate="1986-04-25" collectionCode="ZIN" collectorName="S. V. Melyakh" country="Armenia" location="Alaverdi" pageId="7" pageNumber="458" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<typeStatus id="AD06887AFF9B5F33D3CE55BA2A6E0B5C" box="[224,331,1019,1045]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<specimenCount id="64BBFD51FF9B5F33D27755BA2A4B0B5C" box="[345,366,1019,1045]" pageId="7" pageNumber="458" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="14ACAE1DFF9B5F33D2AE55BA2A970B5C" box="[384,434,1019,1045]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="458" type="Museum">ZIN</collectionCode>
|
||
) of
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D2CA55BA29FB0B5C" authorityName="Abdurakhmanov et Nabozhenko" authorityYear="2011" box="[484,734,1019,1045]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi">
|
||
<emphasis id="40C9EACAFF9B5F33D2CA55BA29FB0B5C" box="[484,734,1019,1045]" italics="true" pageId="7" pageNumber="458">Probaticus medvedevi</emphasis>
|
||
</taxonomicName>
|
||
with labels: Cyrilliс “Арм. ССР Алаверди 25.04.86 Мелях С.В.” (
|
||
<collectingCountry id="0AAA7648FF9B5F33D3C9525E2AB00B70" box="[231,405,1055,1081]" name="Armenia" pageId="7" pageNumber="458">Armenian SSR</collectingCountry>
|
||
,
|
||
<location id="77626003FF9B5F33D28E525E29200B70" LSID="urn:lsid:plazi:treatment:FA1487CEFF9B5F33D3B954CF28780988:77626003FF9B5F33D28E525E29200B70" box="[416,517,1055,1081]" country="Armenia" name="Alaverdi" pageId="7" pageNumber="458">Alaverdi</location>
|
||
,
|
||
<date id="06031018FF9B5F33D13F525F29AC0B70" box="[529,649,1054,1081]" pageId="7" pageNumber="458" value="1986-04-25">
|
||
<collectingDate id="1647E9F0FF9B5F33D13F525F29AC0B70" box="[529,649,1054,1081]" pageId="7" pageNumber="458" value="1986-04-25">25.iv.1986</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="DF48530EFF9B5F33D1E6525E28430B70" box="[712,870,1055,1081]" pageId="7" pageNumber="458">S.V. Melyakh</collectorName>
|
||
), “
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D0A9525E2A950B14" authority="Nabozhenko et Abdurakhmanov 2011" authorityName="Nabozhenko et Abdurakhmanov" authorityYear="2011" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi" subGenus="Pelorinus">
|
||
<emphasis id="40C9EACAFF9B5F33D0A9525E2F240B70" box="[903,1025,1055,1081]" italics="true" pageId="7" pageNumber="458">Probaticus</emphasis>
|
||
(
|
||
<emphasis id="40C9EACAFF9B5F33D73C525E2F5A0B70" box="[1042,1151,1055,1081]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
|
||
)
|
||
<emphasis id="40C9EACAFF9B5F33D7BE525E2E230B70" box="[1168,1286,1055,1081]" italics="true" pageId="7" pageNumber="458">medvedevi</emphasis>
|
||
Nabozhenko et
|
||
<bibRefCitation id="162C4B29FF9B5F33D39F52022A950B14" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[177,432,1091,1117]" pageId="7" pageNumber="458" refId="ref21743" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="720236D8FF9B5F33D3E952272A770BEC" blockId="7.[151,1437,982,1729]" pageId="7" pageNumber="458">
|
||
<emphasis id="40C9EACAFF9B5F33D3E952272A130BC9" bold="true" box="[199,310,1126,1152]" pageId="7" pageNumber="458">Material.</emphasis>
|
||
<materialsCitation id="C2D53C85FF9B5F33D21052262A6B0BEC" ID-GBIF-Occurrence="3828382330" collectorName="Coll. Reitter" pageId="7" pageNumber="458" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="64BBFD51FF9B5F33D21052262A470BC8" box="[318,354,1127,1153]" pageId="7" pageNumber="458" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="64BBFD51FF9B5F33D25E52262AB30BC8" box="[368,406,1127,1153]" pageId="7" pageNumber="458" type="female">2♀</specimenCount>
|
||
: “Kaukasus Leder”, “Allard type”, “
|
||
<collectorName id="DF48530EFF9B5F33D01F522628E70BC8" box="[817,962,1127,1153]" pageId="7" pageNumber="458">Coll. Reitter</collectorName>
|
||
”, “
|
||
<typeStatus id="AD06887AFF9B5F33D0CC52262F760BC8" box="[994,1107,1127,1153]" pageId="7" pageNumber="458" type="paratype">Paratypus</typeStatus>
|
||
1877
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D7B452262EB70BC8" authority="Allard" authorityName="Allard" authorityYear="1876" box="[1178,1426,1127,1153]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
|
||
<emphasis id="40C9EACAFF9B5F33D7B452262E650BC9" box="[1178,1344,1127,1153]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
|
||
Allard
|
||
</taxonomicName>
|
||
” (curator’s label)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="720236D8FF9B5F33D3E952EE2FC909D4" blockId="7.[151,1437,982,1729]" pageId="7" pageNumber="458">
|
||
<emphasis id="40C9EACAFF9B5F33D3E952EE2A340B80" bold="true" box="[199,273,1199,1225]" pageId="7" pageNumber="458">Notes.</emphasis>
|
||
This very rare species was described in 1876 based on the specimen(s) from the collection of Reiche (
|
||
<bibRefCitation id="162C4B29FF9B5F33D65852EE2A260BA4" author="Allard, E." pageId="7" pageNumber="458" pagination="1 - 80" refId="ref21895" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard 1876</bibRefCitation>
|
||
) and redescribed in 1877 using the type specimen(s) and beetles collected by Leder (
|
||
<bibRefCitation id="162C4B29FF9B5F33D79252922E6D0BA4" author="Allard, E." box="[1212,1352,1235,1261]" pageId="7" pageNumber="458" pagination="13 - 268" refId="ref21920" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">Allard 1877</bibRefCitation>
|
||
). Later,
|
||
<bibRefCitation id="162C4B29FF9B5F33D3B952B62A980A58" author="Schneider, O. & Leder, H." box="[151,445,1271,1297]" pageId="7" pageNumber="458" pagination="3 - 262" refId="ref23434" refString="Schneider, O. & Leder, H. (1878) Beitrage zur Kenntniss der kaukasischen Kaferfauna. Verhandlungen des Naturforschendes Vereines in Brunn, 16 (Supplementum), 3 - 262." type="journal article" year="1878">Schneider & Leder (1878)</bibRefCitation>
|
||
again published Allard’s redescription 1877 and added the species’ type locality: “Elisabetthal”. It was a settlement of German colonists, which is now called Asureti (
|
||
<collectingCountry id="0AAA7648FF9B5F33D723535A2F4E0A7C" box="[1037,1131,1307,1333]" name="Georgia" pageId="7" pageNumber="458">Georgia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="1789501FFF9B5F33D758535D2FDB0A7E" box="[1142,1278,1308,1335]" degrees="41" direction="north" minutes="35" orientation="latitude" pageId="7" pageNumber="458" precision="15" seconds="45" value="41.595833">41°35′45″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1789501FFF9B5F33D624535D2EB40A7E" box="[1290,1425,1308,1335]" degrees="44" direction="east" minutes="40" orientation="longitude" pageId="7" pageNumber="458" precision="15" seconds="00" value="44.666668">44°40′00″E</geoCoordinate>
|
||
). Both description and redescription (
|
||
<bibRefCitation id="162C4B29FF9B5F33D10D537E29880A10" author="Allard, E." box="[547,685,1343,1369]" pageId="7" pageNumber="458" pagination="1 - 80" refId="ref21895" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard 1876</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="162C4B29FF9B5F33D198537E29CA0A10" author="Allard, E." box="[694,751,1343,1369]" pageId="7" pageNumber="458" pagination="13 - 268" refId="ref21920" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">1877</bibRefCitation>
|
||
) have differences, and the latter is more consistent with specimens from
|
||
<collectingCountry id="0AAA7648FF9B5F33D23453222A510A34" box="[282,372,1379,1405]" name="Georgia" pageId="7" pageNumber="458">Georgia</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="0AAA7648FF9B5F33D13D5322295D0A34" box="[531,632,1379,1405]" name="Armenia" pageId="7" pageNumber="458">Armenia</collectingCountry>
|
||
, as mentioned by
|
||
<bibRefCitation id="162C4B29FF9B5F33D06953222E280A34" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[839,1293,1379,1405]" pageId="7" pageNumber="458" refId="ref21743" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov & Nabozhenko (2011)</bibRefCitation>
|
||
. In addition, the Russian authors erroneously interpreted species from South
|
||
<collectingCountry id="0AAA7648FF9B5F33D04653C628EF0AE8" box="[872,970,1415,1441]" name="Armenia" pageId="7" pageNumber="458">Armenia</collectingCountry>
|
||
and Nakhchivan as
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D79E53C62E3A0AE9" authorityName="Allard" authorityYear="1876" box="[1200,1311,1415,1440]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
|
||
<emphasis id="40C9EACAFF9B5F33D79E53C62E3A0AE9" box="[1200,1311,1415,1440]" italics="true" pageId="7" pageNumber="458">E. vicinus</emphasis>
|
||
</taxonomicName>
|
||
(described below as
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D22D53EA2ABC0A8C" authority="Nabozhenko, 2022" authorityName="Nabozhenko" authorityYear="2022" box="[259,409,1451,1477]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="kalashiani" status="sp. nov.">
|
||
<emphasis id="40C9EACAFF9B5F33D22D53EA2ABC0A8C" box="[259,409,1451,1477]" italics="true" pageId="7" pageNumber="458">E. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BFA57B1FF9B5F33D28F53EA2ADF0A8C" box="[417,506,1451,1477]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) since Bogachev (1938). As a result, Abdurakhmanov & Nabozhenko described a new species
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D208538E293B0AA0" authorityName="Nabozhenko et Abdurakhmanov" authorityYear="2011" box="[294,542,1487,1513]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi">
|
||
<emphasis id="40C9EACAFF9B5F33D208538E293B0AA0" box="[294,542,1487,1513]" italics="true" pageId="7" pageNumber="458">Probaticus medvedevi</emphasis>
|
||
</taxonomicName>
|
||
, from northern
|
||
<collectingCountry id="0AAA7648FF9B5F33D1FE538E28170AA0" box="[720,818,1487,1513]" name="Armenia" pageId="7" pageNumber="458">Armenia</collectingCountry>
|
||
(
|
||
<figureCitation id="EA862A5DFF9B5F33D06F538E289A0AA0" box="[833,959,1487,1513]" captionStart="FIGURE 6" captionStartId="8.[151,250,1851,1875]" captionTargetBox="[213,1377,185,1824]" captionTargetId="figure-32@8.[208,1380,181,1827]" captionTargetPageId="8" captionText="FIGURE 6. Euboeus vicinus, habitus, labels, details of structure. A = male dorsally (Georgia, Asureti); B = female dorsally (Georgia, Asureti); C = female dorsally (Georgia, Asureti); D = female dorsally (Armenia, Alaverdi), holotype of Probaticus medvedevi; E = labels of non-type specimens from Georgia, Asureti (HNHM); F = labels of the holotype of Probaticus medvedevi; G = male inner sternite VIII, ventrally; H = spiculum gastrale; I = median lobe of aedeagus, ventrally; J = tegmen, ventrally; K = apical piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.6785931" httpUri="https://zenodo.org/record/6785931/files/figure.png" pageId="7" pageNumber="458">Figs 6D, F</figureCitation>
|
||
). Allard examined the specimens of Leder and confirmed that they belong to his species
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D19953B228450945" authorityName="Allard" authorityYear="1876" box="[695,864,1523,1549]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
|
||
<emphasis id="40C9EACAFF9B5F33D19953B228450945" box="[695,864,1523,1549]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
|
||
</taxonomicName>
|
||
, as he wrote in redescription (
|
||
<bibRefCitation id="162C4B29FF9B5F33D7E453B22E7F0944" author="Allard, E." box="[1226,1370,1523,1549]" pageId="7" pageNumber="458" pagination="13 - 268" refId="ref21920" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">Allard 1877</bibRefCitation>
|
||
). The Leder’s specimens (
|
||
<figureCitation id="EA862A5DFF9B5F33D25B505629700978" box="[373,597,1559,1585]" captionStart="FIGURE 6" captionStartId="8.[151,250,1851,1875]" captionTargetBox="[213,1377,185,1824]" captionTargetId="figure-32@8.[208,1380,181,1827]" captionTargetPageId="8" captionText="FIGURE 6. Euboeus vicinus, habitus, labels, details of structure. A = male dorsally (Georgia, Asureti); B = female dorsally (Georgia, Asureti); C = female dorsally (Georgia, Asureti); D = female dorsally (Armenia, Alaverdi), holotype of Probaticus medvedevi; E = labels of non-type specimens from Georgia, Asureti (HNHM); F = labels of the holotype of Probaticus medvedevi; G = male inner sternite VIII, ventrally; H = spiculum gastrale; I = median lobe of aedeagus, ventrally; J = tegmen, ventrally; K = apical piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.6785931" httpUri="https://zenodo.org/record/6785931/files/figure.png" pageId="7" pageNumber="458">Figs 6A–C, E, G–K</figureCitation>
|
||
) are not
|
||
<typeStatus id="AD06887AFF9B5F33D19B505628320978" box="[693,791,1559,1585]" pageId="7" pageNumber="458" type="syntype">syntypes</typeStatus>
|
||
(as marked on the labels) because the original description was based on other specimens from the Reiche collection. The females from
|
||
<collectingCountry id="0AAA7648FF9B5F33D0D6507A2F77091C" box="[1016,1106,1595,1621]" name="Georgia" pageId="7" pageNumber="458">Georgia</collectingCountry>
|
||
are conspecific to the female (
|
||
<typeStatus id="AD06887AFF9B5F33D3B1501E2A210930" box="[159,260,1631,1657]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
|
||
) of
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D21A501E2AEF0930" authorityName="Nabozhenko et Abdurakhmanov" authorityYear="2011" box="[308,458,1631,1657]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi">
|
||
<emphasis id="40C9EACAFF9B5F33D21A501E2AEF0930" box="[308,458,1631,1657]" italics="true" pageId="7" pageNumber="458">E. medvedevi</emphasis>
|
||
</taxonomicName>
|
||
despite the variability. As a result, I propose the following synonymy:
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D7DB501E2A0009D4" authority="Allard, 1876" authorityName="Allard" authorityYear="1876" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="vicinus">
|
||
<emphasis id="40C9EACAFF9B5F33D7DB501E2EB90931" box="[1269,1436,1631,1657]" italics="true" pageId="7" pageNumber="458">Helops vicinus</emphasis>
|
||
<bibRefCitation id="162C4B29FF9B5F33D3B950C22A0009D4" author="Allard, E." box="[151,293,1667,1693]" pageId="7" pageNumber="458" pagination="1 - 80" refId="ref21895" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="B5BD4D5BFF9B5F33D26D50C22F5C09D4" authority="Abdurakhmanov et Nabozhenko, 2011" authorityName="Abdurakhmanov et Nabozhenko" authorityYear="2011" box="[323,1145,1667,1693]" class="Insecta" family="Tenebrionidae" genus="Probaticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="medvedevi" subGenus="Pelorinus">
|
||
<emphasis id="40C9EACAFF9B5F33D26D50C22A9809D4" box="[323,445,1667,1693]" italics="true" pageId="7" pageNumber="458">Probaticus</emphasis>
|
||
(
|
||
<emphasis id="40C9EACAFF9B5F33D2E350C2291F09D4" box="[461,570,1667,1693]" italics="true" pageId="7" pageNumber="458">Pelorinus</emphasis>
|
||
)
|
||
<emphasis id="40C9EACAFF9B5F33D16450C229E509D4" box="[586,704,1667,1693]" italics="true" pageId="7" pageNumber="458">medvedevi</emphasis>
|
||
<bibRefCitation id="162C4B29FF9B5F33D1EB50C22F5C09D4" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[709,1145,1667,1693]" pageId="7" pageNumber="458" refId="ref21743" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov et Nabozhenko, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="40C9EACAFF9B5F33D7AA50C22FC909D4" bold="true" box="[1156,1260,1667,1693]" pageId="7" pageNumber="458">
|
||
<taxonomicNameLabel id="5BFA57B1FF9B5F33D7AA50C22FC909D4" box="[1156,1260,1667,1693]" pageId="7" pageNumber="458">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AA76553FF9B5F33D3E950E728780988" box="[199,861,1702,1729]" pageId="7" pageNumber="458" type="distribution">
|
||
<paragraph id="720236D8FF9B5F33D3E950E728780988" blockId="7.[151,1437,982,1729]" box="[199,861,1702,1729]" pageId="7" pageNumber="458">
|
||
<emphasis id="40C9EACAFF9B5F33D3E950E72A440989" bold="true" box="[199,353,1702,1728]" pageId="7" pageNumber="458">Distribution.</emphasis>
|
||
Southern
|
||
<collectingCountry id="0AAA7648FF9B5F33D2FA50E629140988" box="[468,561,1703,1729]" name="Georgia" pageId="7" pageNumber="458">Georgia</collectingCountry>
|
||
, northern area of
|
||
<collectingCountry id="0AAA7648FF9B5F33D1DD50E6287C0988" box="[755,857,1703,1729]" name="Armenia" pageId="7" pageNumber="458">Armenia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |