231 lines
38 KiB
XML
231 lines
38 KiB
XML
<document id="D5DE97F3CDE454C50638EE084593E37B" ID-CLB-Dataset="88477" ID-DOI="10.11646/zootaxa.5159.4.1" ID-GBIF-Dataset="cffc486a-265f-4d35-a289-a4a54845c77d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6785909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656658261263" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V." docDate="2022" docId="FA1487CEFF965F38D3B9528F292B0BE8" docLanguage="en" docName="zootaxa.5159.4.1.pdf" docOrigin="Zootaxa 5159 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euboeus (Pelorinus) arzanovi Nabozhenko 2022, sp. nov." docType="treatment" docUuid="5803B205-2722-40BA-87C3-7EAC3049ABC6" docUuidSource="ZooBank" docVersion="6" lastPageNumber="463" masterDocId="062DFFB6FF9C5F34D32E56412B250F49" masterDocTitle="A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="461" updateTime="1699353871700" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="921344F9BEF22CFC53907DFEF0F00675">A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan</mods:title>
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<mods:name id="1FBFFCD7AEA91AE6E1E1212FD0714374" type="personal">
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<mods:namePart id="57465888AB3C4F608A91983224BFB8DE">Nabozhenko, Maxim V.</mods:namePart>
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<mods:title id="9A76291DFC74754DF4C3C0A0AA0494B9">Zootaxa</mods:title>
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<mods:part id="DD0BB9C3FC32D6CC1ABE7F674E4BC485">
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<mods:date id="E58EDD9BCB417208FAEC842A96506A38">2022</mods:date>
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<mods:number id="C6E73CA08CE0547CCC95B2EE27EC221B">2022-07-01</mods:number>
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<treatment id="FA1487CEFF965F38D3B9528F292B0BE8" ID-DOI="http://doi.org/10.5281/zenodo.6785915" ID-GBIF-Taxon="196377799" ID-Zenodo-Dep="6785915" LSID="urn:lsid:zoobank.org:act:5803B205-2722-40BA-87C3-7EAC3049ABC6" httpUri="http://treatment.plazi.org/id/FA1487CEFF965F38D3B9528F292B0BE8" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
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<subSubSection id="3AA76553FF965F3ED3B9528F29430BA0" box="[151,614,1230,1257]" pageId="10" pageNumber="461" type="nomenclature">
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<paragraph id="720236D8FF965F3ED3B9528F29430BA0" blockId="10.[151,614,1230,1293]" box="[151,614,1230,1257]" pageId="10" pageNumber="461">
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<heading id="294A81B4FF965F3ED3B9528F29430BA0" bold="true" box="[151,614,1230,1257]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
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<emphasis id="40C9EACAFF965F3ED3B9528F29430BA0" bold="true" box="[151,614,1230,1257]" pageId="10" pageNumber="461">
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<taxonomicName id="B5BD4D5BFF965F3ED3B9528F29240BA0" authority="Nabozhenko, 2022" authorityName="Nabozhenko" authorityYear="2022" box="[151,513,1230,1257]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="arzanovi" status="sp. nov." subGenus="Pelorinus">
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<emphasis id="40C9EACAFF965F3ED3B9528F2A240BA0" bold="true" box="[151,257,1230,1257]" italics="true" pageId="10" pageNumber="461">Euboeus</emphasis>
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||
(
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<emphasis id="40C9EACAFF965F3ED23C528F2AA20BA0" bold="true" box="[274,391,1230,1257]" italics="true" pageId="10" pageNumber="461">Pelorinus</emphasis>
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||
)
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<emphasis id="40C9EACAFF965F3ED2B7528E29240BA0" bold="true" box="[409,513,1231,1257]" italics="true" pageId="10" pageNumber="461">arzanovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5BFA57B1FF965F3ED126528E29430BA0" box="[520,614,1231,1257]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3AA76553FF965F3ED3B952B22BCE0A44" box="[151,235,1267,1293]" pageId="10" pageNumber="461" type="description">
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||
<paragraph id="720236D8FF965F3ED3B952B22BCE0A44" blockId="10.[151,614,1230,1293]" box="[151,235,1267,1293]" pageId="10" pageNumber="461">
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||
(
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<figureCitation id="EA862A5DFF965F3ED3B152B22BC70A44" box="[159,226,1267,1293]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Fig. 8</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3AA76553FF965F3ED3B9537A281B0A30" pageId="10" pageNumber="461" type="materials_examined">
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<paragraph id="720236D8FF965F3ED3B9537A281B0A30" blockId="10.[151,1437,1338,2013]" pageId="10" pageNumber="461">
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<emphasis id="40C9EACAFF965F3ED3B9537A2A630A1D" bold="true" box="[151,326,1338,1365]" pageId="10" pageNumber="461">Type material.</emphasis>
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<materialsCitation id="C2D53C85FF965F3ED260537A281B0A30" ID-GBIF-Occurrence="3828382318" collectingDate="1970-04-28" collectionCode="HNHM" collectorName="Wittmer & v. Bothmer" country="Iran" elevation="2100" location="Marzanabad" municipality="Gachsar" pageId="10" pageNumber="461" specimenCount="2" specimenCount-male="2" stateProvince="Alborz" typeStatus="holotype">
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||
<typeStatus id="AD06887AFF965F3ED260537A2A9C0A1C" box="[334,441,1339,1365]" pageId="10" pageNumber="461" type="holotype">Holotype</typeStatus>
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||
,
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<specimenCount id="64BBFD51FF965F3ED2EB537A2AFF0A1C" box="[453,474,1339,1365]" pageId="10" pageNumber="461" type="male">♂</specimenCount>
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and
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<typeStatus id="AD06887AFF965F3ED13D537A29530A1C" box="[531,630,1339,1365]" pageId="10" pageNumber="461" type="paratype">paratype</typeStatus>
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||
,
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<specimenCount id="64BBFD51FF965F3ED1AC537A29B20A1C" box="[642,663,1339,1365]" pageId="10" pageNumber="461" type="male">♂</specimenCount>
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||
(
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<collectionCode id="14ACAE1DFF965F3ED189537A28210A1C" box="[679,772,1339,1365]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="461">HNHM</collectionCode>
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):
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<collectingCountry id="0AAA7648FF965F3ED036537A286D0A1C" box="[792,840,1339,1365]" name="Iran" pageId="10" pageNumber="461">Iran</collectingCountry>
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,
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<collectingMunicipality id="9266ACA2FF965F3ED07A537A28950A1C" box="[852,944,1339,1365]" pageId="10" pageNumber="461">Gachsar</collectingMunicipality>
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[
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<collectingRegion id="B079F83AFF965F3ED090537A2F740A1C" box="[958,1105,1339,1365]" country="Iran" name="Alborz" pageId="10" pageNumber="461">Alborz Prov.</collectingRegion>
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||
]—
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||
<location id="77626003FF965F3ED75D537A2E250A1C" LSID="urn:lsid:plazi:treatment:FA1487CEFF965F38D3B9528F292B0BE8:77626003FF965F3ED75D537A2E250A1C" box="[1139,1280,1339,1365]" country="Iran" municipality="Gachsar" name="Marzanabad" pageId="10" pageNumber="461" stateProvince="Alborz">Marzanabad</location>
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||
[
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<collectingRegion id="B079F83AFF965F3ED621537A2BF20A30" country="Iran" name="Mazandaran" pageId="10" pageNumber="461">Mazandaran Prov.</collectingRegion>
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||
],
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||
<quantity id="B5459B3DFF965F3ED3C7531F2AA30A30" box="[233,390,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="10" pageNumber="461" unit="m" value="2100.0" valueMax="2500.0" valueMin="1700.0">
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||
<elevation id="F990D1EBFF965F3ED3C7531F2AA30A30" box="[233,390,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="10" pageNumber="461" unit="m" value="2100.0" valueMax="2500.0" valueMin="1700.0">1700–2500 m</elevation>
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||
</quantity>
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||
,
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||
<date id="06031018FF965F3ED2BF531E29220A30" box="[401,519,1375,1401]" pageId="10" pageNumber="461" value="1970-04-28">
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||
<collectingDate id="1647E9F0FF965F3ED2BF531E29220A30" box="[401,519,1375,1401]" pageId="10" pageNumber="461" value="1970-04-28">28.iv.1970</collectingDate>
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||
</date>
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||
(leg.
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||
<collectorName id="DF48530EFF965F3ED169531E29800A30" box="[583,677,1375,1401]" pageId="10" pageNumber="461">Wittmer</collectorName>
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||
,
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<collectorName id="DF48530EFF965F3ED19E531E28170A30" box="[688,818,1375,1401]" pageId="10" pageNumber="461">v. Bothmer</collectorName>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3AA76553FF965F38D3E953C32ABB0C14" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461" type="description">
|
||
<paragraph id="720236D8FF965F3ED3E953C32BEF0AAC" blockId="10.[151,1437,1338,2013]" pageId="10" pageNumber="461">
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||
<emphasis id="40C9EACAFF965F3ED3E953C32A7C0AD5" bold="true" box="[199,345,1410,1436]" pageId="10" pageNumber="461">Description.</emphasis>
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Male. Body slender, elongate, black, with dull shine (
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<figureCitation id="EA862A5DFF965F3ED09153C22F3F0AD4" box="[959,1050,1411,1437]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Fig. 8A</figureCitation>
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). Measurements: Y = 1.47;
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<collectingCountry id="0AAA7648FF965F3ED67B53C22E5C0AD4" box="[1365,1401,1411,1437]" name="Philippines" pageId="10" pageNumber="461">PH</collectingCountry>
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<subScript id="EE39349DFF965F3ED65753D42EA00AEA" attach="left" box="[1401,1413,1429,1443]" fontSize="6" pageId="10" pageNumber="461">w</subScript>
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= 1.44; P
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<subScript id="EE39349DFF965F3ED3C653F82BD10A8E" attach="both" box="[232,244,1465,1479]" fontSize="6" pageId="10" pageNumber="461">w</subScript>
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P
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<subScript id="EE39349DFF965F3ED22D53F82A2D0A8E" attach="left" box="[259,264,1465,1479]" fontSize="6" pageId="10" pageNumber="461">l</subScript>
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= 1.09; E
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<subScript id="EE39349DFF965F3ED25753F82A5B0A8E" attach="both" box="[377,382,1465,1479]" fontSize="6" pageId="10" pageNumber="461">l</subScript>
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E
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<subScript id="EE39349DFF965F3ED2A153F82ABE0A8E" attach="left" box="[399,411,1465,1479]" fontSize="6" pageId="10" pageNumber="461">w</subScript>
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= 1.74; EH
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<subScript id="EE39349DFF965F3ED10F53F829080A8E" attach="left" box="[545,557,1465,1479]" fontSize="6" pageId="10" pageNumber="461">w</subScript>
|
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= 1.94; EP
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<subScript id="EE39349DFF965F3ED18053F8299F0A8E" attach="left" box="[686,698,1465,1479]" fontSize="6" pageId="10" pageNumber="461">w</subScript>
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= 1.34; EP
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<subScript id="EE39349DFF965F3ED01553F828650A8E" attach="left" box="[827,832,1465,1479]" fontSize="6" pageId="10" pageNumber="461">l</subScript>
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= 2.6–2.61. Body length
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<quantity id="B5459B3DFF965F3ED74D53E72FD10A88" box="[1123,1268,1446,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="10" pageNumber="461" unit="mm" value="12.75" valueMax="13.0" valueMin="12.5">12.5–13 mm</quantity>
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, width
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<quantity id="B5459B3DFF965F3ED66653E72BE30AAC" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="10" pageNumber="461" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
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.
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</paragraph>
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||
<paragraph id="720236D8FF965F3ED3E953AE2A1609A8" blockId="10.[151,1437,1338,2013]" pageId="10" pageNumber="461">
|
||
Head (
|
||
<figureCitation id="EA862A5DFF965F3ED23D53AE2A940940" box="[275,433,1519,1545]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Figs 8B, E, F</figureCitation>
|
||
). Anterior margin of epistoma straight (
|
||
<figureCitation id="EA862A5DFF965F3ED05953AE28F50940" box="[887,976,1519,1545]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Fig. 8E</figureCitation>
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). Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head moderately fine and sparse on frons (interpuncture distance near 3 times as long as puncture diameter) and denser on epistoma and genae (interpuncture distance subequal to or near 1.5 times as long as puncture diameter). Ventral side of epicranium coarsely and densely punctured; temples depressed behind eyes in the lower half (
|
||
<figureCitation id="EA862A5DFF965F3ED61A503E2EAA09D0" box="[1332,1423,1663,1689]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Fig. 8B</figureCitation>
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). Antennae comparatively long, with four apical antennomeres, extending beyond base of pronotum, reaching 1/3 of elytral length.
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</paragraph>
|
||
<paragraph id="720236D8FF965F38D3E950AA2E550E54" blockId="10.[151,1437,1338,2013]" lastBlockId="12.[151,1437,151,1185]" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
|
||
Prothorax (
|
||
<figureCitation id="EA862A5DFF965F3ED26B50AA2AC3084C" box="[325,486,1771,1797]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Figs 8B, E–G</figureCitation>
|
||
). Pronotum weakly transverse, with almost equal length and width, widest at middle (
|
||
<figureCitation id="EA862A5DFF965F3ED3B1514E2BDF0860" box="[159,250,1807,1833]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="10" pageNumber="461">Fig. 8E</figureCitation>
|
||
). Lateral margins of pronotum weakly rounded in middle part, straightly narrowed in apical third and widely emarginated in basal quarter; anterior margin trisinuate; base slightly rounded, almost straight, with emargination at middle. Antero-lateral corners not projected, right, pointed at apex; postero-lateral corners right, narrowly rounded at apex. Lateral margins not beaded; anterior margin only with very short bead marginally; disc completely beaded. Disc of pronotum convex, not flattened along lateral margin, without impressions. Puncturation of disc fine and sparse (puncture diameter 2–4 times as short as interpuncture distance), slightly coarser and denser on lateral sides; punctures round (
|
||
<figureCitation id="EA862A5DFF905F38D28A56D6293A0FF8" box="[420,543,151,177]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8E, F</figureCitation>
|
||
). Prosternum with fine and sparse simple puncturation (
|
||
<figureCitation id="EA862A5DFF905F38D7B656D62FD50FF8" box="[1176,1264,151,177]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8B</figureCitation>
|
||
). Outer margin of prothoracic hypomera narrowly flattened only near base. Prohypomera with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background (
|
||
<figureCitation id="EA862A5DFF905F38D72B569E2F450FB0" box="[1029,1120,223,249]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8G</figureCitation>
|
||
). Prosternal process smooth and shiny (only rugose near apex), almost without punctures, slightly convex at apex, not beaded (
|
||
<figureCitation id="EA862A5DFF905F38D7CA57422E410E54" box="[1252,1380,259,285]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8B, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="26C26650FF975F3FD3B9512C2A4A08B8" ID-DOI="http://doi.org/10.5281/zenodo.6785941" ID-Zenodo-Dep="6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1901,1925]" targetBox="[190,1383,184,1872]" targetPageId="11">
|
||
<paragraph id="720236D8FF975F3FD3B9512C2A4A08B8" blockId="11.[151,1436,1901,2033]" pageId="11" pageNumber="462">
|
||
<emphasis id="40C9EACAFF975F3FD3B9512C2A3608CC" bold="true" box="[151,275,1901,1925]" pageId="11" pageNumber="462">FIGURE 8.</emphasis>
|
||
<taxonomicName id="B5BD4D5BFF975F3FD234512C2AF508CC" authority="Nabozhenko, 2022" authorityName="Nabozhenko" authorityYear="2022" box="[282,464,1901,1925]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="arzanovi" status="sp. nov.">
|
||
<emphasis id="40C9EACAFF975F3FD234512C2AF508CC" box="[282,464,1901,1925]" italics="true" pageId="11" pageNumber="462">Euboeus arzanovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="40C9EACAFF975F3FD2F9512C290208CC" bold="true" box="[471,551,1901,1925]" pageId="11" pageNumber="462">
|
||
<taxonomicNameLabel id="5BFA57B1FF975F3FD2F9512C290208CC" box="[471,551,1901,1925]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="720236D8FF905F38D3E957662A6B0EBC" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||
Pterothorax (
|
||
<figureCitation id="EA862A5DFF905F38D27557662AFB0E08" box="[347,478,295,321]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8A, B</figureCitation>
|
||
). Scutellar shield triangle, moderately coarsely punctured. Elytra strongly elongate, widest slightly behind middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (
|
||
<figureCitation id="EA862A5DFF905F38D163572E298C0EC0" box="[589,681,367,393]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8A</figureCitation>
|
||
). Anterior half of mesoventrite strongly wrinkled, pubescent, with more smooth middle (
|
||
<figureCitation id="EA862A5DFF905F38D2A557D22AC40EE4" box="[395,481,403,429]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8B</figureCitation>
|
||
). Mesepisterna coarsely and densely punctured by large foveae; puncturation of mesepimera and metepisterna sparser and finer; metaventrite coarsely and moderately sparsely punctured and pubescent with short setae.
|
||
</paragraph>
|
||
<paragraph id="720236D8FF905F38D3E957BE2A750DE0" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||
Legs (
|
||
<figureCitation id="EA862A5DFF905F38D22057BE2AB50D50" box="[270,400,511,537]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8B, G</figureCitation>
|
||
). Trochanters with short dense reddish setae (but not dense brush), protrochanters with two long setae (
|
||
<figureCitation id="EA862A5DFF905F38D23754622A570D74" box="[281,370,547,573]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8G</figureCitation>
|
||
), meso- and metatrochanters with one long seta. Basal third of femora with the same dense on flex side (
|
||
<figureCitation id="EA862A5DFF905F38D22854062A780D28" box="[262,349,583,609]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Fig. 8B</figureCitation>
|
||
); another surface of femora with sparse puncturation and short setae. Pro- and metatibiae slightly curved, mesotibiae stright. Pro- and mesotarsomeres 1–4 widened, trasverse, pro- and mesotarsomere 1 as wide as apex of protibia.
|
||
</paragraph>
|
||
<paragraph id="720236D8FF905F38D3E954F22E440C70" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||
Abdomen (
|
||
<figureCitation id="EA862A5DFF905F38D26A54F22AD80D84" box="[324,509,691,717]" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8B–D, H–J</figureCitation>
|
||
). Abdominal ventrites with moderately dense, fine puncturation and simple, moderate, short setae (shortly pubescent); ventrite 5 with coarser punctures and longer setae at apex. Genitalia (
|
||
<figureCitation id="EA862A5DFF905F38D63254962BEC0C5C" captionStart="FIGURE 8" captionStartId="11.[151,250,1901,1925]" captionTargetBox="[190,1383,184,1872]" captionTargetId="figure-15@11.[190,1397,181,1877]" captionTargetPageId="11" captionText="FIGURE 8. Euboeus arzanovi sp. nov., habitus, details of structure. A = male, dorsally; B—male, ventrally; C = tegmen, ventrally; D = apical piece of aedeagus, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H = male inner sternite VIII, ventrally; I = spiculum gastrale; J = median lobe of aedeagus with inverted connective membrane, ventrally." figureDoi="http://doi.org/10.5281/zenodo.6785941" httpUri="https://zenodo.org/record/6785941/files/figure.png" pageId="12" pageNumber="463">Figs 8C, D, H–J</figureCitation>
|
||
). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with wide median longitudinal impression; lateral margins straight at middle and straightly narrowed to narrowly rounded apex.
|
||
</paragraph>
|
||
<paragraph id="720236D8FF905F38D3E955022ABB0C14" blockId="12.[151,1437,151,1185]" box="[199,414,835,861]" pageId="12" pageNumber="463">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AA76553FF905F38D3E9552629CA0CEC" pageId="12" pageNumber="463" type="etymology">
|
||
<paragraph id="720236D8FF905F38D3E9552629CA0CEC" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||
<emphasis id="40C9EACAFF905F38D3E955262A6B0CC8" bold="true" box="[199,334,871,897]" pageId="12" pageNumber="463">Etymology.</emphasis>
|
||
The species is named in honour of my first teacher during student times and known specialist on
|
||
<taxonomicName id="B5BD4D5BFF905F38D3B955CA293A0CEC" authority="DrSc. Yury Arzanov" authorityName="DrSc. Yury Arzanov" box="[151,543,907,933]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="family">Curculionidae DrSc. Yury Arzanov</taxonomicName>
|
||
(Rostov-on-Don).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AA76553FF905F38D3E955EF292B0BE8" pageId="12" pageNumber="463" type="diagnosis">
|
||
<paragraph id="720236D8FF905F38D3E955EF292B0BE8" blockId="12.[151,1437,151,1185]" pageId="12" pageNumber="463">
|
||
<emphasis id="40C9EACAFF905F38D3E955EF2AE80C81" bold="true" box="[199,461,942,968]" pageId="12" pageNumber="463">Differential diagnosis.</emphasis>
|
||
The new species is close to
|
||
<taxonomicName id="B5BD4D5BFF905F38D02B55EE284E0C80" authorityName="Nabozhenko" authorityYear="2022" box="[773,875,943,969]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="40C9EACAFF905F38D02B55EE284E0C80" box="[773,875,943,969]" italics="true" pageId="12" pageNumber="463">E. merkli</emphasis>
|
||
</taxonomicName>
|
||
by the structure of the aedeagus (basal piece without rugosity, apical piece with median longitudinal impression) and the narrow, not beaded pronotum with weakly rounded margins, but differs in following characters: the lower half of temples is impressed behind the eye (weakly rounded in
|
||
<taxonomicName id="B5BD4D5BFF905F38D23B525A2A5E0B7C" authorityName="Nabozhenko" authorityYear="2022" box="[277,379,1051,1077]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="40C9EACAFF905F38D23B525A2A5E0B7C" box="[277,379,1051,1077]" italics="true" pageId="12" pageNumber="463">E. merkli</emphasis>
|
||
</taxonomicName>
|
||
); lateral margins of pronotum straight at apical quarter and widely emarginated in basal quarter (regularly rounded in
|
||
<taxonomicName id="B5BD4D5BFF905F38D2A7527E2AD50B10" authorityName="Nabozhenko" authorityYear="2022" box="[393,496,1087,1113]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="40C9EACAFF905F38D2A7527E2AD50B10" box="[393,496,1087,1113]" italics="true" pageId="12" pageNumber="463">E. merkli</emphasis>
|
||
</taxonomicName>
|
||
); prothoracic hypomera with microrugosity and fine sparse smooth punctures (large, coarse, round foveae and smooth interpuncture space between them in
|
||
<taxonomicName id="B5BD4D5BFF905F38D0E252222F130B34" authorityName="Nabozhenko" authorityYear="2022" box="[972,1078,1123,1149]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="40C9EACAFF905F38D0E252222F130B34" box="[972,1078,1123,1149]" italics="true" pageId="12" pageNumber="463">E. merkli</emphasis>
|
||
</taxonomicName>
|
||
); protrochanters with two long setae (one long seta in
|
||
<taxonomicName id="B5BD4D5BFF905F38D2B952C62ADB0BE8" authorityName="Nabozhenko" authorityYear="2022" box="[407,510,1159,1185]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="40C9EACAFF905F38D2B952C62ADB0BE8" box="[407,510,1159,1185]" italics="true" pageId="12" pageNumber="463">E. merkli</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |