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<document id="E74C1CCF07104471EDCA99909AEA0163" ID-DOI="10.11646/zootaxa.1457.1.3" ID-ISSN="1175­5334" ID-Zenodo-Dep="5077414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625630204467" checkinUser="felipe" docAuthor="Webb, M. D. &amp; Viraktamath, C. A." docDate="2007" docId="FA098787A358B039FF3CEE9F8DAF60CE" docLanguage="en" docName="zt01457p055.pdf" docOrigin="Zootaxa 1457 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1457.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Scaphoideus rubroguttatus Matsumura, Nusillard 2007" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="0630FFFFA35BB03CFFABE83F8C3E6104" masterDocTitle="Three new Old World species of the leafhopper genus Scaphoideus Uhler (Hemiptera: Auchenorrhyncha: Cicadellidae)" masterLastPageNumber="55" masterPageNumber="49" pageNumber="52" updateTime="1698993956223" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4A04A257627071A207F4195B489ED619">Three new Old World species of the leafhopper genus Scaphoideus Uhler (Hemiptera: Auchenorrhyncha: Cicadellidae)</mods:title>
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<mods:namePart id="8F7339F624906F7EB7694C2CAC450B95">Webb, M. D.</mods:namePart>
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<mods:namePart id="7A3CDA693A909D70DCC5E79ED016A7AD">Viraktamath, C. A.</mods:namePart>
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<mods:title id="53B4D9906E8859FB465508F797100783">Zootaxa</mods:title>
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<mods:date id="64C9EABBE778E62F60BB47E44A44F271">2007</mods:date>
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<treatment id="FA098787A358B039FF3CEE9F8DAF60CE" ID-DOI="http://doi.org/10.5281/zenodo.5077690" ID-Zenodo-Dep="5077690" LSID="urn:lsid:plazi:treatment:FA098787A358B039FF3CEE9F8DAF60CE" httpUri="http://treatment.plazi.org/id/FA098787A358B039FF3CEE9F8DAF60CE" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52">
<subSubSection id="3ABA651AA358B03FFF3CEE9F8E4667BE" box="[151,632,1696,1722]" pageId="3" pageNumber="52" type="nomenclature">
<paragraph id="721F3691A358B03FFF3CEE9F8E4667BE" blockId="3.[151,632,1696,1757]" box="[151,632,1696,1722]" pageId="3" pageNumber="52">
<heading id="295781FDA358B03FFF3CEE9F8E4667BE" bold="true" box="[151,632,1696,1722]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
<taxonomicName id="B5A04D12A358B03FFF3CEE9F8E4667BE" ID-CoL="79SYV" authority="Matsumura" authorityName="Matsumura, Nusillard" authorityYear="2007" box="[151,632,1696,1722]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="rubroguttatus">
<emphasis id="40D4EA83A358B03FFF3CEE9F8E4667BE" bold="true" box="[151,632,1696,1722]" pageId="3" pageNumber="52">
<emphasis id="40D4EA83A358B03FFF3CEE9F8DE367BE" bold="true" box="[151,477,1696,1722]" italics="true" pageId="3" pageNumber="52">Scaphoideus rubroguttatus</emphasis>
Matsumura
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABA651AA358B03FFF3CEEFC8D5E67D9" box="[151,352,1731,1757]" pageId="3" pageNumber="52" type="description">
<paragraph id="721F3691A358B03FFF3CEEFC8D5E67D9" blockId="3.[151,632,1696,1757]" box="[151,352,1731,1757]" pageId="3" pageNumber="52">
(
<figureCitation id="EA9B2A14A358B03FFF34EEFC8D1367D9" box="[159,301,1731,1757]" captionStart="FIGURES 1016" captionStartId="3.[151,269,1232,1256]" captionTargetBox="[192,1376,204,1211]" captionTargetId="figure-196@3.[167,1405,179,1223]" captionTargetPageId="3" captionText="FIGURES 1016. Scaphoideus species. 1012, S. rubroguttatus, 1011, aedeagus (left lateral and posterior, respectively), 12, style; 1314, S. unimaculata, aedeagus (left lateral and posterior, respectively). 1516, S. quangtriensis, aedeagus, left lateral and posterior view, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5077420" httpUri="https://zenodo.org/record/5077420/files/figure.png" pageId="3" pageNumber="52">Figs 1012</figureCitation>
,
<figureCitation id="EA9B2A14A358B03FFE91EEFC8D6667D9" box="[314,344,1731,1757]" captionStart="FIGURES 1722" captionStartId="4.[151,269,1907,1931]" captionTargetBox="[188,1384,202,1836]" captionTargetId="figure-56@4.[152,1435,157,1881]" captionTargetPageId="4" captionText="FIGURES 1722. Scaphoideus species (habitus). 17, S. unimaculatus (holotype); 18, S. quangtriensis (holotype); 19, S. rubroguttatus; 20, S. dellagiustinai (paratype, France); 21, S. baeiticus (holotype); 22, S. albosignatus (syntype)." figureDoi="http://doi.org/10.5281/zenodo.5077422" httpUri="https://zenodo.org/record/5077422/files/figure.png" pageId="3" pageNumber="52">19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3ABA651AA358B03FFF3CEF2C8E11664E" pageId="3" pageNumber="52" type="reference_group">
<paragraph id="721F3691A358B03FFF3CEF2C8E11664E" blockId="3.[151,1437,1811,1866]" pageId="3" pageNumber="52">
<treatmentCitationGroup id="52B011BFA358B03FFF3CEF2C8E11664E" pageId="3" pageNumber="52">
<treatmentCitation id="F3011080A358B03FFF3CEF2C8E88662E" author="Matsumura, S." box="[151,694,1811,1834]" page="223" pageId="3" pageNumber="52" year="1914">
<taxonomicName id="B5A04D12A358B03FFF3CEF2C8E88662E" ID-CoL="79SYV" authority="Matsumura, 1914: 223" authorityName="Matsumura, Nusillard" authorityPageNumber="223" authorityYear="2007" box="[151,694,1811,1834]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="rubroguttatus">
<emphasis id="40D4EA83A358B03FFF3CEF2C8D89662E" box="[151,439,1811,1834]" italics="true" pageId="3" pageNumber="52">Scaphoideus rubroguttatus</emphasis>
<bibRefCitation id="16314B60A358B03FFE15EF2C8E88662E" author="Matsumura, S." box="[446,694,1811,1834]" pageId="3" pageNumber="52" pagination="165 - 240" refId="ref2990" refString="Matsumura, S. (1914) Die Jassinen und einige neue Acocephalinen japans. Journal of the College of Agriculture, Tohoku Imperial University, 5, 165 - 240." type="journal article" year="1914">Matsumura, 1914: 223</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="F3011080A358B03FFD69EF2C8F87662E" author="Okada, T." box="[706,953,1811,1834]" page="197" pageId="3" pageNumber="52" year="1977">
<bibRefCitation id="16314B60A358B03FFD69EF2C8F87662E" author="Okada, T." box="[706,953,1811,1834]" pageId="3" pageNumber="52" pagination="192 - 198" refId="ref3090" refString="Okada, T. (1977) Redescriptions of four Japanese Scaphoideus species (Homoptera, Cicadellidae). Kontyu, 45, 192 - 198." type="journal article" year="1977">Okada, 1977: 197198</bibRefCitation>
</treatmentCitation>
, figs 2228;
<treatmentCitation id="F3011080A358B03FFBE5EF2C8D35664E" author="Kitbamroong, N. A. &amp; Freytag, P. H." page="28" pageId="3" pageNumber="52" year="1978">
<bibRefCitation id="16314B60A358B03FFBE5EF2C8D35664E" author="Kitbamroong, N. A. &amp; Freytag, P. H." pageId="3" pageNumber="52" pagination="9 - 31" refId="ref2867" refString="Kitbamroong, N. A. &amp; Freytag, P. H. (1978) The species of the genus Scaphoideus (Homoptera: Cicadellidae) found in Thailand, with descriptions of new species. Pacific Insects, 18, 9 - 31." type="journal article" year="1978">Kitbamroong &amp; Freytag, 1978: 2830</bibRefCitation>
</treatmentCitation>
, figs 3738, 135140, 146.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3ABA651AA358B03FFF6DEF40885666C6" pageId="3" pageNumber="52" type="materials_examined">
<paragraph id="721F3691A358B03FFF6DEF40885666C6" blockId="3.[151,1436,1917,1986]" pageId="3" pageNumber="52">
<emphasis id="40D4EA83A358B03FFF6DEF408D82669D" bold="true" box="[198,444,1919,1945]" pageId="3" pageNumber="52">Material examined.</emphasis>
<materialsCitation id="C2C83CCCA358B03FFE6EEF408F5E669D" box="[453,864,1917,1946]" collectingDate="2000-08-16" country="Japan" location="JAPAN" pageId="3" pageNumber="52" specimenCount="1" specimenCount-male="1">
<collectingCountry id="0AB77601A358B03FFE6EEF408E1C669D" box="[453,546,1919,1945]" name="Japan" pageId="3" pageNumber="52">JAPAN</collectingCountry>
:
<specimenCount id="64A6FD18A358B03FFD84EF408E6C669E" box="[559,594,1917,1946]" pageId="3" pageNumber="52" type="male">1♂</specimenCount>
, Honshu,
<date id="061E1051A358B03FFD64EF408F5E669D" box="[719,864,1919,1945]" pageId="3" pageNumber="52" value="2000-08-16">
<collectingDate id="165AE9B9A358B03FFD64EF408F5E669D" box="[719,864,1919,1945]" pageId="3" pageNumber="52" value="2000-08-16">16.viii.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C2C83CCCA358B03FFCC5EF40885D66C6" collectingDate="1990-10-11" collectingDateMax="1993-05-07" collectingDateMin="1990-10-11" collectionCode="BMNH" collectorName="Ryukyus Is &amp; Iriomote &amp; Miyako &amp; Ishigaki &amp; M. Hayashi. &amp; M. Hayashi" country="Japan" pageId="3" pageNumber="52" specimenCount="4" specimenCount-male="4">
<specimenCount id="64A6FD18A358B03FFCC5EF408FAF669E" box="[878,913,1917,1946]" pageId="3" pageNumber="52" type="male">4♂</specimenCount>
, 19,
<collectorName id="DF555347A358B03FFC7CEF408862669D" box="[983,1116,1919,1945]" pageId="3" pageNumber="52">Ryukyus Is</collectorName>
(
<collectorName id="DF555347A358B03FFBC6EF4088EA669D" box="[1133,1236,1919,1945]" pageId="3" pageNumber="52">Iriomote</collectorName>
<date id="061E1051A358B03FFB76EF40896D669D" box="[1245,1363,1919,1945]" pageId="3" pageNumber="52" value="1990-10-11">
<collectingDate id="165AE9B9A358B03FFB76EF40896D669D" box="[1245,1363,1919,1945]" pageId="3" pageNumber="52" value="1990-10-11">11.x.1990</collectingDate>
</date>
,
<collectorName id="DF555347A358B03FFACAEF408CFF66C6" pageId="3" pageNumber="52">Miyako</collectorName>
<date id="061E1051A358B03FFF63EF978D7B66C6" box="[200,325,1960,1986]" pageId="3" pageNumber="52" value="1992-11-22">
<collectingDate id="165AE9B9A358B03FFF63EF978D7B66C6" box="[200,325,1960,1986]" pageId="3" pageNumber="52" value="1992-11-22">22.xi.1992</collectingDate>
</date>
and
<collectorName id="DF555347A358B03FFED4EF978DE366C6" box="[383,477,1960,1986]" pageId="3" pageNumber="52">Ishigaki</collectorName>
<date id="061E1051A358B03FFE4FEF978E7566C6" box="[484,587,1960,1986]" pageId="3" pageNumber="52" value="1993-05-07">
<collectingDate id="165AE9B9A358B03FFE4FEF978E7566C6" box="[484,587,1960,1986]" pageId="3" pageNumber="52" value="1993-05-07">7.v.1993</collectingDate>
</date>
),
<collectorName id="DF555347A358B03FFDCAEF978ECC66C6" box="[609,754,1960,1986]" pageId="3" pageNumber="52">M. Hayashi.</collectorName>
(coll.
<collectorName id="DF555347A358B03FFC96EF978FFB66C6" box="[829,965,1960,1986]" pageId="3" pageNumber="52">M. Hayashi</collectorName>
and
<collectionCode id="14B1AE54A358B03FFC55EF97886566C6" box="[1022,1115,1960,1986]" pageId="3" pageNumber="52">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="26DF6619A35FB038FF3CEF4C897666A9" ID-DOI="http://doi.org/10.5281/zenodo.5077422" ID-Zenodo-Dep="5077422" httpUri="https://zenodo.org/record/5077422/files/figure.png" pageId="4" pageNumber="53" startId="4.[151,269,1907,1931]" targetBox="[188,1384,202,1836]" targetPageId="4">
<paragraph id="721F3691A35FB038FF3CEF4C897666A9" blockId="4.[151,1436,1907,1966]" pageId="4" pageNumber="53">
<emphasis id="40D4EA83A35FB038FF3CEF4C8D60668F" bold="true" box="[151,350,1907,1931]" pageId="4" pageNumber="53">FIGURES 1722.</emphasis>
<taxonomicName id="B5A04D12A35FB038FECFEF4B8E01668F" box="[356,575,1908,1931]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="40D4EA83A35FB038FECFEF4B8DD5668F" box="[356,491,1908,1931]" italics="true" pageId="4" pageNumber="53">Scaphoideus</emphasis>
species
</taxonomicName>
(habitus). 17,
<taxonomicName id="B5A04D12A35FB038FD71EF4B8FBA668F" authorityName="Webb &amp; Viraktamath" authorityYear="2007" box="[730,900,1908,1931]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="unimaculatus">
<emphasis id="40D4EA83A35FB038FD71EF4B8ED9668F" box="[730,743,1908,1931]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FD5FEF4B8FBA668F" box="[756,900,1908,1931]" italics="true" pageId="4" pageNumber="53">unimaculatus</emphasis>
</taxonomicName>
(holotype); 18,
<taxonomicName id="B5A04D12A35FB038FB9BEF4B88E2668F" authorityName="Webb &amp; Viraktamath" authorityYear="2007" box="[1072,1244,1908,1931]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="quangtriensis">
<emphasis id="40D4EA83A35FB038FB9BEF4B8803668F" box="[1072,1085,1908,1931]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FBE0EF4B88E2668F" box="[1099,1244,1908,1931]" italics="true" pageId="4" pageNumber="53">quangtriensis</emphasis>
</taxonomicName>
(holotype); 19,
<taxonomicName id="B5A04D12A35FB038FA23EF4B8D1766AA" authorityName="Matsumura, Nusillard" authorityYear="2007" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="rubroguttatus">
<emphasis id="40D4EA83A35FB038FA23EF4B89AB668F" box="[1416,1429,1908,1931]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FF3CEFA88D1766AA" box="[151,297,1943,1966]" italics="true" pageId="4" pageNumber="53">rubroguttatus</emphasis>
</taxonomicName>
; 20,
<taxonomicName id="B5A04D12A35FB038FEF5EFA88E3266AA" authorityName="Webb &amp; Viraktamath" authorityYear="2007" box="[350,524,1942,1966]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="dellagiustinai">
<emphasis id="40D4EA83A35FB038FEF5EFA88D5566AA" box="[350,363,1943,1966]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FED2EFA88E3266AA" box="[377,524,1943,1966]" italics="true" pageId="4" pageNumber="53">dellagiustinai</emphasis>
</taxonomicName>
(paratype, France); 21,
<taxonomicName id="B5A04D12A35FB038FCA7EFA88FB866AA" box="[780,902,1942,1966]" class="Insecta" family="Cicadellidae" genus="Scaphoideus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="baeiticus">
<emphasis id="40D4EA83A35FB038FCA7EFA88F2766AA" box="[780,793,1943,1966]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FC8DEFA88FB866AA" box="[806,902,1943,1966]" italics="true" pageId="4" pageNumber="53">baeiticus</emphasis>
</taxonomicName>
(holotype); 22,
<taxonomicName id="B5A04D12A35FB038FB98EFA888EB66AA" box="[1075,1237,1942,1966]" class="Insecta" family="Cicadellidae" genus="Platyretus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="albosignatus">
<emphasis id="40D4EA83A35FB038FB98EFA8887E66AA" box="[1075,1088,1943,1966]" italics="true" pageId="4" pageNumber="53">S</emphasis>
.
<emphasis id="40D4EA83A35FB038FBE6EFA888EB66AA" box="[1101,1237,1943,1966]" italics="true" pageId="4" pageNumber="53">albosignatus</emphasis>
</taxonomicName>
(syntype).
</paragraph>
</caption>
<subSubSection id="3ABA651AA35EB039FF6DE8A88DAF60CE" pageId="5" pageNumber="54" type="discussion">
<paragraph id="721F3691A35EB039FF6DE8A88DAF60CE" blockId="5.[151,1436,151,458]" pageId="5" pageNumber="54">
<emphasis id="40D4EA83A35EB039FF6DE8A88D0161B5" bold="true" box="[198,319,151,177]" pageId="5" pageNumber="54">Remarks.</emphasis>
There is some uncertainty as to the identity of this species, originally described from a single female from
<collectingCountry id="0AB77601A35EB039FE85E8FF8DB661DE" box="[302,392,192,218]" name="Taiwan" pageId="5" pageNumber="54">Taiwan</collectingCountry>
. We take the identity of the species from
<bibRefCitation id="16314B60A35EB039FCC1E8FF883561DE" author="Okada, T." box="[874,1035,192,218]" pageId="5" pageNumber="54" pagination="192 - 198" refId="ref3090" refString="Okada, T. (1977) Redescriptions of four Japanese Scaphoideus species (Homoptera, Cicadellidae). Kontyu, 45, 192 - 198." type="journal article" year="1977">Okada (1977)</bibRefCitation>
as first reviser, who examined the female
<typeStatus id="AD1B8833A35EB039FF46E8D78D216006" box="[237,287,232,258]" pageId="5" pageNumber="54">type</typeStatus>
and figured a male specimen from
<collectingCountry id="0AB77601A35EB039FD15E8D78F3A6006" box="[702,772,232,258]" name="Japan" pageId="5" pageNumber="54">Japan</collectingCountry>
. Specimens from the Ryukyus Islands (near Taiwan) figured here, also match this identity. The species was also treated by
<bibRefCitation id="16314B60A35EB039FC1EE92F8917602E" author="Kitbamroong, N. A. &amp; Freytag, P. H." box="[949,1321,272,298]" pageId="5" pageNumber="54" pagination="9 - 31" refId="ref2867" refString="Kitbamroong, N. A. &amp; Freytag, P. H. (1978) The species of the genus Scaphoideus (Homoptera: Cicadellidae) found in Thailand, with descriptions of new species. Pacific Insects, 18, 9 - 31." type="journal article" year="1978">Kitbamroong &amp; Freytag (1978)</bibRefCitation>
who also examined and figured the female
<typeStatus id="AD1B8833A35EB039FD8FE9078E646056" box="[548,602,312,338]" pageId="5" pageNumber="54">type</typeStatus>
. However, the male specimen they figured from
<collectingCountry id="0AB77601A35EB039FB37E907893A6056" box="[1180,1284,312,338]" name="Thailand" pageId="5" pageNumber="54">Thailand</collectingCountry>
could represent a different species as, although the figures lack detail, the aedeagus looks different to that figured here. They also recorded the species from
<collectingCountry id="0AB77601A35EB039FDE0E9B78E9060A6" box="[587,686,392,418]" name="Vietnam" pageId="5" pageNumber="54">Vietnam</collectingCountry>
and the
<collectingCountry id="0AB77601A35EB039FCBEE9B78FA360A6" box="[789,925,392,418]" name="Philippines" pageId="5" pageNumber="54">Philippines</collectingCountry>
. Our specimen from
<collectingCountry id="0AB77601A35EB039FB3DE9B788C760A6" box="[1174,1273,392,418]" name="Vietnam" pageId="5" pageNumber="54">Vietnam</collectingCountry>
is a new species described below.
</paragraph>
</subSubSection>
</treatment>
</document>