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<document id="F371FBD3A79D56501FAE8E24D0B66258" ID-DOI="10.11646/zootaxa.5020.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5223173" ID-ZooBank="4FF882DF-334F-49C8-A576-4192B5F2654C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629388374973" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2021" docId="FA08E75DAA138072FDC0325592A36A29" docLanguage="en" docName="zootaxa.5020.2.1.pdf" docOrigin="Zootaxa 5020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habetia quatrispina Ingrisch 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="245" masterDocId="06319F25AA3F805EFD57366B91686D58" masterDocTitle="Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="245" updateTime="1699071125630" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="87388D0310722B2A111CD7FB271EBD69">Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)</mods:title>
</mods:titleInfo>
<mods:name id="6A686D0D61E13A23F1E4A2D1DF280693" type="personal">
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<mods:namePart id="3937A20DF798C7D99C34D2978E95B774">Ingrisch, Sigfrid</mods:namePart>
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<mods:title id="C2BAD6D0154F74332B823927DE83A59C">Zootaxa</mods:title>
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<mods:date id="0CAAB906B9DD850D1011F10ADB0C55A0">2021</mods:date>
<mods:detail id="37B830D920307C220948CA9F38EBB21A" type="pubDate">
<mods:number id="1F65776FB80A7A7FAABEB88888A70557">2021-08-12</mods:number>
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<mods:number id="E878A06559D32CFC5CB51298170BE470">5020</mods:number>
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<mods:identifier id="54A6961070F16860DD4A42CBFDB4A18C" type="DOI">10.11646/zootaxa.5020.2.1</mods:identifier>
<mods:identifier id="0977BABB82B003532D56EB39AA52C7A2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="FA08E75DAA138072FDC0325592A36A29" ID-DOI="http://doi.org/10.5281/zenodo.5223255" ID-Zenodo-Dep="5223255" LSID="urn:lsid:plazi:treatment:FA08E75DAA138072FDC0325592A36A29" httpUri="http://treatment.plazi.org/id/FA08E75DAA138072FDC0325592A36A29" lastPageNumber="245" pageId="44" pageNumber="245">
<subSubSection id="3ABB05C0AA138072FDC0325590866901" box="[151,494,1086,1113]" pageId="44" pageNumber="245" type="nomenclature">
<paragraph id="721E564BAA138072FDC0325590866901" blockId="44.[151,502,1086,1150]" box="[151,494,1086,1113]" pageId="44" pageNumber="245">
<heading id="2956E127AA138072FDC0325590866901" bold="true" box="[151,494,1086,1113]" fontSize="11" level="1" pageId="44" pageNumber="245" reason="1">
<emphasis id="40D58A59AA138072FDC0325590866901" bold="true" box="[151,494,1086,1113]" pageId="44" pageNumber="245">
<taxonomicName id="B5A12DC8AA138072FDC0325590E26901" ID-CoL="8QJCL" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[151,394,1086,1113]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="245" phylum="Arthropoda" rank="species" species="quatrispina" status="sp. nov.">
<emphasis id="40D58A59AA138072FDC0325590E26901" bold="true" box="[151,394,1086,1113]" italics="true" pageId="44" pageNumber="245">Habetia quatrispina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BE63722AA138072FCC6325490866901" box="[401,494,1087,1113]" pageId="44" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA138072FDC03208909E6926" box="[151,502,1123,1150]" pageId="44" pageNumber="245" type="description">
<paragraph id="721E564BAA138072FDC03208909E6926" blockId="44.[151,502,1086,1150]" box="[151,502,1123,1150]" pageId="44" pageNumber="245">
<figureCitation id="EA9A4ACEAA138072FDC0320891816925" box="[151,233,1123,1149]" captionStart="FIGURE 2" captionStartId="16.[151,250,1848,1872]" captionTargetBox="[151,1436,358,1825]" captionTargetId="figure-102@16.[151,1436,345,1825]" captionTargetPageId="16" captionText="FIGURE 2. Habitus of Habetia species with moderately long ovipositor (AJ, females; K, male): A, H. bivittata sp. nov. (Chimbu Valley); B, H. dentata sp. nov. (Saidor); C, H. elevata sp. nov. (Salamaua); D, H. pinnigera sp. nov. (Madang); E, H. kondiu sp. nov. (Mendi); F, H. simbai sp. nov. (Simbai); G, H. wau sp. nov. (Nami Creek); H, H. sororcula sp. nov. (Sentani); I, H. nigricauda sp. nov. (Popondetta); J, H. quatrispina sp. nov. (Pionierlager); K, H. pilleata sp. nov. holotype (Wau 1200m). All scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223177" httpUri="https://zenodo.org/record/5223177/files/figure.png" pageId="44" pageNumber="245">Figs 2J</figureCitation>
,
<figureCitation id="EA9A4ACEAA138072FDA2320890796925" box="[245,273,1123,1149]" captionStart="FIGURE 4" captionStartId="18.[151,250,1811,1835]" captionTargetBox="[151,1436,184,1787]" captionTargetId="figure-19@18.[151,1436,181,1787]" captionTargetPageId="18" captionText="FIGURE 4. Frons of Habetia species: AC, H. bivittata sp. nov., variation of the black marks (A Goroka male, B Mt. Wilhelm male, C Keltiga female); D, H. dentata sp. nov. female (Saidor); E, H. kondiu sp. nov. female (Kubor Range); F, H. pinnigera sp. nov. female (Madang); G, H. lalibu sp. nov. male (Lalibu); H, H. simbai sp. nov. female (Simbai Valley); I, H. nigricauda sp. nov. female (Popondetta); J, H. quatrispina sp. nov. female (Pionierlager); K, H. wau sp. nov. male (Nami Creek); L, H. sororcula sp. nov. female (Ifar). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5223181" httpUri="https://zenodo.org/record/5223181/files/figure.png" pageId="44" pageNumber="245">4J</figureCitation>
,
<figureCitation id="EA9A4ACEAA138072FC4B320890556925" box="[284,317,1123,1149]" captionStart="FIGURE 6" captionStartId="22.[151,250,1863,1887]" captionTargetBox="[151,1436,417,1839]" captionTargetId="figure-132@22.[151,1436,417,1840]" captionTargetPageId="22" captionText="FIGURE 6. Female subgenital plate of Habetia species in about ventral view: AB, H. bivittata sp. nov. (A Chimbu Valley, B Moife); C, H. nigricauda sp. nov. (Popondetta); D, H. wau sp. nov. (Upper Watut); E, H. simbai sp. nov. (Simbai Valley); F, H. dentata sp. nov. (Saidor); G-H, H. kondiu (G Mt Hagen, H Mendi); I, H. pinnigera sp. nov. (Madang); J, H. elevata sp. nov. (Salamaua); K, H. sororcula sp. nov. (Ifar); L, H. quatrispina sp. nov. (Pionierlager), the arrow points at the lateral spine-like projection. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223185" httpUri="https://zenodo.org/record/5223185/files/figure.png" pageId="44" pageNumber="245">6L</figureCitation>
,
<figureCitation id="EA9A4ACEAA138072FC1E320890046925" box="[329,364,1123,1149]" captionStart="FIGURE 7" captionStartId="23.[151,250,1785,1809]" captionTargetBox="[151,1436,182,1759]" captionTargetId="figure-15@23.[151,1436,181,1761]" captionTargetPageId="23" captionText="FIGURE 7. Female subgenital plate of Habetia species in lateral (BC, EI, MN) and oblique lateral (A, D, JL, O) view: AB, H. bivittata sp. nov. (A Aiyura, B Mt Wilhelm); C, H. quatrispina sp. nov. (Pionierlager); D, H. kondiu sp. nov. (Kubor Range); E, H. simbai sp. nov. (Simbai Valley); FG, H. wau sp. nov. (F Nami Creak, G Wau); H, H. sororcula sp. nov. (Ifar); IJ, H. pinnigera sp. nov. (Madang); K, H. multispinulosa Griffini, 1908 (Rigo Dorom); LM, H. elevata sp. nov. (Salamaua); NO, H. nigricauda sp. nov. (Popondetta). The arrows in A, C and H point at the tip of the dorso-lateral lobes. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223187" httpUri="https://zenodo.org/record/5223187/files/figure.png" pageId="44" pageNumber="245">7C</figureCitation>
, Map 1(16).
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA138072FDC032C092EF69B1" pageId="44" pageNumber="245" type="materials_examined">
<paragraph id="721E564BAA138072FDC032C092EF69B1" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
<materialsCitation id="C2C95C16AA138072FDC032C092EF69B1" collectingDate="1912-04-01" collectingDateMax="1912-06-30" collectingDateMin="1912-04-01" collectionCode="ZMB" collectorName="Burgers &amp; Museum &amp; Naturkunde &amp; Berlin" country="Papua New Guinea" latitude="-4.3166666" location="Pionierlager" longLatPrecision="1306" longitude="141.91667" pageId="44" pageNumber="245" specimenCount="1" stateProvince="East Sepik" typeStatus="holotype">
<emphasis id="40D58A59AA138072FDC032C0901F699C" bold="true" box="[151,375,1194,1221]" pageId="44" pageNumber="245">
<typeStatus id="AD1AE8E9AA138072FDC032C0906C699D" box="[151,260,1195,1221]" pageId="44" pageNumber="245" type="holotype">Holotype</typeStatus>
(female):
</emphasis>
<collectingCountry id="0AB616DBAA138072FCD732C09335699D" box="[384,605,1195,1221]" name="Papua New Guinea" pageId="44" pageNumber="245">Papua New Guinea</collectingCountry>
,
<collectingRegion id="B06598A9AA138072FF3E32C0938D699D" box="[617,741,1195,1221]" country="Papua New Guinea" name="East Sepik" pageId="44" pageNumber="245">East Sepik</collectingRegion>
,
<location id="777E0090AA138072FFA632C09214699D" LSID="urn:lsid:plazi:treatment:FA08E75DAA138072FDC0325592A36A29:777E0090AA138072FFA632C09214699D" box="[753,892,1195,1221]" country="Papua New Guinea" latitude="-4.3166666" longLatPrecision="1306" longitude="141.91667" name="Pionierlager" pageId="44" pageNumber="245" stateProvince="East Sepik">Pionierlager</location>
, (
<geoCoordinate id="1795308CAA138072FEC632C09289699D" box="[913,993,1195,1221]" degrees="4" direction="south" minutes="19" orientation="latitude" pageId="44" pageNumber="245" precision="925" value="-4.3166666">4°19S</geoCoordinate>
,
<geoCoordinate id="1795308CAA138072FEB932C19535699D" box="[1006,1117,1194,1221]" degrees="141" direction="east" minutes="55" orientation="longitude" pageId="44" pageNumber="245" precision="925" value="141.91667">141°55E</geoCoordinate>
),
<date id="061F708BAA138072F92432C09443699D" box="[1139,1323,1195,1221]" pageId="44" pageNumber="245" value="1912-04-01" valueMax="1912-06-30" valueMin="1912-04-01">
<collectingDate id="165B8963AA138072F92432C09443699D" box="[1139,1323,1195,1221]" pageId="44" pageNumber="245" value="1912-04-01" valueMax="1912-06-30" valueMin="1912-04-01">1.iv.30.vi.1912</collectingDate>
</date>
, KAE 52, leg.
<collectorName id="DF54339DAA138072FD9032A4907669B1" box="[199,286,1231,1257]" pageId="44" pageNumber="245">Bürgers</collectorName>
—depository:
<collectorName id="DF54339DAA138072FC9632A4934B69B1" box="[449,547,1231,1257]" pageId="44" pageNumber="245">Museum</collectorName>
für
<collectorName id="DF54339DAA138072FF0632A493B069B1" box="[593,728,1231,1257]" pageId="44" pageNumber="245">Naturkunde</collectorName>
,
<collectorName id="DF54339DAA138072FFB432A4924169B1" box="[739,809,1231,1257]" pageId="44" pageNumber="245">Berlin</collectorName>
(
<collectionCode id="14B0CE8EAA138072FE6E32A4921369B1" box="[825,891,1231,1257]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="44" pageNumber="245" type="Museum">ZMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA138072FD9032999454680D" pageId="44" pageNumber="245" type="diagnosis">
<paragraph id="721E564BAA138072FD9032999454680D" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
<emphasis id="40D58A59AA138072FD903299902A6854" bold="true" box="[199,322,1266,1292]" pageId="44" pageNumber="245">Diagnosis.</emphasis>
The new species is unique for the possession of four spine-like projections of the female subgenital plate: one pair from the end of the upcurved baso-lateral areas, and the other pair from the end of the central disc. Moreover, the dorsal ovipositor valves have the dorsal margin markedly convex but not strongly widened.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA138072FD90333593D26A71" pageId="44" pageNumber="245" type="description">
<paragraph id="721E564BAA138072FD90333592A66899" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
<emphasis id="40D58A59AA138072FD90333590316820" bold="true" box="[199,345,1374,1400]" pageId="44" pageNumber="245">Description.</emphasis>
Coloration of face uniformly pale, of general color; mandibles of same color, only at, in situ hidden, tip black; antennal scrobae light to medium brown (
<figureCitation id="EA9A4ACEAA138072FE6A33E892E668C5" box="[829,910,1411,1437]" captionStart="FIGURE 4" captionStartId="18.[151,250,1811,1835]" captionTargetBox="[151,1436,184,1787]" captionTargetId="figure-19@18.[151,1436,181,1787]" captionTargetPageId="18" captionText="FIGURE 4. Frons of Habetia species: AC, H. bivittata sp. nov., variation of the black marks (A Goroka male, B Mt. Wilhelm male, C Keltiga female); D, H. dentata sp. nov. female (Saidor); E, H. kondiu sp. nov. female (Kubor Range); F, H. pinnigera sp. nov. female (Madang); G, H. lalibu sp. nov. male (Lalibu); H, H. simbai sp. nov. female (Simbai Valley); I, H. nigricauda sp. nov. female (Popondetta); J, H. quatrispina sp. nov. female (Pionierlager); K, H. wau sp. nov. male (Nami Creek); L, H. sororcula sp. nov. female (Ifar). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5223181" httpUri="https://zenodo.org/record/5223181/files/figure.png" pageId="44" pageNumber="245">Fig. 4J</figureCitation>
). Femora with the following number of spines on ventral margins: (1) a 6, p 78; (2) a 78, p 5; (3) a 13, p 1315 (n=1).
</paragraph>
<paragraph id="721E564BAA138072FD9033A0901068BD" blockId="44.[151,1437,1194,1905]" box="[199,376,1483,1509]" pageId="44" pageNumber="245">Male unknown.</paragraph>
<paragraph id="721E564BAA138072FD90338495A06BB9" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
Female. Subgenital plate with convex ventral surface but in lateral areas curved more strongly dorsad; lateral margins convex behind basal area, the marginal lateral areas divide from the disc and form a pair of stout, rounded projections with approaching margins and acute tip; in the further curse the lateral margins then run in a bow to the somewhat elevated central area; the central area divides at end into a pair of apical, spine-like projections that are separated by a wide and straight interspace with subtruncate bottom (
<figureCitation id="EA9A4ACEAA138072FEC3301492856BC1" box="[916,1005,1663,1689]" captionStart="FIGURE 6" captionStartId="22.[151,250,1863,1887]" captionTargetBox="[151,1436,417,1839]" captionTargetId="figure-132@22.[151,1436,417,1840]" captionTargetPageId="22" captionText="FIGURE 6. Female subgenital plate of Habetia species in about ventral view: AB, H. bivittata sp. nov. (A Chimbu Valley, B Moife); C, H. nigricauda sp. nov. (Popondetta); D, H. wau sp. nov. (Upper Watut); E, H. simbai sp. nov. (Simbai Valley); F, H. dentata sp. nov. (Saidor); G-H, H. kondiu (G Mt Hagen, H Mendi); I, H. pinnigera sp. nov. (Madang); J, H. elevata sp. nov. (Salamaua); K, H. sororcula sp. nov. (Ifar); L, H. quatrispina sp. nov. (Pionierlager), the arrow points at the lateral spine-like projection. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223185" httpUri="https://zenodo.org/record/5223185/files/figure.png" pageId="44" pageNumber="245">Figs 6L</figureCitation>
,
<figureCitation id="EA9A4ACEAA138072FEAF301495746BC1" box="[1016,1052,1663,1689]" captionStart="FIGURE 7" captionStartId="23.[151,250,1785,1809]" captionTargetBox="[151,1436,182,1759]" captionTargetId="figure-15@23.[151,1436,181,1761]" captionTargetPageId="23" captionText="FIGURE 7. Female subgenital plate of Habetia species in lateral (BC, EI, MN) and oblique lateral (A, D, JL, O) view: AB, H. bivittata sp. nov. (A Aiyura, B Mt Wilhelm); C, H. quatrispina sp. nov. (Pionierlager); D, H. kondiu sp. nov. (Kubor Range); E, H. simbai sp. nov. (Simbai Valley); FG, H. wau sp. nov. (F Nami Creak, G Wau); H, H. sororcula sp. nov. (Ifar); IJ, H. pinnigera sp. nov. (Madang); K, H. multispinulosa Griffini, 1908 (Rigo Dorom); LM, H. elevata sp. nov. (Salamaua); NO, H. nigricauda sp. nov. (Popondetta). The arrows in A, C and H point at the tip of the dorso-lateral lobes. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223187" httpUri="https://zenodo.org/record/5223187/files/figure.png" pageId="44" pageNumber="245">7C</figureCitation>
). Ovipositor only little longer than body but for one third longer than the hind femur, laterally compressed; behind basal constriction with dorsal margin convex and moderately elevated around mid-length, in apical area straight and narrow (
<figureCitation id="EA9A4ACEAA138072F93C30AC95D46BB9" box="[1131,1212,1735,1761]" captionStart="FIGURE 2" captionStartId="16.[151,250,1848,1872]" captionTargetBox="[151,1436,358,1825]" captionTargetId="figure-102@16.[151,1436,345,1825]" captionTargetPageId="16" captionText="FIGURE 2. Habitus of Habetia species with moderately long ovipositor (AJ, females; K, male): A, H. bivittata sp. nov. (Chimbu Valley); B, H. dentata sp. nov. (Saidor); C, H. elevata sp. nov. (Salamaua); D, H. pinnigera sp. nov. (Madang); E, H. kondiu sp. nov. (Mendi); F, H. simbai sp. nov. (Simbai); G, H. wau sp. nov. (Nami Creek); H, H. sororcula sp. nov. (Sentani); I, H. nigricauda sp. nov. (Popondetta); J, H. quatrispina sp. nov. (Pionierlager); K, H. pilleata sp. nov. holotype (Wau 1200m). All scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223177" httpUri="https://zenodo.org/record/5223177/files/figure.png" pageId="44" pageNumber="245">Fig. 2J</figureCitation>
).
</paragraph>
<paragraph id="721E564BAA138072FD90308093D26A71" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
Measurements (
<specimenCount id="64A79DC2AA138072FCD6308090816A5E" box="[385,489,1771,1798]" pageId="44" pageNumber="245" type="female">1 female</specimenCount>
).—Body w/o wings: 30; pronotum: 8; tegmen: 27; hind femur: 22; antenna: 90; ovipositor length: 33; ovipositor height:
<quantity id="B559FBAEAA138072FF0A316493DE6A71" box="[605,694,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="44" pageNumber="245" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA138072FD90315892A36A29" pageId="44" pageNumber="245" type="etymology">
<paragraph id="721E564BAA138072FD90315892A36A29" blockId="44.[151,1437,1194,1905]" pageId="44" pageNumber="245">
<emphasis id="40D58A59AA138072FD90315890266A15" bold="true" box="[199,334,1843,1869]" pageId="44" pageNumber="245">Etymology.</emphasis>
The new species is named for the presence of four spine-like projections of the female subgenital plate, from the Latin words
<emphasis id="40D58A59AA138072FC87313C93C36A28" box="[464,683,1879,1904]" italics="true" pageId="44" pageNumber="245">quaternio, quattuor</emphasis>
= four and
<emphasis id="40D58A59AA138072FE79313C92036A28" box="[814,875,1879,1904]" italics="true" pageId="44" pageNumber="245">spina</emphasis>
= spine.
</paragraph>
</subSubSection>
</treatment>
</document>