542 lines
70 KiB
XML
542 lines
70 KiB
XML
<document id="CDA518C44AABECD99810D7E98CA0855B" ID-CLB-Dataset="306850" ID-DOI="10.13133/2284-4880/567" ID-GBIF-Dataset="bb480e48-e028-4025-9974-1226184c5839" ID-ISSN="2284-4880" ID-Zenodo-Dep="12763624" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1721241701966" checkinUser="felipe" docAuthor="Perissinotto, Renzo, Beinhundner, Gerhard & Strümpher, Werner" docDate="2021" docId="FA0787D9FFF0FFABFCDDF8FDE1C52730" docLanguage="en" docName="FragEntomol.53.2.367-376.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/567" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.2:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="2" docTitle="Anoplocheilus (Anoplocheilus) clarki Perissinotto, Beinhundner & Strümpher, 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="370" masterDocId="063EFFA1FFF1FFA8FFEDFFE9E270231C" masterDocTitle="Description of a new species of Anoplocheilus MacLeay, 1838 from South Africa, with review of the genus (Coleoptera: Scarabaeidae, Cetoniinae)" masterLastPageNumber="376" masterPageNumber="367" pageNumber="368" updateTime="1734980332779" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="0FFA9C379007FC4CD0164960817BD977" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="80B7548A15C32257E454D162DAD0DB98">
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<mods:title id="EA2E1BE97B6D7D6FC73B82CED8BE48C0">Description of a new species of Anoplocheilus MacLeay, 1838 from South Africa, with review of the genus (Coleoptera: Scarabaeidae, Cetoniinae)</mods:title>
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<mods:name id="BAB520521A52325B931CEC2AD804F776" type="personal">
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<mods:roleTerm id="D7C57F0FACEF7862D29F9A3FD1AACAF4">Author</mods:roleTerm>
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<mods:namePart id="E85A76CA6866AF9B742290502843167B">Perissinotto, Renzo</mods:namePart>
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<mods:affiliation id="AD5AF53BE2682CEFA8D4AE13442ED547">Coastal and Marine Research (CMR), Nelson Mandela University - P. O. Box 77000, Gqeberha 6031, South Africa - renzo. perissinotto @ mandela. ac. za</mods:affiliation>
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<mods:nameIdentifier id="406D61B8759AFF528202EA21BA8594F1" type="email">renzo.perissinotto@mandela.ac.za</mods:nameIdentifier>
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<mods:namePart id="9516146DED905E0C75BFB2BC8A0B9E19">Beinhundner, Gerhard</mods:namePart>
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<mods:affiliation id="B2CBAB05A33695D3CE98B38F730E4FE8">Am Steigberg 25, D- 97502 Euerbach, Germany - g. beinhundner @ t-online. de</mods:affiliation>
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<mods:nameIdentifier id="03DCB9D233D67EC5F077EEAF3F7F2FC6" type="email">g.beinhundner@t-online.de</mods:nameIdentifier>
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<mods:namePart id="3B9CC35C4233B2739B9B18CDD850F9F6">Strümpher, Werner</mods:namePart>
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<mods:affiliation id="FE8064B92EC64D1D6DC3F1487BFDFF00">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa - strumpher @ ditsong. org. za</mods:affiliation>
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<mods:nameIdentifier id="133BDEAD2C60DC9C0DE2D076B26B5FAA" type="email">strumpher@ditsong.org.za</mods:nameIdentifier>
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<mods:title id="AF53E54FC6BB68E3238BA7D320DD3F33">Fragmenta entomologica</mods:title>
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<mods:date id="6C1C2EFBBCD6B8E12A612BBB02F7BDE6">2021</mods:date>
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<mods:number id="8D73D7C480FD767FC535579F25A3297F">2021-11-30</mods:number>
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<mods:number id="656C9705365E934645A0CEC21E9F0F0B">53</mods:number>
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<treatment id="FA0787D9FFF0FFABFCDDF8FDE1C52730" ID-DOI="http://doi.org/10.5281/zenodo.14549193" ID-Zenodo-Dep="14549193" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730" httpUri="http://treatment.plazi.org/id/FA0787D9FFF0FFABFCDDF8FDE1C52730" lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Cetoniinae">
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<subSubSection id="3AB46544FFF0FFA9FCDDF8FDE7EB2430" box="[816,1435,1812,1836]" pageId="1" pageNumber="368" type="nomenclature">
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<paragraph id="721136CFFFF0FFA9FCDDF8FDE7EB2430" blockId="1.[815,1437,1812,1932]" box="[816,1435,1812,1836]" pageId="1" pageNumber="368">
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<taxonomicName id="B5AE4D4CFFF0FFA9FCDDF8FDE6BB2430" ID-CoL="8SSRJ" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[816,1227,1812,1836]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov." subGenus="Anoplocheilus">
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<emphasis id="40DAEADDFFF0FFA9FCDDF8FDE6BB2430" bold="true" box="[816,1227,1812,1836]" pageId="1" pageNumber="368">
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<emphasis id="40DAEADDFFF0FFA9FCDDF8FDE1BF2430" bold="true" box="[816,975,1812,1836]" italics="true" pageId="1" pageNumber="368">Anoplocheilus</emphasis>
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(
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<emphasis id="40DAEADDFFF0FFA9FC33F8FDE60D2430" bold="true" box="[990,1149,1812,1836]" italics="true" pageId="1" pageNumber="368">Anoplocheilus</emphasis>
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)
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<emphasis id="40DAEADDFFF0FFA9FB66F8FDE6BB2430" bold="true" box="[1163,1227,1812,1836]" italics="true" pageId="1" pageNumber="368">clarki</emphasis>
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5BE957A6FFF0FFA9FB3DF8FCE7532430" box="[1232,1315,1813,1836]" pageId="1" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
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(
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||
<figureCitation id="EA952A4AFFF0FFA9FADDF8FDE7082430" box="[1328,1400,1812,1836]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="1" pageNumber="368">Figs. 1</figureCitation>
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,
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<figureCitation id="EA952A4AFFF0FFA9FA68F8FDE7E32430" box="[1413,1427,1812,1836]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="1" pageNumber="368">2</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3AB46544FFF0FFAAFCDDF8DDE1712090" lastPageId="2" lastPageNumber="369" pageId="1" pageNumber="368" type="diagnosis">
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<paragraph id="721136CFFFF0FFAAFCDDF8DDE1712090" blockId="1.[815,1437,1812,1932]" lastBlockId="2.[150,773,180,908]" lastPageId="2" lastPageNumber="369" pageId="1" pageNumber="368">
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<emphasis id="40DAEADDFFF0FFA9FCDDF8DDE1D42450" bold="true" box="[816,932,1844,1868]" pageId="1" pageNumber="368">Diagnosis.</emphasis>
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<taxonomicName id="B5AE4D4CFFF0FFA9FC44F8DDE6C22450" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[937,1202,1844,1868]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
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<emphasis id="40DAEADDFFF0FFA9FC44F8DDE6302450" box="[937,1088,1844,1868]" italics="true" pageId="1" pageNumber="368">Anoplocheilus</emphasis>
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||
(
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||
<emphasis id="40DAEADDFFF0FFA9FBA3F8DDE6162457" box="[1102,1126,1844,1867]" italics="true" pageId="1" pageNumber="368">A.</emphasis>
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||
)
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||
<emphasis id="40DAEADDFFF0FFA9FB99F8DDE6C22450" box="[1140,1202,1844,1868]" italics="true" pageId="1" pageNumber="368">clarki</emphasis>
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</taxonomicName>
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||
can be separated from its closest relative,
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||
<taxonomicName id="B5AE4D4CFFF0FFA9FC1AF8BDE6E82470" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[1015,1176,1876,1900]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
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<emphasis id="40DAEADDFFF0FFA9FC1AF8BDE67E2477" box="[1015,1038,1876,1899]" italics="true" pageId="1" pageNumber="368">A.</emphasis>
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||
(
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||
<emphasis id="40DAEADDFFF0FFA9FBF0F8BDE6442477" box="[1053,1076,1876,1899]" italics="true" pageId="1" pageNumber="368">A.</emphasis>
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||
)
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<emphasis id="40DAEADDFFF0FFA9FBAFF8BCE6E82470" box="[1090,1176,1877,1900]" italics="true" pageId="1" pageNumber="368">rusticus</emphasis>
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</taxonomicName>
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<emphasis id="40DAEADDFFF0FFA9FB75F8BCE6EC2470" box="[1176,1180,1877,1900]" italics="true" pageId="1" pageNumber="368">,</emphasis>
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by differences observed mainly at the level of the clypeus, pronotum, scutellum and aeadeagal parameres (
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<figureCitation id="EA952A4AFFF3FFAAFE55FF5DE06523D0" box="[440,533,180,204]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[816,846,1885,1904]" captionStartId-1="4.[151,182,914,933]" captionStartId-2="5.[136,167,1857,1876]" captionStartId-3="5.[817,848,1857,1876]" captionTargetBox-0="[816,1436,1178,1849]" captionTargetBox-1="[151,771,181,885]" captionTargetBox-2="[127,773,1121,1831]" captionTargetBox-3="[803,1431,1121,1834]" captionTargetId-0="figure-741@3.[813,1439,1168,1872]" captionTargetId-1="figure-727@4.[151,771,179,889]" captionTargetId-2="figure-540@5.[127,765,1121,1843]" captionTargetId-3="figure-542@5.[828,1437,1118,1828]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." captionText-1="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." captionText-2="Fig. 3 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." captionText-3="Fig. 4 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi-0="http://doi.org/10.5281/zenodo.12763626" figureDoi-1="http://doi.org/10.5281/zenodo.12763628" figureDoi-2="http://doi.org/10.5281/zenodo.12763630" figureDoi-3="http://doi.org/10.5281/zenodo.14549189" httpUri-0="https://zenodo.org/record/12763626/files/figure.png" httpUri-1="https://zenodo.org/record/12763628/files/figure.png" httpUri-2="https://zenodo.org/record/12763630/files/figure.png" httpUri-3="https://zenodo.org/record/14549189/files/figure.png" pageId="2" pageNumber="369">Figs 1-4</figureCitation>
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). Firstly, the general clypeal shape is more squarish in
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<taxonomicName id="B5AE4D4CFFF3FFAAFDEBFF3DE0E223F0" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[518,658,212,236]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
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<emphasis id="40DAEADDFFF3FFAAFDEBFF3DE06E23F7" box="[518,542,212,235]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFDC0FF3DE03423F7" box="[557,580,212,235]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFDB9FF3DE0E223F0" box="[596,658,212,236]" italics="true" pageId="2" pageNumber="369">clarki</emphasis>
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||
</taxonomicName>
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||
than in
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||
<taxonomicName id="B5AE4D4CFFF3FFAAFD03FF3DE36B2210" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
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||
<emphasis id="40DAEADDFFF3FFAAFD03FF3DE17523F7" box="[750,773,212,235]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
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||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFF4DFF1DE2C72217" box="[160,183,244,267]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
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||
)
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||
<emphasis id="40DAEADDFFF3FFAAFF25FF1CE36B2210" box="[200,283,245,268]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
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||
</taxonomicName>
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||
, in which the antero-lateral corners are actually smoothly rounded. Also, the denticles on the anterior margin of the clypeus are more raised and developed in
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<taxonomicName id="B5AE4D4CFFF3FFAAFF7AFEBDE3592270" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[151,297,340,364]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
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||
<emphasis id="40DAEADDFFF3FFAAFF7AFEBDE2DF2277" box="[151,175,340,363]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFF2CFEBDE2A82277" box="[193,216,340,363]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFF06FEBDE3592270" box="[235,297,340,364]" italics="true" pageId="2" pageNumber="369">clarki</emphasis>
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||
</taxonomicName>
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||
than in
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||
<taxonomicName id="B5AE4D4CFFF3FFAAFE64FEBDE0412270" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[393,561,340,364]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFE64FEBDE3D02277" box="[393,416,340,363]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFE5EFEBDE3BA2277" box="[435,458,340,363]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
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||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFE30FEBCE0412270" box="[477,561,341,364]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
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||
</taxonomicName>
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and the antero-lateral declivity on each side of the clypeus is substantially more pronounced in the former than in the latter species (
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||
<figureCitation id="EA952A4AFFF3FFAAFF72FE5DE28F22D0" box="[159,255,436,460]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">Figs 2 A</figureCitation>
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,
|
||
<figureCitation id="EA952A4AFFF3FFAAFEE1FE5DE34722D0" box="[268,311,436,460]" captionStart="Fig" captionStartId="5.[817,848,1857,1876]" captionTargetBox="[803,1431,1121,1834]" captionTargetId="figure-542@5.[828,1437,1118,1828]" captionText="Fig. 4 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.14549189" httpUri="https://zenodo.org/record/14549189/files/figure.png" pageId="2" pageNumber="369">4 A</figureCitation>
|
||
). While the pronotal and scutellar surfaces in
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFF5EFE3DE32422F0" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[179,340,468,492]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFF5EFE3DE2BA22F7" box="[179,202,468,491]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFF34FE3DE28022F7" box="[217,240,468,491]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFEEDFE3CE32422F0" box="[256,340,469,492]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
|
||
</taxonomicName>
|
||
are always entirely black, even in specimens with predominatly testaceous elytra (
|
||
<bibRefCitation id="163F4B3EFFF3FFAAFDB7FE1DE2BF2130" author="Holm E. & Marais E." pageId="2" pageNumber="369" refId="ref7872" refString="Holm E., Marais E. 1992. Fruit chafers of Southern Africa (Scarabaeidae: Cetoniini). Ekogilde, Hartbeespoort (South Africa), 326 pp." type="book" year="1992">Holm & Marais 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="163F4B3EFFF3FFAAFF36FDFDE3132130" author="Marais A. P." box="[219,355,532,556]" pageId="2" pageNumber="369" pagination="82 - 83" refId="ref8469" refString="Marais A. P. 1994. Observations on Anoplocheilus rusticus (Gory & Percheron) (Scarabaeidae: Cetoniinae) (true or day flying fruit chafers). Metamorphosis, 5 (2): 82 - 83." type="journal article" year="1994">Marais 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="163F4B3EFFF3FFAAFE82FDFDE0472130" author="Beinhundner G." box="[367,567,532,556]" pageId="2" pageNumber="369" refId="ref7604" refString="Beinhundner G. 2017. The Cetoniinae of Africa. Gerhard Beinhundner, Euerbach, 1199 pp." type="book" year="2017">Beinhundner 2017</bibRefCitation>
|
||
), in the light forms of
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFF5EFDDDE34A2150" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[179,314,564,588]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFF5EFDDDE2BA2157" box="[179,202,564,587]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFF3AFDDDE29F2157" box="[215,239,564,587]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFF11FDDDE34A2150" box="[252,314,564,588]" italics="true" pageId="2" pageNumber="369">clarki</emphasis>
|
||
</taxonomicName>
|
||
both surfaces exhibit some testaceous areas (
|
||
<figureCitation id="EA952A4AFFF3FFAAFF70FDBDE28F2170" box="[157,255,596,620]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Figs 1 A</figureCitation>
|
||
,
|
||
<figureCitation id="EA952A4AFFF3FFAAFEE0FDBDE3482170" box="[269,312,596,620]" captionStart="Fig" captionStartId="5.[136,167,1857,1876]" captionTargetBox="[127,773,1121,1831]" captionTargetId="figure-540@5.[127,765,1121,1843]" captionTargetPageId="5" captionText="Fig. 3 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763630" httpUri="https://zenodo.org/record/12763630/files/figure.png" pageId="2" pageNumber="369">3 A</figureCitation>
|
||
), reminiscent of those observed in similar forms of
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFF16FD9DE00B2190" authority="(Wiedemann, 1823)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[251,635,628,652]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="germari" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFF16FD9DE3622197" box="[251,274,628,651]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFECCFD9DE3482197" box="[289,312,628,651]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFEA5FD9CE3ED2190" box="[328,413,629,652]" italics="true" pageId="2" pageNumber="369">germari</emphasis>
|
||
(
|
||
<bibRefCitation id="163F4B3EFFF3FFAAFE41FD9DE0042190" author="Wiedemann C. R. W." box="[428,628,628,652]" pageId="2" pageNumber="369" refId="ref8836" refString="Wiedemann C. R. W. 1823. Zoologisches Magazin Vol. 2 (1). Akademische Buchhandlung, Kiel, 164 pp." type="book" year="1823">Wiedemann, 1823</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The general body pubescence in
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFE87FD7DE38421B0" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[362,500,660,684]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFE87FD7DE3F221B7" box="[362,386,660,683]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFE7DFD7DE3D821B7" box="[400,424,660,683]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFE5BFD7DE38421B0" box="[438,500,660,684]" italics="true" pageId="2" pageNumber="369">clarki</emphasis>
|
||
</taxonomicName>
|
||
is remarkably thicker and longer than in
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFED8FD5DE3A521D0" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[309,469,692,716]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFED8FD5DE33C21D7" box="[309,332,692,715]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFEB6FD5DE30221D7" box="[347,370,692,715]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFE6FFD5CE3A521D0" box="[386,469,693,716]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
|
||
</taxonomicName>
|
||
, particularly on head vertex and ventral surfaces (
|
||
<figureCitation id="EA952A4AFFF3FFAAFE90FD3DE3AE21F0" box="[381,478,724,748]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Figs 1 B</figureCitation>
|
||
,
|
||
<figureCitation id="EA952A4AFFF3FFAAFE07FD3DE06321F0" box="[490,531,724,748]" captionStart="Fig" captionStartId="5.[136,167,1857,1876]" captionTargetBox="[127,773,1121,1831]" captionTargetId="figure-540@5.[127,765,1121,1843]" captionTargetPageId="5" captionText="Fig. 3 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763630" httpUri="https://zenodo.org/record/12763630/files/figure.png" pageId="2" pageNumber="369">3 B</figureCitation>
|
||
). Finally, the aedeagal parameres of
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFECFFD1DE3DA2010" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[290,426,756,780]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFECFFD1DE3492017" box="[290,313,756,779]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFEA5FD1DE32F2017" box="[328,351,756,779]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFE81FD1DE3DA2010" box="[364,426,756,780]" italics="true" pageId="2" pageNumber="369">clarki</emphasis>
|
||
</taxonomicName>
|
||
are substantially shorter but wider than those of
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFED3FCFDE3AD2030" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[318,477,788,812]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFED3FCFDE3252037" box="[318,341,788,811]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFE89FCFDE30B2037" box="[356,379,788,811]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFE64FCFCE3AD2030" box="[393,477,789,812]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EA952A4AFFF3FFAAFE04FCFDE0352030" box="[489,581,788,812]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">Figs 2 C</figureCitation>
|
||
,
|
||
<figureCitation id="EA952A4AFFF3FFAAFDA2FCFDE0062030" box="[591,630,788,812]" captionStart="Fig" captionStartId="5.[817,848,1857,1876]" captionTargetBox="[803,1431,1121,1834]" captionTargetId="figure-542@5.[828,1437,1118,1828]" captionText="Fig. 4 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.14549189" httpUri="https://zenodo.org/record/14549189/files/figure.png" pageId="2" pageNumber="369">4 C</figureCitation>
|
||
), particularly at the apex where they form a rounded lateral expansion, which in frontal view imparts a width approximately twice as large as that observed in
|
||
<taxonomicName id="B5AE4D4CFFF3FFAAFE54FC9DE0292090" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[441,601,884,908]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF3FFAAFE54FC9DE3A12097" box="[441,465,884,907]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF3FFAAFE0DFC9DE3872097" box="[480,503,884,907]" italics="true" pageId="2" pageNumber="369">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF3FFAAFDE8FC9CE0292090" box="[517,601,885,908]" italics="true" pageId="2" pageNumber="369">rusticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EA952A4AFFF3FFAAFD8AFC9DE0B22090" box="[615,706,884,908]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">Figs 2 E</figureCitation>
|
||
,
|
||
<figureCitation id="EA952A4AFFF3FFAAFD20FC9DE0842090" box="[717,756,884,908]" captionStart="Fig" captionStartId="5.[817,848,1857,1876]" captionTargetBox="[803,1431,1121,1834]" captionTargetId="figure-542@5.[828,1437,1118,1828]" captionText="Fig. 4 – Anoplocheilus (A.) rusticus (Gory & Percheron, 1833). Male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.14549189" httpUri="https://zenodo.org/record/14549189/files/figure.png" pageId="2" pageNumber="369">4 E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB46544FFF3FFABFF7AFC5DE32D2170" lastPageId="3" lastPageNumber="370" pageId="2" pageNumber="369" type="description">
|
||
<paragraph id="721136CFFFF3FFAAFF7AFC5DE0C720D0" blockId="2.[150,774,948,1932]" box="[151,695,948,972]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFF7AFC5DE3AE20D0" bold="true" box="[151,478,948,972]" pageId="2" pageNumber="369">
|
||
Description of
|
||
<typeStatus id="AD15886DFFF3FFAAFED2FC5DE3D120D0" box="[319,417,948,972]" pageId="2" pageNumber="369" type="holotype">holotype</typeStatus>
|
||
male
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="EA952A4AFFF3FFAAFE01FC5DE05F20D0" box="[492,559,948,972]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Figs 1</figureCitation>
|
||
A-C, 2 A-E)
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFF50FC3CE17420F0" blockId="2.[150,774,948,1932]" box="[189,772,980,1004]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFF50FC3CE29520F0" box="[189,229,981,1004]" italics="true" pageId="2" pageNumber="369">Size</emphasis>
|
||
. Total Length =
|
||
<quantity id="B5569B2AFFF3FFAAFE6AFC3DE39420F0" box="[391,484,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="2" pageNumber="369" unit="mm" value="13.3">13.3 mm</quantity>
|
||
; Maximum Width =
|
||
<quantity id="B5569B2AFFF3FFAAFD5DFC3DE17020F0" box="[688,768,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="369" unit="mm" value="7.1">7.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFF50FC1DE08C2610" blockId="2.[150,774,948,1932]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFF50FC1DE2812710" box="[189,241,1012,1036]" italics="true" pageId="2" pageNumber="369">Body</emphasis>
|
||
: Mostly matte with head, pronotum and scutellum predominatly black but elytra testaceous with lines of brown spots on costal and umbonal elevations; dense, round to irregular sculpture on head and pronotum, becoming scattered, shallow and predominantly of horse-shoe
|
||
<typeStatus id="AD15886DFFF3FFAAFDA6FB9CE0072791" box="[587,631,1141,1165]" pageId="2" pageNumber="369">type</typeStatus>
|
||
on scutellum and elytra; with dense and long, pale-yellow to tawny setae on head vertex and around all lateral margins, becoming shorter and scattered on pronotal and elytral declivities and disappearing on their discal areas as well as scutellum (
|
||
<figureCitation id="EA952A4AFFF3FFAAFD8DFB1DE0CE2610" box="[608,702,1268,1292]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Figs. 1 A</figureCitation>
|
||
,
|
||
<figureCitation id="EA952A4AFFF3FFAAFD25FB1DE09F2610" box="[712,751,1268,1292]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">2 A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFF51FAFDE00E25B0" blockId="2.[150,774,948,1932]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFF51FAFDE2862630" box="[188,246,1300,1324]" italics="true" pageId="2" pageNumber="369">Head</emphasis>
|
||
. Entirely black and relatively flat in central region; clypeus broadly squarish in shape, with marked sinuation on apical margin and sharply sloping declivities on antero-lateral margins; anterior margin sharply upturned and forming two pairs of symmetric denticles, with proximal more developed than distal pair (
|
||
<figureCitation id="EA952A4AFFF3FFAAFDA4FA5DE0DE26D0" box="[585,686,1460,1484]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">Fig. 2 A</figureCitation>
|
||
); entire surface covered in coarse dense sculpture; pale-yellow to tawny setae distributed along entire surface, but generally shorter and more scattered on clypeus and frons, becoming much longer and denser on vertex and antennal pedicel; antenna dark brown to black, with club approximately as long as flagellum; thin light setae scattered across flagellum, becoming thicker and denser on pedicel.
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFF50F95CE7232250" blockId="2.[150,774,948,1932]" lastBlockId="2.[830,1453,180,1933]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFF50F95CE35325D0" box="[189,291,1717,1740]" italics="true" pageId="2" pageNumber="369">Pronotum</emphasis>
|
||
. Black and matte, with shiny suprascutellar area exhibiting testaceous, horizontally elongate double macula, not reaching posterior margin; pair of symmetric brown maculae also on lateral declivities anteriad of marginal round angle; with dense round to crescent punctures across entire surface; virtually glabrous on disc but exhibiting short to medium light-yellow setae on lateral and anterior declivities and margins (
|
||
<figureCitation id="EA952A4AFFF3FFAAFBADFF5DE6EA23D0" box="[1088,1178,180,204]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Fig. 1 A</figureCitation>
|
||
); shape broadly octagonal with lateral margins smoothly rounded at centre, antero-lateral margins sharply rounded with pronounced angle, postero-lateral margins smoothly rounded and shifting posteriad, leading then to smooth pre-scutellar arch at middle.
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFC88FEBDE7F12170" blockId="2.[830,1453,180,1933]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFC88FEBDE1A42270" box="[869,980,340,364]" italics="true" pageId="2" pageNumber="369">Scutellum.</emphasis>
|
||
Shiny and black, with two symmetric, longitudinally elongate, testaceous spots on each basal side of dark midline; exhibiting longitudinal incision running parallel to lateral groove from basal margin to middle of total scutellar length; few, shallow horse-shoe to crescent punctures along lateral margins inwards of incision, with occasional short pale setae emerging at centre of punctures; isoscelic triangular in shape with sharp apex; lateral grooves well-developed and deepening towards apex (
|
||
<figureCitation id="EA952A4AFFF3FFAAFAF6FDBDE7052170" box="[1307,1397,596,620]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Fig. 1 A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFC88FD9DE1962750" blockId="2.[830,1453,180,1933]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFC88FD9DE1C62190" box="[869,950,628,652]" italics="true" pageId="2" pageNumber="369">Elytron</emphasis>
|
||
. Testaceous and matte, with dark maculae of variable size spread in longitudinal lines along the length of each costa and on umbones; all costae moderately elevated across entire surface but fading on apical declivity, striae exhibiting sublineate and semicontiguous crescent to horseshoe sculpture; with shiny black to dark brown lining around scutellum and sutural margin; lateral margin reborded and dark, not covering lateral edges of abdominal tergites; humeral callus prominent but apical callus poorly raised; sub-humeral arch with extremely weak sinuation and posterior margin smoothly rounded without projections or upturning at apex; virtually glabrous on disc and humeral callus, but with scattered short and light setae distributed along lateral and apical declivities (
|
||
<figureCitation id="EA952A4AFFF3FFAAFC91FBDDE1AA2750" box="[892,986,1076,1100]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Fig. 1 A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFC8BFBBDE1982610" blockId="2.[830,1453,180,1933]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFC8BFBBDE1BC2770" box="[870,972,1108,1132]" italics="true" pageId="2" pageNumber="369">Pygidium</emphasis>
|
||
. Broadly triangular in shape, with smoothly rounded apex and unevenly domed; with dense but fine subconcentric rugulose sculpture across entire surface; with light-yellow to tawny short setae scattered throughout disc, becoming long and fine along lateral margins and apex (
|
||
<figureCitation id="EA952A4AFFF3FFAAFC6DFB1DE1AA2610" box="[896,986,1268,1292]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="2" pageNumber="369">Fig. 2 B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFAAFC88FAFCE67A2410" blockId="2.[830,1453,180,1933]" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFC88FAFCE1E72630" box="[869,919,1301,1324]" italics="true" pageId="2" pageNumber="369">Legs</emphasis>
|
||
. Black with occasional brown tips, short and robust with typical fossorial characters; tarsal segments of average cetonine length, with apical ones twice as long as preceding units; meso- and metatarsomeres with marked dorso-distal protrusions; tibiae densely sculptured with mid longitudinal ridge and numerous light-yellow setae present mainly on inner margin, becoming progressively longer and denser from protibia to metatibia; protibia broad and tridentate, with teeth blunt and all equally well-developed, but distance between distal and middle teeth shorter than that between middle and proximal teeth; mesotibia with double mid spine on outer carina sharply pointed and spurs elongate, thin and sharp; metatibia with outer carina bearing one blunt but robust distal tooth and two smaller teeth further up, spurs elongate and rather sharp (
|
||
<figureCitation id="EA952A4AFFF3FFAAFC65F91DE1B62410" box="[904,966,1780,1804]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="2" pageNumber="369">Fig. 1</figureCitation>
|
||
A-C).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF3FFABFC88F8FDE0162230" blockId="2.[830,1453,180,1933]" lastBlockId="3.[136,758,180,620]" lastPageId="3" lastPageNumber="370" pageId="2" pageNumber="369">
|
||
<emphasis id="40DAEADDFFF3FFAAFC88F8FDE6772430" box="[869,1031,1812,1836]" italics="true" pageId="2" pageNumber="369">Ventral surface</emphasis>
|
||
. Black and shiny, covered in dense and long light-yellow pubescence, except on femora, central part of abdominal sternites and metasternal region, where setae are short or very scattered; mesosternal lobe extremely reduced, smoothly rounded and not protruding forward; metasternal lobe with anterior portion of median sulcus exhibiting oblong groove of triangular shape; abdominal sternites with flat area at centre (
|
||
<figureCitation id="EA952A4AFFF2FFABFE0EFEFDE0522230" box="[483,546,276,300]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="3" pageNumber="370">Fig. 1</figureCitation>
|
||
B-C).
|
||
</paragraph>
|
||
<paragraph id="721136CFFFF2FFABFF43FEDDE32D2170" blockId="3.[136,758,180,620]" pageId="3" pageNumber="370">
|
||
<emphasis id="40DAEADDFFF2FFABFF43FEDDE3682250" box="[174,280,308,332]" italics="true" pageId="3" pageNumber="370">Aedeagus</emphasis>
|
||
. Parameres compact and dark, with dorsal lobes gradually tapering medially and then expanding slightly towards apico-lateral margin to form smoothly rounded apex, here slightly bending downwards and exhibiting flat tip surface (clearly visible in frontal view,
|
||
<figureCitation id="EA952A4AFFF2FFABFF65FE3DE2BD22F0" box="[136,205,468,492]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="3" pageNumber="370">Fig. 2</figureCitation>
|
||
C-E); inner margin of dorsal lobes remarkably straight, bending outwards only slightly towards central region in dorsal view; ventral lobes generally narrower than dorsal lobes and emerging only towards base in dorsal view (
|
||
<figureCitation id="EA952A4AFFF2FFABFF1CFDBDE3202170" box="[241,336,596,620]" captionStart="Fig" captionStartId="4.[151,182,914,933]" captionTargetBox="[151,771,181,885]" captionTargetId="figure-727@4.[151,771,179,889]" captionTargetPageId="4" captionText="Fig. 2 – Anoplocheilus (A.) clarki sp. nov. Holotype male: A, clypeus; B, pygidium; C, parameres, dorsal view; D, parameres, lateral view; E, parameres, frontal view. Photographs by Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763628" httpUri="https://zenodo.org/record/12763628/files/figure.png" pageId="3" pageNumber="370">Fig. 2 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB46544FFF2FFABFF65FD7DE3E320D0" pageId="3" pageNumber="370" type="etymology">
|
||
<paragraph id="721136CFFFF2FFABFF65FD7DE3E320D0" blockId="3.[136,757,660,972]" pageId="3" pageNumber="370">
|
||
<emphasis id="40DAEADDFFF2FFABFF65FD7DE31121B0" bold="true" box="[136,353,660,684]" pageId="3" pageNumber="370">Derivatio nominis.</emphasis>
|
||
This species is named after Derek Clark, former conservation manager of a number of parks and farms in
|
||
<collectingCountry id="0AB9765FFFF2FFABFECEFD3DE3F221F0" box="[291,386,724,748]" name="Namibia" pageId="3" pageNumber="370">Namibia</collectingCountry>
|
||
,
|
||
<collectingCountry id="0AB9765FFFF2FFABFE7CFD3DE06D21F0" box="[401,541,724,748]" name="South Africa" pageId="3" pageNumber="370">South Africa</collectingCountry>
|
||
and
|
||
<collectingCountry id="0AB9765FFFF2FFABFDB7FD3DE0C721F0" box="[602,695,724,748]" name="Ethiopia" pageId="3" pageNumber="370">Ethiopia</collectingCountry>
|
||
, who promptly brought the specimens retrieved from a farm trough in the Komsberg to the attention of the lead author. During the past three decades, Mr Clark has made a very significant contribution to the entomofauna of
|
||
<collectingCountry id="0AB9765FFFF2FFABFD91FCBDE2D22090" name="South Africa" pageId="3" pageNumber="370">South Africa</collectingCountry>
|
||
, discovering several new species from remote areas in his effort to enhance nature conservation and biodiversity knowledge in the region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB46544FFF2FFABFF65FC1DE0EE2610" pageId="3" pageNumber="370" type="distribution">
|
||
<paragraph id="721136CFFFF2FFABFF65FC1DE0EE2610" blockId="3.[134,757,1012,1292]" pageId="3" pageNumber="370">
|
||
<emphasis id="40DAEADDFFF2FFABFF65FC1DE36A2710" bold="true" box="[136,282,1012,1036]" pageId="3" pageNumber="370">Distribution.</emphasis>
|
||
This is obviously an inland sister species of the strictly coastal
|
||
<taxonomicName id="B5AE4D4CFFF2FFABFE9DFBFDE0612730" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[368,529,1044,1068]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF2FFABFE9DFBFDE3F72737" box="[368,391,1044,1067]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF2FFABFE7BFBFDE3DE2737" box="[406,430,1044,1067]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF2FFABFE50FBFCE0612730" box="[445,529,1045,1068]" italics="true" pageId="3" pageNumber="370">rusticus</emphasis>
|
||
</taxonomicName>
|
||
. The series collected most recently originates from the Roggeveldberge range to the south of Sutherland in the
|
||
<collectingRegion id="B06AF82DFFF2FFABFE51FBBDE0CB2770" box="[444,699,1108,1132]" country="South Africa" name="Northern Cape" pageId="3" pageNumber="370">Northern Cape Province</collectingRegion>
|
||
(
|
||
<figureCitation id="EA952A4AFFF2FFABFD25FBBDE2E52790" captionStart="Fig" captionStartId="6.[153,184,792,811]" captionTargetBox="[151,771,183,771]" captionTargetId="figure-778@6.[151,774,181,775]" captionTargetPageId="6" captionText="Fig. 5 – Known distribution of Anoplocheilus (A.) rusticus (Gory & Percheron, 1833) and A. (A.) clarki sp. nov. in the western South African region (map adapted from www.freeworldmaps.net)." figureDoi="http://doi.org/10.5281/zenodo.12763632" httpUri="https://zenodo.org/record/12763632/files/figure.png" pageId="3" pageNumber="370">Fig. 5</figureCitation>
|
||
). There are also a few other specimens retrieved from old collections and originating from adjacent mountain ranges of the
|
||
<collectingRegion id="B06AF82DFFF2FFABFF3EFB5DE37A27D0" box="[211,266,1204,1228]" country="South Africa" name="Western Cape" pageId="3" pageNumber="370">Cape</collectingRegion>
|
||
Fold Belt, namely the Cederberg, the Koue Bokkeveld, the Hexrivierberge, the Boland Mountains and the Langeberge, all in the
|
||
<collectingRegion id="B06AF82DFFF2FFABFE70FB1CE0EA2610" box="[413,666,1268,1292]" country="South Africa" name="Western Cape" pageId="3" pageNumber="370">Western Cape Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB46544FFF2FFABFF65FADDE3A62450" pageId="3" pageNumber="370" type="materials_examined">
|
||
<paragraph id="721136CFFFF2FFABFF65FADDE3A62450" blockId="3.[135,757,1332,1868]" pageId="3" pageNumber="370">
|
||
<emphasis id="40DAEADDFFF2FFABFF65FADDE3422650" bold="true" box="[136,306,1332,1356]" pageId="3" pageNumber="370">Type material.</emphasis>
|
||
<materialsCitation id="C2C63C92FFF2FFABFED1FADDE0082670" collectingDate="2015-04-01" collectionCode="ISAM" collectorName="Derek Clark" country="South Africa" location="Komsberg" municipality="Komsberg" pageId="3" pageNumber="370" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="AD15886DFFF2FFABFED1FADDE3EF2650" box="[316,415,1332,1356]" pageId="3" pageNumber="370" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="64A8FD46FFF2FFABFE47FADCE3B12650" box="[426,449,1333,1356]" count="1" pageId="3" pageNumber="370" type="male">♂</specimenCount>
|
||
:
|
||
<emphasis id="40DAEADDFFF2FFABFE3DFADDE0142650" bold="true" box="[464,612,1332,1356]" pageId="3" pageNumber="370">
|
||
<collectingCountry id="0AB9765FFFF2FFABFE3DFADDE0142650" box="[464,612,1332,1356]" name="South Africa" pageId="3" pageNumber="370">South Africa</collectingCountry>
|
||
</emphasis>
|
||
, NC,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFD46FADCE2CA2670" country="South Africa" name="Komsberg" pageId="3" pageNumber="370">Komsberg</collectingMunicipality>
|
||
,
|
||
<date id="0610100FFFF2FFABFF2AFABDE3392670" box="[199,329,1364,1388]" pageId="3" pageNumber="370" value="2015-04-01">
|
||
<collectingDate id="1654E9E7FFF2FFABFF2AFABDE3392670" box="[199,329,1364,1388]" pageId="3" pageNumber="370" value="2015-04-01">1 Apr 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFEBBFABDE3AC2670" box="[342,476,1364,1388]" pageId="3" pageNumber="370">Derek Clark</collectorName>
|
||
legit (
|
||
<collectionCode id="14BFAE0AFFF2FFABFDCAFABDE01F2670" box="[551,623,1364,1388]" pageId="3" pageNumber="370">ISAM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="C2C63C92FFF2FFABFD68FABCE3252690" collectingDate="2015-04-01" collectionCode="RPRM" collectorName="Derek Clark" country="South Africa" pageId="3" pageNumber="370" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus id="AD15886DFFF2FFABFD68FABCE0812671" box="[645,753,1365,1389]" pageId="3" pageNumber="370" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="64A8FD46FFF2FFABFF65FA9DE2DB2690" box="[136,171,1396,1420]" count="1" pageId="3" pageNumber="370" type="male">1♂</specimenCount>
|
||
, idem (
|
||
<collectionCode id="14BFAE0AFFF2FFABFF13FA9DE33E2690" box="[254,334,1396,1420]" pageId="3" pageNumber="370">RPRM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFE8FFA9DE35026B0" collectingDate="2015-04-08" collectionCode="RPRM" collectorName="Derek Clark" pageId="3" pageNumber="370" specimenCount="2" specimenCount-male="2">
|
||
<specimenCount id="64A8FD46FFF2FFABFE8FFA9DE3F62690" box="[354,390,1396,1420]" count="2" pageId="3" pageNumber="370" type="male">2♂</specimenCount>
|
||
, ibidem
|
||
<date id="0610100FFFF2FFABFE08FA9DE0132690" box="[485,611,1396,1420]" pageId="3" pageNumber="370" value="2015-04-08">
|
||
<collectingDate id="1654E9E7FFF2FFABFE08FA9DE0132690" box="[485,611,1396,1420]" pageId="3" pageNumber="370" value="2015-04-08">8 Apr 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFD82FA9DE0842690" box="[623,756,1396,1420]" pageId="3" pageNumber="370">Derek Clark</collectorName>
|
||
legit (
|
||
<collectionCode id="14BFAE0AFFF2FFABFF24FA7DE36926B0" box="[201,281,1428,1452]" pageId="3" pageNumber="370">RPRM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFEC2FA7DE05B26D0" collectingDate="1924-10-15" collectionCode="ISAM" collectorName="Versfeld" country="South Africa" location="Ceres Dist" municipality="Cold Bokkeveld" pageId="3" pageNumber="370" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="64A8FD46FFF2FFABFEC2FA7DE32226B0" box="[303,338,1428,1452]" count="1" pageId="3" pageNumber="370" type="male">1♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFE8FFA7DE06626B0" box="[354,534,1428,1452]" pageId="3" pageNumber="370">Cold Bokkeveld</collectingMunicipality>
|
||
,
|
||
<location id="77716014FFF2FFABFDC9FA7DE0EA26B0" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFDC9FA7DE0EA26B0" box="[548,666,1428,1452]" country="South Africa" municipality="Cold Bokkeveld" name="Ceres Dist" pageId="3" pageNumber="370">Ceres Dist</location>
|
||
,
|
||
<date id="0610100FFFF2FFABFD45FA7DE2B026D0" pageId="3" pageNumber="370" value="1924-10-15">
|
||
<collectingDate id="1654E9E7FFF2FFABFD45FA7DE2B026D0" pageId="3" pageNumber="370" value="1924-10-15">15 Oct 1924</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFF25FA5DE35126D0" box="[200,289,1460,1484]" pageId="3" pageNumber="370">Versfeld</collectorName>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFEC3FA5DE30626D0" box="[302,374,1460,1484]" pageId="3" pageNumber="370">ISAM</collectionCode>
|
||
: COL-A066116)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFDD9FA5DE00A2510" collectingDate="1949-12-01" collectionCode="ISAM" collectorName="Ceres C. P." country="South Africa" location="South African Museum Expedition" municipality="Upper Sources Olifants River" pageId="3" pageNumber="370" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="64A8FD46FFF2FFABFDD9FA5DE02C26D0" box="[564,604,1460,1484]" count="1" pageId="3" pageNumber="370" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFD88FA5CE33326F0" pageId="3" pageNumber="370">Upper Sources Olifants River</collectingMunicipality>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFEA2FA3DE3B226F0" box="[335,450,1492,1516]" pageId="3" pageNumber="370">Ceres C.P.</collectorName>
|
||
,
|
||
<date id="0610100FFFF2FFABFE22FA3DE02026F0" box="[463,592,1492,1516]" pageId="3" pageNumber="370" value="1949-12-01">
|
||
<collectingDate id="1654E9E7FFF2FFABFE22FA3DE02026F0" box="[463,592,1492,1516]" pageId="3" pageNumber="370" value="1949-12-01">1 Dec 1949</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="77716014FFF2FFABFDB0FA3DE3162510" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFDB0FA3DE3162510" country="South Africa" municipality="Upper Sources Olifants River" name="South African Museum Expedition" pageId="3" pageNumber="370">South African Museum Expedition</location>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFE95FA1DE3B02510" box="[376,448,1524,1548]" pageId="3" pageNumber="370">ISAM</collectionCode>
|
||
: COL-A027029)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFD65FA1DE3462570" collectingDate="1949-12-01" collectionCode="ISAM" country="South Africa" county="Wit River Valley" location="South African Museum Expedition" municipality="Bains Kloof" pageId="3" pageNumber="370" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="64A8FD46FFF2FFABFD65FA1DE0D92510" box="[648,681,1524,1548]" count="1" pageId="3" pageNumber="370" type="female">1♀</specimenCount>
|
||
[missing head and pronotum],
|
||
<collectingCounty id="9B704E43FFF2FFABFE4CF9FDE02B2530" box="[417,603,1556,1580]" pageId="3" pageNumber="370">Wit River Valley</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFD84F9FDE0802530" box="[617,752,1556,1580]" pageId="3" pageNumber="370">Bains Kloof</collectingMunicipality>
|
||
,
|
||
<date id="0610100FFFF2FFABFF65F9DDE37C2550" box="[136,268,1588,1612]" pageId="3" pageNumber="370" value="1949-12-01">
|
||
<collectingDate id="1654E9E7FFF2FFABFF65F9DDE37C2550" box="[136,268,1588,1612]" pageId="3" pageNumber="370" value="1949-12-01">1 Dec 1949</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="77716014FFF2FFABFEF7F9DDE0E72550" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFEF7F9DDE0E72550" box="[282,663,1588,1612]" country="South Africa" county="Wit River Valley" municipality="Bains Kloof" name="South African Museum Expedition" pageId="3" pageNumber="370">South African Museum Expedition</location>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFD44F9DDE0812550" box="[681,753,1588,1612]" pageId="3" pageNumber="370">ISAM</collectionCode>
|
||
: COL-A027030)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFEABF9BDE0392590" collectingDate="1975-10-05" collectionCode="TMSA" collectorName="MJ Duke" country="South Africa" location="Mitchell's Pass" pageId="3" pageNumber="370" specimenCode="CPH6312" specimenCount="1" specimenCount-female="1" stateProvince="Cape Province" typeStatus="paratype">
|
||
<specimenCount id="64A8FD46FFF2FFABFEABF9BDE3172570" box="[326,359,1620,1644]" count="1" pageId="3" pageNumber="370" type="female">1♀</specimenCount>
|
||
,
|
||
<location id="77716014FFF2FFABFE94F9BDE0532570" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFE94F9BDE0532570" box="[377,547,1620,1644]" country="South Africa" name="Mitchell's Pass" pageId="3" pageNumber="370" stateProvince="Cape Province">Mitchell’s Pass</location>
|
||
,
|
||
<collectingRegion id="B06AF82DFFF2FFABFDDEF9BDE0A72570" box="[563,727,1620,1644]" country="South Africa" name="Western Cape" pageId="3" pageNumber="370">Cape Province</collectingRegion>
|
||
,
|
||
<date id="0610100FFFF2FFABFD0AF9BDE2812590" pageId="3" pageNumber="370" value="1975-10-05">
|
||
<collectingDate id="1654E9E7FFF2FFABFD0AF9BDE2812590" pageId="3" pageNumber="370" value="1975-10-05">5 Oct 1975</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFF12F99CE3152590" box="[255,357,1652,1676]" pageId="3" pageNumber="370">MJ Duke</collectorName>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFE9AF99DE3B72590" box="[375,455,1652,1676]" collectionName="South Africa, Gauteng, Pretoria, Transvaal Museum" pageId="3" pageNumber="370">TMSA</collectionCode>
|
||
:
|
||
<specimenCode id="22089EB4FFF2FFABFE39F99DE0332590" box="[468,579,1652,1676]" pageId="3" pageNumber="370">CPH6312</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFDB5F99DE08425B0" collectingDate="1925-10" collectingDateMax="1925-11-01" collectingDateMin="1925-10" collectionCode="TMSA-" collectorName="Dr H Brauns" country="South Africa" county="Capland" location="Stellenbosch" municipality="Stellenbosch" pageId="3" pageNumber="370" specimenCode="CPH6305" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
|
||
<specimenCount id="64A8FD46FFF2FFABFDB5F99DE0092590" box="[600,633,1652,1676]" count="1" pageId="3" pageNumber="370" type="male">1♂</specimenCount>
|
||
,
|
||
<collectingCounty id="9B704E43FFF2FFABFD79F99DE0802590" box="[660,752,1652,1676]" pageId="3" pageNumber="370">Capland</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFF65F97DE36325B0" box="[136,275,1684,1708]" pageId="3" pageNumber="370">Stellenbosch</collectingMunicipality>
|
||
,
|
||
<date id="0610100FFFF2FFABFEF0F97DE3F225B0" box="[285,386,1684,1708]" pageId="3" pageNumber="370" value="1925-10">
|
||
<collectingDate id="1654E9E7FFF2FFABFEF0F97DE3F225B0" box="[285,386,1684,1708]" pageId="3" pageNumber="370" value="1925-10">Oct 1925</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFE61F97CE06325B0" box="[396,531,1685,1708]" pageId="3" pageNumber="370">Dr H Brauns</collectorName>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFDCDF97DE00525B0" box="[544,629,1684,1708]" pageId="3" pageNumber="370">TMSA-</collectionCode>
|
||
<specimenCode id="22089EB4FFF2FFABFD97F97DE09925B0" box="[634,745,1684,1708]" pageId="3" pageNumber="370">CPH6305</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="C2C63C92FFF2FFABFF6AF95DE0A725F0" collectingDate="1925-11-01" collectingDateMax="1925-11-01" collectingDateMin="1925-10" collectionCode="ISAM" collectorName="Dr H Brauns" country="South Africa" county="Capland" location="Swellendam Dist." municipality="Tradouw Pass" pageId="3" pageNumber="370" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
|
||
1 ind,
|
||
<collectingMunicipality id="9275ACB5FFF2FFABFF24F95DE31025D0" box="[201,352,1716,1740]" pageId="3" pageNumber="370">Tradouw Pass</collectingMunicipality>
|
||
,
|
||
<location id="77716014FFF2FFABFE87F95DE05925D0" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFE87F95DE05925D0" box="[362,553,1716,1740]" country="South Africa" county="Capland" municipality="Tradouw Pass" name="Swellendam Dist." pageId="3" pageNumber="370" stateProvince="Western Cape">Swellendam Dist.</location>
|
||
,
|
||
<date id="0610100FFFF2FFABFDDEF95DE0DF25D0" box="[563,687,1716,1740]" pageId="3" pageNumber="370" value="1925-11-01">
|
||
<collectingDate id="1654E9E7FFF2FFABFDDEF95DE0DF25D0" box="[563,687,1716,1740]" pageId="3" pageNumber="370" value="1925-11-01">1 Nov 1925</collectingDate>
|
||
</date>
|
||
<location id="77716014FFF2FFABFD58F95DE3B125F0" LSID="urn:lsid:plazi:treatment:FA0787D9FFF0FFABFCDDF8FDE1C52730:77716014FFF2FFABFD58F95DE3B125F0" country="South Africa" county="Capland" municipality="Tradouw Pass" name="South African Museum Expedition" pageId="3" pageNumber="370" stateProvince="Western Cape">South African Museum Expedition</location>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFE3EF93DE06B25F0" box="[467,539,1748,1772]" pageId="3" pageNumber="370">ISAM</collectionCode>
|
||
: COL-A027028)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="C2C63C92FFF2FFABFD0BF93DE3A12450" collectingDate="1997-09-22" collectionCode="BMPC" collectorName="R Perissinotto & L Clennell" country="South Africa" location="South Africa" pageId="3" pageNumber="370" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="64A8FD46FFF2FFABFD0BF93DE2EC2410" count="1" pageId="3" pageNumber="370" type="female">1 ♀</specimenCount>
|
||
[missing head and pronotum],
|
||
<collectingCountry id="0AB9765FFFF2FFABFE01F91DE0042410" box="[492,628,1780,1804]" name="South Africa" pageId="3" pageNumber="370">South Africa</collectingCountry>
|
||
WC, Bushmanskloof, on stream sand bank,
|
||
<date id="0610100FFFF2FFABFE13F8FDE0FC2430" box="[510,652,1812,1836]" pageId="3" pageNumber="370" value="1997-09-22">
|
||
<collectingDate id="1654E9E7FFF2FFABFE13F8FDE0FC2430" box="[510,652,1812,1836]" pageId="3" pageNumber="370" value="1997-09-22">22 Sep 1997</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DF5B5319FFF2FFABFD77F8FCE2A12450" pageId="3" pageNumber="370">R Perissinotto</collectorName>
|
||
&
|
||
<collectorName id="DF5B5319FFF2FFABFF1EF8DCE3132450" box="[243,355,1844,1868]" pageId="3" pageNumber="370">L Clennell</collectorName>
|
||
(
|
||
<collectionCode id="14BFAE0AFFF2FFABFE9FF8DDE3B42450" box="[370,452,1844,1868]" pageId="3" pageNumber="370">BMPC</collectionCode>
|
||
)]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB46544FFF2FFABFCDDFF5DE1C52730" pageId="3" pageNumber="370" type="discussion">
|
||
<paragraph id="721136CFFFF2FFABFCDDFF5DE1C52730" blockId="3.[815,1438,180,1132]" pageId="3" pageNumber="370">
|
||
<emphasis id="40DAEADDFFF2FFABFCDDFF5DE1EF23D0" bold="true" box="[816,927,180,204]" pageId="3" pageNumber="370">Remarks.</emphasis>
|
||
Specimens range in size from
|
||
<quantity id="B5569B2AFFF2FFABFB14FF5DE7ED23D0" box="[1273,1437,180,204]" metricMagnitude="-1" metricUnit="m" metricValue="3.0607" metricValueMax="3.3782" metricValueMin="2.7432" pageId="3" pageNumber="370" unit="in" value="12.05" valueMax="13.3" valueMin="10.8">10.8 to 13.3 in</quantity>
|
||
total length and from
|
||
<quantity id="B5569B2AFFF2FFABFBF6FF3DE6ED23F0" box="[1051,1181,212,236]" metricMagnitude="-1" metricUnit="m" metricValue="1.6001999999999998" metricValueMax="1.8034" metricValueMin="1.397" pageId="3" pageNumber="370" unit="in" value="6.3" valueMax="7.1" valueMin="5.5">5.5 to 7.1 in</quantity>
|
||
maximum width. Their elytral colour varies from testaceous with longitudinal lines of dark spots (
|
||
<figureCitation id="EA952A4AFFF2FFABFBE0FEFDE6FE2230" box="[1037,1166,276,300]" captionStart="Fig" captionStartId="3.[816,846,1885,1904]" captionTargetBox="[816,1436,1178,1849]" captionTargetId="figure-741@3.[813,1439,1168,1872]" captionTargetPageId="3" captionText="Fig. 1 – Anoplocheilus (A.) clarki sp. nov.Holotype male:A, dorsal habitus; B, ventral habitus; C, lateral habitus (Photographs by Lynette Clennell)." figureDoi="http://doi.org/10.5281/zenodo.12763626" httpUri="https://zenodo.org/record/12763626/files/figure.png" pageId="3" pageNumber="370">Fig. 1 A, C</figureCitation>
|
||
) to predominantly black with few testaceous to brown spots scattered across the disk. Intermediate forms to these show lines of spots joined together to compose transverse wave-like bands across the entire surface. No entirely black forms have been recorded yet, but in predominantly black forms the testaceous patterns on pronotum and scutellum fade away, and these areas become therefore entirely black. As expected, the only female specimen identified so far exhibits all the typical traits of their
|
||
<taxonomicName id="B5AE4D4CFFF2FFABFC50FDDDE62C2150" baseAuthorityName="Gory & Percheron" baseAuthorityYear="1833" box="[957,1116,564,588]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="rusticus" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF2FFABFC50FDDDE1A42157" box="[957,980,564,587]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF2FFABFC0EFDDDE18A2157" box="[995,1018,564,587]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF2FFABFBE5FDDCE62C2150" box="[1032,1116,565,588]" italics="true" pageId="3" pageNumber="370">rusticus</emphasis>
|
||
</taxonomicName>
|
||
counterparts, namely a slightly broader protibia, blunter but wider metatibial spurs and a more convex area in the middle of abdominal sternites than in males. Also, the denticles on the clypeal apex are more pronounced and recurved in male specimens compared to the females. Adult activity in
|
||
<taxonomicName id="B5AE4D4CFFF2FFABFB33FD3DE71F21F0" authorityName="Perissinotto & Beinhundner & Strümpher" authorityYear="2021" box="[1246,1391,724,748]" class="Insecta" family="Scarabaeidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="clarki" subGenus="Anoplocheilus">
|
||
<emphasis id="40DAEADDFFF2FFABFB33FD3DE68521F7" box="[1246,1269,724,747]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
(
|
||
<emphasis id="40DAEADDFFF2FFABFAEAFD3DE76F21F7" box="[1287,1311,724,747]" italics="true" pageId="3" pageNumber="370">A.</emphasis>
|
||
)
|
||
<emphasis id="40DAEADDFFF2FFABFADCFD3DE71F21F0" box="[1329,1391,724,748]" italics="true" pageId="3" pageNumber="370">clarki</emphasis>
|
||
</taxonomicName>
|
||
has so far been recorded only throughout the southern spring (late September to early December) and in early autumn (April) and, like in its sister species, the life cycle seems to be linked to fine sandy substrata, on the banks of rivers and streams. All specimens collected recently in the Komsberg were retrieved from the same farm trough adjacent to a dry river bed exhibiting substantial sand banks (D. Clark, pers. comm.). A few old specimens also carry unmistakable clues that they were found along river beds (cf. label data above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |