687 lines
111 KiB
XML
687 lines
111 KiB
XML
<document id="5026F4310B9063CC0CCBD87272BC09EC" ID-CLB-Dataset="55814" ID-DOI="10.1080/00222933.2013.791952" ID-GBIF-Dataset="857d6372-de50-463d-8471-717e72bdd3ba" ID-ISSN="1464-5262" ID-Zenodo-Dep="4608237" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615907007564" checkinUser="carolina" docAuthor="Schmidt, Joachim, Wrase, David W. & Sciaky, Riccardo" docDate="2013" docId="FA03878BD34EFF9EFDFEFDB3FBB63768" docLanguage="en" docName="JNATHIST.47.41-42.2671-2689.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (41 - 42)" docSource="http://dx.doi.org/10.1080/00222933.2013.791952" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Kashmirobroscus Schmidt & Wrase & Sciaky 2013, gen. nov." docType="treatment" docUuid="304B2D40-25B9-4FD1-9A55-4CDABAA86101" docUuidSource="ZooBank" docVersion="8" lastPageNumber="2681" masterDocId="063AFFF3D34AFF92FFDEFFF1FFE8331E" masterDocTitle="Description of Kashmirobroscus gen. nov. with two new species from the Northwest Himalaya, and remarks on the East Asian genus Eobroscus Kryzhanovskij, 1951 (Coleoptera: Carabidae: Broscini)" masterLastPageNumber="2689" masterPageNumber="2671" pageNumber="2673" updateTime="1698933453466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E318765F389B6B8352A969EAEBC79944">Description of Kashmirobroscus gen. nov. with two new species from the Northwest Himalaya, and remarks on the East Asian genus Eobroscus Kryzhanovskij, 1951 (Coleoptera: Carabidae: Broscini)</mods:title>
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<mods:namePart id="0D0B9FFB51940F6B6C671A24A11CAF43">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="3449EF22EA727B2A27ED1C41EAA3A24A">Lindenstrasse 31, 18211 Admannshagen, Germany;</mods:affiliation>
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<mods:namePart id="C816E1531001B48AC6B60076445C9BDF">Wrase, David W.</mods:namePart>
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<mods:affiliation id="69083497405498C00A44CF8DEF8755AE">Dunckerstrasse 78, 10437 Berlin, Germany;</mods:affiliation>
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<mods:namePart id="A736266FD9E74A91D41E400DFA0CEF7C">Sciaky, Riccardo</mods:namePart>
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<mods:affiliation id="FA733AA8D0CAFB621D28F90F1B83AB79">Via Fiamma 13, 20129 Milano, Italy</mods:affiliation>
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<mods:title id="74B43ED1B4B6D3F5BBE5426E470D192E">Journal of Natural History</mods:title>
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<mods:date id="DC0E1C85E88C48F8D1D3F164E9A9D8C3">2013</mods:date>
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<mods:title id="C5B7D75507FF0EE391E502B14FF4CE8C">J. Nat. Hist.</mods:title>
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<mods:number id="0087F1456EAEF5DF6AA5D4EEE0589223">2013-09-03</mods:number>
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<mods:number id="10A969AD1A8B4CC440637CDB2D529571">41 - 42</mods:number>
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<mods:identifier id="11530EA50EAE9B4FE028E496B2FAFC51" type="DOI">10.1080/00222933.2013.791952</mods:identifier>
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<treatment id="FA03878BD34EFF9EFDFEFDB3FBB63768" ID-DOI="http://doi.org/10.5281/zenodo.6502425" ID-GBIF-Taxon="194854827" ID-Zenodo-Dep="6502425" LSID="urn:lsid:zoobank.org:act:304B2D40-25B9-4FD1-9A55-4CDABAA86101" httpUri="http://treatment.plazi.org/id/FA03878BD34EFF9EFDFEFDB3FBB63768" lastPageId="12" lastPageNumber="2681" pageId="4" pageNumber="2673">
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<subSubSection id="3AB06516D34EFF96FDFEFDB3FCAF3145" box="[544,839,578,604]" pageId="4" pageNumber="2673" type="nomenclature">
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<paragraph id="7215369DD34EFF96FDFEFDB3FCAF3145" blockId="4.[486,896,547,604]" box="[544,839,578,604]" pageId="4" pageNumber="2673">
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<heading id="295D81F1D34EFF96FDFEFDB3FCAF3145" box="[544,839,578,604]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="2673" reason="2">
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<taxonomicName id="B5AA4D1ED34EFF96FDFEFDB3FD333145" ID-CoL="8LS2N" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[544,731,578,603]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2673" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="40DEEA8FD34EFF96FDFEFDB3FD333145" bold="true" box="[544,731,578,603]" italics="true" pageId="4" pageNumber="2673">Kashmirobroscus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5BED57F4D34EFF96FD3CFDB5FCAF3145" box="[738,839,580,604]" pageId="4" pageNumber="2673" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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<subSubSection id="3AB06516D34EFF96FF0DFD72FBE031A4" pageId="4" pageNumber="2673" type="type_taxon">
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<paragraph id="7215369DD34EFF96FF0DFD72FD0F3184" blockId="4.[211,1032,642,698]" box="[211,743,642,667]" pageId="4" pageNumber="2673">
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<heading id="295D81F1D34EFF96FF0DFD72FD0F3184" box="[211,743,642,667]" fontSize="10" level="3" pageId="4" pageNumber="2673" reason="5">
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<typeStatus id="AD11883FD34EFF96FF0DFD72FEE23185" box="[211,266,643,667]" pageId="4" pageNumber="2673">Type</typeStatus>
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species:
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<taxonomicName id="B5AA4D1ED34EFF96FEB2FD72FD643185" ID-CoL="8LXWP" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[364,652,643,667]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2673" phylum="Arthropoda" rank="species" species="ortrudae" status="sp. nov.">
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<emphasis id="40DEEA8FD34EFF96FEB2FD72FD643185" box="[364,652,643,667]" italics="true" pageId="4" pageNumber="2673">Kashmirobroscus ortrudae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5BED57F4D34EFF96FD4DFD72FD0F3184" box="[659,743,643,666]" pageId="4" pageNumber="2673" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="7215369DD34EFF96FF0DFD53FBE031A4" blockId="4.[211,1032,642,698]" box="[211,1032,674,698]" pageId="4" pageNumber="2673">
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<uri id="063B3A9FD34EFF96FF0DFD53FBE031A4" box="[211,1032,674,698]" pageId="4" pageNumber="2673">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="3AB06516D34EFF96FF0DFCF1FD483060" pageId="4" pageNumber="2673" type="discussion">
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<paragraph id="7215369DD34EFF96FF0DFCF1FE133009" blockId="4.[211,1172,767,895]" box="[211,507,767,791]" pageId="4" pageNumber="2673">
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<heading id="295D81F1D34EFF96FF0DFCF1FE133009" box="[211,507,767,791]" fontSize="10" level="3" pageId="4" pageNumber="2673" reason="5">
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<emphasis id="40DEEA8FD34EFF96FF0DFCF1FE133009" box="[211,507,767,791]" italics="true" pageId="4" pageNumber="2673">Derivation of generic name</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7215369DD34EFF96FF0DFCD6FD483060" blockId="4.[211,1172,767,895]" pageId="4" pageNumber="2673">
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A masculine noun that combines the name of the geographical origin of the new taxon, the Kashmir Himalaya, with the name of the nominotypical genus of the tribe
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<taxonomicName id="B5AA4D1ED34EFF96FF0DFC97FEDB3060" authorityName="Roig-Junent" authorityYear="2000" box="[211,307,870,894]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2673" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
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, the genus
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<taxonomicName id="B5AA4D1ED34EFF96FE70FC99FD743060" authority="Panzer, 1813" authorityName="Panzer" authorityYear="1813" box="[430,668,870,895]" class="Insecta" family="Carabidae" genus="Broscus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2673" phylum="Arthropoda" rank="genus">
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<emphasis id="40DEEA8FD34EFF96FE70FC99FDEC3061" box="[430,516,872,895]" italics="true" pageId="4" pageNumber="2673">Broscus</emphasis>
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Panzer, 1813
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3AB06516D34EFF95FF0DFC35FEB73526" lastPageId="7" lastPageNumber="2676" pageId="4" pageNumber="2673" type="description">
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<paragraph id="7215369DD34EFF96FF0DFC35FEBA30C5" blockId="4.[211,1172,964,1664]" box="[211,338,964,987]" pageId="4" pageNumber="2673">
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<heading id="295D81F1D34EFF96FF0DFC35FEBA30C5" box="[211,338,964,987]" fontSize="10" level="3" pageId="4" pageNumber="2673" reason="5">
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<emphasis id="40DEEA8FD34EFF96FF0DFC35FEBA30C5" box="[211,338,964,987]" italics="true" pageId="4" pageNumber="2673">Description</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="7215369DD34EFF96FF0DFC1AFDE7371D" blockId="4.[211,1172,964,1664]" box="[211,527,1003,1027]" pageId="4" pageNumber="2673">
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Body length:
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<quantity id="B5529B78D34EFF96FEB4FC1AFDE4371D" box="[362,524,1003,1027]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" metricValueMax="1.3599999999999999" metricValueMin="1.16" pageId="4" pageNumber="2673" unit="mm" value="12.6" valueMax="13.6" valueMin="11.6">11.6–13.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="7215369DD34EFF96FF20FBFAFBCB375C" blockId="4.[211,1172,964,1664]" pageId="4" pageNumber="2673">Colour: Body surface reddish brown to piceous, glossy. Appendages reddish brown with mouthparts somewhat lighter and femora and tibiae somewhat darker.</paragraph>
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<paragraph id="7215369DD34EFF96FF20FBBBFEA4363C" blockId="4.[211,1172,964,1664]" pageId="4" pageNumber="2673">Micro-sculpture: Entire surface covered with micro-puncturation (magnification 100 ×). Surface of head with mesh pattern isodiametric, very indistinctly engraved, visible under high magnification (100 ×) except on labrum which has micro-meshes distinctly engraved (magnification 40 ×). Pronotum with mesh pattern somewhat irregular, very shallowly impressed (magnification 100 ×). Elytra with mesh pattern isodiametric, somewhat more deeply impressed than on head and pronotum.</paragraph>
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<paragraph id="7215369DD34EFF97FF20FADBFD633241" blockId="4.[211,1172,964,1664]" lastBlockId="5.[143,1104,167,1658]" lastPageId="5" lastPageNumber="2674" pageId="4" pageNumber="2673">
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Head: Large, moderately convex, somewhat narrower than pronotum, with surface smooth. Eyes relatively small, slightly prominent, with one (exceptionally with two) supraorbital seta at level of hind margin of eye (
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<figureCitation id="EA912A18D34EFF96FCFDFA98FC51369F" box="[803,953,1385,1409]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="4" pageNumber="2673">Figure 3A, C</figureCitation>
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). Tempora distinct, indistinctly swollen, smooth, about as long as length of eye diameter (viewed dorsally), temporal ridge incomplete. Frontal impressions shallow, divergent posteriorly. Vertex with an impunctate transverse impression that is deepened in middle (
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<figureCitation id="EA912A18D34EFF96FC33FA38FB6E36FF" box="[1005,1158,1481,1505]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="4" pageNumber="2673">Figure 3A, C</figureCitation>
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). Mandibles long and slender with seta in scrobe.
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<collectingRegion id="B06EF87FD34EFF96FCD2FA1BFC86351F" box="[780,878,1514,1537]" country="Myanmar" name="Chin" pageId="4" pageNumber="2673">Mentum</collectingRegion>
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with two setae each side and with pair of deep paramedian foveae; apical tooth truncate, shallowly emarginate, hence shallowly bifid; submentum with three setae each side, the middle one longer and broader (
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<figureCitation id="EA912A18D34EFF96FEB1F9B8FE09357F" box="[367,481,1609,1633]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="4" pageNumber="2673">Figure 3B</figureCitation>
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). Ligula in central part swollen, bisetose apically, paraglossa almost as long as ligula. Cardo with one, stipes with two or three setae. Gula with two deep transverse grooves on each side. Antennae relatively long (ninth or tenth antennomere extended to pronotal base), with scape glabrous apart from the dorsoapical seta; pedicel with one or two ventroapical setae; third antennomere glabrous apart from the ring of obligatory apical setae; fourth antennomere apart from the ring of long obligatory apical setae with numerous fine and short setae in apical third; 5th to 11th antennomeres pubescent on all surfaces.
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</paragraph>
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<paragraph id="7215369DD34FFF97FF64FE99FE6F31C3" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">
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Pronotum: Cordate, distinctly narrower than elytra (EW
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<emphasis id="40DEEA8FD34FFF97FCE7FE96FCAD3261" bold="true" box="[825,837,359,383]" italics="true" pageId="5" pageNumber="2674">/</emphasis>
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PW 1.54–1.69), as wide as long or somewhat wider than long (PW
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<emphasis id="40DEEA8FD34FFF97FD52FE76FD703281" bold="true" box="[652,664,391,415]" italics="true" pageId="5" pageNumber="2674">/</emphasis>
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PL 0.99–1.10), widest at apical third, strongly contracted towards base (PW
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<emphasis id="40DEEA8FD34FFF97FDE7FE56FDAD32A1" bold="true" box="[569,581,423,447]" italics="true" pageId="5" pageNumber="2674">/</emphasis>
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PBW 1.50–1.64), dorsally fairly convex. Apical margin somewhat wider than basal margin, with apical angles weakly protruded. Basal margin markedly convex. Lateral margin finely beaded, rounded in anterior four-fifths and rectilinearly divergent towards basal angles; latter obtuse, blunt at tip. Basal portion of epipleuron near basal angles visible in dorsal view (
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<figureCitation id="EA912A18D34FFF97FCEBFDD7FC5C3120" box="[821,948,550,574]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="5" pageNumber="2674">Figures 1A</figureCitation>
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,
|
||
<figureCitation id="EA912A18D34FFF97FC1AFDD7FC0E3120" box="[964,998,550,574]" captionStart="Figure 2" captionStartId="7.[143,210,1380,1402]" captionTargetBox="[145,1104,171,1346]" captionTargetId="figure-5@7.[144,1104,168,1347]" captionTargetPageId="7" captionText="Figure 2. Kashmirobroscus and Eobroscus in dorsal view. (A) Kashmirobroscus mirabilis sp. nov., holotype; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal." figureDoi="http://doi.org/10.5281/zenodo.4608241" httpUri="https://zenodo.org/record/4608241/files/figure.png" pageId="5" pageNumber="2674">2A</figureCitation>
|
||
). Lateral margin each side with three to five setae in apical half and one seta slightly anteriad basal angle. Median line moderately deep, not reaching apical and basal margin; anterior transverse impression indistinct, posterior transverse impression very shallow but distinct, laterobasal foveae narrow, shallow. Base more or less coarsely punctuate or transversely wrinkled.
|
||
</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FD14FC493063" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">Elytra: Long-ovate, flattened on disc; humeri widely rounded, with apical portion of lateral border not reflexed, terminated at level of fifth stria. Parascutellar seta and parascutellar stria absent. Striae on disc slightly impressed, distinctly punctured, almost absent at sides and near apex, intervals on disc weakly convex. Lateral umbilical series consists of one posthumeral, one preapical and two apical setae.</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FC74FCA93083" blockId="5.[143,1104,167,1658]" box="[186,833,901,925]" pageId="5" pageNumber="2674">Hind wings: Reduced to about one-third of elytral length.</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FC54FC123702" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">Sterna: Prosternal process furrowed, not margined; prosternum, pro- and mesepisternum, mesepimeron and metepisternum smooth or, in some specimens, with few shallow punctures. Metepisternum moderately long, with ventral margin approximately 1.5 times longer than anterior margin, moderately narrowed posteriad.</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FBD5FCFE37A2" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">
|
||
Abdomen: Smooth in middle, somewhat wrinkled laterally and with a fine irregular puncturation (
|
||
<figureCitation id="EA912A18D34FFF97FEEDFBB5FE4C3742" box="[307,420,1092,1116]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="5" pageNumber="2674">Figure 1B</figureCitation>
|
||
), sternites IV–VI each with one pair of ambulatory paramedian setae, sternite VI of male apically with a button-like, round elevation at middle (
|
||
<figureCitation id="EA912A18D34FFF97FF46FB75FEC63782" box="[152,302,1156,1180]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="5" pageNumber="2674">Figure 1B, C</figureCitation>
|
||
), last abdominal sternite apically usually with one seta on each side in male, two in female (but chaetotaxy sometimes irregular).
|
||
</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FB34FC683685" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">
|
||
Legs: Slender, with protrochanters bisetose and with mesotrochanters and metatrochanters unisetose. Male protarsomeres very indistinctly dilated, tarsomeres 1 and 2 with adhesive vestiture ventrally; mesotarsomeres without adhesive vestiture. Dorsal side of mesotarsomere and metatarsomere 1 (less on metatarsomeres 2) with a blunt longitudinal carina in middle that is bordered by a distinct longitudinal furrow on each side (
|
||
<figureCitation id="EA912A18D34FFF97FEDBFA92FE7F3665" box="[261,407,1379,1403]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="5" pageNumber="2674">Figure 4J, K</figureCitation>
|
||
). Tarsomere 5 on ventral side with one couple of setae behind middle (
|
||
<figureCitation id="EA912A18D34FFF97FF32FA72FEBE3685" box="[236,342,1411,1435]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="5" pageNumber="2674">Figure 4J</figureCitation>
|
||
, exceptionally with an additional seta in middle).
|
||
</paragraph>
|
||
<paragraph id="7215369DD34FFF97FF64FA52FD0F3564" blockId="5.[143,1104,167,1658]" pageId="5" pageNumber="2674">
|
||
Male genitalia: Sternum IX with sclerotized ring complete, with right margin markedly narrowed (
|
||
<figureCitation id="EA912A18D34FFF97FEA2FA32FE0036C5" box="[380,488,1475,1499]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="5" pageNumber="2674">Figure 5J</figureCitation>
|
||
). Median lobe of aedeagus with basal orifice partially open, with basal third of dorsal surface sclerotized, without basal expansions, but with wing-like expansions on lateral and ventrolateral surfaces (
|
||
<figureCitation id="EA912A18D34FFF97FCF8F9F2FC5D3505" box="[806,949,1539,1563]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="5" pageNumber="2674">Figure 5G, I</figureCitation>
|
||
). Internal sac with sclerite X wide, with a distinct constriction in middle and with two latero-apical projections (
|
||
<figureCitation id="EA912A18D34FFF97FEC3F9B3FE953544" box="[285,381,1602,1626]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="5" pageNumber="2674">Figure 5</figureCitation>
|
||
G–I); sclerite Y present; apical plate composed of free spiculae, without tooth. Both parameres setiferous (
|
||
<figureCitation id="EA912A18D34FFF97FDB0F993FD3E3564" box="[622,726,1634,1658]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="5" pageNumber="2674">Figure 5I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="26D56615D34CFF94FF0DFAA5FC543699" ID-DOI="http://doi.org/10.5281/zenodo.4608239" ID-Zenodo-Dep="4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="6" pageNumber="2675" startId="6.[211,278,1364,1386]" targetBox="[216,1169,169,1331]" targetPageId="6">
|
||
<paragraph id="7215369DD34CFF94FF0DFAA5FC543699" blockId="6.[211,1172,1364,1415]" pageId="6" pageNumber="2675">
|
||
Figure 1.
|
||
<taxonomicName id="B5AA4D1ED34CFF94FEE5FAA5FDD63674" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[315,574,1364,1386]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="2675" phylum="Arthropoda" rank="species" species="ortrudae" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD34CFF94FEE5FAA5FDD63674" box="[315,574,1364,1386]" italics="true" pageId="6" pageNumber="2675">Kashmirobroscus ortrudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D34CFF94FD9AFAA4FD7B3674" box="[580,659,1365,1386]" pageId="6" pageNumber="2675" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7215369DD34CFF94FF20FA5FFD8F36F8" blockId="6.[211,1171,1454,1510]" pageId="6" pageNumber="2675">
|
||
Female genitalia (
|
||
<figureCitation id="EA912A18D34CFF94FE10FA5FFDAD36D8" box="[462,581,1454,1478]" captionStart="Figure 6" captionStartId="13.[143,210,860,882]" captionTargetBox="[148,1104,169,827]" captionTargetId="figure-5@13.[144,1104,169,827]" captionTargetPageId="13" captionText="Figure 6. Female genital tract of Kashmirobroscus and Eobroscus in lateral view. (A) Kashmirobroscus ortrudae sp. nov, paratype, female; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviations: bc, bursa copulatrix; co, common oviduct; gc, gonocoxites, gsc, gonosubcoxites; hs, helminthoid sclerite; sp, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4608249" httpUri="https://zenodo.org/record/4608249/files/figure.png" pageId="6" pageNumber="2675">Figure 6A</figureCitation>
|
||
): Helminthoid sclerite of accessory gland of bursa copulatrix long and thin, U-shaped.
|
||
</paragraph>
|
||
<paragraph id="7215369DD34CFF94FF0DF9EBFE18352C" blockId="6.[211,1171,1562,1690]" box="[211,496,1562,1586]" pageId="6" pageNumber="2675">
|
||
<heading id="295D81F1D34CFF94FF0DF9EBFE18352C" box="[211,496,1562,1586]" fontSize="10" level="3" pageId="6" pageNumber="2675" reason="5">
|
||
<emphasis id="40DEEA8FD34CFF94FF0DF9EBFE18352C" box="[211,496,1562,1586]" italics="true" pageId="6" pageNumber="2675">Phylogenetic relationships</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7215369DD34CFF94FF0DF9B3FDBB3584" blockId="6.[211,1171,1562,1690]" pageId="6" pageNumber="2675">
|
||
Evidence for inclusion of
|
||
<taxonomicName id="B5AA4D1ED34CFF94FE2BF9B3FD593544" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[501,689,1602,1626]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="2675" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD34CFF94FE2BF9B3FD593544" box="[501,689,1602,1626]" italics="true" pageId="6" pageNumber="2675">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D34CFF94FD67F9B2FCF63544" box="[697,798,1603,1627]" pageId="6" pageNumber="2675" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
in the subtribe
|
||
<taxonomicName id="B5AA4D1ED34CFF94FC0EF9B3FBDB3544" authorityName="Hope" authorityYear="1831" box="[976,1075,1602,1626]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="2675" phylum="Arthropoda" rank="subTribe" subTribe="Broscina">Broscina</taxonomicName>
|
||
is based on the following two diagnostic features, which are considered synapomorphies (RoigJuñent 2000;
|
||
<bibRefCitation id="163B4B6CD34CFF94FEB6F973FDAB3584" author="Liebherr JK & Marris JWM & Emberson RM & Syrett P & Roig-Junent S" box="[360,579,1666,1690]" pageId="6" pageNumber="2675" pagination="395 - 414" refId="ref7455" refString="Liebherr JK, Marris JWM, Emberson RM, Syrett P, Roig-Junent S. 2011. Orthoglymma wangapeka gen. n., sp. n. (Coleoptera: Carabidae: Broscini): a newly discovered relict from the Buller Terrane, north-western South Island, New Zealand, corroborates a general pattern of Gondwanan endemism. Syst Entomol. 36: 395 - 414." type="journal article" year="2011">Liebherr et al. 2011</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
<caption id="26D56615D34DFF95FF51FA95FBA73689" ID-DOI="http://doi.org/10.5281/zenodo.4608241" ID-Zenodo-Dep="4608241" httpUri="https://zenodo.org/record/4608241/files/figure.png" pageId="7" pageNumber="2676" startId="7.[143,210,1380,1402]" targetBox="[145,1104,171,1346]" targetPageId="7">
|
||
<paragraph id="7215369DD34DFF95FF51FA95FBA73689" blockId="7.[143,1103,1380,1431]" pageId="7" pageNumber="2676">
|
||
Figure 2.
|
||
<taxonomicName id="B5AA4D1ED34DFF95FF28FA95FE773664" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[246,415,1380,1402]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD34DFF95FF28FA95FE773664" box="[246,415,1380,1402]" italics="true" pageId="7" pageNumber="2676">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED34DFF95FE11FA95FDDC3664" authorityName="Kryzhanovskij" authorityYear="1951" box="[463,564,1380,1402]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD34DFF95FE11FA95FDDC3664" box="[463,564,1380,1402]" italics="true" pageId="7" pageNumber="2676">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
in dorsal view. (A)
|
||
<taxonomicName id="B5AA4D1ED34DFF95FD2BFA95FC113664" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[757,1017,1380,1402]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD34DFF95FD2BFA95FC113664" box="[757,1017,1380,1402]" italics="true" pageId="7" pageNumber="2676">Kashmirobroscus mirabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D34DFF95FC20FA94FBA43664" box="[1022,1100,1381,1402]" pageId="7" pageNumber="2676" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; (B)
|
||
<taxonomicName id="B5AA4D1ED34DFF95FEC5FA70FD7A3689" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[283,658,1409,1431]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD34DFF95FEC5FA70FE113689" box="[283,505,1409,1431]" italics="true" pageId="7" pageNumber="2676">Eobroscus bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD34DFF95FE21FA73FD7A3689" author="Morvan P" box="[511,658,1409,1431]" pageId="7" pageNumber="2676" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, female specimen from Mt Manaslu, Nepal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7215369DD34DFF95FF74FA30FDC536E7" blockId="7.[170,1103,1473,1592]" pageId="7" pageNumber="2676">
|
||
Sclerite X in the internal sac of aedeagal median lobe with upturned latero-apical projections (see
|
||
<figureCitation id="EA912A18D34DFF95FE4FFA10FE0736E7" box="[401,495,1505,1529]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="7" pageNumber="2676">Figure 5</figureCitation>
|
||
G–I).
|
||
</paragraph>
|
||
<paragraph id="7215369DD34DFF95FF74F9F1FEB73526" blockId="7.[170,1103,1473,1592]" pageId="7" pageNumber="2676">
|
||
Helminthoid sclerite of accessory gland of bursa copulatrix U-shaped (see
|
||
<figureCitation id="EA912A18D34DFF95FF02F9D1FEA73526" box="[220,335,1568,1592]" captionStart="Figure 6" captionStartId="13.[143,210,860,882]" captionTargetBox="[148,1104,169,827]" captionTargetId="figure-5@13.[144,1104,169,827]" captionTargetPageId="13" captionText="Figure 6. Female genital tract of Kashmirobroscus and Eobroscus in lateral view. (A) Kashmirobroscus ortrudae sp. nov, paratype, female; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviations: bc, bursa copulatrix; co, common oviduct; gc, gonocoxites, gsc, gonosubcoxites; hs, helminthoid sclerite; sp, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4608249" httpUri="https://zenodo.org/record/4608249/files/figure.png" pageId="7" pageNumber="2676">Figure 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB06516D34DFF9EFF64F993FB923201" lastPageId="12" lastPageNumber="2681" pageId="7" pageNumber="2676" type="discussion">
|
||
<paragraph id="7215369DD34DFF9AFF64F993FEBE354B" blockId="7.[143,1103,1634,1691]" lastBlockId="8.[211,1172,1565,1621]" lastPageId="8" lastPageNumber="2677" pageId="7" pageNumber="2676">
|
||
Within
|
||
<taxonomicName id="B5AA4D1ED34DFF95FEFDF993FE613564" authorityName="Hope" authorityYear="1831" box="[291,393,1634,1658]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="subTribe" subTribe="Broscina">Broscina</taxonomicName>
|
||
,
|
||
<taxonomicName id="B5AA4D1ED34DFF95FE76F993FD8C3564" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[424,612,1634,1658]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD34DFF95FE76F993FD8C3564" box="[424,612,1634,1658]" italics="true" pageId="7" pageNumber="2676">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D34DFF95FDA1F992FD133564" box="[639,763,1635,1659]" pageId="7" pageNumber="2676" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, together with
|
||
<taxonomicName id="B5AA4D1ED34DFF95FC01F993FE913584" authority="Kryzhanovskij, 1951" authorityName="Kryzhanovskij" authorityYear="1951" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2687" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD34DFF95FC01F993FBA73564" box="[991,1103,1634,1658]" italics="true" pageId="7" pageNumber="2676">Eobroscus</emphasis>
|
||
Kryzhanovskij, 1951
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED34DFF95FE6CF972FD523584" authority="LeConte, 1869" authorityName="LeConte" authorityYear="1869" box="[434,698,1666,1690]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2676" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD34DFF95FE6CF972FDE43584" box="[434,524,1667,1690]" italics="true" pageId="7" pageNumber="2676">Zacotus</emphasis>
|
||
LeConte, 1869
|
||
</taxonomicName>
|
||
, seems to form a natural group that is defined by the following two derived character states (
|
||
<bibRefCitation id="163B4B6CD342FF9AFC87F9ECFBCA352B" author="Roig-Junent SA" box="[857,1058,1565,1589]" pageId="8" pageNumber="2677" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="163B4B6CD342FF9AFBECF9ECFEAE354B" author="Liebherr JK & Marris JWM & Emberson RM & Syrett P & Roig-Junent S" pageId="8" pageNumber="2677" pagination="395 - 414" refId="ref7455" refString="Liebherr JK, Marris JWM, Emberson RM, Syrett P, Roig-Junent S. 2011. Orthoglymma wangapeka gen. n., sp. n. (Coleoptera: Carabidae: Broscini): a newly discovered relict from the Buller Terrane, north-western South Island, New Zealand, corroborates a general pattern of Gondwanan endemism. Syst Entomol. 36: 395 - 414." type="journal article" year="2011">Liebherr et al. 2011</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
<caption id="26D56615D342FF9AFF0DFA85FCAD36FF" ID-DOI="http://doi.org/10.5281/zenodo.4608243" ID-Zenodo-Dep="4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="8" pageNumber="2677" startId="8.[211,278,1396,1418]" targetBox="[212,1172,168,1363]" targetPageId="8">
|
||
<paragraph id="7215369DD342FF9AFF0DFA85FCAD36FF" blockId="8.[211,1172,1396,1505]" pageId="8" pageNumber="2677">
|
||
Figure 3. Head in dorsal (A, C, D) and ventral (B) views of
|
||
<taxonomicName id="B5AA4D1ED342FF9AFC97FA85FC1A3694" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[841,1010,1396,1418]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2677" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD342FF9AFC97FA85FC1A3694" box="[841,1010,1396,1418]" italics="true" pageId="8" pageNumber="2677">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED342FF9AFBF6FA85FB653694" authorityName="Kryzhanovskij" authorityYear="1951" box="[1064,1165,1396,1418]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2677" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD342FF9AFBF6FA85FB653694" box="[1064,1165,1396,1418]" italics="true" pageId="8" pageNumber="2677">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
. (A, B)
|
||
<taxonomicName id="B5AA4D1ED342FF9AFEC8FA60FDF036B9" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[278,536,1425,1447]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2677" phylum="Arthropoda" rank="species" species="ortrudae" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD342FF9AFEC8FA60FDF036B9" box="[278,536,1425,1447]" italics="true" pageId="8" pageNumber="2677">Kashmirobroscus ortrudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D342FF9AFDC3FA63FD8336B9" box="[541,619,1426,1447]" pageId="8" pageNumber="2677" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, male; (C)
|
||
<taxonomicName id="B5AA4D1ED342FF9AFCE7FA60FBD536B9" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[825,1085,1425,1447]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2677" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD342FF9AFCE7FA60FBD536B9" box="[825,1085,1425,1447]" italics="true" pageId="8" pageNumber="2677">Kashmirobroscus mirabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D342FF9AFB9CFA63FB7836B9" box="[1090,1168,1426,1447]" pageId="8" pageNumber="2677" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; (D)
|
||
<taxonomicName id="B5AA4D1ED342FF9AFEBCFA5FFD3036DA" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[354,728,1454,1476]" class="Insecta" family="Carabidae" genus="Eobruscus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="2677" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD342FF9AFEBCFA5FFDA936DA" box="[354,577,1454,1476]" italics="true" pageId="8" pageNumber="2677">Eobruscus bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD342FF9AFD99FA5EFD3036DA" author="Morvan P" box="[583,728,1454,1476]" pageId="8" pageNumber="2677" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7215369DD342FF9AFF30F98CFBA435AB" blockId="8.[238,1160,1661,1717]" pageId="8" pageNumber="2677">
|
||
Gula on each side with two deep transverse grooves or oblique foveae (
|
||
<figureCitation id="EA912A18D342FF9AFBD6F98CFB90358B" box="[1032,1144,1661,1685]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="8" pageNumber="2677">Figure 3B</figureCitation>
|
||
). Helminthoid sclerite of the female genitalia thin and long (see
|
||
<figureCitation id="EA912A18D342FF9AFC77F96CFBD735AB" box="[937,1087,1693,1717]" captionStart="Figure 6" captionStartId="13.[143,210,860,882]" captionTargetBox="[148,1104,169,827]" captionTargetId="figure-5@13.[144,1104,169,827]" captionTargetPageId="13" captionText="Figure 6. Female genital tract of Kashmirobroscus and Eobroscus in lateral view. (A) Kashmirobroscus ortrudae sp. nov, paratype, female; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviations: bc, bursa copulatrix; co, common oviduct; gc, gonocoxites, gsc, gonosubcoxites; hs, helminthoid sclerite; sp, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4608249" httpUri="https://zenodo.org/record/4608249/files/figure.png" pageId="8" pageNumber="2677">Figure 6A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF64FF56FBE333E1" blockId="9.[143,1103,167,255]" pageId="9" pageNumber="2678">
|
||
Within this group the East Asian genus
|
||
<taxonomicName id="B5AA4D1ED343FF9BFDA1FF59FD0733DE" authorityName="Kryzhanovskij" authorityYear="1951" box="[639,751,168,192]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFDA1FF59FD0733DE" box="[639,751,168,192]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
and the Northwest Himalayan genus
|
||
<taxonomicName id="B5AA4D1ED343FF9BFF06FF39FE7C33FE" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[216,404,200,224]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD343FF9BFF06FF39FE7C33FE" box="[216,404,200,224]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D343FF9BFE42FF39FDEB33C1" box="[412,515,200,224]" pageId="9" pageNumber="2678" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
have six characters in common that are considered synapomorphies and so, allow for the assumption of a sister-group relationship:
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FED9FBCF3003" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Transverse impression on vertex markedly deep (
|
||
<figureCitation id="EA912A18D343FF9BFD31FED6FC513221" box="[751,953,295,319]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="9" pageNumber="2678">Figure 3A, C, D</figureCitation>
|
||
). Head with vertex smooth seems to represent the plesiomorphic character state as it is developed in most
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE66FE96FDFD3261" authorityName="Roig-Junent" authorityYear="2000" box="[440,533,359,383]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
genera and in the outgroup taxa
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC41FE96FBF33261" box="[927,1051,359,383]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Apotomini">Apotomini</taxonomicName>
|
||
and Melaenini. A transverse impression is also developed in the
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC6EFE76FBFB3281" authorityName="Hope" authorityYear="1831" box="[944,1043,391,415]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="subTribe" subTribe="Broscina">Broscina</taxonomicName>
|
||
genera
|
||
<taxonomicName id="B5AA4D1ED343FF9BFED6FE56FDD032A0" authority="Lindroth, 1961" authorityName="Lindroth" authorityYear="1961" box="[264,568,422,447]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFED6FE56FE6B32A1" box="[264,387,423,447]" italics="true" pageId="9" pageNumber="2678">Broscodera</emphasis>
|
||
Lindroth, 1961
|
||
</taxonomicName>
|
||
(incl.
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD5DFE56FC5D32A0" authority="Deuve, 1990" authorityName="Deuve" authorityYear="1990" box="[643,949,422,447]" class="Insecta" family="Carabidae" genus="Sinobrosculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFD5DFE56FCF332A1" box="[643,795,423,447]" italics="true" pageId="9" pageNumber="2678">Sinobrosculus</emphasis>
|
||
Deuve, 1990
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC14FE59FE4E32C0" authority="Rosenhauer, 1846" authorityName="Rosenhauer" authorityYear="1846" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC14FE59FBA732A1" box="[970,1103,424,447]" italics="true" pageId="9" pageNumber="2678">Broscosoma</emphasis>
|
||
Rosenhauer, 1846
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE38FE36FCDF32C0" authority="Schaum, 1863" authorityName="Schaum" authorityYear="1863" box="[486,823,454,479]" class="Insecta" family="Carabidae" genus="Craspedonotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFE38FE36FD6032C1" box="[486,648,455,479]" italics="true" pageId="9" pageNumber="2678">Craspedonotus</emphasis>
|
||
Schaum, 1863
|
||
</taxonomicName>
|
||
; however, it is distinctly shallower in these taxa than in
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD8EFE16FD2832E1" authorityName="Kryzhanovskij" authorityYear="1951" box="[592,704,487,511]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFD8EFE16FD2832E1" box="[592,704,487,511]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
and in
|
||
<taxonomicName id="B5AA4D1ED343FF9BFCFFFE16FC3532E1" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[801,989,487,511]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD343FF9BFCFFFE16FC3532E1" box="[801,989,487,511]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D343FF9BFC39FE16FBB832E0" box="[999,1104,487,511]" pageId="9" pageNumber="2678" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
In
|
||
<taxonomicName id="B5AA4D1ED343FF9BFEDCFDF6FD703100" authority="LeConte, 1869" authorityName="LeConte" authorityYear="1869" box="[258,664,518,542]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="matthewsii">
|
||
<emphasis id="40DEEA8FD343FF9BFEDCFDF6FE373100" box="[258,479,518,542]" italics="true" pageId="9" pageNumber="2678">Zacotus matthewsii</emphasis>
|
||
LeConte, 1869
|
||
</taxonomicName>
|
||
, which is a supposed sister taxon of
|
||
<taxonomicName id="B5AA4D1ED343FF9BFF02FDD7FEA43120" authorityName="Kryzhanovskij" authorityYear="1951" box="[220,332,550,574]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFF02FDD7FEA43120" box="[220,332,550,574]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="B5AA4D1ED343FF9BFEA1FDD7FDD33120" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[383,571,550,574]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFEA1FDD7FDD33120" box="[383,571,550,574]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
, a deep short longitudinal furrow exists in the middle of the vertex beside a shallow transverse impression. Hence, the markedly deep transverse impression of vertex in
|
||
<taxonomicName id="B5AA4D1ED343FF9BFCDCFD97FC9A3160" authorityName="Kryzhanovskij" authorityYear="1951" box="[770,882,614,638]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFCDCFD97FC9A3160" box="[770,882,614,638]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC4DFD97FBA73160" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[915,1103,614,638]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC4DFD97FBA73160" box="[915,1103,614,638]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
seems to represent a character state that evolved independently from that of the
|
||
<taxonomicName id="B5AA4D1ED343FF9BFED6FD57FE8331A0" authorityName="Hope" authorityYear="1831" box="[264,363,678,702]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="subTribe" subTribe="Broscina">Broscina</taxonomicName>
|
||
genera named above.
|
||
<bibRefCitation id="163B4B6CD343FF9BFDB6FD57FC8C31A0" author="Roig-Junent SA" box="[616,868,678,702]" pageId="9" pageNumber="2678" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent (2000: 27</bibRefCitation>
|
||
, fig. 72) figured misleadingly a transverse impression also on vertex of
|
||
<taxonomicName id="B5AA4D1ED343FF9BFCC2FD37FEF631E3" authority="(Paykull, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" class="Insecta" family="Carabidae" genus="Miscodera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="arctica">
|
||
<emphasis id="40DEEA8FD343FF9BFCC2FD37FC0B31C3" box="[796,995,710,734]" italics="true" pageId="9" pageNumber="2678">Miscodera arctica</emphasis>
|
||
(Paykull, 1798)
|
||
</taxonomicName>
|
||
. Instead, this species has the dorsal surface of head completely smooth and rounded in basal half but has a distinct transverse impression on frons.
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FCD4FCA53040" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Sternite VI of male apically with a button-like elevation at middle (
|
||
<figureCitation id="EA912A18D343FF9BFC75FCD4FBAA3023" box="[939,1090,805,829]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="9" pageNumber="2678">Figure 1B, C</figureCitation>
|
||
). This character is unique within
|
||
<taxonomicName id="B5AA4D1ED343FF9BFDE1FCB4FD743043" authorityName="Roig-Junent" authorityYear="2000" box="[575,668,837,861]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
and outgroups.
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FC94FE8030A3" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Protarsi of male weakly dilated (
|
||
<figureCitation id="EA912A18D343FF9BFDFCFC94FD2A3063" box="[546,706,869,893]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4D, G</figureCitation>
|
||
).
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD06FC97FC5A3063" authorityName="LeConte" authorityYear="1869" box="[728,946,869,893]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="matthewsii">
|
||
<emphasis id="40DEEA8FD343FF9BFD06FC97FC5A3063" box="[728,946,869,893]" italics="true" pageId="9" pageNumber="2678">Zacotus matthewsii</emphasis>
|
||
</taxonomicName>
|
||
possesses the plesiomorphic character state with male protarsomere 1–3 markedly dilated (
|
||
<figureCitation id="EA912A18D343FF9BFF3BFC54FEB030A3" box="[229,344,933,957]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FC34FDDA3605" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Dorsolateral surfaces of tarsi with longitudinal furrows, at least on first hind tarsomere (see
|
||
<figureCitation id="EA912A18D343FF9BFE4EFC15FDEF30E2" box="[400,519,996,1020]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4F</figureCitation>
|
||
, I–K). Tarsi with dorsal side completely smooth (see
|
||
<figureCitation id="EA912A18D343FF9BFECEFBF5FE6C3702" box="[272,388,1028,1052]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4C</figureCitation>
|
||
for
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE66FBF4FD7B3703" authorityName="LeConte" authorityYear="1869" box="[440,659,1029,1053]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="matthewsii">
|
||
<emphasis id="40DEEA8FD343FF9BFE66FBF4FD7B3703" box="[440,659,1029,1053]" italics="true" pageId="9" pageNumber="2678">Zacotus matthewsii</emphasis>
|
||
</taxonomicName>
|
||
) seems to represent the plesiomorphic character state as it is developed in other
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD07FBD5FCDE3722" authorityName="Roig-Junent" authorityYear="2000" box="[729,822,1060,1084]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
(except for
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC15FBD4FBAE3723" authorityName="Lindroth" authorityYear="1961" box="[971,1094,1061,1085]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC15FBD4FBAE3723" box="[971,1094,1061,1085]" italics="true" pageId="9" pageNumber="2678">Broscodera</emphasis>
|
||
</taxonomicName>
|
||
) and outgroups (
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE49FBB5FDFF3742" box="[407,535,1092,1116]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Apotomini">Apotomini</taxonomicName>
|
||
, Melaenini). In contrast to what is developed in
|
||
<taxonomicName id="B5AA4D1ED343FF9BFF02FB95FEA43762" authorityName="Kryzhanovskij" authorityYear="1951" box="[220,332,1124,1148]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFF02FB95FEA43762" box="[220,332,1124,1148]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE57FB95FDAD3762" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[393,581,1124,1148]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFE57FB95FDAD3762" box="[393,581,1124,1148]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
, which have dorsal surfaces of all tarsomeres glabrous (except for the pair of dorsoapical setae on fifth tarsomere), the longitudinal furrows on dorsal surfaces of the tarsomeres of
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC9EFB55FC5337A2" authorityName="Lindroth" authorityYear="1961" box="[832,955,1188,1212]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC9EFB55FC5337A2" box="[832,955,1188,1212]" italics="true" pageId="9" pageNumber="2678">Broscodera</emphasis>
|
||
</taxonomicName>
|
||
are bordered by fine keels which each terminate in a fine seta. It is therefore most likely that longitudinal furrows in
|
||
<taxonomicName id="B5AA4D1ED343FF9BFDF8FB15FD4937E2" authorityName="Lindroth" authorityYear="1961" box="[550,673,1252,1276]" class="Insecta" family="Carabidae" genus="Broscodera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFDF8FB15FD4937E2" box="[550,673,1252,1276]" italics="true" pageId="9" pageNumber="2678">Broscodera</emphasis>
|
||
</taxonomicName>
|
||
evolved independently from those of
|
||
<taxonomicName id="B5AA4D1ED343FF9BFF02FAF5FEA43602" authorityName="Kryzhanovskij" authorityYear="1951" box="[220,332,1284,1308]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFF02FAF5FEA43602" box="[220,332,1284,1308]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="B5AA4D1ED343FF9BFEB1FAF5FDC33602" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[367,555,1284,1308]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFEB1FAF5FDC33602" box="[367,555,1284,1308]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FAD2FBB83662" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Fifth tarsomere with one or two pairs of ventral seate (
|
||
<figureCitation id="EA912A18D343FF9BFCDAFAD2FC8B3625" box="[772,867,1315,1339]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4</figureCitation>
|
||
D–K). Three or more pairs of setae (see
|
||
<figureCitation id="EA912A18D343FF9BFE70FAB2FDE63645" box="[430,526,1347,1371]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="9" pageNumber="2678">Figure 4</figureCitation>
|
||
A–C for
|
||
<taxonomicName id="B5AA4D1ED343FF9BFDAAFAB5FCA53642" authorityName="LeConte" authorityYear="1869" box="[628,845,1348,1372]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="matthewsii">
|
||
<emphasis id="40DEEA8FD343FF9BFDAAFAB5FCA53642" box="[628,845,1348,1372]" italics="true" pageId="9" pageNumber="2678">Zacotus matthewsii</emphasis>
|
||
</taxonomicName>
|
||
) seem to represent the plesiomorphic character state as it is developed in other
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC9BFA92FC4A3665" authorityName="Roig-Junent" authorityYear="2000" box="[837,930,1379,1403]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
and outgroups.
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF74FA72FEB33524" blockId="9.[170,1104,295,1594]" pageId="9" pageNumber="2678">
|
||
Aedeagal median lobe with wing-like lateral expansions in middle portion (
|
||
<figureCitation id="EA912A18D343FF9BFF3BFA52FEAA36A5" box="[229,322,1443,1467]" captionStart="Figure 5" captionStartId="11.[143,210,1350,1372]" captionTargetBox="[144,1103,170,1317]" captionTargetId="figure-5@11.[144,1104,167,1317]" captionTargetPageId="11" captionText="Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of Kashmirobroscus and Eobroscus. (A, B) Eobroscus lutshniki (Roubal, 1928), specimen from Ussurisk, Russia; (C–F) Eobroscus bhutanensis Morvan, 1982 (C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J), Kashmirobroscus ortrudae sp. nov., paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere." figureDoi="http://doi.org/10.5281/zenodo.4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="9" pageNumber="2678">Figure 5</figureCitation>
|
||
A–G, I). Except for some Himalayan species of the genus
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC14FA55FBA736A5" authorityName="Rosenhauer" authorityYear="1846" box="[970,1103,1444,1467]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC14FA55FBA736A5" box="[970,1103,1444,1467]" italics="true" pageId="9" pageNumber="2678">Broscosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="B5AA4D1ED343FF9BFF3BFA35FE9C36C5" authorityName="Habu" authorityYear="1973" box="[229,372,1475,1499]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="species" species="monticola">
|
||
<emphasis id="40DEEA8FD343FF9BFF3BFA35FE9C36C5" box="[229,372,1475,1499]" italics="true" pageId="9" pageNumber="2678">B. monticola</emphasis>
|
||
</taxonomicName>
|
||
group), which have a single sail-like expansion on the ventral surface this character is unique within
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD7DFA13FCE836E4" authorityName="Roig-Junent" authorityYear="2000" box="[675,768,1506,1530]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
and outgroups. Independent evolution of the ventral expansion within a terminal lineage of
|
||
<taxonomicName id="B5AA4D1ED343FF9BFC6EF9F2FBDD3504" authorityName="Rosenhauer" authorityYear="1846" box="[944,1077,1539,1562]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFC6EF9F2FBDD3504" box="[944,1077,1539,1562]" italics="true" pageId="9" pageNumber="2678">Broscosoma</emphasis>
|
||
</taxonomicName>
|
||
is most likely.
|
||
</paragraph>
|
||
<paragraph id="7215369DD343FF9BFF64F992FC5D3584" blockId="9.[143,1104,1634,1690]" pageId="9" pageNumber="2678">
|
||
In addition, the monotypic genus
|
||
<taxonomicName id="B5AA4D1ED343FF9BFD92F992FD4E3564" authorityName="LeConte" authorityYear="1869" box="[588,678,1635,1658]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFD92F992FD4E3564" box="[588,678,1635,1658]" italics="true" pageId="9" pageNumber="2678">Zacotus</emphasis>
|
||
</taxonomicName>
|
||
from North America can be distinguished from
|
||
<taxonomicName id="B5AA4D1ED343FF9BFEF4F973FE723584" authorityName="Kryzhanovskij" authorityYear="1951" box="[298,410,1666,1690]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFEF4F973FE723584" box="[298,410,1666,1690]" italics="true" pageId="9" pageNumber="2678">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="B5AA4D1ED343FF9BFE62F973FD903584" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[444,632,1666,1690]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2678" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD343FF9BFE62F973FD903584" box="[444,632,1666,1690]" italics="true" pageId="9" pageNumber="2678">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
by the following characters:
|
||
</paragraph>
|
||
<caption id="26D56615D340FF98FF0DF9F9FC6635D3" ID-DOI="http://doi.org/10.5281/zenodo.4608245" ID-Zenodo-Dep="4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="10" pageNumber="2679" startId="10.[211,278,1544,1566]" targetBox="[238,1144,162,1511]" targetPageId="10">
|
||
<paragraph id="7215369DD340FF98FF0DF9F9FC6635D3" blockId="10.[211,1172,1544,1741]" pageId="10" pageNumber="2679">
|
||
Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of
|
||
<taxonomicName id="B5AA4D1ED340FF98FE7BF9D4FE143525" authorityName="Roig-Junent" authorityYear="2000" box="[421,508,1573,1595]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
|
||
. (A–C)
|
||
<taxonomicName id="B5AA4D1ED340FF98FD8EF9D7FC5E3525" authority="LeConte, 1869" authorityName="LeConte" authorityYear="1869" box="[592,950,1573,1595]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="species" species="matthewsii">
|
||
<emphasis id="40DEEA8FD340FF98FD8EF9D7FCFC3525" box="[592,788,1573,1595]" italics="true" pageId="10" pageNumber="2679">Zacotus matthewsii</emphasis>
|
||
LeConte, 1869
|
||
</taxonomicName>
|
||
, male specimen from Vancouver, Canada; (D–F)
|
||
<taxonomicName id="B5AA4D1ED340FF98FE2EF9B3FCBB3546" authority="(Roubal, 1928)" baseAuthorityName="Roubal" baseAuthorityYear="1928" box="[496,851,1602,1624]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD340FF98FE2EF9B3FD593546" box="[496,689,1602,1624]" italics="true" pageId="10" pageNumber="2679">Eobroscus lutshniki</emphasis>
|
||
(Roubal, 1928)
|
||
</taxonomicName>
|
||
, male specimen from Ussurisk, Russia; (G–I),
|
||
<taxonomicName id="B5AA4D1ED340FF98FEAEF991FD07356B" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[368,751,1631,1654]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD340FF98FEAEF991FDBA3568" box="[368,594,1632,1654]" italics="true" pageId="10" pageNumber="2679">Eobroscus bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD340FF98FD82F991FD07356B" author="Morvan P" box="[604,751,1631,1653]" pageId="10" pageNumber="2679" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
(G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J)
|
||
<taxonomicName id="B5AA4D1ED340FF98FC79F98CFF1D35AE" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="species" species="ortrudae" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD340FF98FC79F98CFF1D35AE" italics="true" pageId="10" pageNumber="2679">Kashmirobroscus ortrudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D340FF98FF21F96AFEB835AE" box="[255,336,1691,1712]" pageId="10" pageNumber="2679" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, male; (K)
|
||
<taxonomicName id="B5AA4D1ED340FF98FDEFF96BFCD135AE" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[561,825,1690,1712]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2679" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD340FF98FDEFF96BFCD135AE" box="[561,825,1690,1712]" italics="true" pageId="10" pageNumber="2679">Kashmirobroscus mirabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D340FF98FC9CF96AFC7B35AE" box="[834,915,1691,1712]" pageId="10" pageNumber="2679" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="26D56615D341FF99FF51FAB7FE513512" ID-DOI="http://doi.org/10.5281/zenodo.4608247" ID-Zenodo-Dep="4608247" httpUri="https://zenodo.org/record/4608247/files/figure.png" pageId="11" pageNumber="2680" startId="11.[143,210,1350,1372]" targetBox="[144,1103,170,1317]" targetPageId="11">
|
||
<paragraph id="7215369DD341FF99FF51FAB7FE513512" blockId="11.[143,1104,1350,1548]" pageId="11" pageNumber="2680">
|
||
Figure 5. Aedeagal median lobe in left lateral view (A, C, E, G; H shows a cut-out of G) and in dorsal view (B, D, F, I; B and I additionally with parameres), and sternum IX (J) of
|
||
<taxonomicName id="B5AA4D1ED341FF99FF72FA70FEBD3689" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[172,341,1409,1431]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2680" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD341FF99FF72FA70FEBD3689" box="[172,341,1409,1431]" italics="true" pageId="11" pageNumber="2680">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5AA4D1ED341FF99FE55FA70FE183689" authorityName="Kryzhanovskij" authorityYear="1951" box="[395,496,1409,1431]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2680" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD341FF99FE55FA70FE183689" box="[395,496,1409,1431]" italics="true" pageId="11" pageNumber="2680">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
. (A, B)
|
||
<taxonomicName id="B5AA4D1ED341FF99FD96FA70FC453689" authority="(Roubal, 1928)" baseAuthorityName="Roubal" baseAuthorityYear="1928" box="[584,941,1409,1431]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2680" phylum="Arthropoda" rank="species" species="lutshniki">
|
||
<emphasis id="40DEEA8FD341FF99FD96FA70FCE23689" box="[584,778,1409,1431]" italics="true" pageId="11" pageNumber="2680">Eobroscus lutshniki</emphasis>
|
||
(Roubal, 1928)
|
||
</taxonomicName>
|
||
, specimen from Ussurisk, Russia; (C–F)
|
||
<taxonomicName id="B5AA4D1ED341FF99FE56FA6FFD1436AA" authority="Morvan, 1982" authorityName="Morvan" authorityYear="1982" box="[392,764,1438,1460]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2680" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="40DEEA8FD341FF99FE56FA6FFD8F36AA" box="[392,615,1438,1460]" italics="true" pageId="11" pageNumber="2680">Eobroscus bhutanensis</emphasis>
|
||
<bibRefCitation id="163B4B6CD341FF99FDB3FA6EFD1436AA" author="Morvan P" box="[621,764,1438,1460]" pageId="11" pageNumber="2680" pagination="77 - 88" refId="ref7585" refString="Morvan P. 1982. Quatre nouveaux Carabiques du Bhoutan (4 e note) (Coleoptera: Carabidae, Broscini, Pterostichini, Anchomenini). Entomol Basiliensia. 7: 77 - 88." type="journal article" year="1982">Morvan, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
(C, D, specimen from Qingcheng Houshan, China; E, F, specimen from Mt Fansipan, Vietnam); (G–J),
|
||
<taxonomicName id="B5AA4D1ED341FF99FCBBFA4DFF5936F1" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2680" phylum="Arthropoda" rank="species" species="ortrudae" status="sp. nov.">
|
||
<emphasis id="40DEEA8FD341FF99FCBBFA4DFF5936F1" italics="true" pageId="11" pageNumber="2680">Kashmirobroscus ortrudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D341FF99FF66FA28FEEF36F0" box="[184,263,1497,1518]" pageId="11" pageNumber="2680" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, male. Abbreviations: lap, latero-apical projections of internal sac; pl, left paramere; pr, right paramere.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7215369DD341FF99FF74F9B3FCEF3544" blockId="11.[170,790,1602,1690]" box="[170,775,1602,1626]" pageId="11" pageNumber="2680">Dorsal side of body with splendid metallic coloration.</paragraph>
|
||
<paragraph id="7215369DD341FF99FF74F993FCEB3564" blockId="11.[170,790,1602,1690]" box="[170,771,1634,1658]" pageId="11" pageNumber="2680">Fourth antennomere glabrous in basal three-quarters.</paragraph>
|
||
<paragraph id="7215369DD341FF99FF74F973FCFE3584" blockId="11.[170,790,1602,1690]" box="[170,790,1666,1690]" pageId="11" pageNumber="2680">Dorsal side of fifth tarsomere with numerous fine setae.</paragraph>
|
||
<paragraph id="7215369DD346FF9EFF20FF56FB923201" blockId="12.[211,1172,167,287]" pageId="12" pageNumber="2681">
|
||
The character of incomplete temporal ridge, given by
|
||
<bibRefCitation id="163B4B6CD346FF9EFCB9FF56FB8233A1" author="Roig-Junent SA" box="[871,1130,167,191]" pageId="12" pageNumber="2681" pagination="1 - 90" refId="ref7621" refString="Roig-Junent SA. 2000. The subtribes and genera of the tribe Broscini (Coleoptera: Carabidae): cladistic analysis, taxonomic treatment and biogeographical considerations. B Am Mus Nat Hist. 255: 1 - 90." type="journal article" year="2000">Roig-Juñent (2000: 13)</bibRefCitation>
|
||
for separating
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE8FFF39FE4333C1" authorityName="LeConte" authorityYear="1869" box="[337,427,200,223]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFE8FFF39FE4333C1" box="[337,427,200,223]" italics="true" pageId="12" pageNumber="2681">Zacotus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE2BFF39FD8D33FE" authorityName="Kryzhanovskij" authorityYear="1951" box="[501,613,200,224]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFE2BFF39FD8D33FE" box="[501,613,200,224]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
, is not useful, as the ridge is also incomplete in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFF0DFF16FEAB33E1" authorityName="Kryzhanovskij" authorityYear="1951" box="[211,323,231,255]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFF0DFF16FEAB33E1" box="[211,323,231,255]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
species, though the area of interruption is not completely smooth and convex (as in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFEB5FEF9FE2D3201" authorityName="LeConte" authorityYear="1869" box="[363,453,264,287]" class="Insecta" family="Carabidae" genus="Zacotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFEB5FEF9FE2D3201" box="[363,453,264,287]" italics="true" pageId="12" pageNumber="2681">Zacotus</emphasis>
|
||
</taxonomicName>
|
||
), i.e. it is somewhat angled between the upper and lower part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB06516D346FF9EFF0DFEADFD3030C7" pageId="12" pageNumber="2681" type="diagnosis">
|
||
<paragraph id="7215369DD346FF9EFF0DFEADFE56326A" blockId="12.[211,1172,348,985]" box="[211,446,348,372]" pageId="12" pageNumber="2681">
|
||
<heading id="295D81F1D346FF9EFF0DFEADFE56326A" box="[211,446,348,372]" fontSize="10" level="3" pageId="12" pageNumber="2681" reason="5">
|
||
<emphasis id="40DEEA8FD346FF9EFF0DFEADFE56326A" box="[211,446,348,372]" italics="true" pageId="12" pageNumber="2681">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7215369DD346FF9EFF0DFE75FD3030C7" blockId="12.[211,1172,348,985]" pageId="12" pageNumber="2681">
|
||
In
|
||
<taxonomicName id="B5AA4D1ED346FF9EFF2CFE75FE8A3282" authorityName="Kryzhanovskij" authorityYear="1951" box="[242,354,388,412]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFF2CFE75FE8A3282" box="[242,354,388,412]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
the transverse neck constriction is much more sharply incised (as if made with an axe, autapomorphy of
|
||
<taxonomicName id="B5AA4D1ED346FF9EFDF6FE55FD7032A2" authorityName="Kryzhanovskij" authorityYear="1951" box="[552,664,420,444]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFDF6FE55FD7032A2" box="[552,664,420,444]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<figureCitation id="EA912A18D346FF9EFD12FE52FCD732A5" box="[716,831,419,443]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="12" pageNumber="2681">Figure 3D</figureCitation>
|
||
) than in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFC7DFE55FBB732A2" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[931,1119,420,444]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFC7DFE55FBB732A2" box="[931,1119,420,444]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFBBAFE55FEEB32C5" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="EA912A18D346FF9EFECDFE32FE4532C5" box="[275,429,451,475]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="12" pageNumber="2681">Figure 3A, C</figureCitation>
|
||
). In
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE3AFE32FDBC32C5" authorityName="Kryzhanovskij" authorityYear="1951" box="[484,596,451,475]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFE3AFE32FDBC32C5" box="[484,596,451,475]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
longitudinal furrows are developed on dorsal surfaces of all tarsomeres, at least near base (derived character state, see
|
||
<figureCitation id="EA912A18D346FF9EFC28FE12FB6C32E5" box="[1014,1156,483,507]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="12" pageNumber="2681">Figure 4F, I</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE8BFDF2FDF93105" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[341,529,515,539]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFE8BFDF2FDF93105" box="[341,529,515,539]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
longitudinal furrows exist only on dorsolateral surfaces of the first mesotarsomeres and metatarsomeres (
|
||
<figureCitation id="EA912A18D346FF9EFCDEFDD2FC7B3125" box="[768,915,547,571]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="12" pageNumber="2681">Figure 4J, K</figureCitation>
|
||
). The distal portion of the tibiae is longitudinally wrinkled in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFD50FDB2FD163145" authorityName="Kryzhanovskij" authorityYear="1951" box="[654,766,579,603]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFD50FDB2FD163145" box="[654,766,579,603]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
(autapomorphy, see
|
||
<figureCitation id="EA912A18D346FF9EFC33FDB2FBBF3145" box="[1005,1111,579,603]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="12" pageNumber="2681">Figure 4I</figureCitation>
|
||
) and smooth in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE90FD92FDE23165" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[334,522,611,635]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFE90FD92FDE23165" box="[334,522,611,635]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFDCCFD92FD903164" box="[530,632,611,635]" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(plesiomorphic character state, see
|
||
<figureCitation id="EA912A18D346FF9EFBD0FD92FB6B3165" box="[1038,1155,611,635]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="12" pageNumber="2681">Figure 4K</figureCitation>
|
||
). The adhesive vestiture on the ventral side of male mesotarsomeres 1 + 2 is developed in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFEE5FD52FE4331A5" authorityName="Kryzhanovskij" authorityYear="1951" box="[315,427,675,699]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFEE5FD52FE4331A5" box="[315,427,675,699]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
(plesiomorphic character state, see
|
||
<figureCitation id="EA912A18D346FF9EFC82FD53FC1531A4" box="[860,1021,674,698]" captionStart="Figure 4" captionStartId="10.[211,278,1544,1566]" captionTargetBox="[238,1144,162,1511]" captionTargetId="figure-5@10.[238,1144,162,1511]" captionTargetPageId="10" captionText="Figure 4. Protarsi (A, D, G) and mesotarsi (B, E, H) in ventral view, and metatarsi in lateral view (C, F, I–K) of Broscini. (A–C) Zacotus matthewsii LeConte, 1869, male specimen from Vancouver, Canada; (D–F) Eobroscus lutshniki (Roubal, 1928), male specimen from Ussurisk, Russia; (G–I), Eobroscus bhutanensis Morvan, 1982 (G, H, male specimen from Qingcheng Houshan, China; I, female specimen from Mt Manaslu, Nepal); (J) Kashmirobroscus ortrudae sp. nov., paratype, male; (K) Kashmirobroscus mirabilis sp. nov., holotype. Abbreviation: lf, longitudinal furrow on dorsolateral surface of first metatarsomere." figureDoi="http://doi.org/10.5281/zenodo.4608245" httpUri="https://zenodo.org/record/4608245/files/figure.png" pageId="12" pageNumber="2681">Figure 4E, H</figureCitation>
|
||
) and absent in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFF2AFD32FE5831C5" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[244,432,707,731]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFF2AFD32FE5831C5" box="[244,432,707,731]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFE65FD32FDCC31C4" box="[443,548,707,731]" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(autapomorphy). The parascutellar seta is present in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFF0DFD13FEAB31E4" authorityName="Kryzhanovskij" authorityYear="1951" box="[211,323,738,762]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFF0DFD13FEAB31E4" box="[211,323,738,762]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
(plesiomorphic character state) and absent in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFCBBFD13FBC931E4" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[869,1057,738,762]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFCBBFD13FBC931E4" box="[869,1057,738,762]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFBF5FD12FB7C31E4" box="[1067,1172,739,763]" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(autapomorphy). In
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE1AFCF3FD683004" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[452,640,770,794]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFE1AFCF3FD683004" box="[452,640,770,794]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFD52FCF2FD1E3004" box="[652,758,771,795]" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
the number of lateral setae in the pronotal apical half is increased to three to five (autapomorphy) in contrast to a single pair of lateral setae in
|
||
<taxonomicName id="B5AA4D1ED346FF9EFE0EFCB3FDA83044" authorityName="Kryzhanovskij" authorityYear="1951" box="[464,576,834,858]" class="Insecta" family="Carabidae" genus="Eobroscus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="40DEEA8FD346FF9EFE0EFCB3FDA83044" box="[464,576,834,858]" italics="true" pageId="12" pageNumber="2681">Eobroscus</emphasis>
|
||
</taxonomicName>
|
||
(plesiomorphic character state). Additionally, differences of
|
||
<taxonomicName id="B5AA4D1ED346FF9EFEE9FC93FE1B3064" authority="Schmidt & Wrase & Sciaky, 2013" authorityName="Schmidt & Wrase & Sciaky" authorityYear="2013" box="[311,499,866,890]" class="Insecta" family="Carabidae" genus="Kashmirobroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2681" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="40DEEA8FD346FF9EFEE9FC93FE1B3064" box="[311,499,866,890]" italics="true" pageId="12" pageNumber="2681">Kashmirobroscus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BED57F4D346FF9EFE22FC92FD8A3064" box="[508,610,867,891]" pageId="12" pageNumber="2681" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(autapomorphies) are developed in the degree of adaption to the high-altitude environment: body slender (
|
||
<figureCitation id="EA912A18D346FF9EFC91FC73FC223084" box="[847,970,898,922]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="12" pageNumber="2681">Figures 1A</figureCitation>
|
||
,
|
||
<figureCitation id="EA912A18D346FF9EFC09FC73FC113084" box="[983,1017,898,922]" captionStart="Figure 2" captionStartId="7.[143,210,1380,1402]" captionTargetBox="[145,1104,171,1346]" captionTargetId="figure-5@7.[144,1104,168,1347]" captionTargetPageId="7" captionText="Figure 2. Kashmirobroscus and Eobroscus in dorsal view. (A) Kashmirobroscus mirabilis sp. nov., holotype; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal." figureDoi="http://doi.org/10.5281/zenodo.4608241" httpUri="https://zenodo.org/record/4608241/files/figure.png" pageId="12" pageNumber="2681">2A</figureCitation>
|
||
), eyes smaller (
|
||
<figureCitation id="EA912A18D346FF9EFF02FC50FED630A7" box="[220,318,929,953]" captionStart="Figure 3" captionStartId="8.[211,278,1396,1418]" captionTargetBox="[212,1172,168,1363]" captionTargetId="figure-5@8.[212,1172,167,1363]" captionTargetPageId="8" captionText="Figure 3. Head in dorsal (A, C, D) and ventral (B) views of Kashmirobroscus and Eobroscus. (A, B) Kashmirobroscus ortrudae sp. nov., paratype, male; (C) Kashmirobroscus mirabilis sp. nov., holotype; (D) Eobruscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal. Abbreviation: ms, setae of mentum; sms, setae of submentum." figureDoi="http://doi.org/10.5281/zenodo.4608243" httpUri="https://zenodo.org/record/4608243/files/figure.png" pageId="12" pageNumber="2681">Figure 3</figureCitation>
|
||
A–C), elytra flattened (
|
||
<figureCitation id="EA912A18D346FF9EFD8CFC50FD2130A7" box="[594,713,929,953]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="12" pageNumber="2681">Figure 1C</figureCitation>
|
||
), hind-wings reduced, humeri rounded (
|
||
<figureCitation id="EA912A18D346FF9EFF02FC30FEB130C7" box="[220,345,961,985]" captionStart="Figure 1" captionStartId="6.[211,278,1364,1386]" captionTargetBox="[216,1169,169,1331]" captionTargetId="figure-5@6.[212,1172,168,1331]" captionTargetPageId="6" captionText="Figure 1. Kashmirobroscus ortrudae sp. nov., paratype, male. (A) Dorsal view; (B) ventral view; (C) left lateral view. Abbreviation: bt, button-like elevation on sternite VI." figureDoi="http://doi.org/10.5281/zenodo.4608239" httpUri="https://zenodo.org/record/4608239/files/figure.png" pageId="12" pageNumber="2681">Figures 1A</figureCitation>
|
||
,
|
||
<figureCitation id="EA912A18D346FF9EFEB9FC30FE6130C7" box="[359,393,961,985]" captionStart="Figure 2" captionStartId="7.[143,210,1380,1402]" captionTargetBox="[145,1104,171,1346]" captionTargetId="figure-5@7.[144,1104,168,1347]" captionTargetPageId="7" captionText="Figure 2. Kashmirobroscus and Eobroscus in dorsal view. (A) Kashmirobroscus mirabilis sp. nov., holotype; (B) Eobroscus bhutanensis Morvan, 1982, female specimen from Mt Manaslu, Nepal." figureDoi="http://doi.org/10.5281/zenodo.4608241" httpUri="https://zenodo.org/record/4608241/files/figure.png" pageId="12" pageNumber="2681">2A</figureCitation>
|
||
) and metepisterna shortened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AB06516D346FF9EFF0DFBE7FBB63768" pageId="12" pageNumber="2681" type="distribution">
|
||
<paragraph id="7215369DD346FF9EFF0DFBE7FE023730" blockId="12.[211,1172,1046,1142]" box="[211,490,1046,1070]" pageId="12" pageNumber="2681">
|
||
<heading id="295D81F1D346FF9EFF0DFBE7FE023730" box="[211,490,1046,1070]" fontSize="10" level="3" pageId="12" pageNumber="2681" reason="5">
|
||
<emphasis id="40DEEA8FD346FF9EFF0DFBE7FE023730" box="[211,490,1046,1070]" italics="true" pageId="12" pageNumber="2681">Geographical distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7215369DD346FF9EFF0DFBCFFBB63768" blockId="12.[211,1172,1046,1142]" pageId="12" pageNumber="2681">
|
||
<figureCitation id="EA912A18D346FF9EFF0DFBCFFED03748" box="[211,312,1086,1110]" captionStart="Figure 7" captionStartId="14.[211,278,1572,1594]" captionTargetBox="[215,1172,870,1537]" captionTargetId="figure-151@14.[212,1172,866,1540]" captionTargetPageId="14" captionText="Figure 7. Distribution of Kashmirobroscus ortrudae sp. nov. (white square), Kashmirobroscus mirabilis sp. nov. (white triangle) and Eobroscus bhutanensis Morvan, 1982 (white circles) in High Asia." figureDoi="http://doi.org/10.5281/zenodo.4608251" httpUri="https://zenodo.org/record/4608251/files/figure.png" pageId="12" pageNumber="2681">Figure 7</figureCitation>
|
||
shows the geographical distribution. Only two species are known from different localities in the Northwest Himalaya of
|
||
<collectingCountry id="0ABD760DD346FF9EFD24FBAFFCB43768" box="[762,860,1118,1142]" name="Pakistan" pageId="12" pageNumber="2681">Pakistan</collectingCountry>
|
||
(for details see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |