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<document id="72DD0F67C2ECDE6E45F258595649F378" ID-DOI="10.5281/zenodo.198718" ID-GBIF-Dataset="51f3133b-0961-417f-a32b-9dbaf1512e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="198718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460311549789" checkinUser="plazi" docAuthor="Nishikawa, Masaaki, Ikeda, Hiroshi, Kubota, Kohei &amp; Sota, Teiji" docDate="2010" docId="FA01F1186E7C1A564CA0B1F3FA6AFCD3" docLanguage="en" docName="zt02648p031.pdf" docOrigin="Zootaxa 2648" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Silpha carinata" docType="key" docVersion="10" lastPageNumber="15" masterDocId="063889606E721A584C37B164FFF6FFB4" masterDocTitle="Taxonomic redefinition and natural history of the endemic silphid beetle Silpha longicornis (Coleoptera: Silphidae) of Japan, with an analysis of its geographic variation" masterLastPageNumber="31" masterPageNumber="1" pageNumber="15" updateTime="1698254323434" updateUser="plazi">
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<mods:title id="8710AD08DC0BDD16EB70BCA13A230714">Taxonomic redefinition and natural history of the endemic silphid beetle Silpha longicornis (Coleoptera: Silphidae) of Japan, with an analysis of its geographic variation</mods:title>
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<mods:namePart id="49B02CEB5B272EE5A63B7DE4578ACA30">Ikeda, Hiroshi</mods:namePart>
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<mods:namePart id="CC2D0F05D9B8A7E9333DAA83387ACAB5">Kubota, Kohei</mods:namePart>
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<mods:namePart id="F907B340144A14B6C508C2B2D5D6E32F">Sota, Teiji</mods:namePart>
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<subSubSection id="3AB213856E7C1A564CA0B1F3FD89FEAB" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="7217400E6E7C1A564CA0B1F3FD8FFF05" blockId="14.[151,633,151,177]" box="[151,633,151,177]" pageId="14" pageNumber="15">
<heading id="295FF7626E7C1A564CA0B1F3FD8FFF05" bold="true" box="[151,633,151,177]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="40DC9C1C6E7C1A564CA0B1F3FD8FFF05" bold="true" box="[151,633,151,177]" pageId="14" pageNumber="15">
Key to species of
<taxonomicName id="B5A83B8D6E7C1A564D5AB1F3FDD1FF05" box="[365,551,151,177]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="40DC9C1C6E7C1A564D5AB1F3FDD1FF05" bold="true" box="[365,551,151,177]" italics="true" pageId="14" pageNumber="15">Silpha carinata</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="7217400E6E7C1A564CA0B184FD89FEAB" blockId="14.[151,1436,224,287]" pageId="14" pageNumber="15">
Four
<taxonomicName id="B5A83B8D6E7C1A564CE1B184FED7FF4D" box="[214,289,224,249]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="40DC9C1C6E7C1A564CE1B184FED7FF4D" box="[214,289,224,249]" italics="true" pageId="14" pageNumber="15">Silpha</emphasis>
</taxonomicName>
species from Northeast Asia compose the
<taxonomicName id="B5A83B8D6E7C1A564F2AB184FC69FF4D" box="[797,927,224,250]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="40DC9C1C6E7C1A564F2AB184FCDAFF4D" box="[797,812,224,249]" italics="true" pageId="14" pageNumber="15">S</emphasis>
.
<emphasis id="40DC9C1C6E7C1A564F0CB184FC69FF4D" box="[827,927,224,249]" italics="true" pageId="14" pageNumber="15">carinata</emphasis>
</taxonomicName>
group (
<emphasis id="40DC9C1C6E7C1A564FCAB184FBC8FF4D" box="[1021,1086,224,249]" italics="true" pageId="14" pageNumber="15">sensu</emphasis>
<bibRefCitation id="16393DFF6E7C1A564870B184FAE6FF4E" author="Hava" box="[1095,1296,224,250]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A5648BAB184FB34FF4D" box="[1165,1218,224,249]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
). They may be discriminated using the following key:
</paragraph>
</subSubSection>
<subSubSection id="3AB213856E7C1A564CA0B034FA6AFCD3" pageId="14" pageNumber="15" type="key">
<keyStep id="C95C58AB6E7C1A564CA0B034FA6DFE33" pageId="14" pageNumber="15">
<paragraph id="7217400E6E7C1A564CA0B034FA6DFED3" blockId="14.[151,1437,336,871]" box="[151,1435,336,359]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B034FA6DFED3" box="[151,1435,336,359]" pageId="14" pageNumber="15">1 Small impunctate field behind eyes absent................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="7217400E6E7C1A564CA0B014FA6DFE33" blockId="14.[151,1437,336,871]" box="[151,1435,368,391]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B014FA6DFE33" box="[151,1435,368,391]" pageId="14" pageNumber="15">- Small impunctate field behind eyes present ................................................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="C95C58AB6E7C1A564CA0B0F4FA6AFD13" pageId="14" pageNumber="15">
<paragraph id="7217400E6E7C1A564CA0B0F4FA6AFDF3" blockId="14.[151,1437,336,871]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B0F4FA6AFDF3" pageId="14" pageNumber="15">
2 Antennal segment VIII not longer than IX. Median lobe of aedeagus with sides constricted medially (
<bibRefCitation id="16393DFF6E7C1A5648CAB0F4FA6EFE13" author="Portevin" box="[1277,1432,400,423]" pageId="14" pageNumber="15" refString="Portevin, G. (1926) Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini. Encyclopedie Entomologique, (A) 6, 1 - 270." type="journal article" year="1926">Portevin 1926</bibRefCitation>
: Fig. 55;
<bibRefCitation id="16393DFF6E7C1A564D17B0D4FE44FE73" author="Nakane" box="[288,434,432,455]" pageId="14" pageNumber="15" refString="Nakane, T. (1955) The Coleoptera of Japan [22]. Silphidae (continuation). Shin-Konchu, Tokyo, 8 (2), 47 - 51. [In Japanese]" type="journal article" year="1955">Nakane 1955</bibRefCitation>
:
<figureCitation id="EA935C8B6E7C1A564D8BB0D4FE0BFE73" box="[444,509,432,455]" captionStart="FIGURES 1 16" captionStartId="5.[151,269,1727,1751]" captionTargetBox="[159,1436,197,1676]" captionTargetId="figure@5.[151,1436,197,1684]" captionTargetPageId="5" captionText="FIGURES 1 16. Habitus of type specimens and labels (1 7): 1 — Silpha longicornis Portevin, lectotype, Ψ, dorsal view; 2 — same, labels; 3 — Silpha yamatona Kôno, holotype, ♂, dorsal view (photo by S. Shimano); 4 — same, labels (photo by S. Shimano); 5 — Silpha imitator Shibata, holotype, ♂, dorsal view; 6 — same, labels; 7 — same, aedeagus in ventral view. Details of Silpha longicornis Portevin, showing variation of elytron color (8 9), and sculpture (10 13): 8 — black specimen from Mt. Kurikoma, Iwate Pref.; 9 — reddish-brown specimen from Mt. Kasa-ga-take, Nagano Pref. (with aberration of elytral costae); 10 — rugose intervals; 11 — flat intervals; 12 — punctate microsculpture; 13 — smooth microsculpture. Apex of male metatibia, showing terminal projection (14). Prothorax of larvae of Silpha perforata Gebler (15, from Moheji near Hakodate, S. W. Hokkaido) and Silpha longicornis Portevin (16, from Jikkoku-tôge, Nagano Pref.)." httpUri="https://zenodo.org/record/198719/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
;
<bibRefCitation id="16393DFF6E7C1A564E3FB0D4FD4BFE73" author="Hava" box="[520,701,432,455]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A564E7FB0D4FD81FE73" box="[584,631,432,455]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
: Fig. 28); parameres straight. Distribution:
<collectingCountry id="0ABF009E6E7C1A5648BDB0D4FB27FE73" box="[1162,1233,432,455]" name="Russia" pageId="14" pageNumber="15">Russia</collectingCountry>
(from East Sayans through East Siberia to Primorsky Kray, Sakhalin, Moneron Is. and South Kurils (Iturup Is., Kunashir Is., and Shikotan Is.)),
<collectingCountry id="0ABF009E6E7C1A564D06B094FE6DFDB3" box="[305,411,496,519]" name="Mongolia" pageId="14" pageNumber="15">Mongolia</collectingCountry>
,
<collectingCountry id="0ABF009E6E7C1A564D9CB094FE1DFDB3" box="[427,491,496,519]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
(Beijing, Hebei, Heilonjiang, Jiangxi, Jilin, Nei Mongol, Shaanxi, and Shanxi), the Korean Peninsula, Ulleung-do Is. and Jeju-do Is., and
<collectingCountry id="0ABF009E6E7C1A564F34B374FCB6FD93" box="[771,832,528,551]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
(Hokkaido, Rishiri-tô Is., Rebun-tô Is., Teuri-tô Is., Yagishiri-tô Is. and Okushiri-tô Is.)
<emphasis id="40DC9C1C6E7C1A564E23B354FABBFDF3" box="[532,1357,560,583]" italics="true" pageId="14" pageNumber="15">
........................................................................................................
<taxonomicName id="B5A83B8D6E7C1A5648F9B354FABBFDF3" authority="Gebler" authorityName="Gebler" box="[1230,1357,560,583]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="perforata">S. perforata</taxonomicName>
</emphasis>
Gebler
</keyLead>
</paragraph>
<paragraph id="7217400E6E7C1A564CA0B334FA6AFD13" blockId="14.[151,1437,336,871]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B334FA6AFD13" pageId="14" pageNumber="15">
- Antennal segment VIII distinctly longer than IX. Median lobe of aedeagus parallel sided (
<bibRefCitation id="16393DFF6E7C1A5648A7B334FADBFDD3" author="Portevin" box="[1168,1325,592,615]" pageId="14" pageNumber="15" refString="Portevin, G. (1926) Les Grands Necrophages du Globe. Silphini - Necrodini - Necrophorini. Encyclopedie Entomologique, (A) 6, 1 - 270." type="journal article" year="1926">Portevin 1926</bibRefCitation>
: Figs. 58 59;
<bibRefCitation id="16393DFF6E7C1A564CC4B314FE8BFD33" author="Sustek" box="[243,381,624,647]" pageId="14" pageNumber="15" refString="Sustek, Z. (1983) Silpha bilineata Reitter, 1901 and Silpha tatrica Smetana, 1952 - new synonyms of Silpha carinata Herbst, 1783, and some ecological aspects of its intraspecific variability. Annotationes Zoologicae et Botanicae, Bratislava, 153, 1 - 33." type="journal article" year="1983">Šustek 1983</bibRefCitation>
: Figs. 5356;
<bibRefCitation id="16393DFF6E7C1A564E2AB314FD2AFD33" author="Hava" box="[541,732,624,647]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A564E57B314FD64FD33" box="[608,658,624,647]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
: Fig. 31); parameres very sinuate. Distribution: from Europe to Transbaikalia and Central Asia, including
<collectingCountry id="0ABF009E6E7C1A564EBDB3F4FD04FD13" box="[650,754,656,679]" name="Mongolia" pageId="14" pageNumber="15">Mongolia</collectingCountry>
and westernmost
<collectingCountry id="0ABF009E6E7C1A564F85B3F4FC04FD13" box="[946,1010,656,679]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
<emphasis id="40DC9C1C6E7C1A564FC3B3F4FAB8FD13" box="[1012,1358,656,679]" italics="true" pageId="14" pageNumber="15">
..................................
<taxonomicName id="B5A83B8D6E7C1A5648EEB3F4FAB8FD13" authority="Herbst" authorityName="Herbst" box="[1241,1358,656,679]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinata">S. carinata</taxonomicName>
</emphasis>
Herbst
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C95C58AB6E7C1A564CA0B3D4FA6AFCD3" pageId="14" pageNumber="15">
<paragraph id="7217400E6E7C1A564CA0B3D4FA6AFCB3" blockId="14.[151,1437,336,871]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B3D4FA6AFCB3" pageId="14" pageNumber="15">
3 Pronotum densely punctate in its central part. Median lobe with apex hook-shaped in lateral view (
<bibRefCitation id="16393DFF6E7C1A5648C2B3D4FA6EFD73" author="Kusakari" box="[1269,1432,688,711]" pageId="14" pageNumber="15" refString="Kusakari, K. (1993) A preliminary study on Silpha longicornis Portevin. In: Yamaya, B. &amp; Kusakari, K. (Eds.), Insects of Kita Honshu, II. Bunanoki Shuppan, Yamagata, pp. 9 - 17. [In Japanese]" type="book chapter" year="1993">Kusakari 1993</bibRefCitation>
:
<figureCitation id="EA935C8B6E7C1A564CF0B3B4FECDFD53" box="[199,315,720,743]" captionStart="FIGURES 1 16" captionStartId="5.[151,269,1727,1751]" captionTargetBox="[159,1436,197,1676]" captionTargetId="figure@5.[151,1436,197,1684]" captionTargetPageId="5" captionText="FIGURES 1 16. Habitus of type specimens and labels (1 7): 1 — Silpha longicornis Portevin, lectotype, Ψ, dorsal view; 2 — same, labels; 3 — Silpha yamatona Kôno, holotype, ♂, dorsal view (photo by S. Shimano); 4 — same, labels (photo by S. Shimano); 5 — Silpha imitator Shibata, holotype, ♂, dorsal view; 6 — same, labels; 7 — same, aedeagus in ventral view. Details of Silpha longicornis Portevin, showing variation of elytron color (8 9), and sculpture (10 13): 8 — black specimen from Mt. Kurikoma, Iwate Pref.; 9 — reddish-brown specimen from Mt. Kasa-ga-take, Nagano Pref. (with aberration of elytral costae); 10 — rugose intervals; 11 — flat intervals; 12 — punctate microsculpture; 13 — smooth microsculpture. Apex of male metatibia, showing terminal projection (14). Prothorax of larvae of Silpha perforata Gebler (15, from Moheji near Hakodate, S. W. Hokkaido) and Silpha longicornis Portevin (16, from Jikkoku-tôge, Nagano Pref.)." httpUri="https://zenodo.org/record/198719/files/figure.png" pageId="14" pageNumber="15">Figs. 912</figureCitation>
;
<bibRefCitation id="16393DFF6E7C1A564D70B3B4FE0BFD53" author="Hava" box="[327,509,720,743]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A564DB0B3B4FE40FD53" box="[391,438,720,743]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
: Figs. 2930). Apex of male metatibia projected at the inner angle (
<figureCitation id="EA935C8B6E7C1A5648E0B3B4FADEFD53" box="[1239,1320,720,743]" captionStart="FIGURES 1 16" captionStartId="5.[151,269,1727,1751]" captionTargetBox="[159,1436,197,1676]" captionTargetId="figure@5.[151,1436,197,1684]" captionTargetPageId="5" captionText="FIGURES 1 16. Habitus of type specimens and labels (1 7): 1 — Silpha longicornis Portevin, lectotype, Ψ, dorsal view; 2 — same, labels; 3 — Silpha yamatona Kôno, holotype, ♂, dorsal view (photo by S. Shimano); 4 — same, labels (photo by S. Shimano); 5 — Silpha imitator Shibata, holotype, ♂, dorsal view; 6 — same, labels; 7 — same, aedeagus in ventral view. Details of Silpha longicornis Portevin, showing variation of elytron color (8 9), and sculpture (10 13): 8 — black specimen from Mt. Kurikoma, Iwate Pref.; 9 — reddish-brown specimen from Mt. Kasa-ga-take, Nagano Pref. (with aberration of elytral costae); 10 — rugose intervals; 11 — flat intervals; 12 — punctate microsculpture; 13 — smooth microsculpture. Apex of male metatibia, showing terminal projection (14). Prothorax of larvae of Silpha perforata Gebler (15, from Moheji near Hakodate, S. W. Hokkaido) and Silpha longicornis Portevin (16, from Jikkoku-tôge, Nagano Pref.)." httpUri="https://zenodo.org/record/198719/files/figure.png" pageId="14" pageNumber="15">Fig. 14</figureCitation>
). Distribution:
<collectingCountry id="0ABF009E6E7C1A564CC9B394FECDFCB3" box="[254,315,752,775]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
(Honshu northeast of the Kii Mts., Sado Is.)
<emphasis id="40DC9C1C6E7C1A564F2FB394FACDFCB3" box="[792,1339,752,775]" italics="true" pageId="14" pageNumber="15">
...........................................................
<taxonomicName id="B5A83B8D6E7C1A564890B394FACDFCB3" authority="Portevin" authorityName="Portevin" box="[1191,1339,752,775]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longicornis">S. longicornis</taxonomicName>
</emphasis>
Portevin
</keyLead>
</paragraph>
<paragraph id="7217400E6E7C1A564CA0B274FA6AFCD3" blockId="14.[151,1437,336,871]" pageId="14" pageNumber="15">
<keyLead id="C959E33B6E7C1A564CA0B274FA6AFCD3" pageId="14" pageNumber="15">
- Pronotum sparsely punctate in its central part (
<emphasis id="40DC9C1C6E7C1A564E8CB274FD31FC93" box="[699,711,784,807]" italics="true" pageId="14" pageNumber="15">e</emphasis>
.
<emphasis id="40DC9C1C6E7C1A564EFAB274FD2CFC93" box="[717,730,784,807]" italics="true" pageId="14" pageNumber="15">g</emphasis>
.,
<bibRefCitation id="16393DFF6E7C1A564ED8B274FC55FC93" author="Hava" box="[751,931,784,807]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A564F19B274FCAAFC93" box="[814,860,784,807]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
:
<figureCitation id="EA935C8B6E7C1A564F9AB274FC0AFC93" box="[941,1020,784,807]" captionStart="FIGURE 20" captionStartId="9.[151,255,1266,1290]" captionTargetBox="[401,1190,646,1233]" captionTargetId="figure@9.[391,1196,629,1245]" captionTargetPageId="9" captionText="FIGURE 20. Relationship between AMT and PC 1. Females are indicated by open circles and the dashed line, and males are indicated by solid circles and the solid line." httpUri="https://zenodo.org/record/198723/files/figure.png" pageId="14" pageNumber="15">Fig. 20</figureCitation>
). Median lobe gently turned upward in apical part, the apex rounded in lateral view (
<bibRefCitation id="16393DFF6E7C1A564E9EB254FCA8FCF3" author="Hava" box="[681,862,816,839]" pageId="14" pageNumber="15" refString="Hava, J., Schneider, J. &amp; Ruzicka, J. (1999) Four new species of carrion beetles from China (Coleoptera: Silphidae). Entomological Problems, 30, 67 - 83." type="journal article" year="1999">
Háva
<emphasis id="40DC9C1C6E7C1A564EDEB254FCEEFCF3" box="[745,792,816,839]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 1999
</bibRefCitation>
: Figs. 2527, 3843). Apex of male metatibia simple. Distribution:
<collectingCountry id="0ABF009E6E7C1A564D61B234FE60FCD3" box="[342,406,848,871]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
(Shaanxi)
<emphasis id="40DC9C1C6E7C1A564E30B234FB9BFCD3" box="[519,1133,848,871]" italics="true" pageId="14" pageNumber="15">
.................................................................
<taxonomicName id="B5A83B8D6E7C1A564F88B234FB9BFCD3" authority="Hava, Schneider et Ruzicka" authorityName="Hava, Schneider et Ruzicka" box="[959,1133,848,871]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="businskyorum">S. businskyorum</taxonomicName>
</emphasis>
Háva, Schneider et Růžiċka
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
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