treatments-xml/data/F9/7B/5A/F97B5A001A4238420EAF3D6FFD5040C8.xml
2024-06-21 12:58:37 +02:00

298 lines
41 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="8D3A037E32812D903736CD33DC6AFEAD" ID-DOI="10.11646/zootaxa.4019.1.12" ID-GBIF-Dataset="6ef51ae2-1fd9-478d-bcdd-9ced4992551b" ID-ISSN="1175-5326" ID-Zenodo-Dep="253393" ID-ZooBank="710CEEF1-F9D9-469F-9184-DC9903650F4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461260809308" checkinUser="plazi" docAuthor="Hutchings, Pat, Nogueira, João Miguel Matos &amp; Carrerette, Orlemir" docDate="2015" docId="F97B5A001A4238420EAF3D6FFD5040C8" docLanguage="en" docName="zt04019p274.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinothelepus oculeus Hutchings, Nogueira &amp; Carrerette, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="244" masterDocId="054222781A4138460E383919FF984410" masterDocTitle="Telothelepodidae, Thelepodidae and Trichobranchidae (Annelida, Terebelliformia) from Lizard Island, Great Barrier Reef, Australia" masterLastPageNumber="274" masterPageNumber="240" pageNumber="243" updateTime="1698609381483" updateUser="plazi">
<mods:mods id="98F033290981FF330A7A4CDE002CF9A9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87EB156C0BCE6476D6B9CAFD0DE0053B">
<mods:title id="03D52868489B4EFF0A5AECD92A349E9B">Telothelepodidae, Thelepodidae and Trichobranchidae (Annelida, Terebelliformia) from Lizard Island, Great Barrier Reef, Australia</mods:title>
</mods:titleInfo>
<mods:name id="CCFF0098262403A05A75C195D30F3FD4" type="personal">
<mods:role id="F94EC088A4BBA089EA9F7E8DDB8E5B86">
<mods:roleTerm id="9E5CB567233F01E3187BF029EA693D5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15E3E947A1438581A3005C39C09E6921">Hutchings, Pat</mods:namePart>
</mods:name>
<mods:name id="06E4F6350861D29B544E7104AF73BA47" type="personal">
<mods:role id="8137E78EFB6EB6C56C9142A6AE3CF689">
<mods:roleTerm id="C46A813007845D97CB4646697161BDE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE1AD662D82FA4B03BDDB230BDA2C315">Nogueira, João Miguel Matos</mods:namePart>
</mods:name>
<mods:name id="E729B4A33D1F120F59A0A41DF38FFF1F" type="personal">
<mods:role id="EA9208B76527FEF03635FD0B78AC23F7">
<mods:roleTerm id="6BA10E99AF85DD9A4D4CE69034C8E927">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="54DCCF31D9A41CDE57339393D368EEA2">Carrerette, Orlemir</mods:namePart>
</mods:name>
<mods:typeOfResource id="53F128F01CD510293B67E8AADA61D9CA">text</mods:typeOfResource>
<mods:relatedItem id="8A5962BE04C06F62BB6C5FD5EFFC00EA" type="host">
<mods:titleInfo id="05A112C6647C760A6D68A941DC2F71C2">
<mods:title id="4575F51DD58E654B455F412FE9E55FF4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F0E4284E994D30E83B0971294E471065">
<mods:date id="DF8028428A596B3E9194F918258326E1">2015</mods:date>
<mods:detail id="C648CA0E092D18D3E5DEEAF36597D398" type="volume">
<mods:number id="2B7D6DA473FCF0166EEA1A0DE0FCBE43">4019</mods:number>
</mods:detail>
<mods:detail id="BFDF9474C756AE4ECB9FDD481456B504" type="issue">
<mods:number id="DEE20E697A4E96DC501EC9A203741553">1</mods:number>
</mods:detail>
<mods:extent id="C2DCF8DF8624E65B042C78EFDB449D8C" unit="page">
<mods:start id="ED4BDD7CCE9AFF2F6B8A66B46796D5EB">240</mods:start>
<mods:end id="5E9D5900CD835993C03D6B6EACBC880D">274</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7C28D4807936CD417E0E64C21E73CC45">journal article</mods:classification>
<mods:identifier id="EE3605A1DAE89F39FDCF2758FE9E0C59" type="DOI">10.11646/zootaxa.4019.1.12</mods:identifier>
<mods:identifier id="53571CB1AF26DA4A6634ED6867B7FB7B" type="GBIF-Dataset">6ef51ae2-1fd9-478d-bcdd-9ced4992551b</mods:identifier>
<mods:identifier id="6160C1D8B7E3DD58673E1337680EEB83" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EFB7449E1CD9B68B0C8A19035C2E17D4" type="Zenodo-Dep">253393</mods:identifier>
<mods:identifier id="71EAAB92543ED6E27213902568047662" type="ZooBank">710CEEF1-F9D9-469F-9184-DC9903650F4C</mods:identifier>
</mods:mods>
<treatment id="F97B5A001A4238420EAF3D6FFD5040C8" ID-DOI="http://doi.org/10.5281/zenodo.6096722" ID-GBIF-Taxon="119636985" ID-Zenodo-Dep="6096722" LSID="urn:lsid:plazi:treatment:F97B5A001A4238420EAF3D6FFD5040C8" httpUri="http://treatment.plazi.org/id/F97B5A001A4238420EAF3D6FFD5040C8" lastPageId="4" lastPageNumber="244" pageId="3" pageNumber="243">
<subSubSection id="39C8B89D1A4238450EAF3D6FFE9240A2" pageId="3" pageNumber="243" type="nomenclature">
<paragraph id="716DEB161A4238450EAF3D6FFE6C4080" blockId="3.[151,500,1142,1202]" box="[151,500,1142,1168]" pageId="3" pageNumber="243">
<heading id="2A255C7A1A4238450EAF3D6FFE6C4080" bold="true" box="[151,500,1142,1168]" fontSize="11" level="1" pageId="3" pageNumber="243" reason="1">
<emphasis id="43A637041A4238450EAF3D6FFE6C4080" bold="true" box="[151,500,1142,1168]" pageId="3" pageNumber="243">
<taxonomicName id="B6D290951A4238450EAF3D6FFE324080" ID-CoL="4S9ZM" box="[151,426,1142,1168]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="243" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4238450EAF3D6FFE324080" bold="true" box="[151,426,1142,1168]" italics="true" pageId="3" pageNumber="243">Rhinothelepus oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4238450F8A3D6FFE6C4080" box="[434,500,1142,1168]" pageId="3" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="716DEB161A4238450EAF3D80FE9240A2" blockId="3.[151,500,1142,1202]" box="[151,266,1177,1202]" pageId="3" pageNumber="243">
(
<figureCitation id="E9E9F7931A4238450EA73D80FF7F40A2" box="[159,231,1177,1202]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Figs 2</figureCitation>
,
<figureCitation id="E9E9F7931A4238450ECB3D83FE9940A2" box="[243,257,1178,1202]" captionStart="FIGURE 3" captionStartId="6.[151,250,1341,1363]" captionTargetBox="[160,1430,198,1315]" captionTargetId="figure@6.[152,1435,193,1320]" captionTargetPageId="6" captionText="FIGURE 3. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A B. Notochaetae, segments 7 and 14, respectively; C. Tips of notochaetae from posterior row, segment 14; D G. Uncini, segment 8 and three last neuropodia, respectively. Scale bars: A C = 15 µm, D G = 5 µm." httpUri="https://zenodo.org/record/253396/files/figure.png" pageId="3" pageNumber="243">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4238450EAF3DC3FE3041DB" pageId="3" pageNumber="243" type="materials_examined">
<paragraph id="716DEB161A4238450EAF3DC3FE59414F" blockId="3.[151,1437,1242,2016]" pageId="3" pageNumber="243">
<emphasis id="43A637041A4238450EAF3DC3FEDE40E3" bold="true" box="[151,326,1242,1267]" pageId="3" pageNumber="243">
<typeStatus id="AE6955B41A4238450EAF3DC3FF4A40E3" box="[151,210,1242,1267]" pageId="3" pageNumber="243">Type</typeStatus>
material.
</emphasis>
<typeStatus id="AE6955B41A4238450F763DC3FE2340E3" box="[334,443,1242,1267]" pageId="3" pageNumber="243" type="holotype">Holotype</typeStatus>
: AM W.47507, MI QLD 2337, complete specimen, in excellent state of preservation,
<quantity id="B62A46F31A4238450BB63DC3FF5B4106" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="243" unit="mm" value="6.0">6 mm</quantity>
long,
<quantity id="B62A46F31A4238450F353DE6FEFD4106" box="[269,357,1279,1303]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="243" unit="mm" value="0.8">0.8 mm</quantity>
maximum width (segments 1112).
<typeStatus id="AE6955B41A4238450D313DE6FCE84107" box="[777,880,1279,1303]" pageId="3" pageNumber="243" type="paratype">Paratype</typeStatus>
: AM W.44945, MI QLD 2337, 4 mm long, ~
<quantity id="B62A46F31A4238450B403DE7FF5B412A" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="243" unit="mm" value="0.6">0.6 mm</quantity>
wide, 34 segments, incomplete, in relatively good state (broken in two pieces between segments 1314 due to manipulation / dissection).
</paragraph>
<paragraph id="716DEB161A4238450EFF3C73FE3041DB" blockId="3.[151,1437,1242,2016]" pageId="3" pageNumber="243">
<emphasis id="43A637041A4238450EFF3C73FDF24193" bold="true" box="[199,618,1386,1411]" pageId="3" pageNumber="243">Comparative material examined.</emphasis>
<typeStatus id="AE6955B41A4238450C463C73FD7E4193" box="[638,742,1386,1411]" pageId="3" pageNumber="243" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="B6D290951A4238450D1C3C73FB964193" box="[804,1038,1386,1411]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="243" phylum="Annelida" rank="species" species="buku">
<emphasis id="43A637041A4238450D1C3C73FB964193" box="[804,1038,1386,1411]" italics="true" pageId="3" pageNumber="243">Rhinothelepus buku</emphasis>
</taxonomicName>
,
<collectionCode id="17C373D31A4238450A123C72FBF04192" box="[1066,1128,1387,1410]" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="3" pageNumber="243">NTM</collectionCode>
W.009549.
<typeStatus id="AE6955B41A4238450B313C73FAE94193" box="[1289,1393,1386,1411]" pageId="3" pageNumber="243" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="B6D290951A4238450EAF3C97FE0B41B7" box="[151,403,1422,1447]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="243" phylum="Annelida" rank="species" species="lobatus">
<emphasis id="43A637041A4238450EAF3C97FE0B41B7" box="[151,403,1422,1447]" italics="true" pageId="3" pageNumber="243">Rhinothelepus lobatus</emphasis>
</taxonomicName>
, AM W.5234.
<typeStatus id="AE6955B41A4238450C073C97FD3F41B7" box="[575,679,1422,1447]" pageId="3" pageNumber="243" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="B6D290951A4238450CE83C97FC2641B6" box="[720,958,1422,1447]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="243" phylum="Annelida" rank="species" species="macer">
<emphasis id="43A637041A4238450CE83C97FC2641B6" box="[720,958,1422,1447]" italics="true" pageId="3" pageNumber="243">Rhinothelepus macer</emphasis>
</taxonomicName>
, AM W.6783.
<typeStatus id="AE6955B41A4238450A523C97FB4A41B7" box="[1130,1234,1422,1447]" pageId="3" pageNumber="243" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="B6D290951A4238450AC43C97FF6B41DB" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="243" phylum="Annelida" rank="species" species="occabus">
<emphasis id="43A637041A4238450AC43C97FF6B41DB" italics="true" pageId="3" pageNumber="243">Rhinothelepus occabus</emphasis>
</taxonomicName>
AM W.201903.
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4238420EFF3CCFFB5B4571" lastPageId="4" lastPageNumber="244" pageId="3" pageNumber="243" type="description">
<paragraph id="716DEB161A4238420EFF3CCFFB5B4571" blockId="3.[151,1437,1242,2016]" lastBlockId="4.[151,1437,151,1241]" lastPageId="4" lastPageNumber="244" pageId="3" pageNumber="243">
<emphasis id="43A637041A4238450EFF3CCFFEC141FF" bold="true" box="[199,345,1494,1519]" pageId="3" pageNumber="243">Description.</emphasis>
Short bodied species. Transverse prostomium attached to dorsal surface of upper lip; basal part with small eyespots, densely packed in two horizontal rows at each side, terminating dorso-laterally by oblique row of three eyespots; distal part of prostomium low, restricted to base of upper lip, mid-dorsal process elongate, attached to the upper lip basally, distally free (
<figureCitation id="E9E9F7931A4238450CAF3F58FD414249" box="[663,729,1601,1626]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Fig. 2</figureCitation>
AB, F, JK). Buccal tentacles all uniformly cylindrical, of two widths (
<figureCitation id="E9E9F7931A4238450ECF3F7DFEA7426D" box="[247,319,1636,1661]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Fig. 2</figureCitation>
AH, JM), thicker tentacles originating exclusively from both laterals of mid-dorsal process, basally. Peristomium forming lips, continuing dorsally as narrow annulation, with nuchal organs on anterior margin; upper lip large, distinctly longer than wide, slightly convoluted; lower lip expanded, cushion-like, extending across entire ventrum (
<figureCitation id="E9E9F7931A4238450C2D3FD7FDC142F7" box="[533,601,1742,1767]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Fig. 2</figureCitation>
BE, GH, KM). Anterior body highly glandular ventrally, swollen and slightly corrugated; segment 1 conspicuous dorsally and laterally, ventrally covered by expanded lower lip; segment 2 forming complete ring, segments 4 and 5 longer than remaining anterior segments, especially segment 4 (
<figureCitation id="E9E9F7931A4238450EA73E20FF7A4342" box="[159,226,1849,1874]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Fig. 2</figureCitation>
AH, JM). Two pairs of branchiae on segments 23, filamentous, each pair with less than 10 filaments on either side, originating from swollen cushion-like areas of body wall and with narrow mid-dorsal gap inbetween; branchial filaments of segment 2 extending to same level or slightly laterally in relation to notopodia of segment 3, filaments of second pair anterior and dorsal to notopodia (
<figureCitation id="E9E9F7931A4238450D183EBAFCFA43AC" box="[800,866,1955,1980]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="3" pageNumber="243">Fig. 2</figureCitation>
AH, JM). Notopodia extending for 15 segments, until segment 17; notopodia of segments 35 inserted progressively more laterally, then longitudinally aligned; anterior notopodia with slender bayonet-like chaetae in anterior row and narrowly-winged chaetae in posterior row (
<figureCitation id="E9E9F7931A4538420EA739A2FF7944C4" box="[159,225,187,212]" captionStart="FIGURE 3" captionStartId="6.[151,250,1341,1363]" captionTargetBox="[160,1430,198,1315]" captionTargetId="figure@6.[152,1435,193,1320]" captionTargetPageId="6" captionText="FIGURE 3. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A B. Notochaetae, segments 7 and 14, respectively; C. Tips of notochaetae from posterior row, segment 14; D G. Uncini, segment 8 and three last neuropodia, respectively. Scale bars: A C = 15 µm, D G = 5 µm." httpUri="https://zenodo.org/record/253396/files/figure.png" pageId="4" pageNumber="244">Fig. 3</figureCitation>
A); posterior notopodia bearing narrowly-winged chaetae in both rows, wings present from middle third of chaetae in posterior row; chaetae from anterior row about half length of chaetae from posterior row throughout (
<figureCitation id="E9E9F7931A4538420EA7381BFF79450B" box="[159,225,258,283]" captionStart="FIGURE 3" captionStartId="6.[151,250,1341,1363]" captionTargetBox="[160,1430,198,1315]" captionTargetId="figure@6.[152,1435,193,1320]" captionTargetPageId="6" captionText="FIGURE 3. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A B. Notochaetae, segments 7 and 14, respectively; C. Tips of notochaetae from posterior row, segment 14; D G. Uncini, segment 8 and three last neuropodia, respectively. Scale bars: A C = 15 µm, D G = 5 µm." httpUri="https://zenodo.org/record/253396/files/figure.png" pageId="4" pageNumber="244">Fig. 3</figureCitation>
BC). Neuropodia beginning from segment 8, uncini throughout with dorsal button at middle third of base and crest with 2 rows of secondary teeth (
<figureCitation id="E9E9F7931A4538420CB3383CFD49452E" box="[651,721,293,318]" captionStart="FIGURE 3" captionStartId="6.[151,250,1341,1363]" captionTargetBox="[160,1430,198,1315]" captionTargetId="figure@6.[152,1435,193,1320]" captionTargetPageId="6" captionText="FIGURE 3. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A B. Notochaetae, segments 7 and 14, respectively; C. Tips of notochaetae from posterior row, segment 14; D G. Uncini, segment 8 and three last neuropodia, respectively. Scale bars: A C = 15 µm, D G = 5 µm." httpUri="https://zenodo.org/record/253396/files/figure.png" pageId="4" pageNumber="244">Fig. 3</figureCitation>
DG). Elongate nephridial and genital papillae, posterior and longitudinally aligned to bases of notopodia of segments 57. Pygidium smooth (
<figureCitation id="E9E9F7931A4538420A1D3850FBF04571" box="[1061,1128,329,354]" captionStart="FIGURE 2" captionStartId="5.[151,250,1839,1861]" captionTargetBox="[197,1394,184,1812]" captionTargetId="figure@5.[190,1398,180,1819]" captionTargetPageId="5" captionText="FIGURE 2. Rhinothelepus oculeus n. sp., holotype AM W. 47507. A E. Entire worm, left dorso-lateral, left lateral and three progressively more ventral views, from right ventro-lateral to ventral, respectively; F H. Close ups of the anterior end, left dorso-lateral and two progressively closer right ventro-lateral views, respectively; I. Posterior body, right lateral view; J M. Close ups of anterior end, left dorso-lateral, left lateral, right ventro-lateral and ventral views, respectively. Abbreviations: numbers refer to segments, unspecified arrows point to genital papillae, ll = lower lip, P = basal part of prostomium, PP = prostomial process, ul = upper lip. Scale bars: A E = 0.4 mm, F G, I = 0.2 mm, H, J M = 0.15 mm." httpUri="https://zenodo.org/record/253395/files/figure.png" pageId="4" pageNumber="244">Fig. 2</figureCitation>
AE, I).
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4538420EFF3875FAFF4014" pageId="4" pageNumber="244" type="discussion">
<paragraph id="716DEB161A4538420EFF3875FB0645B9" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
<emphasis id="43A637041A4538420EFF3875FEA34595" bold="true" box="[199,315,364,389]" pageId="4" pageNumber="244">Remarks.</emphasis>
Five species of
<taxonomicName id="B6D290951A4538420FD63875FD164595" box="[494,654,364,389]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="genus">
<emphasis id="43A637041A4538420FD63875FD164595" box="[494,654,364,389]" italics="true" pageId="4" pageNumber="244">Rhinothelepus</emphasis>
</taxonomicName>
are known:
<taxonomicName id="B6D290951A4538420D203877FBA44595" authority="Hutchings, 1997" authorityName="Hutchings" authorityYear="1997" box="[792,1084,364,389]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="buku">
<emphasis id="43A637041A4538420D203877FCF64595" box="[792,878,364,389]" italics="true" pageId="4" pageNumber="244">R. buku</emphasis>
<bibRefCitation id="154396E71A4538420D4C3875FBA44595" author="Hutchings" box="[884,1084,364,389]" pageId="4" pageNumber="244" refString="Hutchings, P. A. (1997 a) New species of the family Terebellidae (Polychaeta) from Darwin Harbour Australia. In: Hanley, R." type="book" year="1997" yearSuffix="a">Hutchings, 1997a</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B6D290951A4538420A7E3877FAE14595" authority="Hutchings, 1974" authorityName="Hutchings" authorityYear="1974" box="[1094,1401,364,389]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="lobatus">
<emphasis id="43A637041A4538420A7E3877FB204595" box="[1094,1208,364,389]" italics="true" pageId="4" pageNumber="244">R. lobatus</emphasis>
<bibRefCitation id="154396E71A4538420A863875FAE14595" author="Hutchings" box="[1214,1401,364,389]" pageId="4" pageNumber="244" refString="Hutchings, P. A. (1974) Polychaeta of Wallis Lake, N. S. W. Proceedings of the Linnean Society of New South Wales, 98, 175 - 196." type="journal article" year="1974">Hutchings, 1974</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B6D290951A4538420BBC3877FE3845B8" authority="Hutchings, 1977" authorityName="Hutchings" authorityYear="1977" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="macer">
<emphasis id="43A637041A4538420BBC3877FF4545B8" italics="true" pageId="4" pageNumber="244">R. macer</emphasis>
<bibRefCitation id="154396E71A4538420EDC3889FE3845B8" author="Hutchings" box="[228,416,400,425]" pageId="4" pageNumber="244" refString="Hutchings, P. A. (1977) The Terebelliform polychaeta from Australia, chiefly from Moreton Bay, Queensland. Records of the Australian Museum, 31, 1 - 39. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.31.1977.232" type="journal article" year="1977">Hutchings, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B6D290951A4538420F933888FD7045B8" authority="Hutchings, 1990" authorityName="Hutchings" authorityYear="1990" box="[427,744,400,425]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="occabus">
<emphasis id="43A637041A4538420F933888FDBE45B9" box="[427,550,400,425]" italics="true" pageId="4" pageNumber="244">R. occabus</emphasis>
<bibRefCitation id="154396E71A4538420C143889FD7045B8" author="Hutchings" box="[556,744,400,425]" pageId="4" pageNumber="244" refString="Hutchings, P. A. (1990) Terebellidae (Polychaeta) from the Hong Kong region. In: Moreton, B. (Ed.), Proceedings of the Second International Workshop on the Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 1986. University of Hong Kong, Hong Kong, pp. 377 - 412." type="journal article" year="1990">Hutchings, 1990</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="B6D290951A4538420D1B3888FC0F45B9" box="[803,919,400,425]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420D1B3888FC0F45B9" box="[803,919,400,425]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420DA73888FC4345B8" box="[927,987,401,424]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
described above.
</paragraph>
<paragraph id="716DEB161A4538420EFF38AAFC7E466E" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
Both
<taxonomicName id="B6D290951A4538420F3E38ACFEE145DC" box="[262,377,435,460]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="lobatus">
<emphasis id="43A637041A4538420F3E38ACFEE145DC" box="[262,377,435,460]" italics="true" pageId="4" pageNumber="244">R. lobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="B6D290951A4538420F8B38ACFDA845DC" box="[435,560,435,460]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="occabus">
<emphasis id="43A637041A4538420F8B38ACFDA845DC" box="[435,560,435,460]" italics="true" pageId="4" pageNumber="244">R. occabus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="B6D290951A4538420C8538ACFCAC45DC" box="[701,820,435,460]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420C8538ACFCAC45DC" box="[701,820,435,460]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420D0438ADFCE245DC" box="[828,890,436,460]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
in having a longer prostomial process; a longer and highly convoluted upper lip and shorter deeply corrugated lower lip; in both species segment 1 is forming complete annulation around the body and segment 2 is not continuing ventrally, and both species have many more branchial filaments, ~2030 filaments on each side of pairs (
<bibRefCitation id="154396E71A4538420D753B04FBD84626" author="Nogueira" box="[845,1088,541,566]" pageId="4" pageNumber="244" refString="Nogueira, J. M. M., Hutchings, P. A. &amp; Fukuda, M. V. (2010) Morphology of terebelliform polychaetes (Annelida: Polychaeta: Terebelliformia), with a focus on Terebellidae. Zootaxa, 2460, 1 - 185." type="journal article" year="2010">
Nogueira
<emphasis id="43A637041A4538420DF83B06FC624626" box="[960,1018,541,566]" italics="true" pageId="4" pageNumber="244">et al.</emphasis>
2010
</bibRefCitation>
). In contrast, in
<taxonomicName id="B6D290951A4538420AC43B04FF74464A" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420AC43B04FF74464A" italics="true" pageId="4" pageNumber="244">Rhinothelepus oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420ECE3B5BFEAC4649" box="[246,308,578,601]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
the lower lip is cushion-like and almost smooth, segment 1 is incomplete and segment 2 forms a complete annulation, and there are less than 10 branchial filaments per pair.
</paragraph>
<paragraph id="716DEB161A4538420EFF3B91FE534765" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
<taxonomicName id="B6D290951A4538420EFF3B91FE2D46B0" box="[199,437,648,673]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="macer">
<emphasis id="43A637041A4538420EFF3B91FE2D46B0" box="[199,437,648,673]" italics="true" pageId="4" pageNumber="244">Rhinothelepus macer</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="B6D290951A4538420CB33B90FC9846B1" box="[651,768,648,673]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420CB33B90FC9846B1" box="[651,768,648,673]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420D303B90FCDD46B0" box="[776,837,649,672]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
than to other described species. Both species share a similar morphology of the anterior end, including lips and segments 1 and 2, and number of branchial filaments, however a prostomial process is absent among members of
<taxonomicName id="B6D290951A4538420D783BC9FC3E46F7" box="[832,934,720,743]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="macer">
<emphasis id="43A637041A4538420D783BC9FC3E46F7" box="[832,934,720,743]" italics="true" pageId="4" pageNumber="244">R. macer</emphasis>
</taxonomicName>
and present in
<taxonomicName id="B6D290951A4538420A6F3BC9FB5546F8" box="[1111,1229,719,744]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420A6F3BC9FB5546F8" box="[1111,1229,719,744]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420AEE3BC9FA8D46F7" box="[1238,1301,720,743]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
, and in that species the segments are progressively longer until segments 78, while in
<taxonomicName id="B6D290951A4538420A3A3BEDFBE1471C" box="[1026,1145,755,780]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420A3A3BEDFBE1471C" box="[1026,1145,755,780]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420ABC3BEDFB5C471B" box="[1156,1220,756,779]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
, segments 45 are longer than following ones. In addition, the bayonet-like chaetae of
<taxonomicName id="B6D290951A4538420DA93A0EFC6E473E" box="[913,1014,791,814]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="macer">
<emphasis id="43A637041A4538420DA93A0EFC6E473E" box="[913,1014,791,814]" italics="true" pageId="4" pageNumber="244">R. macer</emphasis>
</taxonomicName>
are much longer, about the same size as chaetae from posterior row, while in
<taxonomicName id="B6D290951A4538420C513A22FD794742" box="[617,737,825,850]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420C513A22FD794742" box="[617,737,825,850]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420CD33A23FCB44741" box="[747,812,826,849]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
, the bayonet-like chaetae are about half the length of chaetae from posterior row.
</paragraph>
<paragraph id="716DEB161A4538420EFF3A99FAFF4014" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
<taxonomicName id="B6D290951A4538420EFF3A99FE3C4789" box="[199,420,896,921]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="buku">
<emphasis id="43A637041A4538420EFF3A99FE3C4789" box="[199,420,896,921]" italics="true" pageId="4" pageNumber="244">Rhinothelepus buku</emphasis>
</taxonomicName>
has a longer prostomial process and a longer convoluted upper lip than
<taxonomicName id="B6D290951A4538420AFF3A9BFAA34789" box="[1223,1339,896,921]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus" status="sp. nov.">
<emphasis id="43A637041A4538420AFF3A9BFAA34789" box="[1223,1339,896,921]" italics="true" pageId="4" pageNumber="244">R. oculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58958A7F1A4538420B7A3A98FAE64788" box="[1346,1406,897,920]" pageId="4" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
In addition, the lower lip of
<taxonomicName id="B6D290951A4538420F833ABCFD8A47AD" box="[443,530,932,957]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="buku">
<emphasis id="43A637041A4538420F833ABCFD8A47AD" box="[443,530,932,957]" italics="true" pageId="4" pageNumber="244">R. buku</emphasis>
</taxonomicName>
forms a convoluted tongue-like projection almost as long as upper lip margins, folded and corrugated, and nephridial and genital papillae are inconspicuous or absent (
<bibRefCitation id="154396E71A4538420AB53ADEFACC47F0" author="Hutchings" box="[1165,1364,967,992]" pageId="4" pageNumber="244" refString="Hutchings, P. A. (1997 a) New species of the family Terebellidae (Polychaeta) from Darwin Harbour Australia. In: Hanley, R." type="book" year="1997" yearSuffix="a">Hutchings 1997a</bibRefCitation>
), with segment 1 conspicuous only ventrally and segment 2 forming a complete ring, with the first pair of branchiae.
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4538420EFF3D17FC7E407D" pageId="4" pageNumber="244" type="etymology">
<paragraph id="716DEB161A4538420EFF3D17FE3A405A" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
<emphasis id="43A637041A4538420EFF3D17FED64037" bold="true" box="[199,334,1038,1063]" pageId="4" pageNumber="244">Etymology.</emphasis>
The specific name “
<taxonomicName id="B6D290951A4538420C743D17FD394037" box="[588,673,1038,1063]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus">
<emphasis id="43A637041A4538420C743D17FD394037" box="[588,673,1038,1063]" italics="true" pageId="4" pageNumber="244">oculeus</emphasis>
</taxonomicName>
” is Latin and refers to the presence of the prostomial eyespots (
<taxonomicName id="B6D290951A4538420EA63D2BFF6D405B" box="[158,245,1074,1099]" class="Polychaeta" family="Terebellidae" genus="Rhinothelepus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="244" phylum="Annelida" rank="species" species="oculeus">oculeus</taxonomicName>
= full of eyes).
</paragraph>
<paragraph id="716DEB161A4538420EFF3D4CFC7E407D" blockId="4.[151,1437,151,1241]" box="[199,998,1109,1134]" pageId="4" pageNumber="244">
<emphasis id="43A637041A4538420EFF3D4CFEB2407E" bold="true" box="[199,298,1109,1134]" pageId="4" pageNumber="244">Habitat.</emphasis>
In amongst coral rubble of patch reefs in shallow waters,
<specimenCount id="67D4209F1A4538420D8C3D4FFC7E407D" box="[948,998,1110,1134]" pageId="4" pageNumber="244" type="generic">
<quantity id="B62A46F31A4538420D8C3D4FFC7A407D" box="[948,994,1110,1134]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="244" unit="m" value="9.0">9 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4538420EFF3D60FE9D40A5" pageId="4" pageNumber="244" type="materials_examined">
<paragraph id="716DEB161A4538420EFF3D60FE9D40A5" blockId="4.[151,1437,151,1241]" pageId="4" pageNumber="244">
<emphasis id="43A637041A4538420EFF3D60FEF34082" bold="true" box="[199,363,1145,1170]" pageId="4" pageNumber="244">
<typeStatus id="AE6955B41A4538420EFF3D60FE9A4082" box="[199,258,1145,1170]" pageId="4" pageNumber="244">Type</typeStatus>
locality.
</emphasis>
Big Vicki's reef,
<geoCoordinate id="14E68DD11A4538420C703D60FD234081" box="[584,699,1145,1169]" direction="south" orientation="latitude" pageId="4" pageNumber="244" precision="15" value="-14.685833">14°41'9&quot;S</geoCoordinate>
,
<geoCoordinate id="14E68DD11A4538420CF53D60FCC64082" box="[717,862,1145,1170]" direction="east" orientation="longitude" pageId="4" pageNumber="244" precision="15" value="145.44194">145°26'31&quot;E</geoCoordinate>
, Lizard Island, Great Barrier Reef, Queensland,
<collectingCountry id="09C5AB861A4538420EAF3D85FE9940A5" box="[151,257,1180,1205]" name="Australia" pageId="4" pageNumber="244">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="39C8B89D1A4538420EFF3DA6FD5040C8" box="[199,712,1215,1241]" pageId="4" pageNumber="244" type="distribution">
<paragraph id="716DEB161A4538420EFF3DA6FD5040C8" blockId="4.[151,1437,151,1241]" box="[199,712,1215,1241]" pageId="4" pageNumber="244">
<emphasis id="43A637041A4538420EFF3DA6FEF840C8" bold="true" box="[199,352,1215,1240]" pageId="4" pageNumber="244">Distribution.</emphasis>
Known only from
<typeStatus id="AE6955B41A4538420C0F3DD8FDFF40C9" box="[567,615,1217,1241]" pageId="4" pageNumber="244">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>