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<document ID-DOI="http://doi.org/10.5281/zenodo.6232002" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743181" ID-ISSN="0024-0672" ID-Zenodo-Dep="6232002" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E2740AE66B6686FEC4FCF4FF1D" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus anas Schilthuizen &amp; Perreau 2008, spec. nov." docType="treatment" docVersion="5" lastPageId="11" lastPageNumber="200" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="10" pageNumber="199" updateTime="1645572769617" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>189</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
<mods:identifier type="ISSN">0024-0672</mods:identifier>
<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6232002" ID-GBIF-Taxon="160743181" ID-Zenodo-Dep="6232002" LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D" httpUri="http://treatment.plazi.org/id/F97A87E2740AE66B6686FEC4FCF4FF1D" lastPageId="11" lastPageNumber="200" pageId="10" pageNumber="199">
<subSubSection box="[458,812,353,377]" pageId="10" pageNumber="199" type="nomenclature">
<paragraph blockId="10.[458,812,353,409]" box="[458,812,353,377]" pageId="10" pageNumber="199">
<heading box="[458,812,353,377]" centered="true" fontSize="10" level="3" pageId="10" pageNumber="199" reason="1">
<taxonomicName authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[458,697,353,377]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="anas" status="spec. nov.">
<emphasis bold="true" box="[458,697,353,377]" italics="true" pageId="10" pageNumber="199">Ptomaphaginus anas</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[703,812,354,377]" pageId="10" pageNumber="199" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[571,700,385,409]" pageId="10" pageNumber="199" type="description">
<paragraph blockId="10.[458,812,353,409]" box="[571,700,385,409]" pageId="10" pageNumber="199">
<heading box="[571,700,385,409]" centered="true" fontSize="10" level="3" pageId="10" pageNumber="199" reason="1">
(
<figureCitation box="[578,688,385,409]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="10" pageNumber="199">figs 16-17</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="199" type="materials_examined">
<paragraph blockId="10.[151,1120,448,751]" pageId="10" pageNumber="199">
Type material.—
<materialsCitation ID-GBIF-Occurrence="2512737961" collectionCode="SBPC, CMNC" collectorName="B. Bendell" country="Malaysia" location="Universiti Malaya Field Station" pageId="10" pageNumber="199" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<typeStatus box="[312,401,449,468]" pageId="10" pageNumber="199">Holotype</typeStatus>
<specimenCount box="[405,423,448,468]" pageId="10" pageNumber="199" type="male">♂</specimenCount>
:
<collectingCountry box="[433,521,449,468]" name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingRegion box="[529,614,449,468]" country="Malaysia" name="Selangor" pageId="10" pageNumber="199">Selangor</collectingRegion>
:
<quantity box="[622,699,449,468]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="10" pageNumber="199" unit="mi" value="15.0">15 miles</quantity>
N of
<collectingRegion box="[749,889,449,468]" country="Malaysia" name="Kuala Lumpur" pageId="10" pageNumber="199">Kuala Lumpur</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A64CDFE64FF09FE15" country="Malaysia" name="Universiti Malaya Field Station" pageId="10" pageNumber="199" stateProvince="Selangor">Universiti Malaya Field Station</location>
, Ulu Gombak, x-xi.1977 leg.
<collectorName box="[485,579,477,497]" pageId="10" pageNumber="199">B. Bendell</collectorName>
(in
<collectionCode box="[614,668,477,496]" pageId="10" pageNumber="199">SBPC</collectionCode>
, due to be deposited at
<collectionCode box="[887,958,477,496]" pageId="10" pageNumber="199">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2512737997" collectionCode="BOR" collectorName="B. Bendell" country="Malaysia" location="Universiti Malaya Field Station" pageId="10" pageNumber="199" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Selangor" typeStatus="paratype">
<typeStatus box="[972,1066,478,497]" pageId="10" pageNumber="199">Paratypes</typeStatus>
same locality as
<typeStatus box="[252,338,506,525]" pageId="10" pageNumber="199">holotype</typeStatus>
:
<specimenCount box="[347,400,505,525]" pageId="10" pageNumber="199" type="male">2 ♂♂</specimenCount>
<specimenCount box="[405,440,505,525]" pageId="10" pageNumber="199" type="female">1 ♀</specimenCount>
(in
<collectionCode box="[477,524,506,525]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="10" pageNumber="199" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2512737952" box="[539,731,505,525]" collectionCode="SBPC" collectorName="B. Bendell" country="Malaysia" isTailEnumeration="true" location="Universiti Malaya Field Station" pageId="10" pageNumber="199" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Selangor" typeStatus="paratype">
<specimenCount box="[539,592,505,525]" pageId="10" pageNumber="199" type="male">3 ♂♂</specimenCount>
<specimenCount box="[597,632,505,525]" pageId="10" pageNumber="199" type="female">1 ♀</specimenCount>
(in
<collectionCode box="[669,724,506,525]" pageId="10" pageNumber="199">SBPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2512737941" box="[740,873,505,525]" collectionCode="FRCS" country="Malaysia" pageId="10" pageNumber="199" specimenCount="1" specimenCount-female="1" stateProvince="Selangor" typeStatus="paratype">
<specimenCount box="[740,774,505,525]" pageId="10" pageNumber="199" type="female">1 ♀</specimenCount>
(in
<collectionCode box="[812,868,506,525]" country="Malaysia" httpUri="http://grbio.org/cool/sxta-ddia" name="Forest Research Centre" pageId="10" pageNumber="199">FRCS</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737938" collectingDate="1977-03-26" collectionCode="MHNG" collectorName="T. Jaccoud" country="Malaysia" location="Cameron Highlands" municipality="Berinchan" pageId="10" pageNumber="199" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="paratype">
<collectingCountry box="[882,970,506,525]" name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingRegion box="[979,1052,506,525]" country="Malaysia" name="Pahang" pageId="10" pageNumber="199">Pahang</collectingRegion>
:
<collectingMunicipality pageId="10" pageNumber="199">Berinchan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A6783FDB3FE43FDCC" box="[207,401,534,553]" country="Malaysia" municipality="Berinchan" name="Cameron Highlands" pageId="10" pageNumber="199" stateProvince="Pahang">Cameron Highlands</location>
,
<date box="[411,506,534,553]" pageId="10" pageNumber="199" value="1977-03-26">
<collectingDate box="[411,506,534,553]" pageId="10" pageNumber="199" value="1977-03-26">26.iii.1977</collectingDate>
</date>
, leg.
<collectorName box="[555,655,534,553]" pageId="10" pageNumber="199">T. Jaccoud</collectorName>
:
<specimenCount box="[663,698,533,553]" pageId="10" pageNumber="199" type="male">1 ♂</specimenCount>
(in
<collectionCode box="[735,810,534,553]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="199" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737950" collectingDate="1978-01-09" collectingDateMax="1978-01-16" collectingDateMin="1978-01-09" collectionCode="SBPC" collectorName="B. Bendell" country="Malaysia" location="Semongak" pageId="10" pageNumber="199" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry box="[825,913,534,553]" name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingCountry box="[922,1006,534,553]" name="Malaysia" pageId="10" pageNumber="199">Sarawak</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A64BBFDB3FB8EFDCC" box="[1015,1116,534,553]" country="Malaysia" name="Semongak" pageId="10" pageNumber="199">Semongak</location>
,
<date box="[151,250,562,581]" pageId="10" pageNumber="199" value="1978-01-09" valueMax="1978-01-16" valueMin="1978-01-09">
<collectingDate box="[151,250,562,581]" pageId="10" pageNumber="199" value="1978-01-09" valueMax="1978-01-16" valueMin="1978-01-09">9-16.i.1978</collectingDate>
</date>
leg.
<collectorName box="[297,396,562,582]" pageId="10" pageNumber="199">B. Bendell</collectorName>
:
<specimenCount box="[406,441,561,581]" pageId="10" pageNumber="199" type="male">1 ♂</specimenCount>
(in
<collectionCode box="[480,535,562,581]" pageId="10" pageNumber="199">SBPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737954" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="CJRZ" country="Malaysia" location="Batu Punggul Resort" pageId="10" pageNumber="199" specimenCount="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[551,639,562,581]" name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingRegion box="[649,708,562,581]" country="Malaysia" name="Sabah" pageId="10" pageNumber="199">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A6582FD96FC47FDA3" box="[718,917,562,582]" country="Malaysia" name="Batu Punggul Resort" pageId="10" pageNumber="199" stateProvince="Sabah">Batu Punggul Resort</location>
env.,
<date box="[973,1116,562,581]" pageId="10" pageNumber="199" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate box="[973,1116,562,581]" pageId="10" pageNumber="199" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24.vi-1.vii.1996</collectingDate>
</date>
, vegetation debris and forest floor litter accumulated around large trees near river:
<specimenCount box="[925,977,590,610]" pageId="10" pageNumber="199" type="male">2 ♂♂</specimenCount>
(in
<collectionCode box="[1012,1066,590,609]" pageId="10" pageNumber="199">CJRZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737971" collectingDate="1987-05-08" collectingDateMax="1987-05-16" collectingDateMin="1987-05-08" collectionCode="MHNG" collectorName="A. Smetana" country="Malaysia" elevation="1500" location="Mt. Kinabalu Park Headquarters" pageId="10" pageNumber="199" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingRegion box="[217,276,619,638]" country="Malaysia" name="Sabah" pageId="10" pageNumber="199">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A6651FDCEFD84FD9B" box="[285,598,619,638]" country="Malaysia" name="Mt. Kinabalu Park Headquarters" pageId="10" pageNumber="199" stateProvince="Sabah">Mt. Kinabalu Park Headquarters</location>
,
<quantity box="[607,675,619,638]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="199" unit="m" value="1500.0">
<elevation box="[607,675,619,638]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="199" unit="m" value="1500.0">1500 m</elevation>
</quantity>
alt.,
<date box="[721,826,619,638]" pageId="10" pageNumber="199" value="1987-05-08" valueMax="1987-05-16" valueMin="1987-05-08">
<collectingDate box="[721,826,619,638]" pageId="10" pageNumber="199" value="1987-05-08" valueMax="1987-05-16" valueMin="1987-05-08">8-16.v.1987</collectingDate>
</date>
,
<collectingMethod box="[835,995,619,638]" pageId="10" pageNumber="199">interception trap</collectingMethod>
, leg.
<collectorName pageId="10" pageNumber="199">A. Smetana</collectorName>
:
<specimenCount box="[205,241,646,666]" pageId="10" pageNumber="199" type="male">1 ♂</specimenCount>
(in
<collectionCode box="[282,357,647,666]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="199" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737947" collectingDate="1987-05-20" collectionCode="MHNG, CMPR" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="700" location="Kibongol" pageId="10" pageNumber="199" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[374,462,647,666]" name="Malaysia" pageId="10" pageNumber="199">Malaysia</collectingCountry>
:
<collectingRegion box="[473,532,647,666]" country="Malaysia" name="Sabah" pageId="10" pageNumber="199">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E2740AE66B6686FEC4FCF4FF1D:740C602F740AE66A6552FD22FDA5FD7F" box="[542,631,647,666]" country="Malaysia" name="Kibongol" pageId="10" pageNumber="199" stateProvince="Sabah">Kibongol</location>
, (
<locationDeviation box="[649,861,647,666]" location="Tambunan" pageId="10" pageNumber="199">
<quantity box="[649,699,647,666]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="199" unit="km" value="7.0">7 km</quantity>
N of Tambunan
</locationDeviation>
),
<quantity box="[879,938,647,666]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="199" unit="m" value="700.0">
<elevation box="[879,938,647,666]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="199" unit="m" value="700.0">700 m</elevation>
</quantity>
alt.,
<date box="[988,1074,647,666]" pageId="10" pageNumber="199" value="1987-05-20">
<collectingDate box="[988,1074,647,666]" pageId="10" pageNumber="199" value="1987-05-20">20.v.1987</collectingDate>
</date>
, leg.
<collectorName box="[151,257,675,694]" pageId="10" pageNumber="199">Burckhardt</collectorName>
&amp;
<collectorName box="[285,330,675,694]" pageId="10" pageNumber="199">Löbl</collectorName>
:
<specimenCount box="[338,390,674,694]" pageId="10" pageNumber="199" type="male">4 ♂♂</specimenCount>
and
<specimenCount box="[437,489,674,694]" pageId="10" pageNumber="199" type="female">2 ♀♀</specimenCount>
(in
<collectionCode box="[524,596,675,694]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="199" type="Museum">MHNG</collectionCode>
and
<collectionCode box="[643,709,675,694]" country="India" name="Centre for Medicinal Plants Research" pageId="10" pageNumber="199" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="10.[151,1120,448,751]" pageId="10" pageNumber="199">
<typeStatus box="[151,337,703,722]" isOtherTypeStatus="true" pageId="10" pageNumber="199">Additional material</typeStatus>
(not included in the type series).— 36 fragmented individuals in preservative from the type locality (
<collectionCode box="[314,363,732,751]" country="USA" lsid="urn:lsid:biocol.org:col:15587" name="Michigan State University" pageId="10" pageNumber="199" type="Herbarium">MSC</collectionCode>
, to be deposited in
<collectionCode box="[546,602,732,751]" pageId="10" pageNumber="199">SBPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="199" type="diagnosis">
<paragraph blockId="10.[151,1120,793,1659]" pageId="10" pageNumber="199">
Diagnosis.— This new species is relatively large and convex. The aedeagus is characteristically shaped (
<figureCitation box="[395,506,825,849]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="10" pageNumber="199">figs 16-17</figureCitation>
): strongly curved, distally tapering into a narrow, flattened, and slightly upturned apex. In general shape reminiscent of the aedeagus of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis italics="true" pageId="10" pageNumber="199">P. murphyi</emphasis>
</taxonomicName>
, which, however, is characterized by a squarish, subapical expansion.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="199" type="description">
<paragraph blockId="10.[151,1120,793,1659]" pageId="10" pageNumber="199">
Description.— Habitus ovoid, relatively convex. Reddish brown to black (though the dark colouration in some specimens may be due to preservation); the tarsi and the edges of thorax and elytra lighter brown. Length
<quantity box="[705,829,987,1011]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.9" metricValueMin="2.3" pageId="10" pageNumber="199" unit="mm" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.3-2.9 mm</quantity>
(n = 9). Antennae slender but not very long, 1.1 times as long as the width of the head (n = 3). Articles 9 and 10 slightly broader than long; article 6 square. Male protarsi moderately dilated: the protibia at its largest width (excluding the lateral spines) is 1.4 times broader than the first article of the protarsus (n = 1). Female protarsi undilated. Thorax 1.65 times as wide as long (n = 3), convex, with the caudal corners distinctly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra short, 1.1 times as long (measured from the caudal tip of the scutellum) as their combined width (n = 3). Elytral apices moderately truncate, identical in both sexes. Aedeagus (
<figureCitation box="[842,949,1246,1270]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="10" pageNumber="199">figs 16-17</figureCitation>
) in dorsal view gradually narrowing towards the apex. The tip is triangular and slightly asymmetric. A broad medial furrow runs over the dorsal side of the aedeagus, and dissolves just short of the apex. In lateral view, the aedeagus is strongly curved, almost semicircular, and distinctly thickened at the point of strongest curvature. In some specimens, the tip of the internal stylet is seen to emerge just ventral of the aedeagal apex. Male genital segment relatively narrow, spoon-shaped, four times as long as wide.
</paragraph>
</subSubSection>
<subSubSection box="[193,1118,1473,1497]" pageId="10" pageNumber="199" type="distribution">
<paragraph blockId="10.[151,1120,793,1659]" box="[193,1118,1473,1497]" pageId="10" pageNumber="199">Distribution.— Apparently widely distributed in the Malay Peninsula and Borneo.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="199" type="discussion">
<paragraph blockId="10.[151,1120,793,1659]" pageId="10" pageNumber="199">
Remarks.— Based on aedeagal shape,
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[617,687,1507,1530]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="anas">
<emphasis box="[617,687,1507,1530]" italics="true" pageId="10" pageNumber="199">P. anas</emphasis>
</taxonomicName>
may be related to several other species from
<collectingRegion box="[211,306,1538,1562]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="199">Sumatra</collectingRegion>
and the Malay Peninsula, viz.,
<taxonomicName authority="Szymczakowski" authorityName="Szymczakowski. Our" authorityYear="1964" box="[650,937,1538,1562]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis box="[650,749,1538,1562]" italics="true" pageId="10" pageNumber="199">P. tarsalis</emphasis>
Szymczakowski
</taxonomicName>
,
<taxonomicName authority="Jeannel" authorityName="Jeannel" authorityYear="1936" box="[946,1115,1538,1562]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="rufus">
<emphasis box="[946,1025,1538,1562]" italics="true" pageId="10" pageNumber="199">P. rufus</emphasis>
Jeannel
</taxonomicName>
,
<taxonomicName authority="Szymczakowski" authorityName="Szymczakowski" authorityYear="1970" box="[151,466,1570,1595]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis box="[151,278,1571,1595]" italics="true" pageId="10" pageNumber="199">P. lacertosus</emphasis>
Szymczakowski
</taxonomicName>
, and
<taxonomicName authority="Szymczakowski." authorityName="Szymczakowski." box="[525,831,1570,1595]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis box="[525,638,1571,1595]" italics="true" pageId="10" pageNumber="199">P. murphyi</emphasis>
Szymczakowski.
</taxonomicName>
The Sumatran
<taxonomicName authority="Szymczakowski" authorityName="Szymczakowski" authorityYear="1964" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis box="[1000,1119,1571,1595]" italics="true" pageId="10" pageNumber="199">P. jacobsoni</emphasis>
Szymczakowski
</taxonomicName>
, known from females only, appears similar to
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[843,913,1604,1627]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="199" phylum="Arthropoda" rank="species" species="anas">
<emphasis box="[843,913,1604,1627]" italics="true" pageId="10" pageNumber="199">P. anas</emphasis>
</taxonomicName>
, but has its caudal thoracal corners not as strongly drawn out.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="200" type="etymology">
<paragraph blockId="11.[151,1119,192,248]" pageId="11" pageNumber="200">
Etymology.—
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[352,398,193,216]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="anas">
<emphasis box="[352,398,193,216]" italics="true" pageId="11" pageNumber="200">anas</emphasis>
</taxonomicName>
, a Latin noun in apposition, meaning ‘duck’. The name refers to the tip of the aedeagus, which has the shape of a duck-bill.
</paragraph>
</subSubSection>
</treatment>
</document>
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