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<mods:title id="3B4FE48F64C685193BA77EA9AA4CB7DD">The chimeric Onobrychis tenoreana (Fabaceae) and lectotypification of the names O. laconica and O. pentelica</mods:title>
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<mods:namePart id="EDE30777F4021EDE5CA7338FBFC65250">Bernardo, Liliana</mods:namePart>
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<mods:affiliation id="A701B0109F37B274EB8C1FE03F5CE196">Department of Biology, Ecology and Geological Sciences, University of Calabria, 87036 Rende (Cosenza), Italy. liliana. bernardo @ unical. it; https: // orcid. org / 0000 - 0002 - 6703 - 6409</mods:affiliation>
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<treatment id="F84087F0FFA6A51C1BB8FAF7AE93FDF6" ID-DOI="http://doi.org/10.5281/zenodo.5483842" ID-GBIF-Taxon="185433428" ID-Zenodo-Dep="5483842" LSID="urn:lsid:plazi:treatment:F84087F0FFA6A51C1BB8FAF7AE93FDF6" httpUri="http://treatment.plazi.org/id/F84087F0FFA6A51C1BB8FAF7AE93FDF6" lastPageId="2" lastPageNumber="242" pageId="0" pageNumber="240">
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<paragraph id="705636E6FFA6A51E1BB8FAF7AC73F84B" blockId="0.[136,1452,1928,2062]" box="[136,557,1928,1954]" pageId="0" pageNumber="240">
<heading id="2B1E818AFFA6A51E1BB8FAF7AC73F84B" bold="true" box="[136,557,1928,1954]" fontSize="11" level="1" pageId="0" pageNumber="240" reason="1">
<emphasis id="429DEAF4FFA6A51E1BB8FAF7AC73F84B" bold="true" box="[136,557,1928,1954]" pageId="0" pageNumber="240">
<emphasis id="429DEAF4FFA6A51E1BB8FAF7AF6FF84B" bold="true" box="[136,305,1928,1954]" italics="true" pageId="0" pageNumber="240">Typification of</emphasis>
<taxonomicName id="B7E94D65FFA6A51E1A08FAF7AC73F84B" ID-CoL="49L24" authorityName="Orph. ex Boissier" authorityYear="1872" box="[312,557,1928,1954]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="240" phylum="Tracheophyta" rank="species" species="laconica">Onobrychis laconica</taxonomicName>
</emphasis>
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<paragraph id="705636E6FFA6A51F1BB8FAD1AF35FE66" blockId="0.[136,1452,1928,2062]" lastBlockId="1.[136,1452,159,399]" lastPageId="1" lastPageNumber="241" pageId="0" pageNumber="240">
<taxonomicName id="B7E94D65FFA6A51E1BB8FAD1AF39F82F" ID-CoL="49L24" authorityName="Orph. ex Boissier" authorityYear="1872" box="[136,359,1966,1990]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="240" phylum="Tracheophyta" rank="species" species="laconica">
<emphasis id="429DEAF4FFA6A51E1BB8FAD1AF39F82F" box="[136,359,1966,1990]" italics="true" pageId="0" pageNumber="240">Onobrychis laconica</emphasis>
</taxonomicName>
was validly published by
<bibRefCitation id="14784B17FFA6A51E194DFAD1AD45F82F" author="Boissier, E." box="[637,795,1966,1990]" pageId="0" pageNumber="240" refId="ref2944" refString="Boissier, E. (1872 - 1873) Flora orientalis, vol. 2. H. George, Genevae, Basileae, Lugduni, 1159 pp." type="book" year="1872">Boissier (1872</bibRefCitation>
1873: 530) through a diagnosis and a description; the habitat and provenance were also provided (“Hab. in siccis regionis subalpinae, mons Malevo Laconiae (Orph), mons Kyllene alt. 6500 (Heldr!), mons Tymphrestus Aetoliae supra Carpenisi alt. 4000 (Samar. et Guicc.!)”.
<bibRefCitation id="14784B17FFA6A51E1F52FA89AB5EF7E7" author="Boissier, E." box="[1122,1280,2038,2062]" pageId="0" pageNumber="240" refId="ref2944" refString="Boissier, E. (1872 - 1873) Flora orientalis, vol. 2. H. George, Genevae, Basileae, Lugduni, 1159 pp." type="book" year="1872">Boissier (1872</bibRefCitation>
1873) reported as synonyms “
<taxonomicName id="B7E94D65FFA7A51F1A16FDDFAFA8FF5E" ID-CoL="9DF8H" authority="Heldr." authorityName="Heldr." box="[294,502,159,183]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="pulchella">
<emphasis id="429DEAF4FFA7A51F1A16FDDFAFF5FF5E" box="[294,427,159,183]" italics="true" pageId="1" pageNumber="241">O. pulchella</emphasis>
Heldr.
</taxonomicName>
in Boiss. Diagn. Ser. II, 6, p. 64, non Schrenk” [i.e.,
<taxonomicName id="B7E94D65FFA7A51F1F09FDDFAB71FF5E" ID-CoL="9DF8H" authority="Heldreich" authorityName="Heldreich" box="[1081,1327,159,183]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="pulchella">
<emphasis id="429DEAF4FFA7A51F1F09FDDFAAE0FF5E" box="[1081,1214,159,183]" italics="true" pageId="1" pageNumber="241">O. pulchella</emphasis>
Heldreich
</taxonomicName>
in
<bibRefCitation id="14784B17FFA7A51F1E64FDE0AEA7FF32" author="Boissier, P. E." pageId="1" pageNumber="241" refId="ref2904" refString="Boissier, P. E. (1859) Diagnoses Plantarum Orientalium novarum, ser. 2, vol. 6. B. Herman &amp; J. - B. Bailliere, Lipsiae &amp; Paris, 148 pp." type="book" year="1859">Boissier (1859: 64)</bibRefCitation>
, and “
<emphasis id="429DEAF4FFA7A51F1A0BFDBBAF8DFF32" box="[315,467,195,219]" italics="true" pageId="1" pageNumber="241">O. Heldreichii</emphasis>
Bge. in litt.” (not validly published according to the Art. 36.1b of ICN).
<taxonomicName id="B7E94D65FFA7A51F1FFDFDBCABF2FF32" ID-CoL="49L24" authorityName="Orph. ex Boissier" authorityYear="1872" box="[1229,1452,195,219]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="laconica">
<emphasis id="429DEAF4FFA7A51F1FFDFDBCABF2FF32" box="[1229,1452,195,219]" italics="true" pageId="1" pageNumber="241">Onobrychis laconica</emphasis>
</taxonomicName>
is therefore a replacement name, due to the existence of the previous and legitimate
<taxonomicName id="B7E94D65FFA7A51F18C8FD97AA89FF16" ID-CoL="9DF8H" authority="Schrenk" authorityName="Schrenk" box="[1016,1239,231,255]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="pulchella">
<emphasis id="429DEAF4FFA7A51F18C8FD97AA25FF16" box="[1016,1147,231,255]" italics="true" pageId="1" pageNumber="241">O. pulchella</emphasis>
Schrenk
</taxonomicName>
in
<bibRefCitation id="14784B17FFA7A51F1FC8FD98AEA6FECA" author="Fischer, F. E. L. &amp; Meyer, C. A." pageId="1" pageNumber="241" refId="ref3087" refString="Fischer, F. E. L. &amp; Meyer, C. A. (1841) Enumeratio Plantarum Novarum vol. 1. G. Fischeri, Petropoli, 113 pp." type="book" year="1841">Fischer &amp; Meyer (1841: 87)</bibRefCitation>
(Arts. 6.11 and 53.3 of ICN). The
<typeStatus id="AF528844FFA7A51F1957FC73ACCBFECD" box="[615,661,268,292]" pageId="1" pageNumber="241">type</typeStatus>
of
<taxonomicName id="B7E94D65FFA7A51F1988FC73AD6AFECA" ID-CoL="49L24" authorityName="Orph. ex Boissier" authorityYear="1872" box="[696,820,267,291]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="laconica">
<emphasis id="429DEAF4FFA7A51F1988FC73AD6AFECA" box="[696,820,267,291]" italics="true" pageId="1" pageNumber="241">O. laconica</emphasis>
</taxonomicName>
is the same of that of the illegitimate
<taxonomicName id="B7E94D65FFA7A51F1FF5FC73ABCFFECA" ID-CoL="9DF8H" authority="Heldr." authorityName="Heldr." box="[1221,1425,267,291]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="pulchella">
<emphasis id="429DEAF4FFA7A51F1FF5FC73AB16FECA" box="[1221,1352,267,291]" italics="true" pageId="1" pageNumber="241">O. pulchella</emphasis>
Heldr.
</taxonomicName>
in Boiss., non Schrenk (Art. 7.4 of ICN). The latter name was published providing a detailed description, habitat, provenance (“Hab. in pascuis regionis abietinae montis
<emphasis id="429DEAF4FFA7A51F1952FC2CADE1FE82" box="[610,959,339,363]" italics="true" pageId="1" pageNumber="241">Tymphresti (Veluchi) Eurytaniae</emphasis>
supra
<emphasis id="429DEAF4FFA7A51F1F38FC2BAA27FE82" box="[1032,1145,340,363]" italics="true" pageId="1" pageNumber="241">Carpenitzi</emphasis>
alt. 4000), and a diagnosis, all in Latin language.
</paragraph>
<caption id="2496666EFFA7A51F1BB8F56CADF7F7C3" ID-DOI="http://doi.org/10.5281/zenodo.5425302" ID-Zenodo-Dep="5425302" box="[136,937,2067,2090]" httpUri="https://zenodo.org/record/5425302/files/figure.png" pageId="1" pageNumber="241" startId="1.[136,229,2067,2089]" targetBox="[289,1299,422,2041]" targetPageId="1">
<paragraph id="705636E6FFA7A51F1BB8F56CADF7F7C3" blockId="1.[136,937,2067,2090]" box="[136,937,2067,2090]" pageId="1" pageNumber="241">
<emphasis id="429DEAF4FFA7A51F1BB8F56CAEA0F7C1" bold="true" box="[136,254,2067,2089]" pageId="1" pageNumber="241">FIGURE 1.</emphasis>
Lectotype of the name
<taxonomicName id="B7E94D65FFA7A51F1AD1F56BACE9F7C0" ID-CoL="49L5P" authorityName="Lacaita" authorityYear="1911" box="[481,695,2068,2089]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="241" phylum="Tracheophyta" rank="species" species="tenoreana">
<emphasis id="429DEAF4FFA7A51F1AD1F56BACE9F7C0" box="[481,695,2068,2089]" italics="true" pageId="1" pageNumber="241">Onobrychis tenoreana</emphasis>
</taxonomicName>
(BM!, without barcode).
</paragraph>
</caption>
<paragraph id="705636E6FFA4A51C1B8DFDDFAE93FDF6" blockId="2.[136,1452,159,543]" pageId="2" pageNumber="242">
We traced the following specimens: FI010913, FI010914, JE00010035, MPU021490, MPU022851, P03077995, P02850986, and P02274509 (duplicates are available at G -Boiss and TAU; see
<bibRefCitation id="14784B17FFA4A51C1F3CFDBCAA86FF32" author="Raptis, D. &amp; Karousou, R. &amp; Hanlidou, E." box="[1036,1240,195,219]" pageId="2" pageNumber="242" pagination="307 - 318" refId="ref3679" refString="Raptis, D., Karousou, R. &amp; Hanlidou, E. (2019) Type specimens in the Herbarium of Aristotle University of Thessaloniki (TAU). Botanika Chronika 22: 307 - 318." type="journal article" year="2019">
Raptis
<emphasis id="429DEAF4FFA4A51C1F6AFDBBAACAFF32" box="[1114,1172,195,219]" italics="true" pageId="2" pageNumber="242">et al.</emphasis>
2019
</bibRefCitation>
). All the examined specimens bear the following original label, printed but in Heldreichs handwriting (P02274509 was hand-copied): “De Heldreich Herbarium Graecum Normale | N. 664.
<taxonomicName id="B7E94D65FFA4A51C19F0FC74ADA7FECA" ID-CoL="9DF8H" authority="Heldr." authorityName="Heldr." authorityYear="2021" box="[704,1017,267,291]" class="Magnoliopsida" family="Fabaceae" genus="Onobrychis" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="242" phylum="Tracheophyta" rank="species" species="pulchella" status="sp. nov.">
<emphasis id="429DEAF4FFA4A51C19F0FC74ADF5FECA" box="[704,939,267,291]" italics="true" pageId="2" pageNumber="242">Onobrychis pulchella</emphasis>
Heldr.
</taxonomicName>
<taxonomicNameLabel id="59AE578FFFA4A51C1F33FC73AA04FECA" box="[1027,1114,268,291]" pageId="2" pageNumber="242" rank="species">nov. sp.</taxonomicNameLabel>
| In pascuis reg. abietinae m. Tymphresti Veluchi hod. Eurytaniae, l.d. Rovia, alt. 4000 supra Carpenisi. D.
<date id="04571026FFA4A51C18DBFC50AA27FEAE" box="[1003,1145,303,327]" pageId="2" pageNumber="242" value="1857-08-08">8 Aug. 1857</date>
. Detexerunt am. Samaritani A Guicciardi”. Each one of these specimens is eligible as
<typeStatus id="AF528844FFA4A51C1830FC2CAD3BFE82" box="[768,869,339,363]" pageId="2" pageNumber="242" type="lectotype">lectotype</typeStatus>
. but FI010913 is the best choice, due to its satisfying conservation status and completeness. In fact, it is represented by three individuals, two of them flowering, the remaining one showing mature fruits (very important for diagnostic purposes). The plant is characterized by appressed hairiness, elliptic-linear leaflets, calyx with brownish hairs, corolla twice longer than calyx, legume a little wider than calyx and with 34 short teeth, so both agreeing with the protologue (
<bibRefCitation id="14784B17FFA4A51C19E9FC9CAD2EFE12" author="Boissier, E." box="[729,880,483,507]" pageId="2" pageNumber="242" refId="ref2944" refString="Boissier, E. (1872 - 1873) Flora orientalis, vol. 2. H. George, Genevae, Basileae, Lugduni, 1159 pp." type="book" year="1872">Boissier 1872</bibRefCitation>
1873) and with the current usage of the name (
<bibRefCitation id="14784B17FFA4A51C1E4DFC9CAE9FFDF6" author="Ball, P. W." pageId="2" pageNumber="242" pagination="41 - 42" refId="ref2480" refString="Ball, P. W. (1968) Onobrychis. In: Heywood, V. H. (Ed.) Flora Europaea. Notulae Systematicae ad Floram Europaeam spectantes. Feddes Repertorium 79: 41 - 42. https: // doi. org / 10.1002 / fedr. 4910790102" type="journal article" year="1968">Ball 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>