494 lines
55 KiB
XML
494 lines
55 KiB
XML
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<mods:title id="3A937C84CC800E3743D80DDF97D94B5A">Taxonomic and nomenclatural changes in three species of Pterostichus Bonelli (Coleoptera: Carabidae) from the Far East</mods:title>
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<mods:namePart id="140DC6ABEC63263BF220B7F005B63EA8">Sasakawa, Kôji</mods:namePart>
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<mods:affiliation id="706A98FCE9B1D5984942269337050140">Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, Yayoi-cho, Inage-ku, Chiba, Japan. ksasa @ chiba-u. jp; https: // orcid. org / 0000 - 0001 - 9246 - 5777</mods:affiliation>
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<mods:nameIdentifier id="4F65F0C1DF05075FA58235BA20BBE533" type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
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<mods:namePart id="69EBEC2867482AB295F644B56CCB125E">Berlov, Oleg</mods:namePart>
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<mods:affiliation id="09399C6CA9DE427EF030828E5D33D040">Laboratory of Experimental Animals, Anti-Plague Institute of Siberia and the Russian Far East, Irkutsk, Russia.</mods:affiliation>
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<mods:namePart id="4FB37348D74C2AA50BE803536AADDE46">Okuzaki, Yutaka</mods:namePart>
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<mods:affiliation id="5ECCCE9680CF7AF11CA0ED3066E8C1B6">Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido, Japan. & Department of General Systems Studies, Graduate School of Arts and Sciences, University of Tokyo, Komaba, Meguro-ku, Tokyo, Japan. yutaka @ g. ecc. u-tokyo. ac. jp; https: // orcid. org / 0000 - 0002 - 9362 - 0078</mods:affiliation>
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<mods:nameIdentifier id="7951905F94ACDC7FB299A9B8B6B42B36" type="email">yutaka@g.ecc.u-tokyo.ac.jp</mods:nameIdentifier>
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<mods:date id="ACE0EAF7E3DE0F1BBBDE61B9C8C94FA9">2020</mods:date>
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<treatment id="F8316A14FE42FFCCFF54C95EB959659B" ID-DOI="http://doi.org/10.5281/zenodo.4448249" ID-GBIF-Taxon="177006701" ID-Zenodo-Dep="4448249" LSID="urn:lsid:plazi:treatment:F8316A14FE42FFCCFF54C95EB959659B" httpUri="http://treatment.plazi.org/id/F8316A14FE42FFCCFF54C95EB959659B" lastPageId="2" lastPageNumber="418" pageId="1" pageNumber="417">
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<taxonomicName id="B798A081FE42FFCFFF54C95EBE63632A" ID-CoL="4Q47R" authority="Morawitz, 1862" authorityName=": Morawitz" authorityYear="1862" box="[151,773,1122,1149]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Petrophilus">
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<emphasis id="42EC0710FE42FFCFFF54C95EBC4B632B" bold="true" box="[151,301,1122,1149]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
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(
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<emphasis id="42EC0710FE42FFCFFEFDC95EBCAC632B" bold="true" box="[318,458,1122,1149]" italics="true" pageId="1" pageNumber="417">Petrophilus</emphasis>
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)
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<emphasis id="42EC0710FE42FFCFFE1FC95FBF5D632B" bold="true" box="[476,571,1123,1149]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
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<bibRefCitation id="1409A6F3FE42FFCFFD82C95EBE63632A" author="Morawitz, A." box="[577,773,1122,1148]" pageId="1" pageNumber="417" pagination="180 - 228" refId="ref6690" refString="Morawitz, A. (1862) Vorlaufige Diagnosen neuer Coleopteren aus Sudost-Sibirien. Melanges Biologiques tires du Bulletin de l'Academie des Sciences de St. - Petersburg, 4, 180 - 228." type="journal article" year="1862">Morawitz, 1862</bibRefCitation>
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<paragraph id="7027DB02FE42FFCFFF54C9BBBD9963F7" blockId="1.[151,773,1122,1185]" box="[151,255,1159,1185]" pageId="1" pageNumber="417">
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<figureCitation id="E8A3C787FE42FFCFFF54C9BBBD8563F7" box="[151,227,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1823,1847]" captionTargetBox="[153,1432,741,1799]" captionTargetId="figure-288@2.[151,1436,741,1799]" captionTargetPageId="2" captionText="FIGURE 1. Differences in male genital structures (top, left posterolateral view; bottom, right ventrolateral view) among subspecies of Pterostichus eximius. Red, green, and blue arrows denote sublobe rv-2 on the endophallus, a swelling adjoining lobe ll on the endophallus, and the dorsal contour at the aedeagal subapex, respectively. Phylogenetic relationships inferred from these characters are shown." figureDoi="http://doi.org/10.5281/zenodo.4401757" httpUri="https://zenodo.org/record/4401757/files/figure.png" pageId="1" pageNumber="417">Figs. 1</figureCitation>
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,
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<figureCitation id="E8A3C787FE42FFCFFF32C9BBBD9963F7" box="[241,255,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1257,1281]" captionTargetBox="[151,1436,694,1233]" captionTargetId="figure-205@3.[151,1436,694,1233]" captionTargetPageId="3" captionText="FIGURE 2. Habitus (A, B), male genitalia (C–F), and female genitalia (G, H) of Pterostichus eximius rishiridakensis ssp. nov. A, dorsal view of holotype male; B, dorsal view of paratype female; C, right lateral view of aedeagus with everted endophallus (holotype male); D, dorsal view of aedeagal apex (paratype male); E, left lateral view of aedeagus with everted endophallus (holotype male); F, left lateral view of right paramere (holotype male); G, dorsal view of right stylomere 2 (paratype female); H, ventral view of right stylomere 2 (paratype female). gp, gonopore; ll, left lateral lobe; lv-1, basal sub-lobe of left ventral lobe; lv-2, apical sub-lobe of left ventral lobe; rv-1, basal sub-lobe of right ventral lobe; rv-2, apical sub-lobe of right ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.4401759" httpUri="https://zenodo.org/record/4401759/files/figure.png" pageId="1" pageNumber="417">2</figureCitation>
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<taxonomicName id="B798A081FE42FFCFFF54C9F7BFFD63B5" ID-CoL="4Q47R" authority=": Morawitz (1862)" authorityName=": Morawitz" authorityYear="1862" box="[151,667,1227,1251]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Steropus">
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<emphasis id="42EC0710FE42FFCFFF54C9F7BC7363B5" box="[151,277,1227,1251]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
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(
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<emphasis id="42EC0710FE42FFCFFEE6C9F7BC1B63B4" box="[293,381,1227,1250]" italics="true" pageId="1" pageNumber="417">Steropus</emphasis>
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)
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<emphasis id="42EC0710FE42FFCFFE4EC9F0BCBC63B5" box="[397,474,1228,1251]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
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:
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<bibRefCitation id="1409A6F3FE42FFCFFE2AC9F7BFFD63B5" author="Morawitz, A." box="[489,667,1227,1251]" pageId="1" pageNumber="417" pagination="180 - 228" refId="ref6690" refString="Morawitz, A. (1862) Vorlaufige Diagnosen neuer Coleopteren aus Sudost-Sibirien. Melanges Biologiques tires du Bulletin de l'Academie des Sciences de St. - Petersburg, 4, 180 - 228." type="journal article" year="1862">Morawitz (1862)</bibRefCitation>
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</taxonomicName>
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: 212
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(original description),
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locality: “Zagan-olui und Tschindansk am Onon” [Transbaikalia, East Siberia].
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<taxonomicName id="B798A081FE42FFCFFF54C834BFF46276" ID-CoL="4Q47R" authority=": Jedlicka (1962)" authorityName="Jedlicka" authorityYear="1962" box="[151,658,1288,1313]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Euryperis">
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<emphasis id="42EC0710FE42FFCFFF54C834BC736276" box="[151,277,1288,1312]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
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(
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<emphasis id="42EC0710FE42FFCFFEE7C835BCEE6276" box="[292,392,1289,1312]" italics="true" pageId="1" pageNumber="417">Euryperis</emphasis>
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)
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<emphasis id="42EC0710FE42FFCFFE55C835BC856276" box="[406,483,1289,1312]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
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:
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<bibRefCitation id="1409A6F3FE42FFCFFE32C834BFF46276" author="Jedlicka, A." box="[497,658,1288,1312]" pageId="1" pageNumber="417" pagination="177 - 346" refId="ref5989" refString="Jedlicka, A. (1962) Monographie des Tribus Pterostichini aus Ostasien (Pterostichini, Trigonotomi, Myadi) (Coleoptera-Ca- rabidae). Entomologische Anhandlungen und Berichte aus dem Staatliche Museum fur Tierkunde in Dresden, 26 (21), 177 - 346." type="journal article" year="1962">Jedlička (1962)</bibRefCitation>
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</taxonomicName>
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||
: 258
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</treatmentCitation>
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;
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<treatmentCitation id="F139FD13FE42FFCFFD0CC834BED46276" author="Lafer, G. S. & Nilsson, A. N. & Kholin, S. K." box="[719,946,1288,1313]" page="16" pageId="1" pageNumber="417" year="1997">
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<bibRefCitation id="1409A6F3FE42FFCFFD0CC834BEEA6277" author="Lafer, G. S. & Nilsson, A. N. & Kholin, S. K." box="[719,908,1288,1313]" pageId="1" pageNumber="417" pagination="13 - 17" refId="ref6435" refString="Lafer, G. S., Nilsson, A. N. & Kholin, S. K. (1997) Additional records and new synonyms of Cicindelidae and Carabidae (Coleoptera) from the lsland of Sakhalin in the Russian Far East. Entomologica Fennica, 8, 13 - 17. https: // doi. org / 10.33338 / ef. 83916" type="journal article" year="1997">
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Lafer
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||
<emphasis id="42EC0710FE42FFCFFCCFC835BE596276" box="[780,831,1288,1312]" italics="true" pageId="1" pageNumber="417">et al.</emphasis>
|
||
(1997)
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</bibRefCitation>
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||
: 16
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||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7027DB02FE42FFCFFF54C81BB9556208" blockId="1.[151,1437,1227,1497]" pageId="1" pageNumber="417">
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<treatmentCitationGroup id="5088FC2CFE42FFCFFF54C81BB9556208" pageId="1" pageNumber="417">
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<treatmentCitation id="F139FD13FE42FFCFFF54C81BBE546269" author="Kirschenhofer, E." box="[151,818,1319,1343]" page="222" pageId="1" pageNumber="417" year="1985">
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<taxonomicName id="B798A081FE42FFCFFF54C81BBE646269" authority=": Kirschenhofer (1985)" authorityName="Kirschenhofer" authorityYear="1985" box="[151,770,1319,1343]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="sachalinensis" subGenus="Steropus">
|
||
<emphasis id="42EC0710FE42FFCFFF54C81BBC736269" box="[151,277,1319,1343]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="42EC0710FE42FFCFFEE6C81BBC1B6268" box="[293,381,1319,1342]" italics="true" pageId="1" pageNumber="417">Steropus</emphasis>
|
||
)
|
||
<emphasis id="42EC0710FE42FFCFFE4FC81BBF736269" box="[396,533,1319,1343]" italics="true" pageId="1" pageNumber="417">sachalinensis</emphasis>
|
||
:
|
||
<materialsCitation id="C0F0D15FFE42FFCFFDE0C81BBE646269" ID-GBIF-Occurrence="3019437305" box="[547,770,1319,1343]" location="Mt. Chekhov" pageId="1" pageNumber="417" specimenCount="1" stateProvince="Sakhalin">
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<bibRefCitation id="1409A6F3FE42FFCFFDE0C81BBE646269" author="Kirschenhofer, E." box="[547,770,1319,1343]" pageId="1" pageNumber="417" pagination="221 - 231" refId="ref6265" refString="Kirschenhofer, E. (1985) Zweineue Pterostichus - Arten aus Sachalin: Pterostichus (Steropus) sachalinensis sp. n. und Pterostichus (Lyperopherus) wellschmiedi sp. n. Entomofauna, 6, 221 - 231." type="journal article" year="1985">Kirschenhofer (1985)</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
: 222
|
||
</treatmentCitation>
|
||
(original description),
|
||
<typeStatus id="AF2365A0FE42FFCFFBDDC814B92C6216" box="[1054,1098,1320,1344]" pageId="1" pageNumber="417">type</typeStatus>
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locality: “Süd-Sachalin, Umgeb. Nowoalexandrowsk, Bergwald (Tschechow-Berg)” [
|
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<location id="75478DD9FE42FFCFFD19C87ABE026208" LSID="urn:lsid:plazi:treatment:F8316A14FE42FFCCFF54C95EB959659B:75478DD9FE42FFCFFD19C87ABE026208" box="[730,868,1350,1374]" name="Mt. Chekhov" pageId="1" pageNumber="417" stateProvince="Sakhalin">Mt. Chekhov</location>
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, southern
|
||
<collectingRegion id="B25C15E0FE42FFCFFC08C87AB94F6208" box="[971,1065,1350,1374]" country="Russia" name="Sakhalin" pageId="1" pageNumber="417">Sakhalin</collectingRegion>
|
||
].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7027DB02FE42FFCFFF54C858BC1762CD" blockId="1.[151,1437,1227,1497]" pageId="1" pageNumber="417">
|
||
<treatmentCitationGroup id="5088FC2CFE42FFCFFF54C858BC1762CD" pageId="1" pageNumber="417">
|
||
<treatmentCitation id="F139FD13FE42FFCFFF54C858BE2A622B" author="Kryzhanovskij, O. L. & Belousov, I. A. & Kabak, I. I. & Kataev, B. M. & Makarov, K. V. & Shilenkov, V. G." box="[151,844,1380,1405]" page="107" pageId="1" pageNumber="417" year="1995">
|
||
<taxonomicName id="B798A081FE42FFCFFF54C858BE7C622A" ID-CoL="4Q47R" authority=": Kryzhanovskij et al. (1995)" authorityName="Kryzhanovskij" authorityYear="1995" box="[151,794,1380,1405]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Petrophilus">
|
||
<emphasis id="42EC0710FE42FFCFFF54C858BC73622A" box="[151,277,1380,1404]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="42EC0710FE42FFCFFEE0C858BCFE622A" box="[291,408,1380,1404]" italics="true" pageId="1" pageNumber="417">Petrophilus</emphasis>
|
||
)
|
||
<emphasis id="42EC0710FE42FFCFFE65C859BC95622A" box="[422,499,1381,1404]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
|
||
:
|
||
<bibRefCitation id="1409A6F3FE42FFCFFDC3C858BE7C622A" author="Kryzhanovskij, O. L. & Belousov, I. A. & Kabak, I. I. & Kataev, B. M. & Makarov, K. V. & Shilenkov, V. G." box="[512,794,1380,1404]" pageId="1" pageNumber="417" refId="ref6365" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V. & Shilenkov, V. G. (1995) A Checklist of the Ground-Beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia, 271 pp." type="book" year="1995">
|
||
Kryzhanovskij
|
||
<emphasis id="42EC0710FE42FFCFFD58C859BFA8622A" box="[667,718,1380,1404]" italics="true" pageId="1" pageNumber="417">et al.</emphasis>
|
||
(1995)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: 107
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="F139FD13FE42FFCFFC95C859B953622A" author="Bousquet, Y." box="[854,1077,1380,1405]" page="505" pageId="1" pageNumber="417" year="2003">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFC95C859B965622B" author="Bousquet, Y." box="[854,1027,1381,1405]" pageId="1" pageNumber="417" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet (2003)</bibRefCitation>
|
||
: 505
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="F139FD13FE42FFCFFBFDC859B87B622B" author="Bousquet, Y." box="[1086,1309,1381,1405]" page="731" pageId="1" pageNumber="417" year="2017">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFBFDC859B98A622B" author="Bousquet, Y." box="[1086,1260,1381,1405]" pageId="1" pageNumber="417" pagination="1810" refId="ref5666" refString="Bousquet, Y. (2017) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, Boston, pp. 675 - 755." type="book chapter" year="2017">Bousquet (2017)</bibRefCitation>
|
||
: 731
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="F139FD13FE42FFCFFAE5C858BC0B62CD" author="Yoshimatsu, S. & Ito, N. & Nakatani, Y. & Yoshitake, H." page="96" pageId="1" pageNumber="417" year="2018">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFAE5C858BC2E62CD" author="Yoshimatsu, S. & Ito, N. & Nakatani, Y. & Yoshitake, H." pageId="1" pageNumber="417" pagination="15 - 191" refId="ref7233" refString="Yoshimatsu, S., Ito, N., Nakatani, Y. & Yoshitake, H. (2018) A list of ground beetles (Insecta: Coleoptera: Caraboidea) in Dr. Kazuo Tanaka Collection preserved in the Insect Museum of Institute for Agro-Environmental Sciences, NARO. Bulletin of the NARO, Agro-Environmental Sciences, 39, 15 - 191. [in Japanese with English summary]" type="journal article" year="2018">
|
||
Yoshimatsu
|
||
<emphasis id="42EC0710FE42FFCFFF04C8B8BD9C62CD" box="[199,250,1411,1435]" italics="true" pageId="1" pageNumber="417">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
: 96
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7027DB02FE42FFCFFF54C89EBE17628F" blockId="1.[151,1437,1227,1497]" pageId="1" pageNumber="417">
|
||
<treatmentCitationGroup id="5088FC2CFE42FFCFFF54C89EBE17628F" pageId="1" pageNumber="417">
|
||
<treatmentCitation id="F139FD13FE42FFCFFF54C89EBEE062EC" author="Kryzhanovskij, O. L. & Belousov, I. A. & Kabak, I. I. & Kataev, B. M. & Makarov, K. V. & Shilenkov, V. G." box="[151,902,1442,1466]" page="107" pageId="1" pageNumber="417" year="1995">
|
||
<taxonomicName id="B798A081FE42FFCFFF54C89EBE3362EC" authority=": Kryzhanovskij et al. (1995)" authorityName="Kryzhanovskij" authorityYear="1995" box="[151,853,1442,1466]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="sachalinensis" subGenus="Petrophilus">
|
||
<emphasis id="42EC0710FE42FFCFFF54C89EBC7362EC" box="[151,277,1442,1466]" italics="true" pageId="1" pageNumber="417">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="42EC0710FE42FFCFFEE0C89EBCFE62EC" box="[291,408,1442,1466]" italics="true" pageId="1" pageNumber="417">Petrophilus</emphasis>
|
||
)
|
||
<emphasis id="42EC0710FE42FFCFFE65C89EBF4962EC" box="[422,559,1442,1466]" italics="true" pageId="1" pageNumber="417">sachalinensis</emphasis>
|
||
:
|
||
<bibRefCitation id="1409A6F3FE42FFCFFDF8C89EBE3362EC" author="Kryzhanovskij, O. L. & Belousov, I. A. & Kabak, I. I. & Kataev, B. M. & Makarov, K. V. & Shilenkov, V. G." box="[571,853,1442,1466]" pageId="1" pageNumber="417" refId="ref6365" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V. & Shilenkov, V. G. (1995) A Checklist of the Ground-Beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia, 271 pp." type="book" year="1995">
|
||
Kryzhanovskij
|
||
<emphasis id="42EC0710FE42FFCFFD15C89FBE6E62EC" box="[726,776,1442,1466]" italics="true" pageId="1" pageNumber="417">et al.</emphasis>
|
||
(1995)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
: 107
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="F139FD13FE42FFCFFC53C89EB9D162EC" author="Berlov, E. & Berlov, O." box="[912,1207,1442,1466]" page="38" pageId="1" pageNumber="417" year="1997">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFC53C89EB9F462EC" author="Berlov, E. & Berlov, O." box="[912,1170,1442,1466]" pageId="1" pageNumber="417" pagination="36 - 46" refId="ref5521" refString="Berlov, E. & Berlov, O. (1997 d) A key to species of the genus Pterostichus (Coleoptera, Carabidae) from Sakhalin Island. Vestnik of the Irkutsk State Agricultural Academy, 5, 36 - 46." type="journal article" year="1997">Berlov & Berlov (1997d)</bibRefCitation>
|
||
: 38
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="F139FD13FE42FFCFFB02C89EB8FA62EC" author="Bousquet, Y." box="[1217,1436,1442,1466]" page="505" pageId="1" pageNumber="417" year="2003">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFB02C89EB80862EC" author="Bousquet, Y." box="[1217,1390,1442,1466]" pageId="1" pageNumber="417" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet (2003)</bibRefCitation>
|
||
: 505
|
||
</treatmentCitation>
|
||
(synonym of
|
||
<taxonomicName id="B798A081FE42FFCFFE8DC8FDBCFD628E" ID-CoL="4Q47R" authority=": Morawitz (1862)" authorityName=": Morawitz" authorityYear="1862" box="[334,411,1473,1496]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Steropus">
|
||
<emphasis id="42EC0710FE42FFCFFE8DC8FDBCFD628E" box="[334,411,1473,1496]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<treatmentCitation id="F139FD13FE42FFCFFE72C8FDBFE8628F" author="Bousquet, Y." box="[433,654,1473,1497]" page="731" pageId="1" pageNumber="417" year="2017">
|
||
<bibRefCitation id="1409A6F3FE42FFCFFE72C8FDBF39628F" author="Bousquet, Y." box="[433,607,1473,1497]" pageId="1" pageNumber="417" pagination="1810" refId="ref5666" refString="Bousquet, Y. (2017) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, Boston, pp. 675 - 755." type="book chapter" year="2017">Bousquet (2017)</bibRefCitation>
|
||
: 731
|
||
</treatmentCitation>
|
||
(synonym of
|
||
<taxonomicName id="B798A081FE42FFCFFCDFC8FDBE0F628E" ID-CoL="4Q47R" authority=": Morawitz (1862)" authorityName=": Morawitz" authorityYear="1862" box="[796,873,1473,1496]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius" subGenus="Steropus">
|
||
<emphasis id="42EC0710FE42FFCFFCDFC8FDBE0F628E" box="[796,873,1473,1496]" italics="true" pageId="1" pageNumber="417">eximius</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="38828889FE42FFCCFF54CB3AB959659B" lastPageId="2" lastPageNumber="418" pageId="1" pageNumber="417" type="materials_examined">
|
||
<paragraph id="7027DB02FE42FFCFFF54CB3ABEDB61DB" blockId="1.[151,1437,1542,2037]" pageId="1" pageNumber="417">
|
||
<emphasis id="42EC0710FE42FFCFFF54CB3ABCED6176" bold="true" box="[151,395,1542,1568]" pageId="1" pageNumber="417">Materials examined.</emphasis>
|
||
<taxonomicName id="B798A081FE42FFCFFE52CB3BBFB36176" authority="eximius" box="[401,725,1543,1569]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="eximius" subSpecies="eximius">
|
||
<emphasis id="42EC0710FE42FFCFFE52CB3BBFB36176" box="[401,725,1543,1569]" italics="true" pageId="1" pageNumber="417">Pterostichus eximius eximius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<materialsCitation id="C0F0D15FFE42FFCFFD21CB3BBC7C6113" ID-GBIF-Occurrence="3019437303" collectingDate="1985-06-05" collectingDateMax="1991-07" collectingDateMin="1985-06-05" collectorName="Sokolov" county="Balyaga" location="Chita Region" municipality="Transbaikalia" pageId="1" pageNumber="417" specimenCount="34" specimenCount-female="12" specimenCount-male="22">
|
||
<specimenCount id="669E108BFE42FFCFFD21CB3BBE736177" box="[738,789,1543,1569]" pageId="1" pageNumber="417" type="male">11♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFCD6CB3BBE536177" box="[789,821,1543,1569]" pageId="1" pageNumber="417" type="female">6♀</specimenCount>
|
||
(
|
||
<specimenCount id="669E108BFE42FFCFFC86CB3ABE0D6177" box="[837,875,1542,1569]" pageId="1" pageNumber="417" type="male">5♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFCA8CB3ABEEA6177" box="[875,908,1542,1569]" pageId="1" pageNumber="417" type="female">2♀</specimenCount>
|
||
,
|
||
<collectingCounty id="9946A38EFE42FFCFFC5BCB3BBE916177" box="[920,1015,1543,1569]" pageId="1" pageNumber="417">Balyaga</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="90434178FE42FFCFFBC3CB3BB9FC6177" box="[1024,1178,1543,1569]" pageId="1" pageNumber="417">Transbaikalia</collectingMunicipality>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFB66CB3BB8636177" box="[1189,1285,1543,1569]" pageId="1" pageNumber="417" value="1991-07">
|
||
<collectingDate id="1462042AFE42FFCFFB66CB3BB8636177" box="[1189,1285,1543,1569]" pageId="1" pageNumber="417" value="1991-07">vii.1991</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DD6DBED4FE42FFCFFACCCB3BB80A6177" box="[1295,1388,1543,1569]" pageId="1" pageNumber="417">Sokolov</collectorName>
|
||
leg. in OB coll.;
|
||
</materialsCitation>
|
||
<materialsCitation id="C0F0D15FFE42FFCFFEE3CB17BDA9613E" ID-GBIF-Occurrence="3019437301" collectingDate="1985-06-05" collectingDateMax="1991-07" collectingDateMin="1985-06-05" collectorName="E. Berlov" county="Balyaga" location="Chita Region" municipality="Transbaikalia" pageId="1" pageNumber="417" specimenCount="34" specimenCount-female="12" specimenCount-male="22">
|
||
<specimenCount id="669E108BFE42FFCFFEE3CB17BC216113" box="[288,327,1579,1605]" pageId="1" pageNumber="417" type="male">6♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFE84CB17BC0E6113" box="[327,360,1579,1605]" pageId="1" pageNumber="417" type="female">4♀</specimenCount>
|
||
,
|
||
<collectorName id="DD6DBED4FE42FFCFFEB0CB17BCDF6113" box="[371,441,1579,1605]" pageId="1" pageNumber="417">Chara</collectorName>
|
||
,
|
||
<location id="75478DD9FE42FFCFFE00CB17BF316113" LSID="urn:lsid:plazi:treatment:F8316A14FE42FFCCFF54C95EB959659B:75478DD9FE42FFCFFE00CB17BF316113" box="[451,599,1579,1605]" county="Balyaga" municipality="Transbaikalia" name="Chita Region" pageId="1" pageNumber="417">Chita Region</location>
|
||
/=
|
||
<collectorName id="DD6DBED4FE42FFCFFDACCB17BE6C6113" box="[623,778,1579,1605]" pageId="1" pageNumber="417">Transbaikalia</collectorName>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFCD7CB16BE196113" box="[788,895,1578,1605]" pageId="1" pageNumber="417" value="1985-06-05">
|
||
<collectingDate id="1462042AFE42FFCFFCD7CB16BE196113" box="[788,895,1578,1605]" pageId="1" pageNumber="417" value="1985-06-05">5.vi.1985</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DD6DBED4FE42FFCFFC48CB17BE936113" box="[907,1013,1579,1605]" pageId="1" pageNumber="417">E. Berlov</collectorName>
|
||
leg. in OB coll.);
|
||
<taxonomicName id="B798A081FE42FFCFFB7ECB17BDA9613E" authorityName="Sasakawa & Berlov & Okuzaki" authorityYear="2020" baseAuthorityName=": Kirschenhofer" baseAuthorityYear="1985" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="eximius" subSpecies="sachalinensis">
|
||
<emphasis id="42EC0710FE42FFCFFB7ECB17BDA9613E" italics="true" pageId="1" pageNumber="417">P. eximius sachalinensis</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="C0F0D15FFE42FFCFFF1ECB73BF2061DB" ID-GBIF-Occurrence="3019437304" collectingDate="1990-06-14" collectingDateMax="2005-07-30" collectingDateMin="1990-06-14" collectionCode="T" collectorName="E. Berlov & Sota" location="Mt. Chekhov" pageId="1" pageNumber="417" specimenCount="28" specimenCount-female="8" specimenCount-male="20" stateProvince="Sakhalin">
|
||
<specimenCount id="669E108BFE42FFCFFF1ECB73BC74613F" box="[221,274,1615,1641]" pageId="1" pageNumber="417" type="male">10♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFED1CB73BC55613F" box="[274,307,1615,1641]" pageId="1" pageNumber="417" type="female">4♀</specimenCount>
|
||
,
|
||
<location id="75478DD9FE42FFCFFEFDCB73BCB5613F" LSID="urn:lsid:plazi:treatment:F8316A14FE42FFCCFF54C95EB959659B:75478DD9FE42FFCFFEFDCB73BCB5613F" box="[318,467,1615,1641]" name="Mt. Chekhov" pageId="1" pageNumber="417" stateProvince="Sakhalin">Mt. Chekhov</location>
|
||
,
|
||
<collectingRegion id="B25C15E0FE42FFCFFE1ECB73BF59613F" box="[477,575,1615,1641]" country="Russia" name="Sakhalin" pageId="1" pageNumber="417">Sakhalin</collectingRegion>
|
||
(
|
||
<specimenCount id="669E108BFE42FFCFFD93CB73BFE1613F" box="[592,647,1615,1641]" pageId="1" pageNumber="417" type="male">10♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFD44CB73BFC3613F" box="[647,677,1615,1641]" pageId="1" pageNumber="417" type="female">3♀</specimenCount>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFD72CB73BE4D613F" box="[689,811,1615,1641]" pageId="1" pageNumber="417" value="1990-06-14">
|
||
<collectingDate id="1462042AFE42FFCFFD72CB73BE4D613F" box="[689,811,1615,1641]" pageId="1" pageNumber="417" value="1990-06-14">14.vi.1990</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="DD6DBED4FE42FFCFFCF5CB73BEC7613F" box="[822,929,1615,1641]" pageId="1" pageNumber="417">E. Berlov</collectorName>
|
||
leg. in OB coll.;
|
||
<specimenCount id="669E108BFE42FFCFFBA2CB73B9E2613F" box="[1121,1156,1615,1641]" pageId="1" pageNumber="417" type="female">1♀</specimenCount>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFB52CB72B875613F" box="[1169,1299,1614,1641]" pageId="1" pageNumber="417" value="2005-07-30">
|
||
<collectingDate id="1462042AFE42FFCFFB52CB72B875613F" box="[1169,1299,1614,1641]" pageId="1" pageNumber="417" value="2005-07-30">30.vii.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="168943C7FE42FFCFFADDCB73B856613F" box="[1310,1328,1615,1641]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="417">T</collectionCode>
|
||
.
|
||
<collectorName id="DD6DBED4FE42FFCFFAF8CB73B80B613F" box="[1339,1389,1615,1641]" pageId="1" pageNumber="417">Sota</collectorName>
|
||
leg. in KS coll.);
|
||
<taxonomicName id="B798A081FE42FFCFFEEBCB4FBF2061DB" authorityName="Sasakawa & Berlov & Okuzaki" authorityYear="2020" box="[296,582,1651,1677]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="eximius" subSpecies="rishiridakensis">
|
||
<emphasis id="42EC0710FE42FFCFFEEBCB4FBF2061DB" box="[296,582,1651,1677]" italics="true" pageId="1" pageNumber="417">P. eximius rishiridakensis</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<specimenCount id="669E108BFE42FFCFFD97CB4FBF1A61DB" box="[596,636,1651,1677]" pageId="1" pageNumber="417" type="male">2♂</specimenCount>
|
||
<specimenCount id="669E108BFE42FFCFFDB8CB4FBFCD61DB" box="[635,683,1651,1677]" pageId="1" pageNumber="417" type="female">10♀</specimenCount>
|
||
, see subspecific section.
|
||
</paragraph>
|
||
<paragraph id="7027DB02FE42FFCFFF04CBAAB92960F8" blockId="1.[151,1437,1542,2037]" pageId="1" pageNumber="417">
|
||
<emphasis id="42EC0710FE42FFCFFF04CBAABFC261E6" bold="true" box="[199,676,1686,1712]" pageId="1" pageNumber="417">Other consubgeneric species examined.</emphasis>
|
||
<taxonomicName id="B798A081FE42FFCFFD6DCBABB90761E4" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1844" box="[686,1121,1687,1714]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="42EC0710FE42FFCFFD6DCBABBED661E6" box="[686,944,1687,1713]" italics="true" pageId="1" pageNumber="417">Pterostichus montanus</emphasis>
|
||
(Motschulsky)
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="669E108BFE42FFCFFBACCBABB9F261E7" box="[1135,1172,1687,1713]" pageId="1" pageNumber="417" type="male">2♂</specimenCount>
|
||
, Mt. Khulugaisha near Mondy, Tunkinskie goltsy Mts. ridge,
|
||
<collectingRegion id="B25C15E0FE42FFCFFD8ACB87BFCA6183" box="[585,684,1723,1749]" country="Russia" name="Buryatiya" pageId="1" pageNumber="417">Buryatia</collectingRegion>
|
||
,
|
||
<quantity id="B76076E7FE42FFCFFD7BCB87BE766183" box="[696,784,1723,1749]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="417" unit="m" value="2300.0">2300 m</quantity>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFCDFCB87BEC16183" box="[796,935,1723,1749]" pageId="1" pageNumber="417" value="1998-07-01" valueMax="1998-07-04" valueMin="1998-07-01">1-4.vii.1998</date>
|
||
, E. Berlov leg. in OB coll.;
|
||
<taxonomicName id="B798A081FE42FFCFFB25CB87BD9061AF" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1962" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="sutschanensis">
|
||
<emphasis id="42EC0710FE42FFCFFB25CB87B8FA6183" box="[1254,1436,1723,1749]" italics="true" pageId="1" pageNumber="417">P. sutschanensis</emphasis>
|
||
Jedlička
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="669E108BFE42FFCFFEC3CBE3BC4261AF" box="[256,292,1759,1785]" pageId="1" pageNumber="417" type="male">3♂</specimenCount>
|
||
(
|
||
<specimenCount id="669E108BFE42FFCFFEF0CBE3BC3E61AF" box="[307,344,1759,1785]" pageId="1" pageNumber="417" type="male">1♂</specimenCount>
|
||
, Lazovsky Pass,
|
||
<collectingRegion id="B25C15E0FE42FFCFFDD6CBE3BF8261AF" box="[533,740,1759,1785]" country="Russia" name="Primorskiy" pageId="1" pageNumber="417">Primorsky Region</collectingRegion>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFD2DCBE2BE0C61AF" box="[750,874,1758,1785]" pageId="1" pageNumber="417" value="2005-08-04">4.viii.2005</date>
|
||
,
|
||
<collectionCode id="168943C7FE42FFCFFCB7CBE3BEE061AF" box="[884,902,1759,1785]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="417">T</collectionCode>
|
||
. Sota leg. in KS coll.;
|
||
<specimenCount id="669E108BFE42FFCFFBBFCBE3B9C461AF" box="[1148,1186,1759,1785]" pageId="1" pageNumber="417" type="male">2♂</specimenCount>
|
||
, Sichote-Alinsky state nature reserve,
|
||
<collectingRegion id="B25C15E0FE42FFCFFE81CA3FBF77604B" box="[322,529,1795,1821]" country="Russia" name="Primorskiy" pageId="1" pageNumber="417">Primorsky Region</collectingRegion>
|
||
,
|
||
<date id="0426FDC2FE42FFCFFDDFCA3FBFF0604B" box="[540,662,1795,1821]" pageId="1" pageNumber="417" value="1998-06-18">18.vi.1998</date>
|
||
, Yundukov leg. in OB coll.). The following species were investigated through
|
||
<bibRefCitation id="1409A6F3FE42FFCFFF34CA1BBF4B6017" author="Sasakawa, K. & Kubota, K." box="[247,557,1830,1857]" pageId="1" pageNumber="417" pagination="389 - 404" refId="ref6803" refString="Sasakawa, K. & Kubota, K. (2005) Cryptic species of the subgenus Morphnosoma Lutshnik (Coleoptera: Carabidae; genus Pterostichus) from Japan. Entomological Science, 8, 389 - 404. https: // doi. org / 10.1111 / j. 1479 - 8298.2005.00139. x" type="journal article" year="2005">Sasakawa & Kubota (2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1409A6F3FE42FFCFFDF8CA1BBF136014" author="Sasakawa, K. & Kubota, K." box="[571,629,1831,1858]" pageId="1" pageNumber="417" pagination="31 - 43" refId="ref6858" refString="Sasakawa, K. & Kubota, K. (2006) Phylogenetic studies of the subgenus Petrophilus Chaudoir (Coleoptera: Carabidae: Pterostichus), with description of a new species sympatric with P. thunbergi Morawitz. Zootaxa, 1357, 31 - 43." type="journal article" year="2006">2006</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="1409A6F3FE42FFCFFD74CA1BBE1B6017" author="Sun, X. & Shi, H. & Sang, W. & Axmacher, J. C." box="[695,893,1831,1857]" pageId="1" pageNumber="417" pagination="95 - 114" refId="ref7055" refString="Sun, X., Shi, H., Sang, W. & Axmacher, J. C. (2018) Two new species of Feroperis Lafer (Carabidae, Pterostichus) from China, with a key to all known Chinese species in this subgenus. ZooKeys, 799, 95 - 114. https: // doi. org / 10.3897 / zookeys. 799.28834" type="journal article" year="2018">
|
||
Sun
|
||
<emphasis id="42EC0710FE42FFCFFD2FCA1BBE406017" box="[748,806,1831,1857]" italics="true" pageId="1" pageNumber="417">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
:
|
||
<taxonomicName id="B798A081FE42FFCFFC49CA1BB9FC6017" authority="(Newman)" baseAuthorityName="Newman" box="[906,1178,1831,1857]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="coracinus">
|
||
<emphasis id="42EC0710FE42FFCFFC49CA1BB9706016" box="[906,1046,1831,1856]" italics="true" pageId="1" pageNumber="417">P. coracinus</emphasis>
|
||
(Newman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFB64CA1BB81D6014" authority="(Dejean)" baseAuthorityName="Dejean" box="[1191,1403,1831,1858]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="findeli">
|
||
<emphasis id="42EC0710FE42FFCFFB64CA1BB86B6017" box="[1191,1293,1831,1857]" italics="true" pageId="1" pageNumber="417">P. findeli</emphasis>
|
||
(Dejean)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFA4BCA1BBCCB6030" authority="(Duftschmid)" baseAuthorityName="Duftschmid" baseAuthorityYear="1812" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="42EC0710FE42FFCFFA4BCA1BBC6C6033" italics="true" pageId="1" pageNumber="417">P. foveolatus</emphasis>
|
||
(Duftschmid)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFE7ACA77BF1B6033" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1962" box="[441,637,1867,1893]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="habui">
|
||
<emphasis id="42EC0710FE42FFCFFE7ACA77BF706033" box="[441,534,1867,1893]" italics="true" pageId="1" pageNumber="417">P. habui</emphasis>
|
||
Jedlička
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFD4ACA77BEDA6030" authority="(Tschitscherine)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1893" box="[649,956,1867,1894]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="jungens">
|
||
<emphasis id="42EC0710FE42FFCFFD4ACA77BF9B6032" box="[649,765,1867,1892]" italics="true" pageId="1" pageNumber="417">P. jungens</emphasis>
|
||
(Tschitschérine)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFC0ACA77B9916033" authority="(Newman)" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[969,1271,1867,1893]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="lachrymosus">
|
||
<emphasis id="42EC0710FE42FFCFFC0ACA77B9126033" box="[969,1140,1867,1893]" italics="true" pageId="1" pageNumber="417">P. lachrymosus</emphasis>
|
||
(Newman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFAC0CA77BDB360DF" authority="Lafer" authorityName="Lafer" authorityYear="1979" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="maichensis">
|
||
<emphasis id="42EC0710FE42FFCFFAC0CA77B8FA6033" box="[1283,1436,1867,1893]" italics="true" pageId="1" pageNumber="417">P. maichensis</emphasis>
|
||
Lafer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFF22CA53BC8060DF" authority="Sun & Shi" authorityName="Sun & Shi" authorityYear="2018" box="[225,486,1903,1929]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="maryseae">
|
||
<emphasis id="42EC0710FE42FFCFFF22CA53BC0160DE" box="[225,359,1903,1928]" italics="true" pageId="1" pageNumber="417">P. maryseae</emphasis>
|
||
Sun & Shi
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFE32CA53BF8A60DC" authority="(Illiger)" baseAuthorityName="Illiger" baseAuthorityYear="1798" box="[497,748,1903,1930]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="melanarius">
|
||
<emphasis id="42EC0710FE42FFCFFE32CA53BFED60DF" box="[497,651,1903,1929]" italics="true" pageId="1" pageNumber="417">P. melanarius</emphasis>
|
||
(Illiger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFD34CA53B97460DF" authority="Sasakawa" authorityName="Sasakawa" box="[759,1042,1903,1929]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="robustistylis">
|
||
<emphasis id="42EC0710FE42FFCFFD34CA53BEFC60DF" box="[759,922,1903,1929]" italics="true" pageId="1" pageNumber="417">P. robustistylis</emphasis>
|
||
Sasakawa
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFBDDCA53B87D60DF" authority="Sun & Shi" authorityName="Sun & Shi" authorityYear="2018" box="[1054,1307,1903,1929]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="silvestris">
|
||
<emphasis id="42EC0710FE42FFCFFBDDCA53B9FB60DF" box="[1054,1181,1903,1929]" italics="true" pageId="1" pageNumber="417">P. silvestris</emphasis>
|
||
Sun & Shi
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFAE4CA53BDB060FB" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="stygicus">
|
||
<emphasis id="42EC0710FE42FFCFFAE4CA53B8FA60DE" box="[1319,1436,1903,1928]" italics="true" pageId="1" pageNumber="417">P. stygicus</emphasis>
|
||
(Say)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFF21CAAFBC8660FB" authority="Morawitz" authorityName="Morawitz" box="[226,480,1939,1965]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="thunbergi">
|
||
<emphasis id="42EC0710FE42FFCFFF21CAAFBC0F60FB" box="[226,361,1939,1965]" italics="true" pageId="1" pageNumber="417">P. thunbergi</emphasis>
|
||
Morawitz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B798A081FE42FFCFFE28CAAFBF8760FB" authority="Sasakawa" authorityName="Sasakawa" authorityYear="2006" box="[491,737,1939,1965]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="tuberifer">
|
||
<emphasis id="42EC0710FE42FFCFFE28CAAFBF0F60FB" box="[491,617,1939,1965]" italics="true" pageId="1" pageNumber="417">P. tuberifer</emphasis>
|
||
Sasakawa
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B798A081FE42FFCFFCD8CAAFB92A60F8" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1850" box="[795,1100,1939,1966]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="uralensis">
|
||
<emphasis id="42EC0710FE42FFCFFCD8CAAFBEF860FB" box="[795,926,1939,1965]" italics="true" pageId="1" pageNumber="417">P. uralensis</emphasis>
|
||
(Motschulsky)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7027DB02FE42FFCCFF04CA8BBEFC67AF" blockId="1.[151,1437,1542,2037]" lastBlockId="2.[151,1437,151,717]" lastPageId="2" lastPageNumber="418" pageId="1" pageNumber="417">
|
||
<emphasis id="42EC0710FE42FFCFFF04CA8BBC776087" bold="true" box="[199,273,1975,2001]" pageId="1" pageNumber="417">Notes.</emphasis>
|
||
<taxonomicName id="B798A081FE42FFCFFEDACA8BBF646086" box="[281,514,1975,2001]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="417" phylum="Arthropoda" rank="species" species="eximius">
|
||
<emphasis id="42EC0710FE42FFCFFEDACA8BBF646086" box="[281,514,1975,2001]" italics="true" pageId="1" pageNumber="417">Pterostichus eximius</emphasis>
|
||
</taxonomicName>
|
||
was originally described based on specimens from Transbaikalia and is currently treated as a monotypic species without any geographical variants that are distinguished taxonomically. This monotypic status is based on work by
|
||
<bibRefCitation id="1409A6F3FE41FFCCFDD2CDABBF8067E7" author="Lafer, G. S. & Nilsson, A. N. & Kholin, S. K." box="[529,742,151,177]" pageId="2" pageNumber="418" pagination="13 - 17" refId="ref6435" refString="Lafer, G. S., Nilsson, A. N. & Kholin, S. K. (1997) Additional records and new synonyms of Cicindelidae and Carabidae (Coleoptera) from the lsland of Sakhalin in the Russian Far East. Entomologica Fennica, 8, 13 - 17. https: // doi. org / 10.33338 / ef. 83916" type="journal article" year="1997">
|
||
Lafer
|
||
<emphasis id="42EC0710FE41FFCCFD95CDABBFF667E7" box="[598,656,151,177]" italics="true" pageId="2" pageNumber="418">et al.</emphasis>
|
||
(1997)
|
||
</bibRefCitation>
|
||
, who examined
|
||
<typeStatus id="AF2365A0FE41FFCCFC61CDABBEB467E7" box="[930,978,151,177]" pageId="2" pageNumber="418">type</typeStatus>
|
||
specimens and the original descriptions of some taxa, and synonymized them with
|
||
<taxonomicName id="B798A081FE41FFCCFDBFCD87BF886782" box="[636,750,187,212]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="eximius">
|
||
<emphasis id="42EC0710FE41FFCCFDBFCD87BF886782" box="[636,750,187,212]" italics="true" pageId="2" pageNumber="418">P. eximius</emphasis>
|
||
</taxonomicName>
|
||
. However, they did not examine the endophallus of the male genitalia, a character that is important in the taxonomy of
|
||
<taxonomicName id="B798A081FE41FFCCFCE2CDE3BEF067AF" authorityName="Latreille" authorityYear="1802" box="[801,918,223,249]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7027DB02FE41FFCCFF04CC3FB959659B" blockId="2.[151,1437,151,717]" pageId="2" pageNumber="418">
|
||
Our comparative study of endophallic and aedeagal structures revealed that populations from the Rishiri-tô and
|
||
<collectingRegion id="B25C15E0FE41FFCCFF04CC1BBC4F6617" box="[199,297,295,321]" country="Russia" name="Sakhalin" pageId="2" pageNumber="418">Sakhalin</collectingRegion>
|
||
islands are both distinguished from the nominotypical population from Transbaikalia. With regard to the endophalli, two morphological differences were found among specimens from the three localities. The first is sub-lobe rv-2 of the right ventral lobe, which is rudimentary in Transbaikalian specimens but distinctly developed in Rishiri-tô and
|
||
<collectingRegion id="B25C15E0FE41FFCCFE9BCCAFBCDC66FB" box="[344,442,403,429]" country="Russia" name="Sakhalin" pageId="2" pageNumber="418">Sakhalin</collectingRegion>
|
||
specimens (
|
||
<figureCitation id="E8A3C787FE41FFCCFD85CCAFBFEF66FB" box="[582,649,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1823,1847]" captionTargetBox="[153,1432,741,1799]" captionTargetId="figure-288@2.[151,1436,741,1799]" captionTargetPageId="2" captionText="FIGURE 1. Differences in male genital structures (top, left posterolateral view; bottom, right ventrolateral view) among subspecies of Pterostichus eximius. Red, green, and blue arrows denote sublobe rv-2 on the endophallus, a swelling adjoining lobe ll on the endophallus, and the dorsal contour at the aedeagal subapex, respectively. Phylogenetic relationships inferred from these characters are shown." figureDoi="http://doi.org/10.5281/zenodo.4401757" httpUri="https://zenodo.org/record/4401757/files/figure.png" pageId="2" pageNumber="418">Fig. 1</figureCitation>
|
||
, red arrows). The second is a swelling adjoining lobe ll (
|
||
<figureCitation id="E8A3C787FE41FFCCFACFCCAFB82966FB" box="[1292,1359,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1823,1847]" captionTargetBox="[153,1432,741,1799]" captionTargetId="figure-288@2.[151,1436,741,1799]" captionTargetPageId="2" captionText="FIGURE 1. Differences in male genital structures (top, left posterolateral view; bottom, right ventrolateral view) among subspecies of Pterostichus eximius. Red, green, and blue arrows denote sublobe rv-2 on the endophallus, a swelling adjoining lobe ll on the endophallus, and the dorsal contour at the aedeagal subapex, respectively. Phylogenetic relationships inferred from these characters are shown." figureDoi="http://doi.org/10.5281/zenodo.4401757" httpUri="https://zenodo.org/record/4401757/files/figure.png" pageId="2" pageNumber="418">Fig. 1</figureCitation>
|
||
, green arrows), which is larger in
|
||
<collectingRegion id="B25C15E0FE41FFCCFE05CC8BBF4E6687" box="[454,552,439,465]" country="Russia" name="Sakhalin" pageId="2" pageNumber="418">Sakhalin</collectingRegion>
|
||
specimens than in Transbaikalian and Rishiri-tô specimens. With regard to the aedeagus, differences were found in the shape of the subapical part among specimens. The dorsal contour of the lateral sides is straight in Rishiri-tô specimens, while in the Transbaikalian and
|
||
<collectingRegion id="B25C15E0FE41FFCCFC2CCCC3B937654F" box="[1007,1105,511,537]" country="Russia" name="Sakhalin" pageId="2" pageNumber="418">Sakhalin</collectingRegion>
|
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specimens the dorsal contour is bent at an obtuse angle in the subapical part, and protrudes dorsally (
|
||
<figureCitation id="E8A3C787FE41FFCCFC74CF1FBE9C656B" box="[951,1018,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1823,1847]" captionTargetBox="[153,1432,741,1799]" captionTargetId="figure-288@2.[151,1436,741,1799]" captionTargetPageId="2" captionText="FIGURE 1. Differences in male genital structures (top, left posterolateral view; bottom, right ventrolateral view) among subspecies of Pterostichus eximius. Red, green, and blue arrows denote sublobe rv-2 on the endophallus, a swelling adjoining lobe ll on the endophallus, and the dorsal contour at the aedeagal subapex, respectively. Phylogenetic relationships inferred from these characters are shown." figureDoi="http://doi.org/10.5281/zenodo.4401757" httpUri="https://zenodo.org/record/4401757/files/figure.png" pageId="2" pageNumber="418">Fig. 1</figureCitation>
|
||
, blue arrows). Thus, specimens from the three localities can be distinguished according to combinations of these endophallic and aedeagal characters. Based on this result, the three populations should now be regarded as separate subspecies. In this taxonomic system, the
|
||
<collectingRegion id="B25C15E0FE41FFCCFF01CFB3BC4265FF" box="[194,292,655,681]" country="Russia" name="Sakhalin" pageId="2" pageNumber="418">Sakhalin</collectingRegion>
|
||
population is reinstated as a subspecies:
|
||
<taxonomicName id="B798A081FE41FFCCFD3DCFB3B97765FF" authorityName="Sasakawa & Berlov & Okuzaki" authorityYear="2020" baseAuthorityName=": Kirschenhofer" baseAuthorityYear="1985" box="[766,1041,655,681]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="subSpecies" species="eximius" status="stat. nov." subSpecies="sachalinensis">
|
||
<emphasis id="42EC0710FE41FFCCFD3DCFB3B97765FF" box="[766,1041,655,681]" italics="true" pageId="2" pageNumber="418">P. eximius sachalinensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="42EC0710FE41FFCCFBD9CFB3B9E165FF" bold="true" box="[1050,1159,655,681]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="59DFBA6BFE41FFCCFBD9CFB3B9E165FF" box="[1050,1159,655,681]" pageId="2" pageNumber="418">stat. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
No scientific name was available for the Rishiri-tô population, so it is described as a new subspecies below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |