597 lines
70 KiB
XML
597 lines
70 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.231.100894" ID-Pensoft-Pub="1314-2003-231-1" ID-Pensoft-UUID="0D762CF99C605B04AF072C867EEB1488" ModsDocID="1314-2003-231-1" checkinTime="1693383686345" checkinUser="pensoft" docAuthor="Knapp, Sandra, Saerkinen, Tiina & Barboza, Gloria E." docDate="2023" docId="F7DA4326A3C2B2C6A21F7AD35A8BA5A4" docLanguage="en" docName="PhytoKeys 231: 1-342" docOrigin="PhytoKeys 231" docPubDate="2023-08-29" docSource="http://dx.doi.org/10.3897/phytokeys.231.100894" docTitle="Solanum cochabambense Bitter, Repert. Spec. Nov. Regni Veg. 10: 553. 1912." docType="treatment" docVersion="1" id="0D762CF99C605B04AF072C867EEB1488" lastPageNumber="1" masterDocId="0D762CF99C605B04AF072C867EEB1488" masterDocTitle="A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)" masterLastPageNumber="342" masterPageNumber="1" pageNumber="1" updateTime="1693383686345" updateUser="pensoft">
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<mods:title id="81C9993F1983B88EF405C98AC22E6616">A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)</mods:title>
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<mods:namePart id="0983ECF97D8D8EDA84D67E1B9F8B4244">Knapp, Sandra</mods:namePart>
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<mods:affiliation id="7173E29097B99332BFA7649282D18030">Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:nameIdentifier id="AFBADA4B0A96DE3CEB81770C04366732" type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="CDE1259B0ABD87709E996C26DA6E8B73">Saerkinen, Tiina</mods:namePart>
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<mods:nameIdentifier id="A9849BAC7EE12E15F0C8D05085C88120" type="ORCID">https://orcid.org/0000-0002-6956-3093</mods:nameIdentifier>
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<mods:affiliation id="BD18FE545EC6B7454F724C31B79A7B8D">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, UK</mods:affiliation>
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<mods:namePart id="AD88671CCFEAD2CCB35966BDEB207E54">Barboza, Gloria E.</mods:namePart>
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<mods:nameIdentifier id="51A62C8620FC2FBDA218851E020EB721" type="ORCID">https://orcid.org/0000-0003-1085-036X</mods:nameIdentifier>
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<mods:affiliation id="8F33F1DF0ACE8587E50D45EA9DD420EB">Instituto Multidisciplinario de Biologia Vegetal (CONICET-Universidad Nacional de Cordoba), Casilla de Correo 495, 5000 Cordoba, Argentina</mods:affiliation>
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<mods:title id="222FA63290B1424DA16BEC55D5FB0716">PhytoKeys</mods:title>
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<mods:date id="BDB59113B6A979CDC177E3A97A46FFF8">2023</mods:date>
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<treatment id="F7DA4326A3C2B2C6A21F7AD35A8BA5A4" LSID="urn:lsid:plazi:treatment:F7DA4326A3C2B2C6A21F7AD35A8BA5A4" httpUri="http://treatment.plazi.org/id/F7DA4326A3C2B2C6A21F7AD35A8BA5A4" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="348FB976C8A1FBFA9AF3B38B282227D1" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="72CC0069F95A4375139EE15DBCDA30AB" pageId="0" pageNumber="1">
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12.
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<taxonomicName id="A63F6FCFB206ED3128E7497F047EB974" ID-CoL="6YTF4" ID-ENA="1426228" authority="Bitter, Repert. Spec. Nov. Regni Veg. 10: 553. 1912." authorityName="Bitter, Repert. Spec. Nov. Regni Veg. 10: 553. 1912." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">Solanum cochabambense Bitter, Repert. Spec. Nov. Regni Veg. 10: 553. 1912.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="588B451FE34EC105656B8F2A792B54AD" pageId="0" pageNumber="1" type="description">
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<paragraph id="BCDD8BCF07D4493E396AE1AD6533D940" pageId="0" pageNumber="1">
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<figureCitation id="A4A744D85307F9F77FC3445314889919" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Representative flowers of South American morelloids A campanulate purple corollas in forked internodal inflorescences in S. fiebrigii B pentagonal white corollas in S. annuum C broadly stellate purple corollas with long-exerted styles in S. pentlandii D broadly stellate white corollas with slightly exerted globose stigmas in S. radicans E deeply stellate pale yellow corollas in S. huayavillense F deeply stellate purple corollas with dark central eye colouration in S. salicifolium G bumblebee visiting and buzzing the anther cone in S. cochabambense H variation in corolla shape, size and colouration within a single individual of S. cochabambense (A Barboza et al. 3548 B Barboza et al. 3495 C Knapp et al. 10248 D Knapp et al. 10277 E Barboza et al. 3531 F Chiarini et al. 819 G Saerkinen et al. 4036 H Knapp et al. 10669). Photos by S. Knapp and T. Saerkinen." figureDoi="10.3897/phytokeys.231.100894.figure3" httpUri="https://binary.pensoft.net/fig/897464" pageId="0" pageNumber="1">Figs 3G, H</figureCitation>
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<figureCitation id="692349BBC4750CE9F0BEBAC6568EB638" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Representative fruits, seeds and stone cells of South American morelloids A bilobed fruits in S. tripartitum with fully mature red berries at the base of the inflorescence and maturing green, yellow, and orange fruits more distally B orange-red fleshy berries of S. corymbosum in highly branched inflorescences C fully mature marbled fruits in S. physalifolium D immature ellipsoid fruits of S. antisuyo E fully mature fruits of S. antisuyo F immature green fruits amongst fully mature purple-black fruits in S. cochabambense G fully mature fruits of S. polytrichostylum H stone cells (also known as sclerotic granules or brachysclerids) found in the fruits of most species of the Morelloid clade (left side of photo) next to the teardrop shaped seeds (right side of photo; S. umalilaense Manoko) I stone cells visible in dried fruits of herbarium specimens (S. triflorum). (A Barboza 3563 B Saerkinen et al. 4604 B C Knapp et al. 10334 D Gonzales 10256 E Gonzales 10256 F Knapp et al. 10363 G Saerkinen et al. 5277 H Nijmegen accession A 24750133 I Podlech 8624 BM 000848286). Photos by P. Gonzales, S. Knapp, and T. Saerkinen." figureDoi="10.3897/phytokeys.231.100894.figure4" httpUri="https://binary.pensoft.net/fig/897465" pageId="0" pageNumber="1">, 4F</figureCitation>
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<figureCitation id="FD868018F2149511E60FB6B9E76725DD" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Solanum cochabambense A flowering branch B detail of adaxial leaf surface C detail of abaxial leaf surface D flower bud E dissected flower F fruiting branch G maturing fruit (A-D Knapp et al. 10341; E Cardenas 5577 F, G Knapp et al. 10339). Illustration by R. Wise." figureDoi="10.3897/phytokeys.231.100894.figure38" httpUri="https://binary.pensoft.net/fig/897499" pageId="0" pageNumber="1">, 38</figureCitation>
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<figureCitation id="4B80FCF3EA44C9C62AD7FA46DC301E61" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Solanum cochabambense A habit B inflorescence with purple corollas C infructescence with white corollas D mature fruits (A, C Knapp et al. 10363 B Knapp 10287 D Barboza et al. 3500). Photos by S. Knapp." figureDoi="10.3897/phytokeys.231.100894.figure39" httpUri="https://binary.pensoft.net/fig/897500" pageId="0" pageNumber="1">, 39</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="430E7ABC6E05AB2537F446B8480DE960" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="5B04899098D23FD7AA359F72B2D9D403" pageId="0" pageNumber="1">
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<taxonomicName id="E2230E2744537257CA00088E28AC8B4B" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum extuspellitum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="extuspellitum">Solanum extuspellitum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 10: 555. 1912. Type. Bolivia. Tarija, 2,300 m, 30 Dec 1903,
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<emphasis id="A5973F6DB1A61AF7E418D58A876FAC64" italics="true" pageId="0" pageNumber="1">K. Fiebrig 2439</emphasis>
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(holotype: B, destroyed [F neg. 2711]; lectotype, designated here: F [V0361919F, acc. # 621247]).
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</paragraph>
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<paragraph id="04425BBC5EE6CD52B11419D18896BFA3" pageId="0" pageNumber="1">
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<taxonomicName id="37C01C43166D663C0119F7579CEDD4F3" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum extuspellitum subsp. subcoeruleum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="extuspellitum" subSpecies="subcoeruleum">Solanum extuspellitum Bitter subsp. subcoeruleum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 10: 556. 1912. Type. Bolivia. Tarija, 2,300 m, 30 Dec 1903,
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<emphasis id="ADFD83C16CC02925F67560F08B080486" italics="true" pageId="0" pageNumber="1">K. Fiebrig 2439</emphasis>
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(holotype: B, destroyed [F neg. 2711]; lectotype, designated here: F [v0361919F, acc. # 621247]).
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</paragraph>
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<paragraph id="6EABAB79AC8D8AC9D99CC91B8DB8A8E1" pageId="0" pageNumber="1">
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<taxonomicName id="D2A1D152D6CFA7E602A8199BE3915CE5" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum lorentzii subsp. var. var. tucumanicum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="lorentzii" subSpecies="var." variety="tucumanicum">Solanum lorentzii Bitter var. tucumanicum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 10: 556. 1912. Type. Argentina.
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<normalizedToken id="0A2041C40F356B2F5BB786AF5BC80F40" originalValue="Tucumán">Tucuman</normalizedToken>
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: sin. loc.,
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<emphasis id="7E18F5E29687CFC822CE2F4B3DBF2904" italics="true" pageId="0" pageNumber="1">P.G. Lorentz & G. Hieronymus 1155</emphasis>
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(holotype: B, destroyed; lectotype, designated by Barboza et al. 2103, pg. 236: CORD [CORD00004238]; isotypes: CORD [CORD00004239, CORD00004240], F [v0073320F, acc. # 50929], K [K000585687], SI [SI003323]).
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</paragraph>
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<paragraph id="A906AC33A71D016071814B9615F2F539" pageId="0" pageNumber="1">
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<taxonomicName id="39571323D18B5F55127FB6AED702D435" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="decachondrum">Solanum decachondrum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 11: 228. 1912. Type. Bolivia. Cochabamba: Cercado, May 1909,
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<emphasis id="E12AC5E80A6B87E2FC85F78AA851CE0C" italics="true" pageId="0" pageNumber="1">O. Buchtien 2411</emphasis>
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(lectotype, designated here: US [00027539, acc. # 700102]; isolectotypes: US [01014170, acc. # 1175973]).
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</paragraph>
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<paragraph id="29C5406166CEF5C6153245A11BFEC0FE" pageId="0" pageNumber="1">
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<taxonomicName id="EDA4592FF57EAC36EC6CDB73F26DE681" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum subsp. var. var. latiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" subSpecies="var." variety="latiusculum">Solanum decachondrum Bitter var. latiusculum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 11: 229. 1912. Type. Bolivia. Cochabamba: Cercado, May 1909,
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<emphasis id="43CE18854E3A616AB0018479B1599C07" italics="true" pageId="0" pageNumber="1">O. Buchtien 2412</emphasis>
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(lectotype, designated here: US [00027538, acc. # 1177823]; isolectotypes: GOET [GOET009219], NY [00139124]).
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</paragraph>
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<paragraph id="4CF047F2E88A92C9A3EFC7144313BF4E" pageId="0" pageNumber="1">
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<taxonomicName id="D2BE30F59ABBF060B402D0A1A9173870" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum subsp. var. var. longiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" subSpecies="var." variety="longiusculum">Solanum decachondrum Bitter var. longiusculum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 11: 229. 1912. Type. Bolivia. Cochabamba: Cercado, May 1909,
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<emphasis id="1B813AE56ACFA8A997A3A156C4F0C015" italics="true" pageId="0" pageNumber="1">O. Buchtien 2411</emphasis>
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(lectotype, designated here: US [01014170, acc. # 1175973]; isolectotype: US [00027539, acc. # 700102]).
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</paragraph>
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<paragraph id="5A36B7E79DC5EBD085D23DE391EC010F" pageId="0" pageNumber="1">
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<taxonomicName id="57ACFB8F06F4CA12D8B1CAEE543B513E" authorityName="Bitter" authorityYear="1916" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum probolospermum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="probolospermum">Solanum probolospermum</taxonomicName>
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Bitter, Bot. Jahrb. Syst. 54, Beibl. 119: 10. 1916. Type. Peru.
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<normalizedToken id="597C31EE86F78AE749351BEB64A0444C" originalValue="Huánuco">Huanuco</normalizedToken>
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: Valle del
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<normalizedToken id="0765595AC78F3CF71B29232FCD30BEB8" originalValue="Río">Rio</normalizedToken>
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Pozuzo encima de Saria, 22 Jul 1913,
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<emphasis id="BE367E8C9C1846F16DC8DA29883D11A2" italics="true" pageId="0" pageNumber="1">A. Weberbauer 6789</emphasis>
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(no herbaria cited; lectotype, designated here: MOL[MOL00005139]; isolectotypes: B, destroyed [F neg. 2682], F [v0043286F, acc. # 647965], GH [01011893], MOL [MOL00005138], US [00027756, acc. # 1444969]).
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</paragraph>
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<paragraph id="68CD511733437022060B15080CD90F6D" pageId="0" pageNumber="1">
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<taxonomicName id="2A619AAFCE3A013129744C87074F4551" authorityName="C.V.Morton" authorityYear="1976" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum lorentzii subsp. var. var. montigenum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="lorentzii" subSpecies="var." variety="montigenum">Solanum lorentzii Bitter var. montigenum</taxonomicName>
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C.V.Morton, Revis. Argentine Sp. Solanum 136. 1976. Type. Argentina.
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<normalizedToken id="C95EAF6EF21D728652AF231C8D29FD56" originalValue="Tucumán">Tucuman</normalizedToken>
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: Dpto. Chicligasta: Estancia Santa Rosa, 8 Jan 1927,
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<emphasis id="2C0922F53B8647920CFD555A30D3F893" italics="true" pageId="0" pageNumber="1">S. Venturi 4760</emphasis>
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(holotype: US [03271889, acc. # 1548937]; isotypes: F [v0073318F, acc. # 695929; v0073319F, acc. # 637505], LP [LP010202, acc. # 010393], MO [MO-2127157, acc. # 960405] S [acc. # R-3117], SI [003322]).
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</paragraph>
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<paragraph id="8160163837A22B10C2AF43580AFEDEED" pageId="0" pageNumber="1">
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<taxonomicName id="CDC972FA663739C86F3C6C0099173423" authorityName="Cabrera" authorityYear="1983" baseAuthorityName="C.V.Morton" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum montigenum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="montigenum">Solanum montigenum</taxonomicName>
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(C.V.Morton) Cabrera, Fl. Prov. Jujuy 8: 435. 1983. Type. Based on
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<taxonomicName id="B3E428A8D96342EE1358187497E8F2F6" authorityName="C.V.Morton" authorityYear="1976" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum lorentzii subsp. var. var. montigenum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="lorentzii" subSpecies="var." variety="montigenum">Solanum lorentzii Bitter var. montigenum</taxonomicName>
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C.V.Morton.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5E38B8E9732E081665A8F1A7E9B5E47" pageId="0" pageNumber="1" type="materials_examined">
|
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<paragraph id="731D33A68906F89392DA69990E5C0F69" pageId="0" pageNumber="1">
|
||
<typeStatus id="75993221B62A50EEF5D4BB38F2ADAA26">Type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DBF2A1B58C1959A377F51836620B52B5" pageId="0" pageNumber="1">
|
||
<materialsCitation id="02317F95601E908A434AEB8A055BBB61" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="M. Bang" country="Bolivia" location="Cochabamba" specimenCount="1" stateProvince="Cochabamba" typeStatus="lectotype">
|
||
<collectingCountry id="466E69B23D944675601D46B1FC445242" name="Bolivia">Bolivia</collectingCountry>
|
||
.
|
||
<collectingRegion id="4587799105A2315073028B0EE93152D3" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
|
||
: Vic.
|
||
<collectingRegion id="A49DF7C46C767193FFE965EA39A37826" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
|
||
, 1891,
|
||
<emphasis id="2FB5BD500BFED8C4277420C4588C07AB" italics="true" pageId="0" pageNumber="1">
|
||
<collectorName id="DD5169DEAA6A3B632992367EBC177F0A">M. Bang</collectorName>
|
||
1151
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="F398416A1F194B7727E26B3FA6BF15BB">lectotype</typeStatus>
|
||
, designated by
|
||
<bibRefCitation id="5B5CF4A854AD851AAB64ACBE16F25867" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. 2013</bibRefCitation>
|
||
, pg. 236: NY [00139097]; isolectotypes: BM [BM000617675], BR [BR0000005538553], CAL [acc.# 316673], E [E00190740], G [G00343347], GH [00077599], MO [MO-503629, acc. # 1815484], NY [00139096],
|
||
</materialsCitation>
|
||
<collectingCountry id="CB721ACEDAF051EB7B3EE8AA98E430B4" name="Philippines">PH</collectingCountry>
|
||
[00030399],
|
||
<collectingCountry id="49545912083481814CD3616C78E910F7" name="United States of America">US</collectingCountry>
|
||
[00610905, acc. # 92001; 00027515, acc. # 1324496], WIS [0256183WIS]).
|
||
</paragraph>
|
||
<caption id="7075CDADE68DAC98C70DA3708A233747" doi="10.3897/phytokeys.231.100894.figure38" httpUri="https://binary.pensoft.net/fig/897499" pageId="0" pageNumber="1" start="Figure 38" startId="F38">
|
||
<paragraph id="A38AE4F4902598510778C636C7DD9FEE" pageId="0" pageNumber="1">
|
||
<emphasis id="E8881364BA4DA6518851812074DE17B8" bold="true" pageId="0" pageNumber="1">Figure 38.</emphasis>
|
||
<taxonomicName id="4BEAB8FAC3851FCE80984A9631094D61" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="17DF1F55FCC3A1CFCA47D77C8E290691" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="79968CC0449A37430339950EFB2CA311" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
flowering branch
|
||
<emphasis id="9B6D99BE74819CD16149AFBD03679E25" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
detail of adaxial leaf surface
|
||
<emphasis id="AAE94EAE700B77ACDF94A3302C8C8C11" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
detail of abaxial leaf surface
|
||
<emphasis id="15E90FFDEA08E27706A9204BBE22724F" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
flower bud
|
||
<emphasis id="B26ECF4EA05344B06A5843C9F7B0C61E" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
dissected flower
|
||
<emphasis id="AA66C2C16837D18A9D483ECCA9B2E8E3" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
fruiting branch
|
||
<emphasis id="060B215B44D6AA69517F95FAE4E9B2EC" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
maturing fruit (
|
||
<emphasis id="70C11C6AF7FABE29E52F9A0C8CFB55EA" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
||
<emphasis id="B572B5F960FDC08C50A5690C72BB06A3" italics="true" pageId="0" pageNumber="1">Knapp et al. 10341</emphasis>
|
||
; E
|
||
<emphasis id="38E2449308B2DF74873DBBB9199F8B8C" italics="true" pageId="0" pageNumber="1">
|
||
<normalizedToken id="D23F355D02AD9D5275ABF5870CF0DE62" originalValue="Cárdenas">Cardenas</normalizedToken>
|
||
5577
|
||
</emphasis>
|
||
<emphasis id="69746D046C772DCDE29AFEAD73280466" bold="true" pageId="0" pageNumber="1">F, G</emphasis>
|
||
<emphasis id="B5571AD5F5D916421CA5BB4FAA4BFD0B" italics="true" pageId="0" pageNumber="1">Knapp et al. 10339</emphasis>
|
||
). Illustration by R. Wise.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="192488980F788A26B20020B8902D23F9" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="C245B95E73120B3643F290F95AA9CB5B" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="91D2DCE39FE02102B60173802C79B18D" pageId="0" pageNumber="1">
|
||
Lax subwoody or woody shrubs, often vine-like with very long stems, to 5 m long, to 3 m if erect. Stems erect or sprawling, terete or slightly angled with tiny spinescent processes along the angles, moderately pubescent with eglandular white simple uniseriate 2-6-celled trichomes to 1 mm long, these soft and spreading; new growth densely white pubescent with eglandular simple uniseriate trichomes like those of the stems; bark of older stems pale brown, glabrescent. Sympodial units difoliate or plurifoliate, the leaves not geminate. Leaves simple or occasionally shallowly toothed, the blades 3.5-16 cm long, 1.5-8 cm wide, variable within an individual plant and always larger on lower stems, elliptic to narrowly elliptic, widest in the lower half, membranous, discolorous; adaxial surfaces sparsely pubescent with soft, spreading, eglandular simple uniseriate trichomes to 1 mm long, like those of the stems, these denser on the veins; abaxial surfaces more densely pubescent with simple uniseriate trichomes, the lamina still visible; principal veins 7-9 pairs, densely pubescent on abaxial surfaces; base acute, somewhat attenuate onto the petiole; margins entire or rarely shallowly toothed, the teeth if present in the basal part of the leaf, ca. 1 mm long, ca. 1.5 mm wide, with acute apices (see
|
||
<emphasis id="85A6F83F63B1454150AE317F4E0D6C63" italics="true" pageId="0" pageNumber="1">Brooke 5125</emphasis>
|
||
, one duplicate entire, one toothed); apex acute to somewhat acuminate; petiole 0.5-2.8 cm long, slightly winged from the decurrent leaf bases in the distal part. Inflorescences internodal or terminating branches, several times branched, 3-13 cm long, with 10-80+ flowers clustered at the branch tips, moderately pubescent with soft, spreading eglandular simple uniseriate trichomes to 1 mm long like those of the stems; peduncle 1.7-10 cm long; pedicels 0.6-1 cm long, 0.5-0.75 mm in diameter at the base, 1-1.5 mm in diameter at the apex, tapering, spreading at anthesis, moderately pubescent like the inflorescence axes, articulated at the base; pedicel scars 0.5-1 mm part at the branched tips. Buds ellipsoid, occasionally somewhat inflated, the corolla strongly exserted from the calyx before anthesis. Flowers 5-merous, cosexual (hermaphroditic). Calyx tube 1-1.5 mm long, conical, the lobes 1.2-2 mm long, 1-1.5 mm wide, narrowly deltate, moderately pubescent with simple uniseriate trichomes like the rest of the plant. Corolla 2-3 cm in diameter, extremely variable through anthesis in size and colour, pale violet to whitish violet, with a pale greenish yellow eye, stellate, lobed 1/3 to 1/2 of the way to the base, the lobes 4-6 mm long, 4-5 mm wide, deltate or broadly deltate, spreading to slightly reflexed at anthesis, glabrous adaxially or with scattered uniseriate trichomes ca. 0.2 mm long at the tips and margins, abaxially densely papillate-puberulent with papillae and simple uniseriate 1-3-celled trichomes to 0.5 mm long along the lobe midveins, tips and margins, the interpetalar tissue glabrous. Stamens equal; filament tube to 0.25 mm long; free portion of the filaments 1-1.5 mm long, pubescent adaxially with densely tangled, transparent weak simple uniseriate trichomes; anthers 3.5-4.5 mm long, 1.2-1.5 mm wide, ellipsoid, yellow, poricidal at the tips, the pores lengthening to slits with age. Ovary conical, glabrous; style 9-10 mm long, straight at anthesis (curved in bud), exserted beyond the anther cone, densely pubescent with transparent simple uniseriate trichomes in the lower half; stigma clavate, somewhat bilobed or capitate, green or dark cream in live plants, the surface minutely papillate. Fruit a globose berry, (0.9)1-1.2 cm in diameter, green and usually maturing purplish black, the pericarp thin, matte, translucent when berry ripe, glabrous; fruiting pedicels 1-1.7 cm long, ca. 1.2 mm in diameter at the base, ca. 2 mm in diameter at the apex, slightly woody, deflexed or spreading, not persistent; fruiting calyx somewhat enlarged, the tube to 2 mm long, the lobes to 2 mm long, appressed to the berry. Seeds 30-50 per berry, 1.5-2 mm long, 1-1.5 mm wide, flattened and teardrop shaped, pale brown to golden tan, the surfaces minutely pitted, the testal cells elongate with sinuate walls. Stone cells 8-12 per berry, 0.5-1 mm in diameter, cream-coloured, distributed throughout the mesocarp. Chromosome number: n = 12 (
|
||
<bibRefCitation id="2ED8D5398463D44C281C2D9B154D38B0" DOI="https://doi.org/10.2307/4117874" author="Edmonds, JM" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="1" pagination="95 - 114" refId="B73" refString="Edmonds, JM, 1972. A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America. Kew Bulletin 27 (1): 95 - 114, DOI: https://doi.org/10.2307/4117874" title="A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America." url="https://doi.org/10.2307/4117874" volume="27" year="1972">Edmonds 1972</bibRefCitation>
|
||
, voucher
|
||
<emphasis id="65E5AA81E2B71982628FBC3739DADDD9" italics="true" pageId="0" pageNumber="1">Steinbach 34</emphasis>
|
||
, as
|
||
<taxonomicName id="5E35276EE2972162C0019E35BA99CB62" family="Solanaceae" kingdom="Plantae" lsidName="S. polytrichostylum subsp. var. var. lorentzii" order="Solanales" pageId="0" pageNumber="1" rank="variety" species="polytrichostylum" subSpecies="var." variety="lorentzii">S. polytrichostylum var. lorentzii</taxonomicName>
|
||
;
|
||
<bibRefCitation id="F9FC00EEEFAAF3B93A7ED22969BB89E0" author="Moscone, EA" journalOrPublisher="Darwiniana" pageId="0" pageNumber="1" pagination="261 - 297" publicationUrl="https://www.jstor.org/stable/23222568" refId="B168" refString="Moscone, EA, 1992. Estudios en cromosomas meioticos en Solanaceae de Argentina. Darwiniana 31: 261 - 297, https://www.jstor.org/stable/23222568" title="Estudios en cromosomas meioticos en Solanaceae de Argentina." url="https://www.jstor.org/stable/23222568" volume="31" year="1992">Moscone 1992</bibRefCitation>
|
||
, voucher
|
||
<emphasis id="5AC73AA66D3325B4237E5EA1F8DDEA08" italics="true" pageId="0" pageNumber="1">Subils et al. 3253</emphasis>
|
||
, as
|
||
<taxonomicName id="0937B87B6A5A4FBCA2E5AAAB1022536E" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="F3750BA16490CD22AA50BE9054625787" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="23297A6C5F411E1865A0514ACDB66A58" DOI="https://doi.org/10.1080/00087114.2013.855389" author="Moyetta, NR" journalOrPublisher="Caryologia" pageId="0" pageNumber="1" pagination="333 - 345" refId="B169" refString="Moyetta, NR, Stiefkensa, LB, Bernardello, G, 2013. Karyotypes of South American species of the Morelloid and Dulcamaroid clades (Solanum, Solanaceae). Caryologia 66 (4): 333 - 345, DOI: https://doi.org/10.1080/00087114.2013.855389" title="Karyotypes of South American species of the Morelloid and Dulcamaroid clades (Solanum, Solanaceae)." url="https://doi.org/10.1080/00087114.2013.855389" volume="66" year="2013">Moyetta et al. 2013</bibRefCitation>
|
||
, voucher
|
||
<emphasis id="6876067D1B095D8411B22820C30DE2C2" italics="true" pageId="0" pageNumber="1">Barboza et al. 2152</emphasis>
|
||
, as
|
||
<taxonomicName id="DE9CFBE72AEF6C0E31582D4AB2F5CEFA" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="64A4DBCE055C54ECF4D15473C49B7B9D" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="358518E1A83F53778B96DE6CF2110E11" doi="10.3897/phytokeys.231.100894.figure39" httpUri="https://binary.pensoft.net/fig/897500" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
|
||
<paragraph id="E5AA8C1060BA7FE95DFDFDDAB3D9F565" pageId="0" pageNumber="1">
|
||
<emphasis id="737D0829CC6EC8F4A6BF5EC74F24A9F3" bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
|
||
<taxonomicName id="7B2F7F2F916FDC89A25E1105FE6C020C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="80864060E72D9174CBE79239B4A3EF55" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CB2E315493E46217494D064EB519A665" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
habit
|
||
<emphasis id="D1195819F16D21704552EDF866AE6F2A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
inflorescence with purple corollas
|
||
<emphasis id="9ECC5ED73CFA3F971CEF80B2FCB875AE" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
infructescence with white corollas
|
||
<emphasis id="12534DD8E1F6B7886C8D156CDA0C572E" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
mature fruits (
|
||
<emphasis id="AB1F8FC2B560B3987322190116DF839C" bold="true" pageId="0" pageNumber="1">A, C</emphasis>
|
||
<emphasis id="4523849688FF22D15DA317030B549115" italics="true" pageId="0" pageNumber="1">Knapp et al.10363</emphasis>
|
||
<emphasis id="5899A17B4840A2C5818B2808463A07DC" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<emphasis id="96BA17BEDF969D874A89D2DE158A9E9C" italics="true" pageId="0" pageNumber="1">Knapp 10287</emphasis>
|
||
<emphasis id="AD043590A2A30D8E647EEA652D6854EB" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<emphasis id="D1F1B356807AF3CCD84AB5F609B8D85D" italics="true" pageId="0" pageNumber="1">Barboza et al. 3500</emphasis>
|
||
). Photos by S. Knapp.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="7F33E89618B42E3B672CBE965D8B2695" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="66C873E2AF3C078F6D9EEF5442F56D35" pageId="0" pageNumber="1">Distribution</paragraph>
|
||
<paragraph id="419039440FFE8AEEFBF1539AE50BBE8A" pageId="0" pageNumber="1">
|
||
<emphasis id="8F34BB05AEDD60D5A021A24E0D215054" bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation id="C074B8DDB3A2BC7C91A2F10A5388A020" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Distribution map of Solanum cochabambense." figureDoi="10.3897/phytokeys.231.100894.figure40" httpUri="https://binary.pensoft.net/fig/897501" pageId="0" pageNumber="1">40</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<taxonomicName id="715B5DD35777EB34B30641BCE80CFC38" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="8E49B1E52CD0AB708D0F5EC5B35706ED" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
occurs from the eastern Andean slopes from northern Peru (Depts. Amazonas, Ancash,
|
||
<normalizedToken id="3C2A4A9DD3D7D1629EDAD596ACBD1C5E" originalValue="Apurímac">Apurimac</normalizedToken>
|
||
, Arequipa, Cajamarca, Cusco, Huancavelica,
|
||
<normalizedToken id="99578A525ED13DC974608062C7BBDEFD" originalValue="Huánuco">Huanuco</normalizedToken>
|
||
,
|
||
<normalizedToken id="7E3E9DEC02E09E0DE2C407A082B739DF" originalValue="Junín">Junin</normalizedToken>
|
||
, La Libertad, Puno, San
|
||
<normalizedToken id="088BE99B5257181CFF87B1F25D29292C" originalValue="Martín">Martin</normalizedToken>
|
||
) throughout the Andean cordillera to Bolivia (Depts. Chuquisaca, Cochabamba, La Paz, Oruro,
|
||
<normalizedToken id="7FE33764BE88D4F5E3EA48DDCA93DA43" originalValue="Potosí">Potosi</normalizedToken>
|
||
, Santa Cruz, Tarija) and northern Argentina (Provs. Jujuy, Salta,
|
||
<normalizedToken id="2E708DE37F508460258180961199D500" originalValue="Tucumán">Tucuman</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<caption id="85B6D3773E35DC7DBABA71D810AA6B81" doi="10.3897/phytokeys.231.100894.figure40" httpUri="https://binary.pensoft.net/fig/897501" pageId="0" pageNumber="1" start="Figure 40" startId="F40">
|
||
<paragraph id="A5970FDCE7492EC0DEFA9606D0FA9518" pageId="0" pageNumber="1">
|
||
<emphasis id="9E89F93198CA2CBBE99FD1C16C875B60" bold="true" pageId="0" pageNumber="1">Figure 40.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="B6326BE8E7D6AFB86C938377EF2C1B6E" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="DA1906ADA60CE657AE2BCE066224C526" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="0D06698768C50EC3A3ADB9E506435AE2" pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph id="745D26C25608671A0B5DC8B7C70F160E" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
|
||
<paragraph id="6D33511D21391F95A306AA87F2C3BA90" pageId="0" pageNumber="1">
|
||
<taxonomicName id="D56005D3EB48EBE3D1D5E03D58237874" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="23F92FCCF4BC75FBA76A9D7532614201" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
grows in a wide variety of middle to high elevation forest types, often at roadsides or in landslips and treefalls, from 150 to 4,120 m; most collections are from elevations above 1,000 m. The single collection from low elevation (
|
||
<emphasis id="8BBF4C82F92D84E2FD1F9BE093F0C4E0" italics="true" pageId="0" pageNumber="1">Roque 295</emphasis>
|
||
from 150 m in
|
||
<normalizedToken id="789F30183E2EE659CFA2BA0F4D43E867" originalValue="Camaná">Camana</normalizedToken>
|
||
, Arequipa, Peru) comes from an area where landslides (
|
||
<normalizedToken id="0CAE5BCC600C671265E43C02B44393A3" originalValue="‘huaicos’">'huaicos'</normalizedToken>
|
||
) are common and perhaps represents seeds washed down from higher elevations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30F3ED19F002F8D23F1F2961FAA54363" pageId="0" pageNumber="1" type="common names and uses">
|
||
<paragraph id="CCDA6F3895875B1B2A7B82842D4105B3" pageId="0" pageNumber="1">Common names and uses.</paragraph>
|
||
<paragraph id="ACB64B67518B7214C6B7E9A2FD1A38B8" pageId="0" pageNumber="1">
|
||
Bolivia. La Paz: chinchi-chinchi (
|
||
<emphasis id="6F4FBBBE0E1EC19D5B930CFFA0F4C879" italics="true" pageId="0" pageNumber="1">Beck 27781</emphasis>
|
||
), cusmayo (
|
||
<emphasis id="75FB54376FBDF649677EB3B5E2AE308E" italics="true" pageId="0" pageNumber="1">Lewis 881659</emphasis>
|
||
). Peru. Ancash: atoqpa
|
||
<normalizedToken id="BBD3BC574422D4A9851C23E4394202B1" originalValue="papán">papan</normalizedToken>
|
||
(papa de zorro) (
|
||
<emphasis id="D1E6F06D295B9CD91C6B4FCC04021909" italics="true" pageId="0" pageNumber="1">Gamarra 662</emphasis>
|
||
); Cusco: ccaya-ccaya (
|
||
<emphasis id="1F21B162085C6003143B07364EFE75F5" italics="true" pageId="0" pageNumber="1">Mexia 8079</emphasis>
|
||
), chinchi-chinchi (
|
||
<emphasis id="9860C605EFF1A0901B3789995650DD92" italics="true" pageId="0" pageNumber="1">Herrera 819</emphasis>
|
||
); muya khaya (
|
||
<emphasis id="6F8181B0BE89DE3E8454151EA0B04FE6" italics="true" pageId="0" pageNumber="1">Franquemont et al. 297</emphasis>
|
||
); qusmayllu (
|
||
<emphasis id="202DD3C903381204783A60BBD9F0B84C" italics="true" pageId="0" pageNumber="1">Franquemont et al. 348</emphasis>
|
||
);
|
||
<normalizedToken id="302AAFE9719E37FF2DA9BBEF2E7E9C25" originalValue="Huánuco">Huanuco</normalizedToken>
|
||
: shopta (
|
||
<emphasis id="03358C377C3ACC9E154FBE9DAB740823" italics="true" pageId="0" pageNumber="1">Weberbauer 6789</emphasis>
|
||
); Puno: chitinqoya (
|
||
<bibRefCitation id="E283F867930AA67E0432ADB00D28E7C0" author="Roersch, C" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="1" refId="B194" refString="Roersch, C, 1994. Plantas Medicinales en el Sur Andino del Peru. 2 vols. Koeltz Scientific Books, Koenigstein." title="Plantas Medicinales en el Sur Andino del Peru. 2 vols. Koeltz Scientific Books, Koenigstein." year="1994">Roersch 1994</bibRefCitation>
|
||
). In the southern Peruvian Quechua community of Chinchero (Cusco, Peru) leaves are used as cattle forage (
|
||
<bibRefCitation id="AEB44D006DAB508B596F3D773739A136" author="Franquemont, C" journalOrPublisher="Sexual Plant Reproduction" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/2643239" refId="B88" refString="Franquemont, C, Franquemont, E, Davis, EW, Plowman, TC, King, SP, Sperling, CR, Neizgoda, C, 1990. The ethnobotany of Chinchero, an Andean community in southern Peru. Fieldiana n.s. 24: 1-126, https://www.biodiversitylibrary.org/page/2643239" title="The ethnobotany of Chinchero, an Andean community in southern Peru. Fieldiana n. s. 24: 1 - 126," url="https://www.biodiversitylibrary.org/page/2643239" year="1990">Franquemont et al. 1990</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4345B3EEFBC8BC3AC063DEE6F9994C93" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="C0C20098B80E5F453A5E1B985C1916BA" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
) and as a wash for the head, especially for hangovers (
|
||
<bibRefCitation id="BA18352CF52F1C198BF84915BB187103" author="Franquemont, C" journalOrPublisher="Sexual Plant Reproduction" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/2643239" refId="B88" refString="Franquemont, C, Franquemont, E, Davis, EW, Plowman, TC, King, SP, Sperling, CR, Neizgoda, C, 1990. The ethnobotany of Chinchero, an Andean community in southern Peru. Fieldiana n.s. 24: 1-126, https://www.biodiversitylibrary.org/page/2643239" title="The ethnobotany of Chinchero, an Andean community in southern Peru. Fieldiana n. s. 24: 1 - 126," url="https://www.biodiversitylibrary.org/page/2643239" year="1990">Franquemont et al. 1990</bibRefCitation>
|
||
, as.
|
||
<taxonomicName id="A50111DF9991898BC2A248D6F7D00DC1" family="Solanaceae" kingdom="Plantae" lsidName="S. glandulosipilosum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="glandulosipilosum">
|
||
<emphasis id="C530D860001B76E45B2B8A9EB4D3586E" italics="true" pageId="0" pageNumber="1">S. glandulosipilosum</emphasis>
|
||
</taxonomicName>
|
||
). In southern Peru more generally leaves of
|
||
<taxonomicName id="BC987F71BFBBF58AAA7395ED52B8475D" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="303C9B0AD722537D8415DCC593D713FC" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
are used medicinally in a tea in the treatment of flu and colds (
|
||
<normalizedToken id="3BB46D9F7901E4439EF397C7D64C8BD8" originalValue="resfrío">resfrio</normalizedToken>
|
||
) and to counter difficulty in urination, and as a macerated plaster to alleviate rheumatic pains (
|
||
<bibRefCitation id="A7F0F9E3CB12BBE080C6EEBE7CB91EE2" author="Roersch, C" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="1" refId="B194" refString="Roersch, C, 1994. Plantas Medicinales en el Sur Andino del Peru. 2 vols. Koeltz Scientific Books, Koenigstein." title="Plantas Medicinales en el Sur Andino del Peru. 2 vols. Koeltz Scientific Books, Koenigstein." year="1994">Roersch 1994</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="02ADBA2513C2237293F60A9005341EF1" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="96E71738E4F69A05C2C5D9F966E49DA0" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2137B20B00985C65FD2A392B3F040ECB" pageId="0" pageNumber="1" type="preliminary conservation status">
|
||
<paragraph id="8BB33B3951469758EA7698EDA5C0CC8A" pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
|
||
<paragraph id="495E8D7E98F5E15FF7ACDEAAC02A8D6D" pageId="0" pageNumber="1">
|
||
<emphasis id="CCC70A33774EDE1809FA7A4EC4690EB5" bold="true" pageId="0" pageNumber="1">
|
||
(
|
||
<bibRefCitation id="84424CBA6AEF0DBE6242C5207EC5AFF8" author="Ibarrola, DA" journalOrPublisher="Nueva Serie" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B116" refString="2022. . https://www.iucnredlist.org/resources/redlistguidelines" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN 2022</bibRefCitation>
|
||
).
|
||
</emphasis>
|
||
Least Concern [LC]. EOO = 7,244,968 km2 [LC]; AOO = 1,132 km2 [VU].
|
||
<taxonomicName id="0D17EB98399E7534AACDE8F25E0F526C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="E4408352CCF6419F3783B54FDE5B5000" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
is a common plant of disturbed areas. Further study may reveal variation that warrants taxonomic distinction, and this preliminary assessment will need revisiting.
|
||
<taxonomicName id="18A268E1315E7861D8E819B5493DD433" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="5261E1AA7E0AE106172E5DB78B57713E" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
occurs within several protected areas across its range (see Supplementary materials).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E11995073C3A4339CBC4A1A78372D215" pageId="0" pageNumber="1" type="discussion">
|
||
<paragraph id="58DCA8D258B176EA922536093D4535CA" pageId="0" pageNumber="1">Discussion.</paragraph>
|
||
<paragraph id="829C0145506D6A4C512E0CAA4BC12DAC" pageId="0" pageNumber="1">
|
||
<taxonomicName id="0EBBE404C6AB727C37C8AD9BE3591401" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="40F5ADFC5207319B0E4F531B6243D01C" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
is one of the most variable and widespread morelloid species in South America.
|
||
<bibRefCitation id="10C610FBD93640929A3F497D29EF8B47" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. (2013)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="C64EBE0FE391BC6C0F4DE24EB349CD16" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="B043C66D87D75C465CE87776E63A06E9" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
in synonymy with
|
||
<taxonomicName id="5B8C0F85E625A565ADD9F418BDE6296F" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="89B09F88967870E5DFFE8BF3CB4860E3" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
, with which it is sympatric in northern Argentina. Further study throughout the range of
|
||
<taxonomicName id="DDBCB21A9BADCA80443E4986A92F37E5" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="1B5F896D7372BC2512077D1E48467EF4" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
confirmed the distinctness of the two species, but individual specimens collected in sympatry can be difficult to identify.
|
||
<taxonomicName id="80A240EA9AB03806C3539C8795EF8B1E" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum cochabambense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cochabambense">
|
||
<emphasis id="2465B15704BCF6E7FE1A71BC0913AED3" italics="true" pageId="0" pageNumber="1">Solanum cochabambense</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="7F370884CD36A691DD88E07FB3BF564F" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="59E944F24D4B11D0FC6855C6BC3581AC" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
in its more highly branched inflorescences (those of
|
||
<taxonomicName id="F93BBD7858F010E9E4288FF941D03F3C" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="D37B70B361FDB8E14A312BD52C5CA7DF" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
are usually only forked), buds that are ellipsoid rather than narrowly ellipsoid and larger corollas and berries. The anthers of
|
||
<taxonomicName id="A5D3E3E9E462E2EAC98EBB57BCB85937" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="AC75EF261B7E501E26C0FF99E6F6D5C5" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
are narrow relative to their length (3.9-5 mm long and 0.6-1 mm wide in
|
||
<taxonomicName id="DB216824B9124164C8605888B93BC926" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="5A4DA93F053AE92DF565F946EFB13171" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
versus 3.5-4 mm long and 0.9-1.2 mm wide in
|
||
<taxonomicName id="6AE2E82B42AAEAFC2FCA53B6EC139282" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="37F3E7B20941B1FBE4701207010AC972" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
) but this character can be difficult to see in the absence of comparative material. The berries of
|
||
<taxonomicName id="D4022E4E6B28320FD4B7E0FADEBC0D5D" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="C62B953D59325C5D9927E976D9D29494" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
are larger (1-1.2 cm in diameter) than those of
|
||
<taxonomicName id="493B5CB554088F95A80B8F10D13D855E" family="Solanaceae" kingdom="Plantae" lsidName="S. aloysiifolium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="aloysiifolium">
|
||
<emphasis id="20E912011426FEB260C63C0417BD7AA4" italics="true" pageId="0" pageNumber="1">S. aloysiifolium</emphasis>
|
||
</taxonomicName>
|
||
(0.5-0.6 cm in diameter), with similar numbers of stone cells.
|
||
</paragraph>
|
||
<paragraph id="29A417CF299BED8166737A741EEBE630" pageId="0" pageNumber="1">
|
||
In Bolivia
|
||
<taxonomicName id="EF4A2C086382F88D16B4B4B1C77F3BE8" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="0CAB49FCEC694F03F1DE9B0348F481CA" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
is partially sympatric with and morphologically very similar to
|
||
<taxonomicName id="84F1104C9308219A0E162D3070325571" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
|
||
<emphasis id="A0728EBBE43DD32AB1B504E6CC5E367F" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="65229FD1708927073D2C096E6CCF4F29" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
|
||
<emphasis id="4C4BC41E7C6DF41DA3BCE5804BDE9DFB" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
|
||
</taxonomicName>
|
||
differs in its possession of dendritic trichomes, while
|
||
<taxonomicName id="66682CCF47820EC2C81BA94F69FA4E0D" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="78217377B409F60F6B018505B790BC5C" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
has only simple trichomes.
|
||
</paragraph>
|
||
<paragraph id="5BB236CC526F993B60ACB2BE420A8B64" pageId="0" pageNumber="1">
|
||
In the northern part of its range,
|
||
<taxonomicName id="3150884B9B6BED5C891993F794D61921" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="0B46A1004BCFA19E69BBCAE8558F7124" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
can be confused with
|
||
<taxonomicName id="36A88F248F1ED7333B69F65F9D57FB69" family="Solanaceae" kingdom="Plantae" lsidName="S. arequipense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="arequipense">
|
||
<emphasis id="40192F71FC198E75F50D476F3DACC4C3" italics="true" pageId="0" pageNumber="1">S. arequipense</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="45153B8F22BB32B2FA09C1D640DA52A2" family="Solanaceae" kingdom="Plantae" lsidName="S. juninense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="juninense">
|
||
<emphasis id="0DED5759013CC6491BBA312C7CB9FECE" italics="true" pageId="0" pageNumber="1">S. juninense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CB1B8FE786CE325989A1E90E152D9C6" family="Solanaceae" kingdom="Plantae" lsidName="S. interandinum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="interandinum">
|
||
<emphasis id="C259B7E70AB59166F98BD26FB27783F8" italics="true" pageId="0" pageNumber="1">S. interandinum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="E779BC8D7F464CB3A73D32DAB49F1AD8" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum juninense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="juninense">
|
||
<emphasis id="0C5935142807DD86E4E10D7C0A053608" italics="true" pageId="0" pageNumber="1">Solanum juninense</emphasis>
|
||
</taxonomicName>
|
||
differs in its possession of glandular trichomes whereas
|
||
<taxonomicName id="6C66A312ECF35DC57D4FD0546ED56C6A" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="8E26ED761F2A44F104349D834CED8B1B" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
is always eglandular.
|
||
<taxonomicName id="D789002C545DF161291FA219D985D81D" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum arequipense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="arequipense">
|
||
<emphasis id="641481143FF04EA02886B88E432B6DE4" italics="true" pageId="0" pageNumber="1">Solanum arequipense</emphasis>
|
||
</taxonomicName>
|
||
has blunt-tipped calyx lobes, anthers 2.5-3 mm long and a strongly capitate stigma, while
|
||
<taxonomicName id="17251E59D9560A93C4AD5BF7D80A6A24" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="206829353475675E6CE39275E03C66EE" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
has long-triangular calyx lobes with acute apices, anthers 3.5-4 mm long and a clavate to only somewhat capitate stigma. The calyx lobes of
|
||
<taxonomicName id="CDC6F007014468CCAF606AA258D169C3" family="Solanaceae" kingdom="Plantae" lsidName="S. interandinum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="interandinum">
|
||
<emphasis id="BC3321324E8313EC1AD940CDBD228EE1" italics="true" pageId="0" pageNumber="1">S. interandinum</emphasis>
|
||
</taxonomicName>
|
||
are longer and more pointed than those of
|
||
<taxonomicName id="D95EF1A420CAF5432CC1A4E2B3A2FAD7" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="FD2EE04AD200DC1E53F17FDF5F73C9BF" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
, and the flowers are smaller (0.8-1.4(1.8) cm in diameter versus 2-3 cm in diameter in
|
||
<taxonomicName id="6F607CB4E6728959E8367DC5F4EFA851" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="D092A472BC566F06002BD896CD61CB2B" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1AF3CE2B88DF800D8175A773EEC163A9" pageId="0" pageNumber="1">
|
||
The extreme variability seen across the range of
|
||
<taxonomicName id="D26EB884ACABDB15291EEB79F7941B28" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="4DD40BDD82588DFBDDBA37D6961757BE" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
may indicate there are several distinct species contained within our rather broad circumscription. In some cases, duplicate collections from the same locality show that variation is present within a single population, which has helped us to recognise this group of specimens as a morphologically variable single species: an example of such variation is leaf margins varying from entire to toothed in duplicates of
|
||
<emphasis id="8179EFFFEAD1F3FF279A994B22B425C6" italics="true" pageId="0" pageNumber="1">Brooke 5125</emphasis>
|
||
. Similarly, variation in corolla shape and size was evident in the field in some populations, as well as inflorescence structure (e.g.,
|
||
<emphasis id="43105A297F5FAB959B348963250FFC7F" italics="true" pageId="0" pageNumber="1">Knapp et al. 10391, Knapp et al. 10392, Knapp et al. 10393, Knapp et al. 10669</emphasis>
|
||
). Variation in other characters such as indumentum, calyx lobe shape and size, and other characters may represent fixed differences between populations, but based on our study of the specimens available across geographic space, we circumscribe this as a single highly variable species. Future studies at the population level throughout the range will be important to identify potential taxonomically recognisable segregates in this species.
|
||
</paragraph>
|
||
<paragraph id="172B5E64ED35523A9D08CB3E77DFF22C" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="8DD95EDFB1133DAF84808C3534B3FAD7" DOI="https://doi.org/10.1002/fedr.19120103305" author="Bitter, G" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="0" pageNumber="1" pagination="529 - 565" refId="B16" refString="Bitter, G, 1912a. Solana nova vel minus cognita I. Repertorium Specierum Novarum Regni Vegetabilis 10 (33-36): 529 - 565, DOI: https://doi.org/10.1002/fedr.19120103305" title="Solana nova vel minus cognita I." url="https://doi.org/10.1002/fedr.19120103305" volume="10" year="1912 a">Bitter (1912a)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="5B90ADAB4CFEBFCFD39859BAAEEE35C0" family="Solanaceae" kingdom="Plantae" lsidName="S. extuspellitum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="extuspellitum">
|
||
<emphasis id="40D8F643BE1B0590312DA1354AE6ABAB" italics="true" pageId="0" pageNumber="1">S. extuspellitum</emphasis>
|
||
</taxonomicName>
|
||
and its variety
|
||
<emphasis id="5C647F2E40FF9087CE8DBDB9E29B2DD4" italics="true" pageId="0" pageNumber="1">subcoerulum</emphasis>
|
||
using the same collection (
|
||
<emphasis id="EF435BED5019B1C5FF9087AB95AA55D8" italics="true" pageId="0" pageNumber="1">Fiebrig 2439</emphasis>
|
||
), citing "p. pt. herb. Berol.!" in each protologue. The sheet in B (now destroyed but photographed as F neg. 2711) has two stems, one with the label "
|
||
<taxonomicName id="74F73F92131C4830BFE848BEE8258E7D" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum extuspellitum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="extuspellitum">Solanum extuspellitum</taxonomicName>
|
||
n. sp." and the other with the label "Solanum (extuspellitum subsp.) subcoeruleum Bitt." - this latter suggesting he had originally considered naming the latter at the specific level. The stems differ only in pubescence density, and both are referrable to
|
||
<taxonomicName id="585C0ED2B4D6F3E4CB0A728FB9806A2B" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
|
||
<emphasis id="60102FEE57E99C4D7BD9D99A60F133A0" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
|
||
</taxonomicName>
|
||
. It is impossible to tell from which stem the fragment held in F came, so we are using it as the lectotype for both names. We have found no other duplicates of
|
||
<emphasis id="21C4E0172C3FE1B5F39E7D92982F3C9B" italics="true" pageId="0" pageNumber="1">Fiebrig 2439</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B8064F414342E67C6A2A4B653A39A2EA" pageId="0" pageNumber="1">
|
||
Later that same year (
|
||
<bibRefCitation id="1B63C92DC7902DFCA5EF757FB212F971" DOI="https://doi.org/10.1002/fedr.19120110917" author="Bitter, G" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="0" pageNumber="1" pagination="202 - 237" refId="B18" refString="Bitter, G, 1912c. Solana nova vel minus cognita III. Repertorium Specierum Novarum Regni Vegetabilis 11 (9-15): 202 - 237, DOI: https://doi.org/10.1002/fedr.19120110917" title="Solana nova vel minus cognita III." url="https://doi.org/10.1002/fedr.19120110917" volume="11" year="1912 c">Bitter 1912c</bibRefCitation>
|
||
), he described
|
||
<taxonomicName id="F271E8922E6DEB5F8472F6B848144CBB" family="Solanaceae" kingdom="Plantae" lsidName="S. decachondrum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="decachondrum">
|
||
<emphasis id="7F2DD1DA699001B28851EC3F843B9D48" italics="true" pageId="0" pageNumber="1">S. decachondrum</emphasis>
|
||
</taxonomicName>
|
||
and its varieties
|
||
<taxonomicName id="47C7F696B72B626A674BABBF15248F23" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. longiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="longiusculum">
|
||
<emphasis id="21758D28873C09F843BAF696AC806E9E" italics="true" pageId="0" pageNumber="1">Solanum decachondrum longiusculum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="244547CD832997309A092706763CD78D" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. latiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="latiusculum">
|
||
<emphasis id="716D64BDB189614E9E7D96545B3BE7E5" italics="true" pageId="0" pageNumber="1">Solanum decachondrum latiusculum</emphasis>
|
||
</taxonomicName>
|
||
using two collections of Otto Buchtien (
|
||
<emphasis id="A33FA0252F98946518A0C751AD9C1081" italics="true" pageId="0" pageNumber="1">Buchtien 2411</emphasis>
|
||
and
|
||
<emphasis id="C39DD02F3757D400201FCBF80AEC0980" italics="true" pageId="0" pageNumber="1">2412</emphasis>
|
||
), citing no herbarium but indicating with
|
||
<normalizedToken id="31593F12FE93B1B529D338E344554A85" originalValue="“!”">"!"</normalizedToken>
|
||
that he had seen them. Otto
|
||
<normalizedToken id="EEC07B1A24CFA9F4F2F1089FC549A63E" originalValue="Buchtien’s">Buchtien's</normalizedToken>
|
||
private herbarium that Bitter cited was donated to the Smithsonian (US) in the 1920s (
|
||
<bibRefCitation id="F1CB92D7810AEFD22E426EB447AE0F6F" author="Morton, CV" journalOrPublisher="Plant Science Bulletin" pageId="0" pageNumber="1" pagination="1 - 4" publicationUrl="https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" refId="B167" refString="Morton, CV, Stern, WL, 1966. The history of the United States national herbarium. Plant Science Bulletin 12 (2): 1 - 4, https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" title="The history of the United States national herbarium." url="https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" volume="12" year="1966">Morton and Stern 1966</bibRefCitation>
|
||
), so lectotypes for names based on
|
||
<normalizedToken id="3A9C5E943063CAEBF281B812A1618BEA" originalValue="Buchtien’s">Buchtien's</normalizedToken>
|
||
collections should be in US. Bitter cited both numbers in the protologue of the species, then used
|
||
<emphasis id="83398FB5CF77901C38E5D5936EFA430F" italics="true" pageId="0" pageNumber="1">Buchtien 2411</emphasis>
|
||
for var.
|
||
<taxonomicName id="8E4C99C9382F34951CB58D1FC1C61BF7" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. longiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="longiusculum">Solanum decachondrum longiusculum</taxonomicName>
|
||
and
|
||
<emphasis id="C6B5A88EE4981800326EF30B2C5CD36E" italics="true" pageId="0" pageNumber="1">Buchtien 2412</emphasis>
|
||
for var.
|
||
<taxonomicName id="6FDA94A22FDAB7C6F112E3F22EDC54F5" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. latiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="latiusculum">Solanum decachondrum latiusculum</taxonomicName>
|
||
. None of the duplicates of these collections we have seen has annotations in
|
||
<normalizedToken id="2D00F3B56051157E44E8EE7383490ADA" originalValue="Bitter’s">Bitter's</normalizedToken>
|
||
hand, but one duplicate in US (01014170, acc. # 1175973) is annotated as var.
|
||
<taxonomicName id="75584A3A027E9DD5A0BDAF2703A1D8F8" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. longuisculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="longuisculum">Solanum decachondrum longuisculum</taxonomicName>
|
||
by Buchtien and is here designated the lectotype of var.
|
||
<taxonomicName id="4EECBDE5E0FC415A9AC7C522E88FDFD9" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. longuisculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="longuisculum">Solanum decachondrum longuisculum</taxonomicName>
|
||
. The other duplicate of
|
||
<emphasis id="95A88898C15BA700CF5ED19102E60A39" italics="true" pageId="0" pageNumber="1">Buchtien 2411</emphasis>
|
||
at US (00027539, acc. # 700102) is designated as the lectotype of
|
||
<taxonomicName id="082298AEDF00309351371A3AE711AC25" family="Solanaceae" kingdom="Plantae" lsidName="S. decachondrum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="decachondrum">
|
||
<emphasis id="AE9D275FFBACB192FA8C3066911772DF" italics="true" pageId="0" pageNumber="1">S. decachondrum</emphasis>
|
||
</taxonomicName>
|
||
. We lectotypify var.
|
||
<taxonomicName id="AF51F97462B13F7CBE3F5B9879E8E901" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. latiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="latiusculum">Solanum decachondrum latiusculum</taxonomicName>
|
||
with the US duplicate (000275538, acc. # 1177823) of
|
||
<emphasis id="070D09583D08AEB48A9F0822E6C83120" italics="true" pageId="0" pageNumber="1">Buchtien 2412</emphasis>
|
||
, as it is the best preserved with both flowers and fruits and is annotated as var.
|
||
<taxonomicName id="7056BB83C4F7613207DB091864D8D26B" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum decachondrum var. latiusculum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="decachondrum" variety="latiusculum">Solanum decachondrum latiusculum</taxonomicName>
|
||
in
|
||
<normalizedToken id="72BF372CF1C3779166494B523D79F2F2" originalValue="Bitter’s">Bitter's</normalizedToken>
|
||
hand.
|
||
</paragraph>
|
||
<paragraph id="3E2E0D70A5022ACE996D9804F46D3DAB" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="AB800748255E648E381F286934D8B44B" author="Bitter, G" journalOrPublisher="Beiblatt" pageId="0" pageNumber="1" pagination="5 - 17" refId="B22" refString="Bitter, G, 1916. Solanaceae andinae. Botanisches Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 54. Beiblatt 119: 5 - 17" title="Solanaceae andinae. Botanisches Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 54." volume="119" year="1916">Bitter (1916)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="23D6DC490054CF8E44FE3D70CBFA933C" family="Solanaceae" kingdom="Plantae" lsidName="S. probolospermum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="probolospermum">
|
||
<emphasis id="48B46D447E3B8AA8C2594233B4CD10C8" italics="true" pageId="0" pageNumber="1">S. probolospermum</emphasis>
|
||
</taxonomicName>
|
||
citing
|
||
<emphasis id="9257035C8C508F49FEA040106CE9D934" italics="true" pageId="0" pageNumber="1">Weberbauer 6789</emphasis>
|
||
but without citing a herbarium. Many duplicates of this collection number have been preserved, and we select the better preserved of the two duplicates of
|
||
<emphasis id="5AEAD37E1E3E0059627B1AF4C14FE6CF" italics="true" pageId="0" pageNumber="1">Weberbauer 6789</emphasis>
|
||
in the herbarium of the Universidad Nacional Agraria La Molina (MOL00005139) as the lectotype.
|
||
<normalizedToken id="5BED27D0B3EFA2BCAA03B5A0541AFA2F" originalValue="Weberbauer’s">Weberbauer's</normalizedToken>
|
||
original personal herbarium is held in MOL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |